taxonID	type	description	language	source
4429BC236B2259F4B02291D556706BDF.taxon	description	Figs 16, 17, 18	en	George, Kai Horst, Tönjes, Lukas, Yurtdaş, Songül (2025): That “ darned Dorsiceratus case ” (Copepoda, Harpacticoida, Cletodidae T. Scott) — an attempt of a systematic approach, including the description of three new species. Zoosystematics and Evolution 101 (1): 173-221, DOI: 10.3897/zse.101.139354
4429BC236B2259F4B02291D556706BDF.taxon	etymology	Etymology. the epitheton andeep refers to the Expeditions ANtarctic benthic DEEP-sea biodiversity (ANDEEP) I + II of RV POLARSTERN (cruise PS 61) that had taken place in 2002 to the Scotia Arc, the Weddell Sea, and the Antarctic peninsula.	en	George, Kai Horst, Tönjes, Lukas, Yurtdaş, Songül (2025): That “ darned Dorsiceratus case ” (Copepoda, Harpacticoida, Cletodidae T. Scott) — an attempt of a systematic approach, including the description of three new species. Zoosystematics and Evolution 101 (1): 173-221, DOI: 10.3897/zse.101.139354
4429BC236B2259F4B02291D556706BDF.taxon	description	Description of female. The body shape (not drawn) corresponds to the generic diagnosis with the pronounced frontal peak, the formation of cuticular processes on the cphth and the first three free thoracic segments, as well as the structure of the extremities. With a body length of 799 µm, Do. andeep sp. nov. is, however, the largest described representative of the genus Dorsiceratus to date. Rostrum (Fig. 16 A) constricted, about 1.6 times longer than broad, cleft at its tip and bearing 2 sensilla and 1 long tube pore. FR (Fig. 17 A) about seven times as long as wide, bearing 7 setae: setae I and II insert on the outer margin halfway along the FR; seta I is tiny and bare, with the bipinnate seta II located right next to it; seta III is bare and inserts subapically on the outer side; seta V is the longest, located apically and flanked externally by seta IV and internally by seta VI, both of which are small and bare; seta VII is biarticulated and arises subapically on the dorsal side of the FR. A 1 (Fig. 16 B) 4 - segmented, segments 1 – 3 of about equal length, segment 4 only about half as long as the preceding ones. First segment anteriorly with a row of long spinules. All setae bare. Setal formula: I – 1; II – 6; III – 6 + aes; IV – 9 + aes.	en	George, Kai Horst, Tönjes, Lukas, Yurtdaş, Songül (2025): That “ darned Dorsiceratus case ” (Copepoda, Harpacticoida, Cletodidae T. Scott) — an attempt of a systematic approach, including the description of three new species. Zoosystematics and Evolution 101 (1): 173-221, DOI: 10.3897/zse.101.139354
55E704A905305D6580A9B45223931B53.taxon	description	Figs 12, 13, 14, 15	en	George, Kai Horst, Tönjes, Lukas, Yurtdaş, Songül (2025): That “ darned Dorsiceratus case ” (Copepoda, Harpacticoida, Cletodidae T. Scott) — an attempt of a systematic approach, including the description of three new species. Zoosystematics and Evolution 101 (1): 173-221, DOI: 10.3897/zse.101.139354
55E704A905305D6580A9B45223931B53.taxon	etymology	Etymology. The epitheton of Dorsiceratus denizae sp. nov. is fondly dedicated to SY’s daughter, Miss Deniz Yurtdaş. Gender: feminine.	en	George, Kai Horst, Tönjes, Lukas, Yurtdaş, Songül (2025): That “ darned Dorsiceratus case ” (Copepoda, Harpacticoida, Cletodidae T. Scott) — an attempt of a systematic approach, including the description of three new species. Zoosystematics and Evolution 101 (1): 173-221, DOI: 10.3897/zse.101.139354
55E704A905305D6580A9B45223931B53.taxon	description	Description of female. Habitus (Fig. 12) slender and cylindrical, with inconspicuous boundary between pro- and urosome. Body length of holotype from rostral tip to end of FR 686 µm (mean 717 µm, range: 686 – 747 µm, n = 3), cphth and body somites dorsally with some sensilla and tube pores. Frontal peak on cphth well-developed, with anterior cephalothoracic setular tufts (ACST). Cphth and first three free thoracic somites dorsally with slender spinulose cuticular processes (CLDP, DT 1 – DT 3), that bear a sensillum at their tips and are accompanied by another sensillum at their bases. Rostrum fused with cphth, strongly constricted, slightly longer than broad and cleft at its tip, with 2 sensilla and 1 tube pore. Last thoracic and first abdominal somite fused into a GDS; former segmental demarcation still indicated by few dorsal spinules and pair of sensilla. Free thoracic body somites laterally with setular tufts. P 2 – P 6 - bearing somites dorsally with 1 tube pore. First abdominal somite (= posterior half of GDS) and the following two abdominal somites dorsally with row of flexible setules on posterior margin; first and second abdominal somites additionally with 2 pairs of sensilla. Penultimate somite dorsally with tube pore on posterior margin. Telson shorter than broad, slightly trapezoidal from dorsal view. Anal operculum basally flanked by a pair of sensilla; apically with long spinules, which increase in length from the outer edges towards the centre. FR (Fig. 13 A) approximately 7 times longer than basal width, slender, equipped with 7 setae. Setae I and II located at mid-length of ramus; seta I minute, bare; seta II much longer, bipinnate; seta III (broken in Fig. 13 A) arising subapically; setae IV and VI small and bare, inserting apically on outer and inner edges, respectively; seta V longest; seta VII (broken in Fig. 13 A) positioned dorsally at subapical part of FR. FR additionally with 1 short broad and 1 slender tube pore on the ventral apical margin. A 1 (Fig. 13 B) 4 - segmented. First segment longest, with long spinules on anterior margin, distally with 1 biplumose seta. Second segment reaching 2 / 3 of first segment, with 6 setae. Third segment almost as long as first, with row of spinules on posterior margin and with 3 bare and 1 bipinnate setae along anterior margin; moreover, subapically, 1 seta and 1 strong aes arise from acrothek, accompanied by 2 bare setae. Fourth segment smallest, with 10 setae (4 of which broken) and 1 aes. Setal formula: I – 1, II – 6, III – 7 + aes, IV – 10 + aes.	en	George, Kai Horst, Tönjes, Lukas, Yurtdaş, Songül (2025): That “ darned Dorsiceratus case ” (Copepoda, Harpacticoida, Cletodidae T. Scott) — an attempt of a systematic approach, including the description of three new species. Zoosystematics and Evolution 101 (1): 173-221, DOI: 10.3897/zse.101.139354
E55D0386BA4653A9B04FAC0BC0B0CD8F.taxon	type_taxon	Type species. Dorsiceratus octocornis Drzycimski, 1967.	en	George, Kai Horst, Tönjes, Lukas, Yurtdaş, Songül (2025): That “ darned Dorsiceratus case ” (Copepoda, Harpacticoida, Cletodidae T. Scott) — an attempt of a systematic approach, including the description of three new species. Zoosystematics and Evolution 101 (1): 173-221, DOI: 10.3897/zse.101.139354
D6AB22D2EA4653668E42E8856E054A5E.taxon	description	Figs 3, 4, 5, 6, 7, 8, 9, 10, 11	en	George, Kai Horst, Tönjes, Lukas, Yurtdaş, Songül (2025): That “ darned Dorsiceratus case ” (Copepoda, Harpacticoida, Cletodidae T. Scott) — an attempt of a systematic approach, including the description of three new species. Zoosystematics and Evolution 101 (1): 173-221, DOI: 10.3897/zse.101.139354
D6AB22D2EA4653668E42E8856E054A5E.taxon	etymology	Etymology. the epitheton karinae is given in fond dedication to LT’s mother, Mrs. Karin Tönjes (Apen, Germany). Gender: feminine.	en	George, Kai Horst, Tönjes, Lukas, Yurtdaş, Songül (2025): That “ darned Dorsiceratus case ” (Copepoda, Harpacticoida, Cletodidae T. Scott) — an attempt of a systematic approach, including the description of three new species. Zoosystematics and Evolution 101 (1): 173-221, DOI: 10.3897/zse.101.139354
D6AB22D2EA4653668E42E8856E054A5E.taxon	description	Description of female. Habitus (Fig. 3 A) slender and cylindrical, with inconspicuous boundary between pro- and urosome. Body length of holotype from rostral tip to end of FR 547 µm (mean 476 µm, range: 422 – 547 µm, n = 4), cphth and body somites dorsally with some sensilla and tube pores. Cphth and first three free thoracic somites dorsally with slender spinulose cuticular processes (CLDP, DT 1 – DT 3) that bear a sensillum at their tips and are accompanied by another sensillum at their bases. Rostrum as in the male (Fig. 11 A). Last thoracic and first abdominal somite fused into a GDS; former segmental demarcation still indicated by dorsal row of small spinules and pair of sensilla. Free thoracic body somites laterally with setular tufts, P 6 - bearing somite (= anterior half of GDS) additionally with tube pore. P 2 – P 6 - bearing somites dorsally also with 1 tube pore. First abdominal somite (= posterior half of GDS) and the following two abdominal somites dorsally with row of spinules at posterior margin; first and second abdominal somites additionally with pair of sensilla. Telson shorter than broad, slightly trapezoidal in shape from dorsal view. Anal operculum with some long spinules at its apical margin and basally flanked by a pair of sensilla. FR (Fig. 3 B) cylindrical and slender, about four times longer than broad, with 6 setae: seta I completely lost; seta II arising at outer margin halfway along the ramus; seta III broken in all eight specimens, but insertion clearly discernible, with a short tube pore next to it; seta IV minute, arising apically at the outer edge; seta V longest, located apically; seta VI as small as IV, positioned apically at inner edge of the ramus; seta VII arising dorsally at the end of the ramus. A 1 (Fig. 4 A) 4 - segmented, segments 1 and 3 of nearly the same length, second segment smaller, fourth segment about half the length of the third segment. First segment with row of long and flexible spinules at anterior margin and with 1 seta (broken in Fig. 4 A) apically at anterior edge; second segment with 7 setae; third segment with 4 setae along anterior margin, at distal margin with 2 short setae and 1 aes that is accompanied by 1 long seta arising from acrothek; fourth segment with 11 setae, six of which biarticulated, and 2 setae accompanying an aes. Setal formula: I – 1; II – 7; III – 7 + aes; IV – 11 + aes. A 2 (Fig. 4 B, B *) with allobasis that carries 2 abexopodal setae but lacks an exopod; endopod with row of spinules proximally on the anterior and distally on the posterior margin; at its distal half it bears 4 long spinules and 3 setae, one of which bipinnate and the others bare. At its posterior edge, the endopod bears 2 cuticular frills (Fig. 4 B *), the first one armed with spinules, apically with 6 elements: 1 unipinnate spine and 1 longer unipinnate seta, plus 3 long geniculate setae (one broken in Fig. 4 B), the outermost fused to a tiny bare seta (broken in Fig. 4 B). Md (Fig. 5 A) produced into long gnathobase that ends in 2 strong and bicuspidate teeth and 1 less strong element that ends in four fine tips; apart from that, the gnathobase lacks any ornamentation. Mandibular palp 1 - segmented, with 1 basal (Fig. 5 A: seta no. 2), 1 exopodal (6), and 3 endopodal setae (3 – 5); basal seta no. 1 completely reduced (Fig. 5 A: asterisk *). Mxl (Fig. 5 B) with praecoxal arthrite bearing 7 strong apical spines, one of which unipinnate, and 2 surface setae. Coxa basally with few long spinules and apically with 1 unipinnate and 1 bare seta. Basis, endo- and exopod fused, carrying 3 lateral setae and 3 apical spines, one of which unipinnate. Mx (Fig. 5 C) syncoxa with 2 endites and few long spinules along distal margin. Both endites of similar shape, not fused to syncoxa, and carrying each 2 bare setae and 1 strong unipinnate spine fused with the segment. Allobasis distinct, also with long spinules at distal margin and produced into strong bare claw; additionally, with 2 bare setae and 1 bare spine. Endopod small and knob-like, equipped with 2 bare setae. Mxp (Fig. 5 D) slender; syncoxa apically with row of spinules and 1 bare seta; basis about 1.6 times longer than syncoxa, with row of long spinules at outer margin; endopod produced into bare, lean claw longer than basis and accompanied by 1 tiny seta at its base. P 1 – P 4 with transversely elongated bases that present a row of long spinules at their anterior margins and 1 outer seta (broken in all corresponding figures); P 2 – P 4 bases additionally with long tube pore. Intercoxal sclerites (exemplified by that from the P 2) long and bow-like. Endopods 2 - segmented (except male P 3 endopod; see below), exopods P 2 – P 4 3 - segmented. P 1 (Fig. 6 A) basis additionally with 1 bare inner seta; enp 1 without setae or spinules, enp 2 with few long spinules at inner and outer margins and with 2 apical bare setae, the inner one being slightly longer than the outer one. Exopod 2 - segmented, exp 1 carrying 1 long bipinnate outer seta; exp 2 more than 2 times longer than exp 1, outer unipinnate seta arising at distal third, 4 geniculate bare setae inserting apically. P 2 – P 4 (Figs 6 B, 7 A, 8 A) with enp 1 being unarmed and much shorter than enp 2; P 2 enp 2 (Fig. 6 B) with some spinules at inner and outer margins and 1 biplumose apical seta that exceeds the length of the whole exopod. P 3 and P 4 endopods (Figs 7 A, 8 A) without spinules but bearing 2 apical setae: the inner one short and bare, the outer one bipinnate and about 4 times longer than the inner one in the P 3 (Fig. 7 A), whilst biplumose and nearly 5 times longer than the inner one in the P 4 (Fig. 8 A). Exopods with slender segments of increasing length from exp 1 to exp 3. All segments except P 4 exp 3 with rows of spinules at inner and / or outer margin; exp 1 with long outer seta, exp 2 with 1 outer and 1 inner seta, the latter bipinnate (P 2), bare (P 3), or biplumose (P 4). Exp 3 as depicted with 2 uni- or bipinnate outer setae, as well as with 2 apical setae (unipinnate in P 2, bare in P 3, and biplumose in P 4); P 2 and P 4 additionally with 1 inner seta (uniplumose in P 2, biplumose in P 4), P 3 with 2 bare inner setae. P 2 and P 3 with long tube pore located between the outer elements. Setation of P 1 – P 4 as given in Table 3. P 5 (Fig. 9 A) with slender basoendopod that bears a row of long spinules along anterior margin and a long bare outer seta arising from short, square setophore and accompanied by long tube pore; endopodal lobe completely incorporated into basis, with 2 bare setae and 1 tube pore. Exopod long and slender, approximately 1.5 times longer than basoendopod, with 1 outer unipinnate and 1 inner bare seta, 1 bare subapical seta shifted to the posterior surface, and 2 apical bipinnate setae. P 6 and GF (Fig. 9 B). P 6 small, lobes fused to a narrow clasp-like structure that carries 1 bare seta on each outer side; close to the gonopore, that “ clasp ” bulges out slightly, being strengthened by a pair of sclerotised “ ribs ”.	en	George, Kai Horst, Tönjes, Lukas, Yurtdaş, Songül (2025): That “ darned Dorsiceratus case ” (Copepoda, Harpacticoida, Cletodidae T. Scott) — an attempt of a systematic approach, including the description of three new species. Zoosystematics and Evolution 101 (1): 173-221, DOI: 10.3897/zse.101.139354
D6AB22D2EA4653668E42E8856E054A5E.taxon	description	Description of male. Male resembling female in most aspects, except in the shape of the A 1, P 3, and P 4 endopods and P 5. Habitus (Fig. 10) as in female but smaller, slenderer, and lacking GDS. Body length from rostral tip of PT 6 to end of FR 376 µm (mean: 374 µm, range: 352 – 401 µm, n = 4). Rostrum (Fig. 11 A) fused with cphth, strongly constricted, slightly longer than broad, and cleft at its tip, with 2 sensilla; tube pore not recognisable in any of the four males. A 1 (Fig. 11 B) consisting of 6 segments, chirocer, geniculation between fifth and sixth segments. The first two segments longest, the third and sixth segments slightly shorter than the fifth one, which is slightly swollen; acrothek not discernible; the fourth segment minute, unarmoured. Setal formula: I – 1; II – 7; III – 4; IV – 0; V – 9 + aes; VI – 9 + aes. P 3 exopod as in female. Endopod 3 - segmented (Fig. 7 B), enp 1 tiny and bare, enp 2 more than three times longer than enp 1, ending on inner apical edge in cuspidal apophysis that slightly overtops the apical margin of enp 3. The latter is as long as enp 1 and bears 2 apical bare setae, the outer one being approximately seven times longer than the inner one. P 4 exopod as in female. Endopod 2 - segmented (Fig. 8 B), enp 1 small, nearly squarish, and bare. Enp 2 about 3.4 times longer than enp 1, apically with 2 setae, the inner one bare, the outer one approximately 3.4 times longer than the inner seta and biplumose. Subapically on the outer margin arises 1 bipinnate spine. P 5 (Fig. 9 C) generally as in female but smaller. Basoendopod more compact than that of female, without spinulose row; endopodal area with 2 bare setae and small tube pore. Exopod only slightly longer than basoendopod, without spinulose ornamentation, setation as in female, likewise with 1 tube pore.	en	George, Kai Horst, Tönjes, Lukas, Yurtdaş, Songül (2025): That “ darned Dorsiceratus case ” (Copepoda, Harpacticoida, Cletodidae T. Scott) — an attempt of a systematic approach, including the description of three new species. Zoosystematics and Evolution 101 (1): 173-221, DOI: 10.3897/zse.101.139354
