identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
5EF2E4AADF3E59F5B67C8047302166B1.text	5EF2E4AADF3E59F5B67C8047302166B1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hanseniella chocoita Montes-Rodríguez & Holguin & Parra-Gómez & Marchant 2026	<div><p>Hanseniella chocoita sp. nov.</p><p>Figs 14, 15, 16, 17, 18, 19</p><p>Type material.</p><p>Holotype • male (CTNI -10574 a), Colombia: Santander, Girón, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.1628&amp;materialsCitation.latitude=6.9867" title="Search Plazi for locations around (long -73.1628/lat 6.9867)">Vereda Chocoita</a>, manual collection in pineapple farming, 850 m a. s. l., 6.9867, -73.1628, 15 - IX- 2020. J. Montes-Rodríguez . Paratypes • 2 females, 1 male (CTNI -10574 b-d) same data as for holotype; • 2 females, 1 male (CTNI 10412 a-c) Colombia: Santander, Girón, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.1628&amp;materialsCitation.latitude=6.9867" title="Search Plazi for locations around (long -73.1628/lat 6.9867)">Vereda Chocoita</a>, manual collection in pineapple farming, 850 m a. s. l., 6.9867, -73.1628, 15 - VIII- 2020. J. Montes-Rodríguez .</p><p>Diagnosis.</p><p>Hanseniella chocoita sp. nov. shares the same pattern of macrochaetae on tergites 2–4, 6–7, and 9 with H. aculeata Jupeau, 1954, H. afromontana Scheller, 1954, H. armigera Scheller, 1961, H. caldaria (Hansen, 1903), H. colombiana Juberthie-Jupeau &amp; Réveillet, 1997, H. conisetosa Scheller, 1971, H. echinata Adam &amp; Burtel, 1956, H. ghanensis Belfield, 1988, H. guimaraensis Scheller, 2007, H. hortulana Scheller, 1971, H. incompta Scheller, 1971, H. ivorensis Juberthie-Jupeau &amp; Kehe, 1978, H. lebrijana sp. nov. Montes, Parra-Gómez, Holguín &amp; Marchant, 2025, H. lucifuga Scheller, 1961, H. milloti Aubrey &amp; Masson, 1953, H. modesta Aubrey &amp; Masson, 1953, H. montana Scheller, 1971, H. orientalis (Hansen, 1903), H. remyi Aubrey &amp; Masson, 1953, H. similis Scheller, 1961 and H. unguiculata (Hansen, 1903) . It differs from them by the following combination of characters: body length 3.4–4.4; distinct central rod on head; first maxillary palp with the apex divided into three tips, central tip broader than flanking ones; surface of antennomeres 4 and 5 with scaly pattern mixed with micropubescence; antennae with three whorls maximum, simple setae of varying size and small trifurcate organs; apical antennomere with usual stalk organ, plus a smaller one; first rudimentary tergite with four or five setae; posterior margin of tergite 2 slightly convex, in tergite 3 mostly straight, in tergites 13 or 14 sightly concave; anterior margin of all tergites glabrous, with pattern of transverse arched mixed with micropubescence; sclerites near coxal sacs with simple setae; first pair of legs with posterior claw straight and subulate, and frontal setae short and lanceolate; 12 th pair of legs with anterior claw with proximal half thick, posterior portion moderately arched and relatively broad, tip acuminate; styli with two setae, with smaller setae arising from a tiny conical protuberance at the apex of styli; cerci setose, bearing simple setae and with a significant basal portion glabrous and with micropubescence. It can also be differentiated from the most similar species by markedly different size and shape of claws of the 12 th pair of legs in H. chocoita sp. nov., while similar in size in H. aculeata . A greater number of setae on the cerci (75–106), while H. montana has fewer than 20. The length of the cerci of H. hortulana and H. armigera is 4 × longer than wide, while in H. chocoita sp. nov. it is less than 3.17. Hanseniella similis in the equiangular trapezoidal shape of the third tergite, whereas in H. chocoita sp. nov. it is subrectangular. It differs from Hanseniella milloti in the shape of its cerci, which is relatively shorter and has fewer and longer setae than H. chocoita sp. nov. and differs from H. incompta in the tip of the maxillary palp, which is divided in H. chocoita sp. nov. and continuous in H. incompta .</p><p>Description.</p><p>Length of body without antennae and cerci 3.4 (3.4–4.4) mm, antenna 1.15–1.37 (1.15–1.71). Head. Slightly longer than wider, 1.01 (1.01–1.17) × broader than long, frontal margin convex, lateral margin at point of articulation smooth, posterior margin straight with rounded posterolateral angles (Fig. 14 A). Central rod ovoid posteriorly (Fig. 14 D). Dorsal surface smooth without micropubescence or microsculpture, except for the anterior area of the head between the antennae, which has a scale-like cuticle. Dorsal surface covered by straight setae not significantly different, few large setae ~ 2.0 × longer than normal setae and 0.50 × the width of the first antennomer, arranged in 3 + 3 anteriorly of antennal insertion, four on the anterior margin of the head, four near the anterior margin and three laterally on each side behind the rounded Tömösváry’s organ. Area around Tömösváry’s organ granular in texture (Fig. 14 C). Each anterior plate of the second maxillae with two proximal setae. External-distal corner of these plates with four or five sets of sensilla with typical chandelier shape decreasing in size proximally and two elongated setae inserted on conic protuberances. Three terminal protuberances with 1–3 setae plus a single large distal sensillum, the two medial protuberances with a contiguous tiny tooth (Fig. 15). First maxillary palp conical with the apex divided into three tips, central tip broader than flanking ones (Fig. 14 B).</p><p>Antennae. (Fig. 16). Long, 0.33–0.40 × the length of the body, composed of 20–24 antennomers. Surface with scaly pattern mixed with micropubescence present from first to 4–5 antennomers, gradually disappearing as micropubescence become more abundant; from segment 6 onwards, pattern no longer visible and micropubescence uniform. First antennomere 1.59 × wider than longer, with a whorl of setae initially with three outer setae, from the second antennomere onwards with 7–10 setae. Second antennomere 1.84 × wider than longer. Second whorl with two types of setae: (i) Small inner setae, three on first antennomere and gradually decreasing in number until disappearing on distal antennomers. (ii) larger setae appear on the intermediate segments with three setae and gradually increasing up to ten setae towards the last antennomeres. Third whorl starts in the middle segments with one seta and ends in the last segments with two. An additional intermediate seta between whorls 2 and 3 present on the penultimate and antepenultimate antennomers. Apical antennomere spherical, apex bearing a large sensory organ borne from a small protuberance, composed of a central stalk which yields five slightly longer spiniform processes. One additional similar organ smaller and without protuberance.</p><p>Tergites. (Fig. 17, Table 8) Cuticle with the area anterior to setae with pattern of transverse arches mixed with micropubescence. First tergite rudimentary, with only four or five central setae. From third tergite onwards: 0.33–0.5 anterior surface without setae, pattern of transverse arches imperceptible in the area with setae. Four types of setae of different length present on tergites: (i) small and similar in size, covering the tergal surface; (ii) ~ 2 × as large as the anterior type, sparsely present on the posterior margin; (iii) ~ 1.5 × larger than the large marginal ones, located on the posterolateral corners except on tergites 1, 14, and 15; (iv) Macrochaetae in segments 2–4, 6, 7, and 9, ~ 1.5 × larger than the anterior type.</p><p>Legs. (Figs 18, 19 H, I) First pair of legs composed of four segments. First podomere subrectangular, ~ 1.3 × longer than wide, with poorly defined scaly pattern, scattered micropubescence and ~ 3 posteroventral setae. Second podomere subrectangular, ~ 1.4 × longer than wide, most of the posterior surface with a scaly pattern and scattered micropubescence that becomes denser in the distal area, with ~ 9 setae on posterior face and four ventral setae. Third podomere subrectangular, 1.15 × longer than wide, weak hexagonal pattern and micropubescence, with ~ 1 posterodorsal and four dorsal setae. Fourth podomere elongated, ~ 3.8 × longer than wide, covered uniformly with micropubescence, with ~ 13 setae arranged in five rows (only ten visible in Fig. 18). Anterior claw basally thick, medially arched, ~ 2 × as large as the posterior, posterior claw straight and subulate, frontal setae short and lanceolate, ~ 0.5 length of the anterior claw.</p><p>12 th pair of legs with all podomeres bearing micropubescence in anterior view. First podomere subtriangular, ~ 1.5 × wider than long, with three or four anteroventral setae. Second podomere subrectangular, ~ 1.3 × longer than wide, anterior face with ~ 14 setae of similar size and four ventral setae. Third podomere trapezoidal, ~ 1.03 × longer than wide, with ~ four on anterior face and seven dorsal setae in two rows. Fourth podomere subrectangular, ~ 2 × longer than wide, with ~ 4 setae on anterior face, seven dorsal setae in two rows, two or three ventral setae and five setae on posterior face (not visible in Fig. 18), the posterior and dorsal setae larger than the others. Fifth podomere elongated, ~ 4.4 × longer than wide, with ~ 21 setae arranged in six longitudinal rows (only 11 visible in Fig. 18). Claws different in size, proximal half of anterior claw thicker than the distal half, posterior portion moderately arched and relatively broad, posterior claw arched and slender than anterior claw, frontal seta lanceolate, about half the length of the anterior claw. Ventral surface. (Fig. 19 A, D) Membranous and mostly granular in appearance, apart from the coxal sacs and styli. Sclerites near coxal sacs presents on every pair of legs except the first. Smallest near the second and last pair of legs, elongated with ~ 3–5 setae. Rest of sclerites subtriangular with ~ 8–33 setae. Last segment ventrally with micropubescence and transverse arch pattern, with ~ 18 setae. Last segment covering partially the base of the cerci ventrally. Coxal sacs. (Fig. 19 A, D). Fully developed only near the bases of legs 3–9, composed of two semicircular sclerites, surface bearing micropubescence, distal sclerite relative to the base leg with ~ 3–5 setae, proximal sclerite with two or three. Leg pairs 2, 10, and 11 with sclerites bearing ~ 2–5 setae. Sclerites of leg pair 2 oval-shaped, on pairs 10 and 11 subtriangular. Male organs. (Fig. 19 A, B) Simple, two contiguous semicircular sclerites covered with micropubescence, each with a row of conical setae directed medially and three or four outer setae. Styli. (Fig. 19 C, D) Elongated with micropubescence, apically with a large conical seta along with another smaller seta. The small seta arises from a tiny conical protuberance at the apex of the styli. Larger seta 2.5 × the size of the small one. Sense calicles. (Fig. 19 E) Pit margin surrounded by simple and some branched setae. Sensory seta with a wide base, inserted in the middle of the cavity, as long as the cerci in length. Cerci. (Fig. 19 F, G) Conical and with micropubescence, 3.08–3.17 longer than broad. Surface covered by 75–76 (75–116) setae, distal ones slightly longer than proximal. 15 % basal portion glabrous, apical 20 % portion also glabrous, but with only 15 % without any micropubescence. Apical setae 0.65–0.66 × the width of cerci. Accompanying seta 0.3 × the apical seta.</p><p>Distribution.</p><p>This species has only been collected in the type locality in the Chocoita district of the municipality of Girón in the department of Santander in Colombia.</p><p>Etymology.</p><p>The specific epithet chocoita refers to the locality, where the type material was collected. Feminine, in agreement with the gender of the genus Hanseniella .</p><p>Remarks.</p><p>The correct identification of Hanseniella chocoita sp. nov. requires careful examination under microscope due to subtle differences in the chaetotaxy of the antennae with Hanseniella cf. unguiculata . Hanseniella chocoita sp. nov. was collected only during the rainy season when the soil was more humid.</p></div>	https://treatment.plazi.org/id/5EF2E4AADF3E59F5B67C8047302166B1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Montes-Rodríguez, José Mauricio;Holguin, Claudia M.;Parra-Gómez, Antonio;Marchant, Sergio	Montes-Rodríguez, José Mauricio, Holguin, Claudia M., Parra-Gómez, Antonio, Marchant, Sergio (2026): Morphological and molecular identification of symphylans (Myriapoda, Symphyla) from Colombian pineapple crops, with descriptions of two new species. ZooKeys 1268: 281-324, DOI: 10.3897/zookeys.1268.159237
CDD0231D77855AC0A47AD642B296ED1D.text	CDD0231D77855AC0A47AD642B296ED1D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hanseniella lebrijana Montes-Rodríguez & Holguin & Parra-Gómez & Marchant 2026	<div><p>Hanseniella lebrijana sp. nov.</p><p>Figs 7, 8, 9, 10, 11, 12, 13</p><p>Type material.</p><p>Holotype • female (CTNI -10573 a), Colombia: Santander, Lebrija, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.2033&amp;materialsCitation.latitude=7.079" title="Search Plazi for locations around (long -73.2033/lat 7.079)">Vereda la Puente</a>, manual collection in pineapple farming, 1297 m a. s. l., 7.0790, -73.2033, 15 - XI- 2019, J. Montes-Rodríguez . Paratypes • 3 females, 3 males (CTNI -10573 b-g) same data as for holotype; 1 female (CTNI -10411 a) Colombia: Santander, Lebrija, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.2033&amp;materialsCitation.latitude=7.079" title="Search Plazi for locations around (long -73.2033/lat 7.079)">Vereda la Puente</a>, manual collection in pineapple farming, 1297 m a. s. l., 7.0790, -73.2033, 15 - IX- 2019, J. Montes-Rodríguez ; • 1 male (CTNI -10411 a) Colombia: Santander, Girón, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.1628&amp;materialsCitation.latitude=6.9867" title="Search Plazi for locations around (long -73.1628/lat 6.9867)">Vereda Chocoita</a>, manual collection in pineapple farming, 850 m a. s. l., 6.9867, -73.1628, 15 - IX- 2020. J. Montes-Rodríguez .</p><p>Type locality.</p><p>Lebrija, Santander, Colombia</p><p>Diagnosis.</p><p>Hanseniella lebrijana sp. nov. shares the same pattern of macrochaetae on tergites 2–4, 6–7 and 9 with H. aculeata Jupeau, 1954, H. afromontana Scheller, 1954, H. armigera Scheller, 1961, H. caldaria (Hansen, 1903), H. chocoita sp. nov. Montes, Parra-Gómez, Holguín &amp; Marchant, 2025, H. colombiana Juberthie-Jupeau &amp; Réveillet, 1997, H. conisetosa Scheller, 1971, H. echinata Adam &amp; Burtel, 1956, H. ghanensis Belfield, 1988, H. guimaraensis Scheller, 2007, H. hortulana Scheller, 1971, H. incompta Scheller, 1971, H. ivorensis Juberthie-Jupeau &amp; Kehe, 1978, H. lucifuga Scheller, 1961, H. milloti Aubrey &amp; Masson, 1953, H. modesta Aubrey &amp; Masson, 1953, H. montana Scheller, 1971, H. orientalis (Hansen, 1903), H. remyi Aubrey &amp; Masson, 1953, H. similis Scheller, 1961 and H. unguiculata (Hansen, 1903) . It differs from them by the following combination of characters: body length 2.9–3.9 mm; distinct central rod on head; first maxillary palp conical and simple; antennae with two whorls maximum, simple setae of varying size and small trifurcate organs; apical antennomere with usual stalked organ, plus a smaller one; first rudimentary tergite with two setae; posterior margin of tergite 2 slightly convex, in tergite 3 mostly straight, in tergites 13 and 14 slightly concave; anterior margin of all tergites glabrous, with micropubescence; sclerites near coxal sacs with simple setae; second podomere of first leg pair with two ventral setae distinctly stronger and larger than the others; claws of 12 th pair of legs arched and subequal in overall length, with acuminated tips; styli with two setae, larger seta 2.5 × the size of the small one; cerci setose, bearing simple setae and with a significant basal portion glabrous and with micropubescence. It can also be differentiated from the most similar species by: the presence of the central rod in the head, which is absent in H. guimaraensis, and in the relative length of the third podomere or femur of the 12 th leg, which is very short in H. guimaraensis . H. colombiana, H. echinata, and H. ghanensis also shows similarity to H. lebrijana sp. nov. but can be easily distinguished by the claws on leg 12, which are noticeably different in size, while they are similar in size in H. lebrijana sp. nov. H. afromontana differs in size, measuring 4.1–9.6 mm, while H. lebrijana measures 2.9–3.9 mm. The cerci in H. lucifuga are 3.7 × longer than it is wide, whereas in H. lebrijana sp. nov. it is 2.4–3.2 × longer than wide. H. caldaria and H. orientalis have three whorls of setae on the intermediate antennomeres, while H. lebrijana does not. Finally, in H. orientalis, the proximal and distal setae of the cerci are the same length, while in H. lebrijana sp. nov. the distal setae are slightly longer.</p><p>Description.</p><p>Length of body without antennae and cerci 3.3 (2.9–3.9) mm, antenna 1.26 (1.26–1.31) mm. Head. Slightly longer than wider, 1.08 (1.06–1.16) × broader than long, frontal margin convex, lateral margin at point of articulation moderately smooth, posterior margin straight with rounded posterolateral angles. Central rod posteriorly ovoid. Dorsal surface smooth without micropubescence or microsculpture, except for the anterior area of the head between the antennae, which has a scale-like cuticle (Fig. 7 B). Dorsal surface covered by straight setae not significantly different, few large setae ~ 2.5 × longer than normal setae and 0.68 × the width of the first antennomer, arranged in 3 + 3 anteriorly of antennal insertion (Fig. 7 A), two near the anterior head margin, and three laterally on each side behind the rounded Tömösváry’s organ (~ 21 microns wide) (Fig. 7 A). Each anterior plate of second maxillae with three proximal setae. External-distal corner of these plates with four sets of sensilla with typical chandelier shape decreasing in size proximally and two elongated setae inserted on conic protuberances, the most posterodistal one with a contiguous tiny tooth. Three terminal protuberances with 1–3 setae plus one large distal sensillum, all with a contiguous tiny tooth. First maxillary palp conical (Fig. 8).</p><p>Antennae. (Fig. 9). Long, 0.26–0.35 × the length of the body, composed of 25–32 antennomers. Surface of each antennomere with micropubescence except the first, which has a scale-like cuticle (Fig. 9 D). First antennomere ~ 1.8 × wider than longer, with a single distal whorl of five setae. Second antennomere ~ 1.4 × wider than longer. From second antennomere and on a single whorl of eight thick and long setae, varying between 7–9, absent on last antennomere. Thick setae from the distal whorl accompanied by thinner and shorter setae from segments 2 and 3, initially with one, gradually and slowly increasing in number up to four or five in the penultimate antennomere. Small trifurcate organ present from antennomeres 2 and 3 onwards. Secondary whorl of setae begins at antennomere 5 with two setae and gradually increases to five or six in the intermediate antennomers and only three in the last ones. Apical antennomere spherical, apex bearing a large sensory organ borne from a small protuberance, composed of a central stalk which yields five slightly longer spiniform processes. One additional similar organ smaller and without protuberance (Fig. 9).</p><p>Tergites. (Figs 10, 11, Table 7) Cuticle with smooth surface, covered in micropubescence. First tergite rudimentary, with only two central setae and very sparse pubescence in a scaly pattern. From second tergite onwards: 0.33–0.5 anterior surface without setae, only with micropubescence or microtrichiae; 0.1 anterior surface of the tergites with microtrichiae in a scaly arrangement and with a well-marked irregular line. Four types of setae of different length present on tergites: (i) small and similar in size, covering the tergal dorsal surface; (ii) ~ 2 × as large as the anterior type, sparsely present on the posterior margin; (iii) ~ 1.5 × larger than the large marginal ones, located on the posterolateral corners except on tergites 1, 14, and 15; (iv) Macrochaetae in segments 2–4, 6–7, and 9, ~ 2.0 × larger than the anterior type. Tergites 6, 9, and 12 longest (less rectangular and squarer) and more setose.</p><p>Legs. (Figs 12, 13 C). First pair of legs composed of four segments. First podomere subrectangular, ~ 1.4 × longer than wide, covered with micropubescence and ~ 4 posteroventral setae. Second podomere trapezoidal, ~ 1.3 × longer than wide, with scale-like cuticle, surface with micropubescence on posterior face with 13 or 14 setae and three ventral setae, two of them stronger and larger than the rest. Third podomere subtrapezoidal, ~ 1.4 × longer than wide, with ~ 4 posterodorsal and three slightly larger dorsal setae. Fourth podomere elongated, ~ 4.8 × longer than wide, covered uniformly with microtrichiae, with ~ 17–19 setae arranged in six rows (only 15 visible in the Fig. 12), ventral and distal ones larger. Anterior claw mostly straight, ~ 2 × as large as the posterior, posterior claw slender and slightly arched, frontal setae short, ~ 0.5 × the length of the anterior claw. 12 th pair of legs with all podomeres with the anterior surface uniformly covered with micropubescence. First podomere subtriangular, ~ 1.4 × wider than long, with ~ 3 setae on anterior face. Second podomere subtriangular, ~ 1.3 × longer than wide, anterior face with ~ 12–14 setae of similar size and two ventral setae. Third podomere trapezoidal, ~ 1.1 × wider than long, with ~ 4–6 anterior and 7–8 dorsal setae in two rows, dorsal setae larger. Fourth podomere subrectangular, ~ 1.8 × longer than wide, with ~ 5 anterior setae, four or five posterior setae, two or three ventral setae, six or seven dorsal setae, the dorsal and posterior setae larger than the others, nearly twice as large. Fifth podomere elongated, ~ 4.4 × longer than wide, with ~ 22-24 setae arranged in six longitudinal rows. Claws thick, arched, and subequal in size, frontal seta ~ 1 / 2 the length of the claws.</p><p>Ventral surface. Membranous and mostly granular in appearance, apart from the coxal sacs and styli. Sclerite near coxal sacs presents on every pair of legs except the first, of varying size with ~ 3–17 setae. Smallest near the second and last pair of legs. Last segment ventrally with micropubescence and ~ 18 setae. Last segment covering partially the base of the cerci ventrally. Coxal sacs. (Fig. 13 B). Fully developed only near the bases of legs 3–9, composed of two sclerites resembling the stomata of a plant, surface bearing micropubescence, distal sclerite relative to the base leg with ~ 4 setae, proximal sclerite with ~ 2. On pairs of legs 10 and 11, instead of a coxal sac, there is an elongated small sclerite with ~ 2-3 setae, and on leg 2, sclerite is circular with ~ 4-5 setae. Male organs (Fig. 13 B) simple, two contiguous semicircular sclerites covered with micropubescence, each with a row of inconspicuous setae directed medially, plus a few scattered hairs outside these rows.</p><p>Styli. (Fig. 13 D) Elongated with micropubescence, apically with a large conical seta along with another smaller seta. Larger seta 2.5 × size of small one. Sense calicles. (Fig. 13 E) Pit margin surrounded by simple and some branched setae. Sensory seta with a wide base, inserted in the middle of the cavity, as long as the cerci in length. Cerci. (Fig. 13 A, C) Conical and with micropubescence, 2.4–3.2 × longer than broad. Surface covered by 81 (60–99) setae, distal ones slightly longer than proximal. 20-25 % basal portion glabrous but with micropubescence, apical 15 % portion glabrous without any micropubescence. Apical setae 0.59–0.74 × width of cerci. Smaller outer seta 0.33 as long as the apical seta.</p><p>Distribution.</p><p>Currently only reported in the municipalities of Girón and Lebrija in the department of Santander in Colombia.</p><p>Etymology.</p><p>The specific epithet lebrijana refers to the municipality of Lebrija, Santander, Colombia, where the type material was collected. The name is formed as an adjective derived from the toponym, in the feminine form, in agreement with the gender of the genus Hanseniella .</p><p>Remarks.</p><p>This species has also been detected in paprika crops [ Capsicum annuum (L.) ( Solanaceae)] (Montes-Rodríguez, pers. obs.).</p></div>	https://treatment.plazi.org/id/CDD0231D77855AC0A47AD642B296ED1D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Montes-Rodríguez, José Mauricio;Holguin, Claudia M.;Parra-Gómez, Antonio;Marchant, Sergio	Montes-Rodríguez, José Mauricio, Holguin, Claudia M., Parra-Gómez, Antonio, Marchant, Sergio (2026): Morphological and molecular identification of symphylans (Myriapoda, Symphyla) from Colombian pineapple crops, with descriptions of two new species. ZooKeys 1268: 281-324, DOI: 10.3897/zookeys.1268.159237
E06765372D4151B29B81DC66F9329E74.text	E06765372D4151B29B81DC66F9329E74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hanseniella undefined-5	<div><p>Hanseniella sp. 5</p><p>Material studied.</p><p>• 1 female, Colombia: Santander, Girón, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.1628&amp;materialsCitation.latitude=6.9867" title="Search Plazi for locations around (long -73.1628/lat 6.9867)">Vereda Chocoita</a>, manual collection in pineapple farming, 850 m a. s. l., 6.9867, -73.1628, 15 - IX- 2020. J. Montes-Rodríguez, GICho 01-1 (field code) .</p><p>Distribution.</p><p>This species has only been collected in the Chocoita district of the municipality of Girón in the department of Santander in Colombia.</p><p>Remarks.</p><p>This is the first species of the genus Hanseniella reported in Neotropics with the posterior margin of the last tergite with a median projection or expansion between the cerci (Fig. 20 C). This morphological characteristic is also present in H. brachycerca, described in New Zealand by Adam and Burtel (1956). The taxonomic key proposed by Scheller and Adis (2002) for identifying Neotropical genera uses this characteristic to differentiate between the genera Hanseniella and Scopoliella, assuming that all Hanseniella species in the Neotropics had a straight or almost straight posterior margin on the last tergite. With the presence of Hanseniella sp. 5, this key was adjusted by incorporating other characteristics of Scopoliella (See below).</p></div>	https://treatment.plazi.org/id/E06765372D4151B29B81DC66F9329E74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Montes-Rodríguez, José Mauricio;Holguin, Claudia M.;Parra-Gómez, Antonio;Marchant, Sergio	Montes-Rodríguez, José Mauricio, Holguin, Claudia M., Parra-Gómez, Antonio, Marchant, Sergio (2026): Morphological and molecular identification of symphylans (Myriapoda, Symphyla) from Colombian pineapple crops, with descriptions of two new species. ZooKeys 1268: 281-324, DOI: 10.3897/zookeys.1268.159237
DFC07F954BDD52EC9080828D12BCFD14.text	DFC07F954BDD52EC9080828D12BCFD14.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hanseniella unguiculata Hansen 1903	<div><p>Hanseniella cf. unguiculata Hansen, 1903</p><p>Material studied.</p><p>• 3 females, 2 males (CTNI 10308 a-e), Colombia: Santander, Lebrija, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.1714&amp;materialsCitation.latitude=7.1978" title="Search Plazi for locations around (long -73.1714/lat 7.1978)">Vereda La Aguada de Ceferino</a>, manual collection in pineapple farming, 975 m a. s. l., 7.1978, -73.1714, 15 - X- 2019, J. Montes-Rodríguez ; • 1 female, 1 male, Colombia: Santander, Lebrija, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.1714&amp;materialsCitation.latitude=7.1978" title="Search Plazi for locations around (long -73.1714/lat 7.1978)">Vereda La Aguada de Ceferino</a>, manual collection in pineapple farming, 975 m a. s. l., 7.1978, -73.1714, 15 - X- 2019, J. Montes-Rodríguez, LELaAgu 01-9 (Field code) ; • 1 female, 1 male (CTNI 10410 a, d) Colombia: Santander, Girón, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.1628&amp;materialsCitation.latitude=6.9867" title="Search Plazi for locations around (long -73.1628/lat 6.9867)">Vereda Chocoita</a>, manual collection in pineapple farming, 850 m a. s. l., 6.9867, -73.1628, 15 - IX- 2020, J. Montes-Rodríguez ; • 1 female, 1 male, (CTNI 10410 b, c) Colombia: Santander, Lebrija, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.2033&amp;materialsCitation.latitude=7.079" title="Search Plazi for locations around (long -73.2033/lat 7.079)">Vereda la Puente</a>, manual collection in pineapple farming, 1297 m a. s. l., 7.0790, -73.2033, 15 - XI- 2019, J. Montes-Rodríguez .</p><p>Distribution.</p><p>This species was collected in the municipalities of Villanueva, Girón, and Lebrija. It was the dominant species in pineapple crops in the department of Santander. The similarity of their sequences to those published by Salazar-Moncada et al. (2015) suggests that this species has a wide distribution in Colombia.</p><p>Remarks.</p><p>Specimens from Santander clearly exhibit the key diagnostic characters traditionally used to identify this species, namely the shape of the claw on leg 12 and macrochaetal pattern. However, Hansen’s (1903) original description of specimens from Venezuela is rather brief, and the accompanying illustrations are insufficient to confirm the identification with complete certainty. Edwards (1959) subsequently provided more detailed documentation, including measurements of tergites, lateral macrochaetae, and new illustrations, although based on material from England. The morphological ranges reported by Edwards (1959) generally overlap with those observed in the Santander specimens, and the illustrations are reasonably consistent with the new material. Nevertheless, in the absence of a modern redescription of the type specimens, we refrain from definitively assigning the Santander material to H. unguiculata . Despite this taxonomic uncertainty, the species appears strongly associated with pineapple cultivation in Santander, where it was the dominant symphylan in sampled crops. Records from Hawaii, where it was the only Hanseniella species reported in pineapple cultivation (Scheller 1961), further support a clear ecological relationship with this host plant. Its broad geographic distribution and occurrence across multiple continents have made it one of the most frequently cited symphylan species in the literature; however, its biogeographic origin and native or introduced status in Colombia and other parts of the world remain unresolved.</p></div>	https://treatment.plazi.org/id/DFC07F954BDD52EC9080828D12BCFD14	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Montes-Rodríguez, José Mauricio;Holguin, Claudia M.;Parra-Gómez, Antonio;Marchant, Sergio	Montes-Rodríguez, José Mauricio, Holguin, Claudia M., Parra-Gómez, Antonio, Marchant, Sergio (2026): Morphological and molecular identification of symphylans (Myriapoda, Symphyla) from Colombian pineapple crops, with descriptions of two new species. ZooKeys 1268: 281-324, DOI: 10.3897/zookeys.1268.159237
4FE47C71F8F850ED932CD65E43422BAD.text	4FE47C71F8F850ED932CD65E43422BAD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Symphylella undefined-1	<div><p>Symphylella sp. 1</p><p>Material studied.</p><p>• 1 female, 1 male (CTNI -10414 - 10415) Colombia: Santander, Lebrija, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.2033&amp;materialsCitation.latitude=7.079" title="Search Plazi for locations around (long -73.2033/lat 7.079)">Vereda la Puente</a>, manual collection in pineapple farming, 1297 m a. s. l., 7.0790, -73.2033, 15 - XI- 2019, J. Montes-Rodríguez ; • 1 male Colombia: Santander, Lebrija, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.2033&amp;materialsCitation.latitude=7.079" title="Search Plazi for locations around (long -73.2033/lat 7.079)">Vereda la Puente</a>, manual collection in pineapple farming, 1297 m a. s. l., 7.0790, -73.2033, 15 - IX- 2019. J. M. Montes-Rodríguez, LELaPue- 10 (field code) ; • 2 females Colombia: Santander, Lebrija, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.1714&amp;materialsCitation.latitude=7.1978" title="Search Plazi for locations around (long -73.1714/lat 7.1978)">Vereda La Aguada de Ceferino</a>, manual collection in pineapple farming, 975 m a. s. l., 7.1978, -73.1714, 15 - X- 2019, J. Montes-Rodríguez, LELaAgu 01 - 11-12 (field code) .</p><p>Distribution.</p><p>This species has only been collected in the municipality of Lebrija in the department of Santander in Colombia.</p><p>Remarks.</p><p>Among the species reported in the Neotropics, this could be most closely related to Symphylella neotropica Hansen, 1903 . The similarities are in the head, specifically in the central rod, with the anterior part conspicuous but narrow and the frontal branches are thin. The conformation of the antennal setae is also similar to S. neotropica, with the first whorl having larger setae than the second.</p><p>The original description describes the cerci as having a moderate number of setae, with six or seven protruding on the lower half of the outer surface, while the remaining setae are inclined. In Symphylella sp. 1, there are a similar number of setae, but the erect setae are not significantly different in length from the rest and tend to be in the distal half of the cerci (Fig. 20 A). Furthermore, the shape of tergites II and III resembles those described by Hansen (1903) for S. neotropica . However, Symphylella sp. 1 has only short setae in contrast of the several long marginal setae present on tergite 3 (= scutum 2 according to the original interpretation of Hansen [1903]) in S. neotropica .</p></div>	https://treatment.plazi.org/id/4FE47C71F8F850ED932CD65E43422BAD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Montes-Rodríguez, José Mauricio;Holguin, Claudia M.;Parra-Gómez, Antonio;Marchant, Sergio	Montes-Rodríguez, José Mauricio, Holguin, Claudia M., Parra-Gómez, Antonio, Marchant, Sergio (2026): Morphological and molecular identification of symphylans (Myriapoda, Symphyla) from Colombian pineapple crops, with descriptions of two new species. ZooKeys 1268: 281-324, DOI: 10.3897/zookeys.1268.159237
3290447E788F513A8FD681565C4A3501.text	3290447E788F513A8FD681565C4A3501.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Symphylella undefined-2	<div><p>Symphylella sp. 2</p><p>Material studied.</p><p>• 2 females, 1 male, Colombia: Santander, Girón, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.1628&amp;materialsCitation.latitude=6.9867" title="Search Plazi for locations around (long -73.1628/lat 6.9867)">Vereda Chocoita</a>, manual collection in pineapple farming, 850 m a. s. l., 6.9867, -73.1628, 15 - IX- 2020, J. Montes-Rodríguez, GICho 01-15 - 17 (field code) .</p><p>Distribution.</p><p>This species has only been collected in the Chocoita district of the municipality of Girón in the department of Santander in Colombia.</p><p>Remarks.</p><p>This species does not resemble any of the species of the genus described in the Neotropics. The cerci has a very distinct dorsolateral seta and a row of three erect setae lateroventrally (Fig. 20 B).</p></div>	https://treatment.plazi.org/id/3290447E788F513A8FD681565C4A3501	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Montes-Rodríguez, José Mauricio;Holguin, Claudia M.;Parra-Gómez, Antonio;Marchant, Sergio	Montes-Rodríguez, José Mauricio, Holguin, Claudia M., Parra-Gómez, Antonio, Marchant, Sergio (2026): Morphological and molecular identification of symphylans (Myriapoda, Symphyla) from Colombian pineapple crops, with descriptions of two new species. ZooKeys 1268: 281-324, DOI: 10.3897/zookeys.1268.159237
