identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
90278E13FFE9FF9657C5FD9F494E9273.text	90278E13FFE9FF9657C5FD9F494E9273.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dichogaster (Diplothecodrilus) bolaui (Michaelsen 1891)	<div><p>Dichogaster (Diplothecodrilus) bolaui (Michaelsen, 1891)</p><p>Benhamia bolavi Michaelsen, 1891: 9; Reynolds &amp; Cook 1976: 80.</p><p>Dichogaster bolaui Michaelsen, 1913a: 401; Ljungström 1972: 6; Reynolds &amp; Reinecke 1976: 8; Zicsi, 1998: 95.</p><p>Dichogaster (Diplothecodrilus) bolaui: Csuzdi, 1995: 102, 2000: 60, 2010: 120 (for complete synonymy list); Plisko, 2010: 292.</p><p>Distribution type: Peregrine</p><p>Distribution in Brazil: Present in 15 Brazilian states (Brown &amp; Fragoso, 2007, Sousa et al. 2020b)</p><p>MARANHÃO: Centro Novo do Maranhão: LB_ ANL000271, 57 adults, in plinthosol in a pasture, (3°40'25.49"S 46°47'20.48"W), April, 2016, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL000274, 5 adults, in plinthosol in a pasture, (3°55'3.55"S 46°45'49.55"W), April, 2017, L.M. Hernández-García, G.X. Rousseau, colls.; Itinga do Maranhão: LB_ ANL000272, 129 adults, in argisol in a pasture, (4°5'43.00"S 46°52'20.00"W), April, 2017, L.M. Hernández-García, G.X. Rousseau, colls. LB_ANL000273, 14 juveniles, in plinthosol in a logged forest, (3°55'23.40"S 46°45'57.04"W), April, 2017, L.M. Hernández-García, G.X. Rousseau, colls. (Figure 2)</p></div>	https://treatment.plazi.org/id/90278E13FFE9FF9657C5FD9F494E9273	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sousa, Sandriel Costa;Hernández-García, Luis Manuel;Beckman, Ruana Aretha Farias Santiago;Rousseau, Guillaume Xavier	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE9FF9657C5FB4C4EDE902B.text	90278E13FFE9FF9657C5FB4C4EDE902B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dichogaster (Diplothecodrilus) saliens (Beddard 1893)	<div><p>Dichogaster (Diplothecodrilus) saliens (Beddard, 1893)</p><p>Microdrilus saliens Beddard, 1892b: 683; Reynolds &amp; Cook, 1976: 166K</p><p>Dichogaster crawi Eisen, 1900: 228; Michaelsen, 1913b: 418K</p><p>Dichogaster saliens Ljungstrom, 1972: 6; Reynolds &amp; Reinecke, 1976: 7; Zicsi, 1998: 71; Dlamini, 2002: 62; Horn et al. K, 2007: 175K</p><p>Dichogaster (Diplothcecodrilus) saliens Csuzdi, 1995: 108, 2000: 66, 2010: 116; Plisko, 2010: 292K</p><p>Dichogaster (Diplothcecodrilus) saliens in saliens group Csuzdi, 1996: 358, 2010: 108K</p><p>Distribution type: Exotic</p><p>Distribution in Brazil: Present in 14 Brazilian states (Brown &amp; Fragoso, 2007)</p><p>MARANHÃO: Caxias (Righi, 1972). (Figure 2)</p></div>	https://treatment.plazi.org/id/90278E13FFE9FF9657C5FB4C4EDE902B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sousa, Sandriel Costa;Hernández-García, Luis Manuel;Beckman, Ruana Aretha Farias Santiago;Rousseau, Guillaume Xavier	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE9FF9757C5F8D44FE49770.text	90278E13FFE9FF9757C5F8D44FE49770.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eudrilus eugeniae (Kinberg 1867)	<div><p>Eudrilus eugeniae (Kinberg, 1867)</p><p>Lumbricus eugeniae Kinberg, 1867: 98 K</p><p>Eudrilus decipiens Perrier, 1871: 1176 K</p><p>Eudrilus lacazii Perrier, 1872: 75 K</p><p>Distribution type: Exotic (Brown &amp; Fragoso, 2007)</p><p>Distribution in Brazil: Present in 10 Brazilian states (Brown &amp; Fragoso, 2007; This study)</p><p>MARANHÃO: Caxias: LB_ ANL000375, 5 adults, in <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.302776&amp;materialsCitation.latitude=-2.5297222" title="Search Plazi for locations around (long -44.302776/lat -2.5297222)">Vermicomposting</a>, (4°51'32"S 43°21'10"W), February, 2019, L.M. Hernández-García.; Raposa: LB_ ANL000376, 4 adults, in soil next to vermicomposting, (2°25'44"S 44°06'34"W), May, 2022, L.M. Hernández-García.; São Luís: LB_ ANL000376, 10 adults, in Vermicomposting, (2°31'47"S 44°18'10"W), May, 2025, L.M. Hernández-García. (Figure 2)</p></div>	https://treatment.plazi.org/id/90278E13FFE9FF9757C5F8D44FE49770	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sousa, Sandriel Costa;Hernández-García, Luis Manuel;Beckman, Ruana Aretha Farias Santiago;Rousseau, Guillaume Xavier	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE8FF9757C5FC7F4E2D92D0.text	90278E13FFE8FF9757C5FC7F4E2D92D0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Holoscolex dossanti Hernandez-Garcia, Rousseau & James 2018	<div><p>Holoscolex dossanti Hernández-García, Rousseau &amp; James 2018</p><p>Distribution type: Native.</p><p>Distribution in Brazil: Maranhão: Centro Novo do Maranhão, Gurupi Biological Reserve (Hernandez et al. 2018a, This study).</p><p>MARANHÃO: Centro Novo do Maranhão: LB_ANL000275, 1 adult, in plinthosol in a logged forest, (3°40’50.06”S 46°46’12.95”W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls. LB_ANL000276, 2 adults, in plinthosol in an old-growth forest, (3°41’27.68”S 46°46’2.88”W), April, 2016, L.M. Hernández-García, G.X. Rousseau, colls. (Figure 2)</p></div>	https://treatment.plazi.org/id/90278E13FFE8FF9757C5FC7F4E2D92D0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sousa, Sandriel Costa;Hernández-García, Luis Manuel;Beckman, Ruana Aretha Farias Santiago;Rousseau, Guillaume Xavier	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE8FF9057C5FAE34D64936D.text	90278E13FFE8FF9057C5FAE34D64936D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Holoscolex fernandoi Hernandez-Garcia, Dos Santos, Rousseau & James 2018	<div><p>Holoscolex fernandoi Hernández-García, Dos Santos, Rousseau &amp; James 2018</p><p>Distribution type: Native.</p><p>Distribution in Brazil: Maranhão: Centro Novo do Maranhão, Gurupi Biological Reserve, Itinga (Hernandez et al. 2018a, This study).</p><p>MARANHÃO: Centro Novo do Maranhão: LB_ ANL000277, 9 adults, in plinthosol in an old-growth forest, (3°41'11.69"S 46°45'34.87"W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000278— 12 juveniles and LB_ ANL000279, 8 adults, in plinthosol in an old-growth forest, (3°41'16.35"S 46°45'34.73"W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000280— 6 juveniles and LB_ ANL000281, 8 adults, in plinthosol in a logged forest, (3°40'50.95"S 46°46'13.06"W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000282— 9 juveniles and LB_ ANL000283, 12 adults, in plinthosol in a logged forest, (3°40'59.74"S 46°46'1.11"W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000284— 6 adults and LB_ ANL000285, 2 juveniles, in plinthosol in an old-growth forest, (3°41'27.21"S 46°45'6.19"W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000286— 7 juveniles and LB_ ANL000287, 6 adults, in plinthosol in an old-growth forest, (3°41'24.87"S 46°45'13.87"W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000288— 1 juvenile and LB_ ANL000289, 7 adults, in plinthosol in a logged forest, (3°41'10.36"S 46°45'56.36"W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000290— 12 adults and LB_ ANL000291, 23 juveniles, in plinthosol in a logged forest, (3°41'20.33"S 46°46'1.16"W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000292— 1 adult and LB_ ANL000293, 1 juvenile, in plinthosol in a pasture, (3°40'25.49"S 46°47'20.47"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000294— 4 adults and LB_ ANL000295, 1 juvenile, in plinthosol in a pasture, (3°37'59.90"S 46°48'57.10"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000296, 4 adults, in plinthosol in an intermediate secondary forest, (3°41'56.61"S 46°45'57.81"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000297, 7 adults, in plinthosol in an intermediate secondary forest, (3°41'57.30"S 46°46'4.80"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000298, 9 adults, in plinthosol in an intermediate secondary forest, (3°42'7.90"S 46°45'42.63"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000299— 10 juveniles and LB_ ANL000300, 12 adults, in plinthosol in an old-growth forest,(3°41'26.11"S 46°46'14.11"W), April2016, L.M.Hernández-García, G.X.Rousseau, colls . LB_ ANL 000301— 27 juveniles, in plinthosol in an old-growth forest, (3°41'18.50"S 46°45'56.20"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000302— 2 juveniles and LB_ ANL000303, 11 adults, in plinthosol in an old-growth forest, (3°41'27.72"S 46°46'2.26"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000304— 29 adults and LB_ ANL000305, 61 juveniles, in plinthosol in an old-growth forest, (3°41'21.00"S 46°46'24.20"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000306— 5 adults and LB_ ANL000307, 5 juveniles, in plinthosol in an intermediate secondary forest, (3°42'18.22"S 46°45'47.40"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000308— 12 juveniles and LB_ ANL000309, 10 adults, in plinthosol in an intermediate secondary forest, (3°42'7.3"S 46°45'36.1"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . ; Itinga do Maranhão: LB_ ANL000310, 7 adults, in latosol in a pasture, (4°2'20.00"S 46°56'57.01"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000311, 1 adult, in latosol in an old-growth forest, (4°2'28.80"S 46°54'30.80"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000312, 2 adults, in latosol in an old-growth forest, (4°2'3.64"S 46°5'3.80"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000313, 3 adults, in latosol in an old-growth forest, (4°2'12.66"S 46°2'12.20"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000314— 9 juveniles and LB_ ANL000315, 1 adult, in argisol in an old-growth forest, (4°5'57.60"S 46°5'57.60"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000316— 5 adults and LB_ ANL000317, 3 juveniles, in argisol in an old-growth forest, (4°6'4.00"S 46°54'58.00"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000318, 2 juveniles, in argisol in a pasture, (4°4'20.98"S 46°52'59.78"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000319, 1 adult, in argisol in an intermediate secondary forest, (4°4'50.30"S 46°52'51.30"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000320, 2 juveniles, in argisol in an intermediate secondary forest, (4°4'31.00"S 46°4'31.00"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000321— 3 juveniles and LB_ ANL000322, 7 adults, in plinthosol in a logged forest, (3°55'23.60"S 46°45'57.04"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000323— 3 adults and LB_ ANL000324, 4 juveniles, in plinthosol in a logged forest, (3°58'59.00"S 46°47'56.80"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000325— 5 adults and LB_ ANL000326, 1 juvenile, in plinthosol in a logged forest, (3°55'3.55"S 46°45'49.55"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls. (Figure 2)</p></div>	https://treatment.plazi.org/id/90278E13FFE8FF9057C5FAE34D64936D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sousa, Sandriel Costa;Hernández-García, Luis Manuel;Beckman, Ruana Aretha Farias Santiago;Rousseau, Guillaume Xavier	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE8FF9757C5FE0B4D2794BC.text	90278E13FFE8FF9757C5FE0B4D2794BC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyperiodrilus africanus Beddard 1891	<div><p>Hyperiodrilus africanus Beddard, 1891</p><p>Distribution type: Exotic (Brown &amp; Fragoso, 2007)</p><p>Distribution in Brazil: Maranhão: São Luís (Sousa et al. 2020b). Pernambuco: Ponta de Pedras (Righi, 1972). (Figure 2)</p></div>	https://treatment.plazi.org/id/90278E13FFE8FF9757C5FE0B4D2794BC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sousa, Sandriel Costa;Hernández-García, Luis Manuel;Beckman, Ruana Aretha Farias Santiago;Rousseau, Guillaume Xavier	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFEFFF9157C5FA3F4F4D97C5.text	90278E13FFEFFF9157C5FA3F4F4D97C5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Holoscolex alatus Hernandez-Garcia, Burgos, Rousseau & James 2018	<div><p>Holoscolex alatus Hernández-García, Burgos, Rousseau &amp; James 2018</p><p>Distribution type: Native.</p><p>Distribution in Brazil: Maranhão: Centro Novo do Maranhão, Gurupi Biological Reserve, Itinga (Hernandez et al. 2018a, This study).</p><p>MARANHÃO: Centro Novo do Maranhão: LB_ ANL000327, 3 adults, in plinthosol in a logged forest, (3°40’50.95”S 46°46’13.06”W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000328, 1 adult, in plinthosol in a logged forest, (3°41’20.35”S 46°46’1.16”W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000329— 1 adult and LB_ ANL000330, 10 juveniles, in plinthosol in a logged forest, (3°40’25.50”S 46°47’20.46”W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000331— 15 adults and LB_ ANL000332, 7 juveniles, in plinthosol in a logged forest, (3°40’22.00”S 46°47’46.00”W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000333— 13 adults and LB_ ANL000334, 12 juveniles, in plinthosol in a logged forest, (3°40’17.20”S 46°47’54.90”W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000335, 4 adults, in plinthosol in a logged forest, (3°40’8.20”S 46°47’54.20”W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000336— 4 juveniles and LB_ ANL000337, 7 adults, in plinthosol in a logged forest, (3°40’7.20”S 46°47’42.50”W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000338, 2 juveniles, in plinthosol in a logged forest, (3°36’59.90”S 46°48’57.50”W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000339, 2 juveniles, in plinthosol in an intermediate secondary forest, (3°41’57.30”S 46°46’04.80”W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000340, 2 adults, in plinthosol in an intermediate secondary forest, (3°42’18.20”S 46°45’47.40”W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls.; Itinga do Maranhão: LB_ ANL000341, 6 juveniles, in argisol in a pasture, (4°5'43.00"S 46°52'21.10"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000342, 3 juveniles, in argisol in an intermediate secondary forest, (4°4'32.00"S 46°52'49.00"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls. (Figure 2)</p></div>	https://treatment.plazi.org/id/90278E13FFEFFF9157C5FA3F4F4D97C5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sousa, Sandriel Costa;Hernández-García, Luis Manuel;Beckman, Ruana Aretha Farias Santiago;Rousseau, Guillaume Xavier	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFEEFF9157C5FD9349279505.text	90278E13FFEEFF9157C5FD9349279505.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glossodrilus antunesi (Righi 1971)	<div><p>Glossodrilus antunesi (Righi, 1971)</p><p>Distribution type: Native.</p><p>Distribution in Brazil: Amapá: Serra do Navio (Righi, 1971); Maranhão: Caxias (Righi, 1972); Roraima: Ilha de Maracá, Estrada Boa Vista—Bonfim, km 100 (Hamoui &amp; Donatelli, 1983; Righi,1990). (Figure 2)</p></div>	https://treatment.plazi.org/id/90278E13FFEEFF9157C5FD9349279505	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sousa, Sandriel Costa;Hernández-García, Luis Manuel;Beckman, Ruana Aretha Farias Santiago;Rousseau, Guillaume Xavier	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFEEFF9157C5FC534D8491ED.text	90278E13FFEEFF9157C5FC534D8491ED.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Righiodrilus gurupi Hernandez-Garcia, Rousseau & James 2017	<div><p>Righiodrilus gurupi Hernández-García, Rousseau &amp; James 2017</p><p>Distribution type: Native.</p><p>Distribution in Brazil: Maranhão: Centro Novo do Maranhão, Gurupi Biological Reserve (Dos Santos et al. 2017), Alcântara (This study).</p><p>MARANHÃO: Centro Novo do Maranhão: LB_ ANL 000351— 3 juveniles and LB_ ANL000352, 4 adults, in plinthosol in an old-growth forest, (3°41’11.71”S 46°45’34.91”W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000353— 2 adults and LB_ ANL000354, 2 juveniles, in plinthosol in an old-growth forest, (3°41’16.35”S 46°45’34.74”W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000355, 1 juvenile, in plinthosol in a logged forest, (3°40’59.74”S 46°46’1.49”W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000356, 1 adult, in plinthosol in an old-growth forest, (3°41’24.87”S 46°45’13.00”W), March 2015, L.M.Hernández-García,G.X. Rousseau, colls . LB_ ANL000357, 3adults,in plinthosol in a logged forest, (3°41’10.76”S 46°45’56.54”W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000358, 1 adult, in plinthosol in a logged forest, (3°41’20.33”S 46°46’1.16”W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000362, 2 adults, in plinthosol in an intermediate secondary forest, (3°41’57.30”S 46°46’4.80”W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000363, 1 adult, in plinthosol in an old-growth forest, (3°41’26.10”S 46°46’14.20”W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000364, 8 adults, in plinthosol in an old-growth forest, (3°41’18.50”S 46°45’57.20”W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000365, 7 adults, in plinthosol in an old-growth forest, (3°41’27.7”S 46°46’2.7”W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000370, 6 adults, in plinthosol in a logged forest, (3°55’3.55”S 46°45’49.97”W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls.; Alcântara: LB_ ANL 000359— 4 adults and LB_ ANL000360, 2 juveniles, in latosol in a young secondary forest, (2°22'12.99"S 44°26'40.30"W), May 2015, J.E.B. Guerrero, L.M. Hernández-García, colls . LB_ ANL000361, 5 adults, in latosol in an intermediate secondary forest, (2°21'56.30"S 44°27'10.50"W), May 2015, J.E.B. Guerrero, L.M. Hernández-García, colls . LB_ ANL 000366— 2 juveniles and LB_ ANL000367, 1 adult, in latosol in a young secondary forest, Alcântara (2°21'51.10"S 44°27'16.40"W), June 2016, J.E.B. Guerrero, L.M. Hernández-García, colls . LB_ ANL000368, 1 adult, in latosol in a logged forest, (2°21'57.00"S 44°27'15.98"W), June 2016, J.E.B. Guerrero, L.M. Hernández-García, colls . LB_ ANL000369, 1 juvenile, in latosol in a young secondary forest, (2°22'3.10"S 44°26'42.10"W), June 2016, J.E.B. Guerrero, L.M. Hernández-García, colls . LB_ ANL000371, 1 juvenile, in latosol in a young secondary riparian forest, (2°20'55.50"S 44°27'56.40"W), June 2013, A.R. Piedade. (Figure 2)</p></div>	https://treatment.plazi.org/id/90278E13FFEEFF9157C5FC534D8491ED	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sousa, Sandriel Costa;Hernández-García, Luis Manuel;Beckman, Ruana Aretha Farias Santiago;Rousseau, Guillaume Xavier	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFEDFF9257C5FD0E490995F9.text	90278E13FFEDFF9257C5FD0E490995F9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arraia nelmae Hernandez-Garcia & James 2018	<div><p>Arraia nelmae Hernández-García &amp; James 2018</p><p>Distribution type: Native.</p><p>Distribution in Brazil: Maranhão: Rosário (Hernández-García et al. 2018b, This study)</p><p>MARANHÃO: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.153717&amp;materialsCitation.latitude=-2.8539915" title="Search Plazi for locations around (long -44.153717/lat -2.8539915)">Rosário</a>: LB_ANL000252, 14 juveniles, in argisol in an advanced AFS, (2°51'4.04"S 44°8'51.42"W), June 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls. LB_ ANL000253, 1 juvenile, in argisol in a young AFS, (2°51'14.37"S 44°9'13.37"W), June 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls. (Figure 2)</p></div>	https://treatment.plazi.org/id/90278E13FFEDFF9257C5FD0E490995F9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sousa, Sandriel Costa;Hernández-García, Luis Manuel;Beckman, Ruana Aretha Farias Santiago;Rousseau, Guillaume Xavier	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFEDFF9257C5FBCA4F989181.text	90278E13FFEDFF9257C5FBCA4F989181.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Brasilisia punki Hernandez-Garcia & James 2018	<div><p>Brasilisia punki Hernández-García &amp; James 2018</p><p>Distribution type: Native.</p><p>Distribution in Brazil: Maranhão:Alcântara (Hernández-García et al. 2018b, This study), São José de Ribamar (This study).</p><p>MARANHÃO: Alcântara: LB_ ANL000254, 5adults,in gleysol in a logged forest,(2°25'3.40"S 44°25'20.00"W), June 2010, P. R.S. Silva, G.X. Rousseau, colls . LB_ ANL 000255— 3 juveniles andLB_ ANL000256, 9 adults, in gleysol in a young secondary forest, (2°25'25.50"S 44°25'31.60"W), June 2010, P. R.S. Silva, G.X. Rousseau, colls . LB_ ANL000257, 7 adults, in gleysol in a grassland, (2°25'26.9"S 44°25'30.7"W), June 2010, P. R.S. Silva, G.X. Rousseau, colls . LB_ ANL000258, 18 juveniles, in gleysol in a logged forest, (2°24'59.15"S 44°25'30.36"W), Jule 2011, P. R.S. Silva, K.H.M. Muchavisoy, colls . LB_ ANL 000259— 2 juveniles an LB_ ANL000260, 3 adults, in gleysol in an intermediate secondary forest, (2°24'58.18"S 44°25'28.40"W), Jule 2011, P. R.S. Silva, K.H.M. Muchavisoy, colls . LB_ ANL 000261— 12 adults and LB_ ANL000262, 2 juveniles, in gleysol in an intermediate secondary forest, (2°25'33.19"S 44°25'26.59"W), Jule 2011, P. R.S. Silva, K.H.M. Muchavisoy, colls . LB_ ANL000264, 1 juvenile, in latosol in a slash-burn, (2°21'26.1"S 44°28'41.1"W), May 2015, J.E.B. Guerrero, L.M. Hernández-García, colls . LB_ ANL000265, 3 juveniles, in latosol in a logged forest, (2°21'58.60"S 44°27'16.50"W), May 2015, J.E.B. Guerrero, L.M. Hernández-García, colls . LB_ ANL000266, 16 juveniles, in latosol in a young secondary forest, (2°22'13.00"S 44°26'40.3"W), May 2015, J.E.B. Guerrero, L.M. Hernández-García, colls . LB_ ANL000267, 1 juvenile, in latosol in an intermediate secondary forest, (2°21'56.30"S 44°27'10.51"W), May 2015, J.E.B. Guerrero, L.M. Hernández-García, colls . LB_ ANL000268, 4 juveniles, in latosol in a young secondary forest, (2°21'51.10"S 44°27'16.40"W), June 2016, J.E.B. Guerrero, L.M. Hernández-García, colls . LB_ ANL 000269— 3 adults and LB_ ANL000270, 13 juveniles, in latosol in a young secondary forest, (2°22'3.1"S 44°26'42.1"W), June 2016, J.E.B. Guerrero, L.M. Hernández-García, colls.; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.134476&amp;materialsCitation.latitude=-2.6497889" title="Search Plazi for locations around (long -44.134476/lat -2.6497889)">São José de Ribamar</a>: LB_ ANL000263, 1 juvenile, in argisol in an old-growth forest, (2°38'59.24"S 44°8'4.1"W), January 2014, J.A.M. Gutiérrez. (Figure 2)</p></div>	https://treatment.plazi.org/id/90278E13FFEDFF9257C5FBCA4F989181	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sousa, Sandriel Costa;Hernández-García, Luis Manuel;Beckman, Ruana Aretha Farias Santiago;Rousseau, Guillaume Xavier	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFEDFF9257C5FF624F86940D.text	90278E13FFEDFF9257C5FF624F86940D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Righiodrilus tocantinensis subsp. tocantinensis (Righi 1972)	<div><p>Righiodrilus tocantinensis tocantinensis (Righi, 1972)</p><p>Distribution type: Native.</p><p>Distribution in Brazil: Maranhão: Caxias (This study), São Luís (Sousa et al. 2020b). Pará: Cametá, Mocajuba (Righi, 1972), Itupiranga, Marabá, Tucuruí (Righi, 1975). Paraíba: João Pessoa (Righi, 1975). Mato Grosso: Pontes e Lacerda (Righi, 1975).</p><p>MARANHÃO: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.46508&amp;materialsCitation.latitude=-4.912197" title="Search Plazi for locations around (long -43.46508/lat -4.912197)">Caxias</a>: LB_ ANL000372, 15 juveniles, in neosol in a logged riparian forest, (4°53’33.60”S 43°25’36.37”W), February 2019, A.M. Sousa, L.M. Hernández-García, colls . LB_ ANL000373, 2 juveniles, in neosol in a logged riparian forest, (4°53’27.92”S 43°25’32.98”W), February 2019, A.M. Sousa, L.M. Hernández-García, colls . LB_ ANL000374, 8 juveniles, in neosol in an old-growth forest, (4°54’43.91”S 43°27’54.28”W), February 2019, A.M. Sousa, L.M. Hernández-García, colls. (Figure 2)</p></div>	https://treatment.plazi.org/id/90278E13FFEDFF9257C5FF624F86940D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sousa, Sandriel Costa;Hernández-García, Luis Manuel;Beckman, Ruana Aretha Farias Santiago;Rousseau, Guillaume Xavier	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFECFF9357C5F8FA4D5991AD.text	90278E13FFECFF9357C5F8FA4D5991AD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Andiorrhinus (Turedrilus) barrosoi Hernandez-Garcia & James 2018	<div><p>Andiorrhinus (Turedrilus) barrosoi Hernández-García &amp; James 2018</p><p>Distribution type: Native.</p><p>Distribution in Brazil: Maranhão:Alcântara, Centro Novo do Maranhão, Itinga, Rosário, São Luís (Hernández-García et al. 2018c)</p></div>	https://treatment.plazi.org/id/90278E13FFECFF9357C5F8FA4D5991AD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sousa, Sandriel Costa;Hernández-García, Luis Manuel;Beckman, Ruana Aretha Farias Santiago;Rousseau, Guillaume Xavier	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFECFF9357C5FD9E4F8B90A9.text	90278E13FFECFF9357C5FD9E4F8B90A9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anteus (Turedrilus) miricuri Hernandez-Garcia & James 2018	<div><p>Andiorhinus (Turedrilus) miricuri Hernández-García &amp; James 2018</p><p>Distribution type: Native.</p><p>Distribution in Brazil: Maranhão: Centro Novo do Maranhão (This study), Itinga do Maranhão (Hernández-García et al. 2018c, This study).</p><p>MARANHÃO: Centro Novo do Maranhão: LB_ ANL000230, 9 juveniles, in latosol in a logged forest, (3°59’52.12”S 46°50’8.77”W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000232, 2 juveniles, in plinthosol in an old-growth forest, (3°41’27.19”S 46°45’6.91”W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000233, 3 juveniles, in plinthosol in an intermediate secondary forest, (3°41’56.70”S 46°45’57.80”W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000234, 4 juveniles, in plinthosol in an intermediate secondary forest, (3°41’57.30”S 46°46’4.80”W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000235, 2 juveniles, in plinthosol in an old-growth forest, (3°41’26.10”S 46°46’14.20”W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000236, 2 juveniles, in plinthosol in an old-growth forest, (3°41’27.70”S 46°46’2.70”W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls.; Itinga do Maranhão: LB_ ANL000231, 6 juveniles, in latosol in an intermediate secondary forest, (4°2'3.80"S 46°54'19.80"W), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-46.79911&amp;materialsCitation.latitude=-3.9997222" title="Search Plazi for locations around (long -46.79911/lat -3.9997222)">Jule</a>, 2011, P. R. S. Silva, K.H.M. Muchavisoy, colls . LB_ ANL000238, 3 juveniles, in latosol in a pasture, (4°2'35.30"S 46°55'56.80"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000239, 6 juveniles, in latosol in an intermediate secondary forest, (4°1'53.70"S 46°52'58.30"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000240, 2 juveniles, in latosol in a pasture, (4°2'17.99"S 46°54'45.50"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000241, 1 juvenile, in latosol in an old-growth forest, (4°2'28.80"S 46°54'30.80"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000242, 8 juveniles, in latosol in an old-growth forest, (4°2'3.80"S 46°54'19.80"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000243, 3 juveniles, in argisol in an old-growth forest, (4°6'4.00"S 46°6'4.00"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000244, 1 juvenile, in argisol in an intermediate secondary forest, (4°4'50.4"S 46°4'50.3"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000245, 6 juveniles, in latosol in a logged forest, (3°59'57.69"S 46°49'43.80"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000246, 8 juveniles, in plinthosol in a logged forest, (3°55'23.60"S 46°45'57.04"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000247, 1 juvenile, in plinthosol in a logged forest, (3°58'50.00"S 46°47'9.99"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000248, 4 juveniles, in plinthosol in a logged forest, (3°59'59.00"S 46°47'56.80"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls. (Figure 2)</p></div>	https://treatment.plazi.org/id/90278E13FFECFF9357C5FD9E4F8B90A9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sousa, Sandriel Costa;Hernández-García, Luis Manuel;Beckman, Ruana Aretha Farias Santiago;Rousseau, Guillaume Xavier	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFECFF9357C5FF2A4D27979D.text	90278E13FFECFF9357C5FF2A4D27979D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liodrilus mendesi Righi 1994	<div><p>Liodrilus mendesi Righi, 1994</p><p>Distribution type: Native.</p><p>Distribution in Brazil: Maranhão: Caxias, São Luís (Sousa, 2020b). Paraíba: João Pessoa (Righi, 1994). (Figure 2)</p></div>	https://treatment.plazi.org/id/90278E13FFECFF9357C5FF2A4D27979D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sousa, Sandriel Costa;Hernández-García, Luis Manuel;Beckman, Ruana Aretha Farias Santiago;Rousseau, Guillaume Xavier	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE3FF9C57C5FE664DB89429.text	90278E13FFE3FF9C57C5FE664DB89429.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atatina arimikuriaka Hernandez-Garcia & James 2024	<div><p>Atatina arimikuriaka Hernández-García &amp; James 2024</p><p>Distribution type: Native.</p><p>Distribution in Brazil: Maranhão: Centro Novo do Maranhão, Gurupi Biological Reserve (Hernández-García et al. 2024). (Figure 2)</p></div>	https://treatment.plazi.org/id/90278E13FFE3FF9C57C5FE664DB89429	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sousa, Sandriel Costa;Hernández-García, Luis Manuel;Beckman, Ruana Aretha Farias Santiago;Rousseau, Guillaume Xavier	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE3FF9C57C5FD7A4DB8952D.text	90278E13FFE3FF9C57C5FD7A4DB8952D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atatina arimikuritea Hernandez-Garcia 2024	<div><p>Atatina arimikuritea Hernández-García 2024</p><p>Distribution type: Native.</p><p>Distribution in Brazil: Maranhão: Centro Novo do Maranhão, Gurupi Biological Reserve (Hernández-García et al. 2024). (Figure 2)</p></div>	https://treatment.plazi.org/id/90278E13FFE3FF9C57C5FD7A4DB8952D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sousa, Sandriel Costa;Hernández-García, Luis Manuel;Beckman, Ruana Aretha Farias Santiago;Rousseau, Guillaume Xavier	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE3FF9C57C5FF624FD39755.text	90278E13FFE3FF9C57C5FF624FD39755.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atatina Righi 1971	<div><p>Genus Atatina Righi, 1971</p><p>To date only two species of Atatina were recorded to Maranhão: Atatina arimikuriaka and Atatina amirikuritea (Hernández-García et al. 2024) . The accession codes LB000001–LB000010 were recodified as presented in the current model (LB_ANL) of the Maranhão Invertebrate Collection (CISMA).</p></div>	https://treatment.plazi.org/id/90278E13FFE3FF9C57C5FF624FD39755	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sousa, Sandriel Costa;Hernández-García, Luis Manuel;Beckman, Ruana Aretha Farias Santiago;Rousseau, Guillaume Xavier	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE3FF9C57C5FC264E2D9389.text	90278E13FFE3FF9C57C5FC264E2D9389.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pontoscolex (Pontoscolex) awa Sousa & Hernandez-Garcia 2020	<div><p>Pontoscolex (Pontoscolex) awa Sousa &amp; Hernández-García 2020</p><p>Distribution type: Native.</p><p>Distribution in Brazil: Maranhão: Bom Jardim* (Sousa et al. 2020b), Itinga do Maranhão (This study).</p><p>MARANHÃO: Itinga do Maranhão: LB_ ANL000343, 2 adults, in latosol in an old-growth forest, (4°02’34.50”S 46°54’36.10”W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000344, 1 adult, in argisol in an old-growth forest, (4°06’04.00”S 46°54’59.00”W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000345 — 3 adults and LB_ ANL000346, 4 juveniles, in argisol in a pasture, (4°4’20.79”S 46°52’59.80”W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000347— 8 adults and LB_ ANL000348, 6 juveniles, in argisol in an intermediate secondary forest, (4°4’50.30”S 46°52’37.40”W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000349— 2 juveniles and LB_ ANL000350, 1 adult, in argisol in an intermediate secondary forest, (4°4’32.00”S 46°52’49.00”W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls. (Figure 2)</p></div>	https://treatment.plazi.org/id/90278E13FFE3FF9C57C5FC264E2D9389	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sousa, Sandriel Costa;Hernández-García, Luis Manuel;Beckman, Ruana Aretha Farias Santiago;Rousseau, Guillaume Xavier	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE3FF9857C5FA1A49439165.text	90278E13FFE3FF9857C5FA1A49439165.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pontoscolex (Pontoscolex) corethrurus (Muller 1857)	<div><p>Pontoscolex (Pontoscolex) corethrurus (Müller, 1857)</p><p>Lumbricus corethrurus Muller, 1857: 113 .</p><p>Pontoscolex arenicola Schmarda, 1861: 11 .</p><p>Eurydame insignis Kinberg, 1867: 101 .</p><p>Urochaeta hystrix Perrier, 1872; 142.</p><p>Urochaeta corethrura Rosa, 1889: 126 .</p><p>Titanus insignis Vaillant, 1889: 95 .</p><p>Urochaeta australiensis Beddard, 1891: 278 .</p><p>Pontoscolex corethrurus Cognetti de Martiis, 1904: 5, 1906: 170; Michaelsen, 1918: 234; Zicsi &amp; Csuzdi, 1987: 274, 1988: 217; Zicsi, 1995: 602, 2002: 120.</p><p>Pontoscolex hawaiensis Beddard, 1895: 660 .</p><p>Pontoscolex (Pontoscolex) corethrurus Righi, 1984 a: 163 .</p><p>Distribution type: Peregrine.</p><p>Distribution in Brazil: Present in 17 Brazilian states (Brown &amp; Fragoso, 2007; This study)</p><p>MARANHÃO: Alcântara: LB_ ANL 000039— 8 adults and LB_ ANL000040, 4 juveniles, in gleysol in an intermediate secondary forest, (2°25'33.19"S 44°25'26.59"W), July 2011, P. R.S. Silva, K.H.M. Muchavisoy, colls . LB_ ANL 000077— 8 adults and LB_ ANL000078, 11 juveniles, in latosol in an advanced secondary riparian forest, (2°21'32.60"S 44°27'43.30"W), November 2012, S.P. Triana, M.L.C. Zelarayán, colls . LB_ ANL 000079— 10 adults and LB_ ANL000080, 72 juveniles,in latosol in an advanced secondary riparian forest,(2°21'32.60"S 44°27'43.30"W), June 2013, A. R. Piedade. LB_ ANL000081, 8 juveniles, in latosol in an intermediate secondary riparian forest, (2°21'5.50"S 44°27'51.40"W), November 2012, S.P. Triana, M.L.C. Zelarayán, colls . LB_ ANL 000082— 4 adults and LB_ ANL000083, 3 juveniles, in latosol in an intermediate secondary riparian forest, (2°21'5.50"S 44°27'51.40"W), June 2013, A. R. Piedade. LB_ ANL000084, 3 juveniles, in latosol in a young secondary riparian forest, (2°21'2.10"S 44°27'56.50"W), November 2012, S.P. Triana, M.L.C. Zelarayán, colls . LB_ ANL000085, 35 juveniles, in latosol in an advanced secondary riparian forest, (2°21'37.80"S 44°27'16.50"W), November 2012, S.P. Triana, M.L.C. Zelarayán, colls . LB_ ANL 000086— 9 adults and LB_ ANL000087, 46 juveniles, in latosol in an advanced secondary riparian forest, (2°21'37.80"S 44°27'16.50"W), June 2013, A. R. Piedade. LB_ ANL000088, 22 juveniles, in latosol in an intermediate secondary riparian forest, (2°21'37.10"S 44°27'36.10"W), November 2012, S.P. Triana, M.L.C. Zelarayán, colls . LB_ ANL 000089— 7 adults and LB_ ANL000090, 43 juveniles, in latosol in an intermediate secondary riparian forest, (2°21'37.10"S 44°27'36.10"W), June 2013,A. R. Piedade. LB_ ANL 000091— 1 adult and LB_ ANL000092, 4 juveniles, in latosol in a young secondary riparian forest, (2°21'35.70"S 44°27'26.70"W), June 2013, A. R. Piedade. LB_ ANL000093, 35 juveniles, in latosol in an advanced secondary riparian forest, (2°21'39.20"S 44°26'30.10"W), November 2012, S.P. Triana, M.L.C. Zelarayán, colls . LB_ ANL 000094— 2 adults and LB_ ANL000095, 45 juveniles, in latosol in an advanced secondary riparian forest, (2°21'39.20"S 44°26'30.10"W), June 2013, A. R. Piedade. LB_ ANL 000096— 6 adults and LB_ ANL000097, 31 juveniles, in latosol in an intermediate secondary riparian forest, (2°21'43.90"S 44°26'36.00"W), June 2013, A. R. Piedade. LB_ ANL 000098— 11 juveniles and LB_ ANL000099, 2 juveniles, in latosol in a young secondary riparian forest, (2°21'42.00"S 44°26'31.90"W), June 2013, A. R. Piedade. LB_ ANL000100, 1 adult, in latosol in a manihot crop, (2°21'37.90"S 44°26'54.60"W), June 2013, A. R. Piedade. LB_ ANL000101, 65 juveniles, in latosol in a logged advanced secondary riparian forest, (2°20'54.00"S 44°28'46.10"W), November 2012, S.P. Triana, M.L.C. Zelarayán, colls . LB_ ANL 000102— 10 adults and LB_ ANL000103, 67 juveniles, in latosol in a logged advanced secondary riparian forest, (2°20'54.00"S 44°28'46.10"W), June 2013, A. R. Piedade. LB_ ANL 000104— 2 juveniles and LB_ ANL000105, 7 juveniles, in latosol in an intermediate secondary riparian forest, (2°20'47.20"S 44°28'29.90"W), June 2013, A. R. Piedade. LB_ ANL 000106— 4 juveniles and LB_ ANL000107, 4 juveniles, in latosol in a young secondary riparian forest, (2°20'52.60"S 44°28'39.60"W), June 2013, A. R. Piedade. LB_ ANL000108, 44 juveniles, in latosol in a logged advanced secondary riparian forest, (2°20'53.4"S 44°28'50.7"W), November 2012, S.P. Triana, M.L.C. Zelarayán, colls . LB_ ANL 000109— 3 adults and LB_ ANL000110, 42 juveniles, in latosol in a logged advanced secondary riparian forest, (2°20'53.4"S 44°28'50.7"W), June 2013, A. R. Piedade. LB_ ANL000111, 62 juveniles, in latosol in an intermediate secondary riparian forest, (2°20'55.50"S 44°28'54.30"W), November 2012, S.P. Triana, M.L.C. Zelarayán, colls . LB_ ANL 000112— 2 adults and LB_ ANL000113, 44 juveniles, in latosol in an intermediate secondary riparian forest, (2°20'55.50"S 44°28'54.30"W), June 2013, A. R. Piedade. LB_ ANL 000114— 1 juvenile and LB_ ANL000115, 1 juvenile, in latosol in a young secondary riparian forest, (2°20'55.70"S 44°28'56.40"W), June 2013, A. R. Piedade. LB_ ANL000116, 3 juveniles, in latosol in a manihot crop, (2°20'52.50"S 44°28'50.00"W), November 2012, S.P. Triana, M.L.C. Zelarayán, colls . LB_ ANL 000117— 1 adult and LB_ ANL000118, 46 juveniles, in latosol in a logged advanced secondary riparian forest, (2°20'56.0"S 44°29'0.96"W), November 2012, S.P. Triana, M.L.C. Zelarayán, colls . LB_ ANL 000119— 1 adult and LB_ ANL000120, 56 juveniles, in latosol in a logged advanced secondary riparian forest, (2°20'56.0"S 44°29'0.96"W), June 2013, A. R. Piedade. LB_ ANL000121, 25 juveniles, in latosol in an intermediate secondary riparian forest, (2°20'52.71"S 44°29'1.33"W), November 2012, S.P. Triana, M.L.C. Zelarayán, colls . LB_ ANL 000122— 1 adult and LB_ ANL000123, 15 juveniles, in latosol in an intermediate secondary riparian forest, (2°20'52.71"S 44°29'1.33"W), June 2013, A. R. Piedade. LB_ ANL000124, 4 juveniles, in latosol in a young secondary riparian forest, (2°20'38.64"S 44°29'10.6"W), November 2012, S.P. Triana, M.L.C. Zelarayán, colls . LB_ ANL 000125— 1 adult and LB_ ANL000126, 39 juveniles, in latosol in a young secondary riparian forest, (2°20'38.64"S 44°29'10.6"W), June 2013, A. R. Piedade. LB_ ANL000127, 2 adults, in latosol in a manihot crop <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.485596&amp;materialsCitation.latitude=-2.3433387" title="Search Plazi for locations around (long -44.485596/lat -2.3433387)">Crop</a>, (2°20'36.02"S 44°29'8.14"W), June 2013, A. R. Piedade. LB_ ANL 000179— 16 juveniles and LB_ ANL000180, 1 adult, in latosol in a fallow, (2°21'27.90"S 44°28'40.70"W), May 2015, J.E.B. Guerrero, L.M. Hernández-García, colls . LB_ ANL000181, 2 juveniles, in latosol in a young secondary forest, (2°21'52.60"S 44°27'14.10"W), May 2015, J.E.B. Guerrero, L.M. Hernández-García, colls . LB_ ANL000182, 5 juveniles, in latosol in an intermediate secondary forest, (2°21'56.30"S 44°27'10.51"W), May 2015, J.E.B. Guerrero, L.M. Hernández-García, colls . LB_ ANL 000183— 13 adults and LB_ ANL000184, 12 juveniles, in latosol in a young secondary forest, (2°21'29.90"S 44°28'39.90"W), May 2015, J.E.B. Guerrero, L.M. Hernández-García, colls . LB_ ANL 000185— 22 juveniles and LB_ ANL000186, 16 adults, in latosol in an intermediate secondary forest, (2°21'30.00"S 44°28'32.23"W), May 2015, J.E.B. Guerrero, L.M. Hernández-García, colls . LB_ ANL000187, 5 juveniles, in latosol in a young secondary forest, (2°22'26.70"S 44°27'43.00"W), May 2015, J.E.B. Guerrero, L. M. Hernández-García, colls . LB_ ANL 000188— 3 adults and LB_ ANL000189, 5 juveniles, in latosol in a logged forest, (2°24'2.70"S 44°27'40.80"W), May 2015, J.E.B. Guerrero, L.M. Hernández-García, colls . LB_ ANL 000190— 3 adults and LB_ ANL000191, 7 juveniles, in latosol in an advanced secondary forest, (2°19'17.40"S 44°28'28.20"W), May 2015, J.E.B. Guerrero, L.M. Hernández-García, colls . LB_ ANL 000206— 77 juveniles and LB_ ANL000207, 9 adults, in latosol in a young secondary forest, (2°21'21.90"S 44°28'44.60"W), June 2016, J.E.B. Guerrero, L.M. Hernández-García, colls . LB_ ANL000208, 7 juveniles, in latosol in a young secondary forest, (2°21'51.10"S 44°27'16.40"W), June 2016, J.E.B. Guerrero, L.M. Hernández-García, colls . LB_ ANL000209, 1 adult, in latosol in a logged forest, (2°21'56.99"S 44°27'15.98"W), June 2016, J.E.B. Guerrero, L.M. Hernández-García, colls . LB_ ANL 000210— 21 juveniles and LB_ ANL000211, 6 adults, in latosol in an advanced secondary forest, (2°21'2.90"S 44°25'48.60"W), June 2016, J.E.B. Guerrero, L.M. Hernández-García, colls . LB_ ANL 000212— 6 juveniles and LB_ ANL000213, 7 adults, in latosol in an advanced secondary forest, (2°20'57.70"S 44°25'42.00"W), June 2016, J.E.B. Guerrero, L.M. Hernández-García, colls . LB_ ANL000214, 6 juveniles, in latosol in a logged forest, (2°24'5.80"S 44°27'44.40"W), June 2016, J.E.B. Guerrero, L.M. Hernández-García, colls . LB_ ANL000215, 1 juvenile, in latosol in a young secondary forest, (2°22'3.10"S 44°26'42.10"W), June 2016, J.E.B. Guerrero, L.M. Hernández-García, colls.; Caxias: LB_ ANL000011, 7 juveniles, in neosol in an old-growth riparian forest, (4°53'27.17"S 43°25'13.87"W), February 2019, A.M. Sousa, L.M. Hernández-Garcia, colls . LB_ ANL 000012— 137 juveniles and LB_ ANL000013, 3 adults, in neosol in an old-growth riparian forest, (4°53'26.45"S 43°25'20.82"W), February 2019, A.M. Sousa, L.M. Hernández-Garcia, colls . LB_ ANL000014, 16 juveniles, in neosol in a logged riparian forest, (4°53'33.58"S 43°25'36.37"W), February 2019, A.M. Sousa, L.M. Hernández-Garcia, colls . LB_ ANL 000015— 45 juveniles and LB_ ANL000016, 2 adults, in neosol in a logged riparian forest, (4°53'27.92"S 43°25'32.98"W), February 2019, A.M. Sousa, L.M. Hernández-Garcia, colls . LB_ ANL 000017— 48 juveniles and LB_ ANL000018, 3 adults, in neosol in a logged riparian forest, (4°53'21.88"S 43°25'29.14"W), February 2019, A.M. Sousa, L.M. Hernández-Garcia, colls . LB_ ANL000019, 23 juveniles, in neosol in a logged riparian forest, (4°53'14.71"S 43°25'31.15"W), February 2019, A.M. Sousa, L.M. Hernández-Garcia, colls . LB_ ANL 000024— 28 juveniles and LB_ ANL000025, 8 adults, in neosol in an old-growth forest, (4°53'59.03"S 43°28'15.53"W), February 2019, A.M. Sousa, L.M. Hernández-Garcia, colls . LB_ ANL000028, 2 juveniles, in neosol in an old-growth forest, (4°54'24.84"S 43°27'58.54"W), February 2019, A.M. Sousa, L.M. Hernández-Garcia, colls . LB_ ANL 000029— 17 juveniles and LB_ ANL000030, 11 adults, in neosol in a logged forest, (4°53'51.28"S 43°28'15.06"W), February 2019, A.M. Sousa, L.M. Hernández-Garcia, colls . LB_ ANL000031, 28 juveniles, in plinthosol in a pasture, (4°49'16.72"S 43°20'28.81"W), February 2019, A.M. Sousa, L.M. Hernández-Garcia, colls . LB_ ANL 000032— 45 juveniles and LB_ ANL000033, 17 adults, in plinthosol in a logged riparian forest, (4°49'9.08"S 43°20'31.90"W), February 2019, A.M. Sousa, L.M. Hernández-Garcia, colls . LB_ ANL 000034— 40 juveniles and LB_ ANL000035, 10 adults, in plinthosol in a logged riparian forest, (4°49'5.42"S 43°20'33.6"W), February 2019, A.M. Sousa, L.M. Hernández-Garcia, colls . LB_ ANL 000037— 9 adults and LB_ ANL000038, 34 juveniles, in plinthosol in a logged riparian forest, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.342667&amp;materialsCitation.latitude=-4.818172" title="Search Plazi for locations around (long -43.342667/lat -4.818172)">Caxias</a> (4°49'5.42"S 43°20'33.60"W), February 2019, A.M. Sousa, L.M. Hernández-Garcia, colls. ; Centro Novo do Maranhão: LB_ ANL000154, 1 juvenile, in plinthosol in an old-growth forest, (3°41'11.71"S 46°45'34.91"W), June 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls . LB_ ANL000155, 4 adults, in plinthosol in an old-growth forest, (3°41'11.71"S 46°45'34.91"W), March 2015, L.M. Hernández-García. LB_ ANL000156, 8 juveniles, in plinthosol in an old-growth forest, (3°41'16.35"S 46°45'34.74"W), June 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls . LB_ ANL000157, 1 adult, in plinthosol in an old-growth forest, (3°41'14.03"S 46°45'34.35"W), June 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls . LB_ ANL 000158— 4 juveniles and LB_ ANL000159, 2 adults, in plinthosol in a logged forest, (3°40'50.06"S 46°46'12.95"W), June 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls . LB_ ANL 000160— 1 juvenile and LB_ ANL000161, 1 adult, in plinthosol in a logged forest, (3°40'50.06"S 46°46'12.95"W), March 2015, L.M. Hernández-García. LB_ ANL000162, 4 juveniles, in plinthosol in a logged forest, (3°40'56.10"S 46°46'8.78"W), June 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls . LB_ ANL000176, 2 adults, in plinthosol in an old-growth forest, (3°41'24.87"S 46°45'13.87"W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000177— 5 adults and LB_ ANL000178, 5 juveniles, in plinthosol in a logged forest, (3°41'10.40"S 46°45'56.76"W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000193, 1 adult, in plinthosol in a pasture, (3°40'25.30"S 46°47'20.50"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000194, 3 adults, in plinthosol in a pasture, (3°40'22.00"S 46°47'46.60"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000195— 6 adults and LB_ ANL000196, 8 juveniles, in plinthosol in a pasture, (3°40'8.20"S 46°47'54.20"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000197— 156 juveniles and LB_ ANL000198, 84 adults, in plinthosol in an intermediate secondary forest, (3°42'9.80"S 46°45'51.10"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000199— 26 adults and LB_ ANL000200, 46 juveniles, in plinthosol in an intermediate secondary forest, (3°42'7.90"S 46°45'42.60"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000201, 3 adults, in plinthosol in an old-growth forest, (3°41'26.10"S 46°46'14.20"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000202, 3 adults, in plinthosol in an old-growth forest, (3°41'27.74"S 46°46'2.68"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000203— 51 adults and LB_ ANL000204, 58 juveniles, in plinthosol in an intermediate secondary forest, (3°42'18.22"S 46°45'47.60"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000205, 2 adults, in plinthosol in an intermediate secondary forest, (3°42'7.3"S 46°45'37.1"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls.; Itinga do Maranhão: LB_ ANL000036, 8 adults, in latosol in a logged forest, Itinga do Maranhão (4°03'20.92"S 46°55'39.29"W), June 2011, P. R.S. Silva, K.H.M. Muchavisoy, colls . LB_ ANL 000216— 6 juveniles and LB_ ANL000217, 3 adults, in latosol in an old-growth forest, (4°2'3.80"S 46°54'20.90"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000218— 59 adults and LB_ ANL000219, 88 juveniles, in latosol in an old-growth forest, (4°2'12.20"S 46°54'23.70"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000220, 3 adults, in argisol in an old-growth forest, (4°6'4.00"S 46°54'59.00"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL000221, 8 adults, in argisol in a pasture, (4°4'20.00"S 46°4'20.78"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000222— 12 juveniles and LB_ ANL000223, 7 adults, in argisol in a pasture, (4°5'43.00"S 46°52'21.00"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000224— 309 juveniles and LB_ ANL000225, 106 adults, in argisol in an intermediate secondary forest, (4°5'7.40"S 46°52'9.40"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000226— 6 adults and LB_ ANL000227, 3 juveniles, in argisol in an intermediate secondary forest, (4°4'50.40"S 46°4'50.30"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls . LB_ ANL 000228— 9 adults and LB_ ANL000229, 5 juveniles, in argisol in an intermediate secondary forest, (4°4'32.00"S 46°5'31.00"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls.; Rosário: LB_ ANL 000128— 8 adults and LB_ ANL000129, 82 juveniles, in argisol in an advanced secondary forest, (2°52'21.93"S 44°7'38.72"W), May 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls . LB_ ANL 000130— 15 adults and LB_ ANL000131, 43 juveniles, in an argisol in a logged advanced secondary riparian forest, (2°49'55.61"S 44°7'56.01"W), May 2014, J.A.M.Gutiérrez, E.G. Cardozo, colls . LB_ ANL 000132— 15adults and LB_ ANL000133, 64 juveniles, in argisol in an advanced secondary riparian forest, (2°51'11.76"S 44°9'2.63"W), May 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls . LB_ ANL 000134— 7 adults and LB_ ANL000135, 35 juveniles, in argisol in an intermediate secondary forest, (2°51'40.42"S 44°8'0.4"W), May 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls . LB_ ANL 000136— 11 adults and LB_ ANL000137, 25 juveniles, in argisol in an intermediate secondary forest, (2°51'34.83"S 44°8'25.15"W), May 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls . LB_ ANL 000138— 10 adults and LB_ ANL000139, 54 juveniles, in argisol in an advanced secondary forest, (2°49'5.28"S 44°8'20.39"W), May 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls . LB_ ANL 000140— 8 adults and LB_ ANL000141, 21 juveniles, in argisol in a young secondary forest, (2°51'46.88"S 44°8'32.26"W), May 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls . LB_ ANL 000142— 11 adults and LB_ ANL000143, 33 juveniles, in argisol in an intermediate secondary forest, (2°51'10.96"S 44°9'30.14"W), June 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls . LB_ ANL 000144— 4 adults and LB_ ANL000145, 31 juveniles, in argisol in an intermediate secondary forest, (2°51'17.18"S 44°9'16.68"W), June 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls . LB_ ANL 000146— 3 adults and LB_ ANL000147, 12 juveniles, in argisol in a young secondary forest, (2°51'2.67"S 44°8'57.33"W), June 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls . LB_ ANL 000148— 42 juveniles and LB_ ANL000149, 4 adults, in argisol in an advanced AFS, (2°51'4.04"S 44°8'52.01"W), June 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls . LB_ ANL 000150— 13 juveniles and LB_ ANL000151, 6 adults, in argisol in a young AFS, (2°51'25.47"S 44°8'3.19"W), June 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls . LB_ ANL 000152— 58 juveniles and LB_ ANL000153, 11 adults, in argisol in an advanced AFS, (2°51'25.47"S 44°8'3.19"W), June 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls.; São Luís: LB_ ANL 000020— 8 adults and LB_ ANL000021, 25 juveniles, in latosol in an intermediate secondary forest, (2°35'15.00"S 44°12'43.00"W), April 2011, P. R.S. Silva, K.H.M. Muchavisoy, colls . LB_ ANL 000022— 1 adult and LB_ ANL000023, 2 juveniles, in latosol in an intermediate secondary forest, (2°35'15.00"S 44°12'32.00"W), April 2011, P. R.S. Silva, K.H.M. Muchavisoy, colls . LB_ ANL 000026— 5 adults and LB_ ANL000027, 17 juveniles, in latosol in an intermediate secondary forest, (2°35'30.99"S 44°12'33.00"W), April 2011, P. R.S. Silva, K.H.M. Muchavisoy, colls . LB_ ANL000058, 1 adult, in latosol in a young secondary forest, São Luís (2°35'25.89"S 44°12'37.10"W), Jule 2013, J.A.M. Gutiérrez. LB_ ANL 000071— 4 adultes and LB_ ANL000072, 7 juveniles, in latosol in an intermediate secondary forest, (2°35'11.19"S 44°12'31.36"W), January 2014, J.A.M. Gutiérrez.; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.208714&amp;materialsCitation.latitude=-2.5864415" title="Search Plazi for locations around (long -44.208714/lat -2.5864415)">Saõ José de Ribamar</a>: LB_ ANL 000073— 7 adults and LB_ ANL000074, 10 juveniles, in argisol in an old-growth forest, (2°38'38.96"S 44°8'17.66"W), January 2014, J.A.M. Gutiérrez. LB_ ANL 000075 — 4 adults and LB_ ANL000076, 12 juveniles, in argisol in an old-growth forest, (2°38'59.24"S 44°8'4.10"W), January 2014, J.A.M. Gutiérrez. LB_ ANL000192, 13 juveniles, in argisol in an old-growth riparian forest, (2°38'42.76"S 44°8'19.99"W), January 2016, G.X. Rousseau.; PARÁ: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.164448&amp;materialsCitation.latitude=-2.3941665" title="Search Plazi for locations around (long -48.164448/lat -2.3941665)">Tomé Açú</a>: LB_ ANL000041 — 12 adults and LB_ ANL000042, 125 juveniles, in latosol in an old-growth forest, (2°23'39.00"S 48°9'52.00"W), July 2011, P.R.S. Silva, K.H.M. Muchavisoy, colls. LB_ ANL 000043— 3 adults and LB_ ANL000044, 20 juveniles, in latosol in a logged forest, (2°30'06.00"S 48°17'26.00"W), July2011,P. R.S. Silva, K.H.M.Muchavisoy,colls . LB_ ANL 000045— 1 adult and LB_ ANL000046, 24 juveniles, in latosol in an intermediate secondary forest, (2°24'46.00"S 48°11'51.00"W), Jule 2011, P. R.S. Silva, K.H.M. Muchavisoy, colls . LB_ ANL 000047— 1 adult and LB_ ANL000048, 65 juveniles, in latosol in an intermediate secondary forest, (2°24'7.00"S 48°11'48.0"W), Jule 2011, P. R.S. Silva, K.H.M. Muchavisoy, colls . LB_ ANL000049, 55 juveniles, in latosol in an intermediate secondary forest, (2°24'6.9984"S 48°11'48.0012"W), Jule 2011, P. R.S. Silva, K.H.M. Muchavisoy, colls . LB_ ANL 000050— 7 adults and LB_ ANL000051, 71 juveniles, in latosol in an intermediate secondary forest, Tomé Açu (2°24'29.04"S 48°11'52.32"W), Jule 2011, P. R.S. Silva, K.H.M. Muchavisoy, colls . LB_ ANL 000052— 14 adults and LB_ ANL000053, 70 juveniles, in latosol in an intermediate secondary forest, (2°31'7.6"S 48°16'32.7"W), August 2014, J.A.M. Gutiérrez, G.X. Rousseau, colls . LB_ ANL 000054— 11 adults and LB_ ANL000055, 59 juveniles, in plinthosol in an intermediate AFS, (2°11'55.94"S 48°9'17.15"W), August 2014, J.A.M. Gutiérrez, G.X. Rousseau, colls . LB_ ANL 000056— 4 adults and LB_ ANL000057, 47 juveniles, in latosol in an intermediate AFS, (2°32'38.37"S 48°16'26.67W), August 2014, J.A.M. Gutiérrez, G.X. Rousseau, colls . LB_ ANL 000059— 6 adults and LB_ ANL000060, 37 juveniles, in latosol in a young AFS, (2°20'58.38"S 48°15'45.83"W), Jule 2011, P. R.S. Silva, G.X. Rousseau, colls . LB_ ANL 000061— 16 adults and LB_ ANL000062, 100 juveniles, in latosol in a young AFS, (2°20'56.76"S 48°15'42.4"W), Jule 2011, P. R.S. Silva, G.X. Rousseau, colls . LB_ ANL 000063— 9 adults and LB_ ANL000064, 86 juveniles, in latosol in a young AFS, (2°20'55.46"S 48°15'40.5"W), Jule 2011, P. R.S. Silva, G.X. Rousseau, colls . LB_ ANL 000065— 10 adults and LB_ ANL000066, 83 juveniles, in latosol in a young AFS, (2°24'12"S 48°14'49"W), Jule 2011, P. R.S. Silva, G.X. Rousseau, colls . LB_ ANL 000067— 8 adults and LB_ ANL000068, 101 juveniles, in latosol in a young AFS, (2°24'15.00"S 48°14'51.00"W), Jule 2011, P. R.S. Silva, G.X. Rousseau, colls . LB_ ANL 000069— 17 adults and LB_ ANL000070, 109 juveniles, in latosol in a young AFS, (2°24'12.00"S 48°14'55.20"W), Jule 2011, P. R.S. Silva, G.X. Rousseau, colls . LB_ ANL 000163— 21 juveniles and LB_ ANL000164, 5 adults, in latosol in a logged forest, (2°24'40.82"S 48°11'34.12"W), August 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls . LB_ ANL 000165— 53 juveniles and LB_ ANL000166, 3adults,in latosol in an intermediate secondary forest,(2°24'48.65"S 48°11'34.38"W), August 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls . LB_ ANL000167, 45 juveniles, in plinthosol in a young secondary forest, (2°12'33.87"S 48°9'28.66"W), August 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls . LB_ ANL 000168— 45 juveniles and LB_ ANL000169, 3 adults, in plinthosol in a young secondary forest, (2°12'34.27"S 48°9'23.68"W), August 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls . LB_ ANL 000170— 70 juveniles and LB_ ANL000171, 7 adults, in plinthosol in an intermediate secondary forest, (2°11'59.71"S 48°8'45.96"W), August 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls . LB_ ANL 000172— 75 juveniles and LB_ ANL000173, 20 adults, in plinthosol in an intermediate secondary forest, (2°11'59.59"S 48°8'58.62"W), August 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls . LB_ ANL 000174— 36 juveniles and LB_ ANL000175, 8 adults, in plinthosol in an intermediate secondary forest, (2°11'55.68"S 48°9'3.06"W), August 2014, J.A.M. Gutiérrez, E.G. Cardozo, colls. (Figure 1)</p></div>	https://treatment.plazi.org/id/90278E13FFE3FF9857C5FA1A49439165	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sousa, Sandriel Costa;Hernández-García, Luis Manuel;Beckman, Ruana Aretha Farias Santiago;Rousseau, Guillaume Xavier	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE6FF9957C5FB7548239312.text	90278E13FFE6FF9957C5FB7548239312.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhinodrilus antonioi Hernandez-Garcia & Sousa 2020	<div><p>Rhinodrilus antonioi Hernández-García &amp; Sousa, 2020</p><p>Distribution type: Native.</p><p>Distribution in Brazil: Maranhão: Caxias (Sousa et al. 2020b). (Figure 2)</p></div>	https://treatment.plazi.org/id/90278E13FFE6FF9957C5FB7548239312	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sousa, Sandriel Costa;Hernández-García, Luis Manuel;Beckman, Ruana Aretha Farias Santiago;Rousseau, Guillaume Xavier	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE6FF9957C5FA554F2190DE.text	90278E13FFE6FF9957C5FA554F2190DE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Urobenus brasiliensis Benham 1886	<div><p>Urobenus brasiliensis Benham, 1886</p><p>Urobenus brasiliensis Benham, 1886: 82 .</p><p>Anteus brasiliensis (Benham, 1886): Rosa, 1896: 108.</p><p>Rhinodrilus brasiliensis: Brown &amp; Fragoso 2007: 446.</p><p>Distribution type: Native.</p><p>Distribution in Brazil: Present in 10 Brazilian states (Brown &amp; Fragoso, 2007, Sousa et al. 2020b)</p><p>Maranhão: Caxias (Sousa et al. 2020b). (Figure 2)</p></div>	https://treatment.plazi.org/id/90278E13FFE6FF9957C5FA554F2190DE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sousa, Sandriel Costa;Hernández-García, Luis Manuel;Beckman, Ruana Aretha Farias Santiago;Rousseau, Guillaume Xavier	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE6FF9957C5F8E94D8291D2.text	90278E13FFE6FF9957C5F8E94D8291D2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Urobenus petrerei Righi 1985	<div><p>Urobenus petrerei Righi, 1985</p><p>Distribution type: Native.</p><p>Distribution in Brazil: Maranhão: São Luís (Sousa et al. 2020b). Pará: Bagagem, near the Tocantins River (Righi, 1985). (Figure 2)</p></div>	https://treatment.plazi.org/id/90278E13FFE6FF9957C5F8E94D8291D2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sousa, Sandriel Costa;Hernández-García, Luis Manuel;Beckman, Ruana Aretha Farias Santiago;Rousseau, Guillaume Xavier	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFFBFF8557C5FC5A49B39748.text	90278E13FFFBFF8557C5FC5A49B39748.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Crassiclitellata	<div><p>Key to the earthworm species recorded in the sampled localities</p><p>1. Prostate present................................................................................... 2</p><p>1. Prostate absent.................................................................................... 8</p><p>2. Two pairs of intraclitellar prostates................................................. Dichogaster (D.) bolaui</p><p>2. One pair of intraclitellar prostates..................................................................... 3</p><p>3. Gizzard absent...................................................................... Liodrilus mendesi</p><p>3. Gizzard present................................................................................... 4</p><p>4. Intestinal gizzard (segments 16–22)................................................ Hyperiodrilus africanus</p><p>4. Pre-clitellar gizzard................................................................................ 5</p><p>5. Two pre-clitellar gizzards........................................................ Dichogaster (D.) saliens</p><p>5. One pre-clitellar gizzard............................................................................ 6</p><p>6. Gizzard in segment 6.................................................................... Arraia nelmae</p><p>6. Gizzard in segment 5............................................................................... 7</p><p>7. Spermathecae pretesticular.............................................................. Brasilisia punki</p><p>7. Spermathecae intraclitellar............................................................ Eudrilus eugeniae</p><p>8. One pair of calciferous glands in 11 or 12.............................................................. 9</p><p>8. More than one pair of calciferous glands.............................................................. 14</p><p>9. Two pairs of testes................................................................................ 10</p><p>9. One pair of testes................................................................................. 12</p><p>10. Tubercula pubertatis flat and wing-shaped............................................... Hosloscolex alatus</p><p>10. Tubercula pubertatis of another shape................................................................ 11</p><p>11. Spermathecae in segments 7 and 8................................................... Holoscolex fernandoi</p><p>11. Spermathecae in segments 8 and 9................................................... Holoscolex dossantosi</p><p>12. Intraclitellar spermathecae.......................................................... Righiodrilus Gurupi</p><p>12. Pre-clitellar spermathecae.......................................................................... 13</p><p>13. Calciferous glands with calcium reservoir.......................................... Righiodrilus tocantinensis</p><p>13. Calciferous glands without calcium reservoir........................................... Glossodrilus antunesi</p><p>14. Trabecular calciferous gland present................................................................. 15</p><p>14. Trabecular calciferous gland absent.................................................................. 16</p><p>15. Calciferous glands subdivided......................................................... Urobenus petrerei</p><p>15. Calciferous glands not subdivided................................................... Urobenus brasiliensis</p><p>16. Intramural lamellar calciferous glands................................................................ 17</p><p>16. Extramural calciferous glands....................................................................... 18</p><p>17. Spermathecae in segments 5–7....................................................... Atatina arimikuriaka</p><p>17. Spermathecae in segment 9.......................................................... Atatina arimikuritea</p><p>18. One pair of testes................................................................................. 19 18. More than one pair of testes........................................................................ 20</p><p>19. Quincunx setae at tail....................................................... Pontoscolex (P.) corethrurus</p><p>19. Regular setae at tail................................................................ Pontoscolex (P.) awa</p><p>20. Calciferous glands tubular type ....................................................... Rhinodrilus antonioi</p><p>20. Calciferous glands lamellar type ..................................................................... 21</p><p>21. Length greater than 20 cm ...................................................... Andiorhinus (T.) miricuri</p><p>21. Length less than 20 cm ........................................................ Andiorrhinus (T.) barrosoi</p></div>	https://treatment.plazi.org/id/90278E13FFFBFF8557C5FC5A49B39748	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sousa, Sandriel Costa;Hernández-García, Luis Manuel;Beckman, Ruana Aretha Farias Santiago;Rousseau, Guillaume Xavier	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
