identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
8C7BE1229E16FFE276F01DAC2DE5FB73.text	8C7BE1229E16FFE276F01DAC2DE5FB73.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caenohalictus Cameron 1903	<div><p>Keys to the species of Caenohalictus from southern Brazil</p><p>Females</p><p>1. Body length 5–7.5 mm; epistomal lobe approximately obtuse and not projected over the clypeus (Fig. 1A); maxillary palpi relatively short, apical palpomere about 1 MOD or less (Fig. 1B)............................................ 2</p><p>- Body length 7.5–11 mm; epistomal lobe approximately straight and projected over the clypeus (Fig. 1B); maxillary palpi relatively long, apical palpomere longer than 1 MOD (Fig. 1D)............................................. 8</p><p>2. Body dark olivaceus with few cyaneus reflections, those reflections are strong on gena (Fig. 8A–D); mesoscutum and metapostnotum granulate (Fig. 8C)...................................... Caenohalictus cyanolucens sp. nov.</p><p>- Body bright metallic green or blue; mesoscutum and metapostnoum polished or lineolate or areolate............... 3</p><p>3. Metapostnotum with transverse striation, forming parallel lines or semicircles (Figs. 4C and 12C); scutellum very sparsely punctate, except close to the edges, weakly lineolate or polished; scape and pedicel apices with small faint yellow spot.. 4</p><p>- Metapostnotum with longitudinal or irregular ornamentation, never forming transverse striation; scutellum sparsely punctate, strongly lineolate or polished; scape and pedicel apices concolor with remaining integument, without yellow spot............................................................................................ 5</p><p>4. Metapostnotum transverse striae parallel to weakly curved (Fig. 12C); clypeus, supraclypeal area and scutellum polished between punctures (Fig. 12 B–C); mesoscutum lineolate, pubescence short and branched (Fig. 12C); terga with metallic reflections throughout the disc; sterna with green metallic reflections; clypeal apical dark band occupying approximately one-half of its length............................................. Caenohalictus incertus (Schrottky, 1902)</p><p>- Metapostnotum transverse striae strongly curved, semicircular (Fig. 4C); clypeus, supraclypeal area and scutellum lineolate-areolate (Fig. 4B–C); mesoscutum micro granulate-areolate anteriorly, lineolate posteriorly, pubescence long and branched (Fig. 4C); terga with metallic reflections only at the base and sides, darkened on the disc; sterna without metallic reflections; clypeal apical dark band occupying approximately one-third of its length.. Caenohalictus arcuatus sp. nov.</p><p>5. Mesoscutum coriaceous and sparsely punctate along the median line, polished and punctate at sides (Fig. 6C); terga and mesepisternum with dark blue reflections (Fig. 6D); fore coxa, trochanter, and femur brown, without strong metallic reflections; flagellomeres dark brown; sterna without metallic reflections; head pubescence mostly black (Fig. 6B)..................................................................... Caenohalictus curticeps (Vachal, 1903)</p><p>- Mesoscutum entirely lineolate to areolate, sparsely, or very sparsely punctate; terga and mesepisternum without dark blue reflections; fore coxa, trochanter, and femur with strong metallic reflections; flagellomeres yellowish anteriorly; sterna with green metallic reflections; head pubescence yellowish.................................................... 6</p><p>6. Scape metallic green or cupreous (Fig. 24B); clypeal apical black band short (Fig. 24B); head and mesoscutum with dense, yellow long branched setae (Fig. 24B–C); propodeal lateral surface granulate-areolate to granulate-lineolate.......................................................................... Caenohalictus tesselatus (Moure, 1940)</p><p>-. Scape black (Fig. 10B, 14B), sometimes with only faint metallic reflections; clypeal apical black band occupying to one third to half of its length (Fig. 10B, 14B); head and mesoscutum with sparse, pale short branched setae (Fig. 10C, 14C); propodeum lateral surface mostly rugulose............................................................. 7</p><p>7. Mesoscutum sparsely punctate, including at sides of parapsidal lines; scutellum areolate and sparsely punctate (Fig. 10C); tarsi light brown to yellow (Fig. 10A); head as long as wide (HW/ HL: 1.05); mesepisternum and metanotum rugulose (Fig. 10C).............................................................. Caenohalictus implexus Moure, 1950</p><p>- Mesoscutum sparsely punctate on disc, becoming punctate at sides of parapsidal lines; scutellum weakly coriaceous and sparsely to densely punctate (Fig. 14C); tarsi dark brown (Fig. 14A); head wider than long (HW/ HL: 1.13); mesepisternum and metanotum rugose (Fig. 14C)............................................ Caenohalictus leticiae sp. nov.</p><p>8. Eyes not enlarged in relation to the face, lower interocular distance 3 to 4 times the eye lower width; antennal socket not retracted in a concavity; maxillary palpomeres maximum size 1 MOD (Fig. 1B,D); scutellum and metanotum slightly convex.......................................................................................... 9</p><p>- Eyes enlarged in relation to the face, lower interocular distance about 2.5 times the eye lower width (Fig. 16B, 22C); antennal socket retracted into the face in a concavity; maxillary palpomeres length&gt; 1 (Fig. 16A); scutellum and metanotum flattened........................................................................................ 11</p><p>9. Body mostly black, with few metallic reflections (Fig. 21); forecoxa brown; antennae brown; hind tibia and tarsi with dark setae; propodeal pit strongly ridged................................. Caenohalictus schulthessi (Vachal, 1903)</p><p>- Body with extensive green or blue metallic reflections; forecoxa bright green; F 3–10 gradually light brown or yellow anteriorly; hind tibia and tarsi setae whitish or yellowish; propodeal pit carinate or weakly ridged................. 10</p><p>10. Body length 8–10 mm; head wider than longer (HW/ HL = 1.09); scutellum densely punctate and smooth; propodeal pit enclosed, weak carinae almost parallel (Fig. 2B); S 1 with strong metallic reflections; terga green or blue or cupreous, dorso-posterior areas with strong metallic reflections..................... Caenohalictus palumbes (Vachal, 1903)</p><p>- Body length 7–7.5 mm; head with elongate aspect, as long as wide (HW/ HL = 1.02); scutellum sparsely punctate and coriaceous; propodeal pit not enclosed, weak ridges divergent above (Fig. 2A); S 1 with weak metallic reflections; terga olivaceous green, dorso-posterior areas with weak metallic reflections............ Caenohalictus parviformis sp. nov.</p><p>11. Propodeal pit enclosed by strong carinae, carinae arched outwards (Fig. 2C); maxillary palpomeres about 1.2–3 times MOD (Fig. 16A); head longer than wide (HW/ HL: 0.96) (Fig. 16B); head concavity below the eye line in lateral view........................................................................... Caenohalictus longipalpis sp. nov.</p><p>- Propodeal pit enclosed by weak carinae, carinae not extending outwards (Fig. 2D); maxillary palpomeres measuring approximately 0.7–1.5x the MOD; head rounded, wider than long (HW/ HL: 1.06) (Fig. 22B); head concavity a little below or close to the eye line in lateral view...................................... Caenohalictus sesquipalpis sp. nov</p><p>Males (males of C. parviformis sp. nov. and C. schulthessi are not known)</p><p>1. Body size 6 mm or less; mesoscutum and the lateral sides of metapostnotum polished or lineolate or areolate; clypeus almost flat, apical area with median depression, yellow marking occupying less than 1/3 its length (as Fig. 11B)...... 2</p><p>- Body size 6 to 10 mm; mesoscutum and the lateral sides of metapostnotum granulate; clypeus convex, yellow marking occupying about 1/2 its length (as Fig. 9B)............................................................. 7</p><p>2. Scutellum with a median longitudinal sulcus with dense punctation; forecoxa dark brown; tarsi brown; antennae black, short (not reaching tegulae), not crenulate; mesoscutum coriaceous and sparsely punctate along the median line, polished and punctate at sides (Fig. 7C).............................................................. C. curticeps</p><p>- Scutellum without median longitudinal sulcus; forecoxa metallic green; tarsi yellow; antennae with yellow markings anteriorly, long (reaching tegulae) and with weakly crenulated flagellomeres; mesoscutum polished or entirely lineolate to areolate, sparsely or very sparsely punctate (Fig. 5C; 11C; 13C; 15C; 25C).................................... 3</p><p>3. Metapostnotum with transverse striation, forming parallel lines or semicircles; metanotum polished or coriaceous (Figs. 13C, 5C)......................................................................................... 4</p><p>- Metapostnotum with longitudinal or irregular ornamentation, never forming transverse striation; metanotum rugulose or coriaceous-rugulose (Figs. 11C, 15C, 25C).............................................................. 5</p><p>4. Metapostnotum transverse striae parallel to weakly curved (Fig. 13C); head as long as wide (HW/ HL: 1); clypeus and supraclypeal mostly polished; terga with metallic reflections throughout the disc....................... C. incertus</p><p>- Metapostnotum transverse striae strongly curved, semicircular (Fig. 5C); head slightly wider than longer (HW/ HL: 1.1); clypeus and supraclypeal area lineolate; terga with metallic reflections only at the base, darkened on the disc........................................................................................... C. arcuatus sp. nov.</p><p>5. Scape metallic green or cupreous; clypeal apical yellow band short (Fig. 25B); S 1 bright green, the remaining sterna light brown to black with green metallic reflections................................................. C. tesselatus</p><p>-. Scape black, sometimes with only faint metallic reflections; clypeal apical yellow band about one third of clypeal length (as in Fig. 15B); S 1 brown to black, metallic reflections when present, only at base, remaining sterna dark brown to black, without metallic reflections.......................................................................... 6</p><p>6. Flagellomeres 2–11 homogeneously covered by intermixed trichodea and placodea sensillae (Fig. 3A); mesoscutum entirely very sparsely punctate (Fig. 11C); propodeum lateral surface lineolate, sparsely punctate; eye setae length approximately 0.7 MOD ............................................................................... C. implexus</p><p>- Flagellomeres 2–11 with anterior large patches of sensilla placodea (Fig. 3B); mesoscutum punctate at sides of parapsidal lines and posterior margin (Fig. 15C); propodeum lateral surface mostly polished and punctate; eye setae length approximately 1 MOD .............................................................. C. leticiae sp. nov.</p><p>7. Scape with metallic reflections; F 2 longer than F 1 and pedicel combined (Fig. 19B); sterna black; metapostnotum rugose; metanotum areolate................................................................................ 8</p><p>- Scape black, without metallic reflections; F 2 as long as F 1 and pedicel combined (Fig. 17B); sterna light to dark brown; metapostnotum longitudinally striate; metanotum granulate or polished....................................... 9</p><p>8. Body size 8–10 mm; flagellomeres 2–11 homogeneously covered by intermixed trichodea and placodea sensillae (Fig. 3C); mandible with preapical tooth; gena concolor to head (Fig. 19A); terga green or blue, with a brown marginal zone; S 1 with the same color as remainder of integument................................................... C. palumbes</p><p>- Body size approximately 7 mm; flagellomeres 2–11 with anterior large patches of sensilla placodea (Fig. 3D); mandible without preapical tooth; gena with strong blue reflections (Fig. 9A), contrasting with the opaque green head; terga dark brown; S 1 black with a weak blue reflection......................................... C. cyanolucens sp. nov.</p><p>9. Maxillary palpomeres about 2.5 times MOD; head and mesosoma bright green or blue; eye setae about 1 MOD (Fig. 17B); mesoscutum granulate anteriorly to areolate on posterior two thirds; S 6 with weak invagination (Fig. 26B)................................................................................. Caenohalictus longipalpis sp. nov</p><p>- Maxillary palpomeres about 0.7–1.5 times MOD; head and mesosoma dark with little metallic reflections; eye setae about 0.7 MOD (Fig. 23B); mesoscutum entirely areolate; S 6 strongly invaginated and enclosed by a membrane (Fig. 26A)....................................................................... Caenohalictus sesquipalpis sp. nov</p></div>	https://treatment.plazi.org/id/8C7BE1229E16FFE276F01DAC2DE5FB73	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cappola, Victor;Gonçalves, Rodrigo B.	Cappola, Victor, Gonçalves, Rodrigo B. (2026): A review of Caenohalictus species from southern Brazil (Apoidea, Caenohalictini). Zootaxa 5782 (3): 401-448, DOI: 10.11646/zootaxa.5782.3.1, URL: https://doi.org/10.11646/zootaxa.5782.3.1
8C7BE1229E11FFEF76F019F42A35FD2E.text	8C7BE1229E11FFEF76F019F42A35FD2E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caenohalictus arcuatus Cappola & Gonçalves 2026	<div><p>1. Caenohalictus arcuatus sp. nov.</p><p>(Figs 1A–B, 4–5, 27G–I, 30A)</p><p>Diagnosis. This species is similar to C. incertus by the transverse striation on metapostnotum, but in C. arcuatus sp. nov. the striae are strongly curved (Fig. 4C) and this new species also has a strong body ornamentation (as areolate head and lineolate mesoscutum) when compared with C. incertus (more polished). It can be distinguished by both sexes having smaller size; integument color turquoise; terga with metallic area occupying only the anterior portion, with darkened area without reflexes, marginal area greenish brown (Fig. 4C, 5C); clypeus and supraclypeal coriaceous. Females have the clypeus apical darkened stripe occupying around one third of its area (Fig. 4B). Males have the head wider than long (Fig. 5B).</p><p>Description</p><p>Female: Body length x̄ =6.78 (N=5). ITD x̄ =1.30 (N=5). UID x̄ =1.05 (N=5). LID x̄ =0.92 (N=5). CLL x̄ =0.36 (N=5). HL x̄ =1.50 (N=5). HW x̄ =1.71 (N=5). Head. Bluish green to turquoise. Labrum: black; apical elevation present, but small; base rugose. Mandible: black with apical amber spot occupying 1/3 of the mandible total length; preapical teeth present. Maxillary palpomeres length: 0.5 the size of MOD or less. Clypeus: punctate to sparsely punctate (I=PD; I&gt;PD); coriaceous; long, thin and sparse setae in the middle (3 MOD), and denser and thicker setae on apex; apical dark band less than 1/2 clypeal length, with a golden reflection on edge. Epistomal lobe: angle obtuse and not projected onto the clypeus. Supraclypeal area: sparsely punctate (I&gt;PD); surface lineolate between punctation. Parocular area: whitish setae, plumose and short to long (0.7–2 MOD). Antennae: scape black; flagellomeres black, yellowish anteriorly. Frons: slightly punctate, blending with the lineolate sculpture. Eye setae: short (0.5–0.7 MOD). Gena: plumose setae, short to long (0.5–4x MOD); micro lineolate with inconspicuous and sparse punctation; bluish green, with blue reflections. Mesosoma. Mesoscutum: sparsely punctate (I&gt;PD); microlineolate. Scutellum: microlineolate; very sparsely punctured on the disc, more densely posteriorly and to the sides. Mesepisternum: areolate on the hypoepimeral area and rugose on the preepisternum and below the scrobal sulcus. Metanotum: polished surface; densely to sparsely punctate; with a few sparse branched setae laterally. Metepisternum: coriaceous. Legs: coxa to femur dark brown, with green metallic reflections, tibia brown or yellow and tarsus yellow, with yellowish plumose setae. Metapostnotum: medium depression occupying most of the dorsal area; transverse striae lineations with semicircle shape. Propodeum: sparse and thin long plumose setae; pit carinate. Metasoma. Terga: green or bluish green anteriorly; black on disc; marginal area dark brown with few dark green reflections. Sterna: S1 dark metallic green; S2–S6 brownish; thin and sparse plumose setae.</p><p>Male: Body length x̄ =6.09 (N=5). ITD x̄ =1.09 (N=5). UID x̄ =1.00 (N=5). LID x̄ =0.70 (N=5). CLL x̄ =0.36 (N=5). HL x̄ =1.35 (N=5). HW x̄ =1.54 (N=5). Head. Metallic green, bluish on vertex, with goldish near the antennal sockets and below. Labrum:black, with apical elevation. Mandible: black; amber apical spot occupying around 1/3 of its length or a little more, sometimes with a yellow spot on the middle; preapical teeth present, but small. Maxillary palpomeres length: 0.5–1.0x MOD. Clypeus: punctate; coriaceous, polished near the disc yellow spot; black apical band occupying around 1/3 of the clypeal length, with a dark edge separating it from the metallic area. Epistomal lobe: angle obtuse and not projected onto clypeus. Supraclypeal area: coriaceous; sparsely punctate to punctate. Paraocular area:pubescence white, short and plumose, thicker on the frons, around the antennal socket and near the apex of clypeus, thinner elsewhere. Antennae: scape black with dark green reflections; flagellomeres black to light brown or yellow anteriorly, with a strong yellow spot on the anterior portion of F1, sometimes extending to more flagellomeres; crenulate, F2 longer than each F3– F10. Frons: coriaceous-rugulose; punctation inconspicuous among ornamentation. Eye setae: size 0.7–1.0x MOD. Gena: turquoise to goldish green below; coriaceous to polished; slightly punctate to densely punctate; long and thick white plumose setae. Mesosoma. Integument color: turquoise, with some goldish green on the sides of scutellum and on the metapostnotum. Mesoscutum: sparsely to very sparsely punctate, a little denser at the edges; coriaceous to polished, areolate on a few areas. Scutellum: polished, weakly lineolate on the edges; very sparsely punctate, crowded punctate near the edges. Mesepisternum: rugulose. Metanotum: coriaceous-punctate. Metepisternum: lineolate to rugulose. Legs: fore and hind coxa metallic green, middle coxa black, fore leg trochanter and femur metallic green, middle and hind leg trochanters and femora black with metallic green reflections, fore tibia and tarsi yellow, middle and hind tibiae dark brown with a few yellow spots, tarsi yellow, pretarsus brown; setae white to yellow. Metapostnotum: median depression occupying most of the dorsal area; transverse striae curved, semi-circle shaped; coriaceous to polished posteriorly. Propodeum: with dense, short and thin plumose setae; pit not carinate, enclosed, parallel. Metasoma. Terga: dark blue anteriorly; black on the disc; dark brown on the marginal area. Sterna: S1 black to dark brown, with green metallic reflections, dark brown elsewhere. Genitalia. Gonobase: wider than long, width equal to gonocoxite length. Volsella: enlarged. Gonapophysis: ventral prong as long as 3/4 of gonapophysis length; dorsal flanges small, starting near the apex, as wide as 1/2 of gonapophysis; with dorsal crest and dorsal ridge. Gonocoxite: longitudinally striate. Gonostylus: VA very reduced, inconspicuous; VP absent; MGL enlarged, globose, coriaceous; IGP small, almost inconspicuous; OGP with a few small setae on apex.</p><p>Etymology. Latin adjective arcuatus, meaning “arched” or “curved like a bow,” and refers to the distinctive curved striae present on the dorsum of the metapostnotum.</p><p>Distribution. Brazil: Paraná.</p><p>Type material. Holotype. Brazil, PR, DZUP. ♀, “ DZUP 605176 ”, “Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.993057&amp;materialsCitation.latitude=-25.23139" title="Search Plazi for locations around (long -49.993057/lat -25.23139)">Parque\ Estadual de Vila Velha</a>,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.993057&amp;materialsCitation.latitude=-25.23139" title="Search Plazi for locations around (long -49.993057/lat -25.23139)">Área I</a>, 25º13’53’’S,\ 49º59’35’’W, 1.IX.2023,\ Gonçalves, Graf, Pundek ” . Paratype. Brazil, PR, DZUP. ♀, “ DZUP 540104 ”,”Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.27548&amp;materialsCitation.latitude=-25.1393" title="Search Plazi for locations around (long -50.27548/lat -25.1393)">Ponta\ Grossa</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.27548&amp;materialsCitation.latitude=-25.1393" title="Search Plazi for locations around (long -50.27548/lat -25.1393)">P1 GIII</a>,\ 25.1393ºS, 50.27548ºW \ 26.VIII.2022, L.V.Graf,\ rede entomológica ”, ♀, “ DZUP 540105 ”, “ Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.27548&amp;materialsCitation.latitude=-25.1393" title="Search Plazi for locations around (long -50.27548/lat -25.1393)">Parque\ Estadual de Vilha Velha</a>,\ 25º14’S 49º59’W,\ 16.iii.2002, G. Melo &amp; R.B.\ Gonçalves ”, ♂, “ DZUP 540106 ”, “ Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.27548&amp;materialsCitation.latitude=-25.1393" title="Search Plazi for locations around (long -50.27548/lat -25.1393)">Parque\ Estadual de Vilha Velha</a>,\ 25º14’S 49º59’W,\ 23.xi.2001, G.A.R. Melo ”, ♀, “ PEVV0002 ”, “ Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.27548&amp;materialsCitation.latitude=-25.1393" title="Search Plazi for locations around (long -50.27548/lat -25.1393)">Parque\ Estadual de Vila Velha</a>,\ 25º14’S, 49º59’W,\ 05.x.2002, G. Melo,\ R. Gonçalves, A. Aguiar ”, ♀, “ PEVV0003 ”, “ Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.27548&amp;materialsCitation.latitude=-25.1393" title="Search Plazi for locations around (long -50.27548/lat -25.1393)">Parque\ Estadual de Vila Velha</a>,\ 25º14’S, 49º59’W,\ 05.x.2002, G. Melo,\ R. Gonçalves, A. Aguiar ”, ♀, “ PEVV0373 ”, “ Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.27548&amp;materialsCitation.latitude=-25.1393" title="Search Plazi for locations around (long -50.27548/lat -25.1393)">Parque\ Estadual de Vila Velha</a>,\ 25º14’S, 49º59’W,\ 12.xii.2002, G.A.R. Melo &amp;\ R.B. Gonçalves ”, ♀, “ PEVV2726 ”, “ Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.27548&amp;materialsCitation.latitude=-25.1393" title="Search Plazi for locations around (long -50.27548/lat -25.1393)">Parque\ Estadual de Vila Velha</a>,\ 25º14’S, 49º59’W,\ 17.viii.2004, G. Melo,\ R. Gonçalves ”, ♂, “ PEVV1446 ”, “ Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.27548&amp;materialsCitation.latitude=-25.1393" title="Search Plazi for locations around (long -50.27548/lat -25.1393)">Parque\ Estadual de Vila Velha</a>,\ 25º14’S, 49º59’W,\ 17.v.2003, Melo e\ Gonçalves ”, ♂, “ PEVV0784 ”, “ Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.27548&amp;materialsCitation.latitude=-25.1393" title="Search Plazi for locations around (long -50.27548/lat -25.1393)">Parque\ Estadual de Vila Velha</a>,\ 25º14’S, 49º59’W,\ 07.ii.2003, G. Melo &amp;\ R. Gonçalves ”, ♂, “ PEVV1454 ”, “ Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.27548&amp;materialsCitation.latitude=-25.1393" title="Search Plazi for locations around (long -50.27548/lat -25.1393)">Parque\ Estadual de Vila Velha</a>,\ 25º14’S, 49º59’W,\ 6.ix.2003, G.A.R. Melo \ &amp; R.B. Gonçalves ”, ♂, “ PEVV1494 ”, “ Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.27548&amp;materialsCitation.latitude=-25.1393" title="Search Plazi for locations around (long -50.27548/lat -25.1393)">Parque\ Estadual de Vila Velha</a>,\ 25º14’S, 49º59’W,\ 6.ix.2003, G.A.R. Melo \ &amp; R.B. Gonçalves ”, ♂, “ DZUP 606150 ”, “ Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.27548&amp;materialsCitation.latitude=-25.1393" title="Search Plazi for locations around (long -50.27548/lat -25.1393)">Parque\ Estadual de Vila Velha</a>,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.27548&amp;materialsCitation.latitude=-25.1393" title="Search Plazi for locations around (long -50.27548/lat -25.1393)">Área I</a>, 25º13’53’’S,\ 49º59’35’’W, 09.I.2024,\ Gonçalves, Graf, Pundek ”, ♂,“ DZUP605146 ”, “ Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.27548&amp;materialsCitation.latitude=-25.1393" title="Search Plazi for locations around (long -50.27548/lat -25.1393)">Parque\ Estadual de Vila Velha</a>,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.27548&amp;materialsCitation.latitude=-25.1393" title="Search Plazi for locations around (long -50.27548/lat -25.1393)">Área I</a>, 25º13’53’’S,\ 49º59’35’’W, 1.IX.2023,\ Gonçalves, Graf, Pundek ”, ♂, “ DZUP 605191 ”, “ Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.27548&amp;materialsCitation.latitude=-25.1393" title="Search Plazi for locations around (long -50.27548/lat -25.1393)">Parque\ Estadual de Vila Velha</a>,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.27548&amp;materialsCitation.latitude=-25.1393" title="Search Plazi for locations around (long -50.27548/lat -25.1393)">Área I</a>, 25º13’53’’S,\ 49º59’35’’W, 1.IX.2023,\ Gonçalves, Graf, Pundek ” .</p></div>	https://treatment.plazi.org/id/8C7BE1229E11FFEF76F019F42A35FD2E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cappola, Victor;Gonçalves, Rodrigo B.	Cappola, Victor, Gonçalves, Rodrigo B. (2026): A review of Caenohalictus species from southern Brazil (Apoidea, Caenohalictini). Zootaxa 5782 (3): 401-448, DOI: 10.11646/zootaxa.5782.3.1, URL: https://doi.org/10.11646/zootaxa.5782.3.1
8C7BE1229E1CFFEC76F01E812DECFB6E.text	8C7BE1229E1CFFEC76F01E812DECFB6E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caenohalictus curticeps (Vachal 1903)	<div><p>2. Caenohalictus curticeps (Vachal, 1903)</p><p>(Figs. 6, 7, 27A–C, 30B)</p><p>Halictus (Paragapostemon) curticeps Vachal, 1903 . Type data: holotype ♂ Type locality: Brazil. Naturhistorisches Museum in Wien. Examined through photographs.</p><p>Caenohalictus mourei Almeida &amp; Laroca, 2005 . Type data: holotype ♀ Type locality: Brazil, Paraná, Curitiba. Museu de Zoologia, Universidade de São Paulo. Examined. New synonym.</p><p>Diagnosis. Both sexes can be easily distinguished from small Caenohalictus species by the mesoscutum sculpture, being coriaceous and sparsely punctate along the median line, polished and punctate at sides (Fig. 6C, 7C). Additional features are the legs dark brown, slightly lighter on the tarsi. Females have dark blue reflections on terga and mesepisternum, and dark setae along the body, especially on the head (Fig. 6A). Males have a non-crenulate, short antennae (not reaching tegula, with F2–11 with similar length while in other species the antennae are crenulated and F2 usually elongate in relation to F3–11), and the scutellum has a median longitudinal sulcus with dense punctation.</p><p>Redescription</p><p>Female: Body length x̄ =7.20 (N=5). ITD x̄ =1.34 (N=5). UID x̄ =1.07 (N=5). LID x̄ =1.02 (N=5). CLL x̄ =0.39 (N=5). HL x̄ =1.57 (N=5). HW x̄ =1.80 (N=5). Head. Metallic green. Labrum: black; apical elevation present, but small; base rugose. Mandible: mostly black, with its apex dark amber and with a yellowish median stripe; preapical teeth present. Maxillary palpomeres length: 0.5 the size of MOD less. Clypeus: punctate (I=PD); slightly coriaceous; long and thin setae (3.5 MOD), with a comb of very short setae in the apex; apical dark band occupying less than 1/2 its length, with a golden reflection edge. Epistomal lobe: angle obtuse and not projected onto the clypeus. Supraclypeal area: sparsely punctulate (I&gt;PD); polished to coriaceous inferiorly. Paraocular area: setae dark to brownish and short. Antennae: scape and flagellomeres black. Frons: slightly punctate, blending with the lineolate sculpture. Eye setae: short (0.5–0.7 MOD). Gena: metallic turquoise; thin and sparse long plumose setae (2–4 MOD); slightly micro lineolate to polished; punctate. Mesosoma. Mesoscutum: sparsely punctate at the disc and anteriorly, densely punctated elsewhere; coriaceous at the disc and anteriorly, polished elsewhere. Scutellum: polished between punctation; very sparsely punctated in the disc, more densely posteriorly and to the sides; on the middle with a longitudinal crowded punctation. Mesepisternum: rugose to coriaceous ventrally. Metanotum: polished surface; densely punctated. Metepisternum: rugose. Legs: dark brown, with slightly lighter tarsi; fore leg with yellowish setae; hind leg with brownish setae. Metapostnotum: medium depression occupying most of the dorsal area; longitudinally sculptured, areolate-rugose, irregularly branched. Propodeum: sparse and thin long plumose setae; pit carinate, not enclosed above. Metasoma. Terga: green or bluish green anteriorly; black on the disc; with a small area between the black and marginal area green bluish green; marginal area dark brown. S1 dark brown. S2–S6 yellowish, brownish or dark brown; plumose setae, a little denser at S2 and S3.</p><p>Male: Body length x̄ =6.48 (N=5). ITD x̄ =1.18 (N=5). UID x̄ =0.93 (N=5). LID x̄ =0.68 (N=5). CLL x̄ =0.32 (N=5). HL x̄ =1.38 (N=5). HW x̄ =1.55 (N=5). Head. Metallic green turquoise. Labrum: yellow. Mandible: black; amber apical spot occupying around 1/3 of its length; lacking preapical teeth. Maxillary palpomeres length: 0.5– 0.7x MOD. Clypeus: punctate to densely punctate; polished to coriaceous above, near the upper edges; yellow spot occupying around 1/3 to 1/4 of the clypeal area, with a dark edge separating it from the metallic area. Epistomal lobe: angle approximately orthogonal, weakly projected onto the clypeus. Supraclypeal area: finely coriaceous; densely punctate. Parocular area: thin, short and brown to white plumose setae. Antennae: scape and flagellomeres black; not crenulate, F2 as long as each F3–F10. Eye setae: small (0.5–0.7x MOD). Gena: metallic blue or turquoise; coriaceous-rugulose; slightly punctate; sparse, long and thick plumose setae. Mesosoma. Integument color: turquoise, with blue metallic green reflections. Mesoscutum: sparsely punctate to punctate anteriorly and on the disc, densely punctate to crowded elsewhere; areolate to lineolate anteriorly and sometimes on the middle of the disc, polished elsewhere. Scutellum: polished; sparsely punctate on the disc to densely punctate elsewhere, on the middle with a longitudinally crowded punctation. Mesepisternum: rugose on the preepisternum, rugulose elsewhere. Metanotum: rugulose; slightly punctate. Metepisternum: rugose to areolate below. Legs: dark brown. Metapostnotum: median depression occupying most of the dorsal area; longitudinally rugose, branched; areolate dorsally, areolate-rugulose to rugulose sideward. Propodeum: sparse, short and thin plumose setae; pit not carinate, not enclosed above. Metasoma. Terga: entirely brown, with strong metallic blue reflections, almost looking like the real color. Sterna: black to dark brown. Genitalia. Gonobase: wider than long, width 2/3 of the gonocoxite length. Volsella: as long as 3/4 the gonapophysis, longer than wide. Gonapophysis: ventral prong reaching 3/4 of gonapophysis length; dorsal flanges starting at the middle of gonapophysis, with 1.5x length of gonapophysis; dorsal crest present. Gonocoxite: longitudinally striate. Gonostylus: VA with a crest on the lower edge; VP concave, merged with VA, with a crest on the middle; MGL reniform; IGP with a thick basal setose lobule; OGP with four long branched setae on apex.</p><p>Variation. Most of the females are green with a bluish metasoma, but some specimens are entirely blue or bluish-green. Males have bluish reflections.</p><p>Comments. Vachal (1903) described Halictus (Paragapostemon) curticeps based on a male specimen that was later included in Caenohalictus by Michener (1969b). More recently Caenohalictus mourei was described by Almeida &amp; Laroca (2005) based on a female. After the analysis of images of the holotype of H. curticeps and the holotype of C. mourei, they are considered as synonyms as the following characteristics are present in both sexes: antennae completely darkened; mesoscutum coriaceous anteriorly, polished elsewhere, with dense punctation, sparser on the disc; legs completely dark brown; sterna without metallic reflexes.</p><p>Distribution. Brazil: Espírito Santo, Paraná, Rio Grande do Sul, Santa Catarina, São Paulo.</p><p>Examined material. Brazil, ES, DZUP, ♀ “ DZUP 539911 ”, ”STA. TEREZA - E. STO\ BRASIL 2-6/I/68\ C. &amp; C. T. Elias leg.”, ♂, “ DZUP 539912 ”, ”Sta. TERESA - ES \ Brasil 12-X-64\ C. ELIAS leg”, ♂, “ DZUP 539913 ”, ”Sta. leopoldina - ES \ Brasil 4 - III - 64\ C. ELIAS, - LEG ”, ♂, ” DZUP 539914 ”, ”Sta. leopoldina - ES \ Brasil 4 - III - 64\ C. ELIAS, - LEG ”, ♂, ” DZUP 539915 ”, ”Sta. leopoldina - ES \ Brasil 4 - III - 64\ C. ELIAS, - LEG ”, ♀, ” DZUP 539916 ”, ”STA. TEREZA - E. STO\ BRASIL 23-31/ V/67 \ C. &amp; C. T. Elias leg.”, ♀, “ DZUP 539917 ”, “Sta. leopoldina - ES \ Brasil 4 - III - 64\ C. ELIAS, - LEG ”, ♂, “ DZUP 539918 ”, “ SANTA TERESA - ES \ BRASIL 14/4/1966 \ C. Tadeu Elias lg”, ♂, “ DZUP 539919 ”, “Sta. leopoldina - ES \ Brasil 4 - III - 64\ C. ELIAS, - LEG ”, ♂, “ DZUP 539920 ”, “ SANTA TERESA - ES \ BRASIL 27/9/1967 \ C. T. &amp; C.Elias lg”, ♂,” DZUP 539921 ”, “ SANTA TERESA - ES \ BRASIL 27/9/1967 \ C. T. &amp; C.Elias lg”, ♂, “ DZUP 539922 ”, “ SANTA TERESA - ES \ BRASIL / 18- 23/XII/1967 \ C. T. &amp; C.Elias ”, ♂, “ DZUP 539923 ”, “Sta. leopoldina - ES \ Brasil 4 - III - 64\ C. ELIAS, - LEG ”, ♂, “ DZUP 539924 ”, “CARIACICA - E. SANTO \ BRASIL 10-15/IV/67\ C. Elias leg.”, ♂, “ DZUP 540165 ”, “ SANTA TERESA - ES \ BRASIL 1823/XII /1967\ C. T. &amp; C. Elias ”, ♂, “ DZUP 540166 ”, “ SANTA TERESA - ES \ BRASIL - 22/11/1967 \ C. T. &amp; C. Elias leg”, ♀, “ DZUP 540167 ”, “ SANTA TERESA - ES \ BRASIL 1823/XII /1967\ C. T. &amp; C. Elias ”, ♀, “ DZUP 540168 ”, “ SANTA TERESA - ES \ BRASIL 18-23/XII/1967 \ C. T. &amp; C. Elias ”. PR, DZUP, ♂, “ DZUP 539881 ”, “CASTRO - PR - BRASIL\ 28/IV/73\ S. Laroca leg.”, ♂, “ DZUP 539882 ”, “CASTRO - PR - BRASIL \ 28/IV/73\ S. Laroca leg.”, ♂, “ DZUP 539883 ”, “Brasil, Paraná,\ Piraquara, Mananciais \ da Serra, 17.i.2004,\ G. Melo &amp; A. Aguiar ”, ♂, “ DZUP 539884 ”, “Brasil, Paraná,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.966667&amp;materialsCitation.latitude=-25.483334" title="Search Plazi for locations around (long -48.966667/lat -25.483334)">Piraquara</a>,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.966667&amp;materialsCitation.latitude=-25.483334" title="Search Plazi for locations around (long -48.966667/lat -25.483334)">Mananciais da Serra</a>,\ 8.ii.2003, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.966667&amp;materialsCitation.latitude=-25.483334" title="Search Plazi for locations around (long -48.966667/lat -25.483334)">Melo</a> ”, ♀, “ DZUP 539885 ”, “Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.966667&amp;materialsCitation.latitude=-25.483334" title="Search Plazi for locations around (long -48.966667/lat -25.483334)">Piraquara</a>,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.966667&amp;materialsCitation.latitude=-25.483334" title="Search Plazi for locations around (long -48.966667/lat -25.483334)">Mananciais da Serra</a>,\ 25º29’S 48º58’W,\ 24.iv.2002, G. AR. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.966667&amp;materialsCitation.latitude=-25.483334" title="Search Plazi for locations around (long -48.966667/lat -25.483334)">Melo</a> ”, ♀, “ DZUP 539886 ”, “ Brasil, Paraná,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.966667&amp;materialsCitation.latitude=-25.483334" title="Search Plazi for locations around (long -48.966667/lat -25.483334)">Piraquara</a>,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.966667&amp;materialsCitation.latitude=-25.483334" title="Search Plazi for locations around (long -48.966667/lat -25.483334)">Mananciais da Serra</a>,\ 06.ii.2007,\ A. Aguiar ”, ♀, “ DZUP 539887 ”, “Brasil, Paraná,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.966667&amp;materialsCitation.latitude=-25.483334" title="Search Plazi for locations around (long -48.966667/lat -25.483334)">Piraquara</a>,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.966667&amp;materialsCitation.latitude=-25.483334" title="Search Plazi for locations around (long -48.966667/lat -25.483334)">Mananciais da Serra</a>,\ 06.ii.2007,\ A. Aguiar ”, ♀, “ DZUP 539888 ”, “Brasil, Paraná,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.966667&amp;materialsCitation.latitude=-25.483334" title="Search Plazi for locations around (long -48.966667/lat -25.483334)">Piraquara</a>,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.966667&amp;materialsCitation.latitude=-25.483334" title="Search Plazi for locations around (long -48.966667/lat -25.483334)">Mananciais da Serra</a>,\ 06.ii.2007,\ A. Aguiar ”, ♂, “ DZUP 539890 ”, “CURITIBA-PARANÁ\ BRASIL- 23/9/1955 \ Moure &amp; Michener”, ♀, “ DZUP 539891 ”, “Curitiba - Pr.\ 20.III.87\ C. Mareze, leg”, ♀, “ DZUP 539892 ”, “Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.966667&amp;materialsCitation.latitude=-25.483334" title="Search Plazi for locations around (long -48.966667/lat -25.483334)">Campo</a> \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.966667&amp;materialsCitation.latitude=-25.483334" title="Search Plazi for locations around (long -48.966667/lat -25.483334)">Largo</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.966667&amp;materialsCitation.latitude=-25.483334" title="Search Plazi for locations around (long -48.966667/lat -25.483334)">Rio Verde</a>, 900m,\ 25º25’S 49º27’W,\ 09.x.2005, G. Melo ”, ♀, “ DZUP 539893 ”, “ Brasil, Paraná, Campo\ Largo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.966667&amp;materialsCitation.latitude=-25.483334" title="Search Plazi for locations around (long -48.966667/lat -25.483334)">Rio Verde</a>, 900m,\ 25º25’S 49º27’W,\ 09.x.2005, G. Melo ”, ♀, “ DZUP 539894 ”, “Brasil, Paraná, Campo\ Largo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.966667&amp;materialsCitation.latitude=-25.483334" title="Search Plazi for locations around (long -48.966667/lat -25.483334)">Rio Verde</a>, 900m,\ 25º25’S 49º27’W,\ 09.x.2005, G. Melo ”, ♀, “ DZUP 539895 ”, “Brasil, Paraná,\ Guarapuava,\ Pq. das Araucárias,\ 23.x.2003, G. Melo ”, ♀, “ DZUP 539896 ”, “Brasil, Paraná,\ Guarapuava,\ Pq. das Araucárias,\ 23.x.2003, G. Melo ”, ♀, “ DZUP 539897 ”, “ Brasil, Paraná,\ Guarapuava,\ Pq. das Araucárias,\ 23.x.2003, G. Melo ”, ♀, “ DZUP 539898 ”, “Brasil, Paraná,\ Guarapuava,\ Pq. das Araucárias,\ 23.x.2003, G. Melo ”, ♀, “ DZUP 539907 ”, “Brasil, PR, São Mateus\ do Sul, Fazenda Durgo\ 13.X.2009 \ Rafael Kamke leg.\\ Ponto amostral\ Fazenda Durgo\ 11:00 - 12:00\\ CJS 8139”, ♀, “ DZUP 539908 ”, “Brasil, PR, São Mateus\ do Sul, Fazenda Durgo\ 13.X.2009 \ Rafael Kamke leg.\\ Ponto amostral\ Fazenda Durgo\ 11:00 - 12:00\\ CJS 8138”, ♀, “ DZUP 539909 ”, “ Brasil, PR, São Mateus\ do Sul, Fazenda Durgo\ 13.X.2009 \ Rafael Kamke leg.\\ Ponto amostral\ Fazenda Durgo\ 11:00 - 12:00\\ CJS 8140”, ♂, “ DZUP 540157 ”, “CASTRO - PR - BRASIL\ 27/IV/73\ S. Laroca leg.”. RS, DZUP, ♀, “ DZUP 540158 ”, “Brasil, RS, Canguçu\ Propriedade 4 Orgânica\ Mata Nativa\ 31º29’34.81”S, 52º42’49.84”W \ 25.i.2023 \ Kurz, M.”.SC, DZUP, ♀, “ DZUP 539889 ”, “Brasil, Santa Catarina,\ Pomerode, 21.iv.2007,\ Melo, Aguiar, Martins &amp;\ Gonçalves”, ♂, “ DZUP 539899 ”, “1960\ Brasilien \ Nova Teutonia \ 27º11’ S 52º 23’ L\ Fritz Plaumann ”, ♂, “ DZUP 539900 ”, “1950\ Brasilien\ Nova Teutonia\ 27º11’ S 52º 23’ L\ Fritz Plaumann”, ♂, “ DZUP 539901 ”, “ Nova Teutonia,\ Santa Caterina,\ Brazil I-1952 \ L. E. Plaumann ”, ♀, “ DZUP 539902 ”, “ N. Teutônia - SC\ Brasil 28 X-1955 \ A. Maller, leg”, ♀, “ DZUP 539903 ”, “ N. Teutônia. SC\ Br. 9/I/1956 \ Plaumann leg”, ♂, “ DZUP 539904 ”, “ N. Teutônia. SC \ Br. 9/I/1956 \ Plaumann leg”, ♂, “ DZUP 539905 ”, “ N. Teutônia. SC\ Br. 9/I/1956 \ Plaumann leg”, ♂, “ DZUP 539906 ”, “N. TEUTONIA - SC\ BRASIL- 24 /I/957\ F. Plaumann leg”. SP, DZUP, ♂, “ DZUP 539910 ”, “Ribeirao Pires,\ Sao Paulo, Brazil \ VI - 1954 \ N.L.H.KRAOSS” .</p></div>	https://treatment.plazi.org/id/8C7BE1229E1CFFEC76F01E812DECFB6E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cappola, Victor;Gonçalves, Rodrigo B.	Cappola, Victor, Gonçalves, Rodrigo B. (2026): A review of Caenohalictus species from southern Brazil (Apoidea, Caenohalictini). Zootaxa 5782 (3): 401-448, DOI: 10.11646/zootaxa.5782.3.1, URL: https://doi.org/10.11646/zootaxa.5782.3.1
8C7BE1229E1FFFE976F019412AA5F883.text	8C7BE1229E1FFFE976F019412AA5F883.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caenohalictus cyanolucens Cappola & Gonçalves 2026	<div><p>3. Caenohalictus cyanolucens sp. nov.</p><p>(Figs. 3D, 8–9, 26C, 29D–F, 31B)</p><p>Diagnosis. This species can be separated from other species by the combination of short maxillary palpomeres and obtuse epistomal angle with the mesoscutum and metapostnotum granulate. Another diagnostic feature is the bright metallic blue gena, which contrasts with the rest of the integument (Fig. 8A, 9A). The following features are relevant to species diagnosis: a deep, areolate mesepisternal sulcus and male flagellomeres 2–11 with anterior large patches of placodea sensillae (Fig. 3D).</p><p>Description</p><p>Female: Body length x̄ =8.24 (N=2). ITD x̄ =1.44 (N=2). UID x̄ =1.22 (N=2). LID x̄ =1.18 (N=2). CLL x̄ =0.34 (N=2). HL x̄ =1.65 (N=2). HW x̄ =2.00 (N=2). Head. Black, with a few olivaceous spots, around the antennal socket, on the upper portion of the clypeus, on the upper inner orbit of the eye and around the occiput. Labrum: black; apical elevation presente; base rugose. Mandible: black; apical amber spot occupying 1/5 of the mandible total length; mandible apex truncated, lacking preapical teeth. Maxillary palpomeres length: 0.7 MOD or less. Clypeus: punctate to densely punctate (I=PD; i&lt;pd); granulate-areolate to granulate-lineolate; long (2–4 MOD), thin and very sparse plumose setae; middle of the apical dark band going to almost one half of the clypeus, with a golden and purple reflection on edges. Epistomal lobe: angle obtuse and not projected onto the clypeus. Supraclypeal area: sparsely punctated (I&gt;PD); areolate to micro lineolate. Parocular area: yellow to white setae, plumose and long (2–3 MOD), thick around the antennal socket and thin elsewhere.Antennae: scape and flagellomeres black to dark brown. Frons: slightly punctate, mixing with the areolate ornamentation. Eye setae: very short, almost inconspicuous (0.3–0.1 MOD). Gena: slightly punctate; sparse, long and thick plumose setae; rugose-lineolate; metallic blue above, dark green below. Mesosoma. Mesoscutum: sparsely to very sparsely punctate, with a few punctate to densely punctate on the edges; granulate-areolate to granulate-lineolate. Scutellum: micro lineolate; very sparsely punctate. Mesepisternum: areolate on the hypoepimeral area and rugose to rugulose on the preepisternum, lineolate-rugulose below the scrobal sulcus. Metanotum: rugulose-punctulate. Metepisternum: areolate to micro lineolate. Legs: black to dark brown, tarsi light brown with a few yellowish spots; setae yellow and brown. Metapostnotum: medium depression occupying most of the dorsal area; longitudinal striae, areolate-rugose, irregularly; granulate-areolate. Propodeum: pit not carinate, not enclosed. Metasoma. Terga: black on the disc, with dark green reflections near the marginal area; marginal area brown, with a strong white reflection on the apex. Sterna: S1 dark brown; S2–S6 brown to light brown on the pre gradular area and posterior marginal area, dark brown on the post gradular area; dense and thin plumose setae on the post gradular area, glabrous on the pre gradular area.</p><p>Male: Body length x̄ =7.78 (N=3). ITD x̄ =1.27 (N=4). UID x̄ =1.12 (N=4). LID x̄ =0.76 (N=4). CLL x̄ =0.38 (N=4). HL x̄ =1.58 (N=4). HW x̄ =1.79 (N=4). Head. Goldish green below the antennal sockets and dark turquoise above. Labrum: yellow. Mandible: black; amber apical spot occupying around 1/3 of its length. Maxillary palpomeres length: 0.5–1.0x MOD. Clypeus: punctate to sparsely punctate; coriaceous to areolate; yellow spot occupying around 1/2 of the clypeal area, with a dark edge separating it from the metallic area. Epistomal lobe: angle obtuse and not projected onto the clypeus. Supraclypeal area: coriaceous to areolate; punctate to sparsely punctate. Parocular area: yellow to white plumose, thick and long setae, on the clypeus a little thinner. Antennae: scape black with very weakly blue reflections, sometimes with a few yellow spots; flagellomeres black to yellow anteriorly, except on the last ones, that are dark brown anteriorly, can vary from F7–11 to F10–11, crenulate, F2 longer than each F3– F10, flagellomeres 2–11 with anterior large patches of sensilla placodea. Frons: areolate-rugulose; inconspicuous punctation, blending with the sculpture. Eye setae: 0.7–1.0x MOD. Gena: metallic blue to turquoise; rugose; slightly punctate; sparse, long and thick plumose setae. Mesosoma. Integument color: dark green on the pronotum, mesoscutum, scutellum and metanotum, sometimes with some dark blue tones except on the scutellum; dark blue on the median depression of metapostnotum, on the sides and ventrally of mesosoma. Mesoscutum: punctate to sparsely punctate; granulate-areolate to coriaceous. Scutellum: lineolate to coriaceous; sparsely punctated to very sparsely punctated. Mesepisternum: areolate-rugulose to lineolate. Metanotum: coriaceous; slightly and sparsely punctate. Metepisternum: areolate-rugulose to lineolate. Legs: coxa black with dark bluish green reflections, trochanter and femur black with green reflections, fore leg tibia dark brown to yellow, yellow elsewhere, middle leg tibia dark brown with yellow apex, yellow elsewhere, hind leg tibia black, yellow elsewhere; legs with white to yellow plumose setae. Metapostnotum: median depression occupying most of the dorsal area; striae areolate-rugose; granulate-areolate. Propodeum: with sparse, short and thick plumose setae; pit not carinate, not enclosed above, sometimes weakly. Metasoma. Terga: mostly black, with a few dark green tones, mostly anteriorly; marginal area dark brown. Sterna: S1 dark brown, with dark blue reflections, dark brown elsewhere. Genitalia. Gonobase: wider than long, as long as 2/3 the gonocoxite. Volsella: as long as 1/2 the gonapophysis, as wide as long. Gonapophysis: ventral prong present; dorsal flanges starting almost at the apex, as wide as 2x the gonapophysis length; dorsal outer tooth present; dorsal crest present; dorsal depressions and ridge absent. Gonocoxite: longitudinally striate. Gonostylus: VA long lobe, thicker at the base to the middle, and thinner from the middle to the apex, with small setae on its apex; VP small lobe merged with VA, directed upwards; MGL reniform to arcuated, concave; IGP with a basal setose lobule, with thick and small setae; OGP with 5–6 long branched setae on apex.</p><p>Etymology. Cyanolucens is compounded from the Greek cyanos (blue) and the Latin lucens (shining, gleaming), referring to the metallic blue shining gena, which contrasts with the predominantly green integument on the remainder of the body.</p><p>Variation. There are a few female specimens that have black integument and sometimes with the blue reflections being almost imperceptible.</p><p>Distribution. Brazil: Paraná, Rio Grande do Sul, Santa Catarina, São Paulo.</p><p>Type material. Holotype. Brazil, PR, DZUP. ♂, “ DZUP 540123 ”, “Brasil, Paraná,\ Palmas, REVIS,\ 26.557ºS 51.542ºW,\ 24.I.2013, A.Pereira-\ Rocha, Arm. Malaise ”. Paratype. Brazil, PR, DZUP. ♂, “ DZUP 540122 ”, “Brasil, Paraná,\ Palmas, REVIS,\ 26.557ºS 51.542ºW,\ 24.I.2013, A.Pereira-\ Rocha, Arm. Malaise ”, ♂, “ DZUP 547051 ”, “ Brasil, Paraná, Curitiba,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.28276&amp;materialsCitation.latitude=-25.378448" title="Search Plazi for locations around (long -49.28276/lat -25.378448)">Parque Tanguá</a>,\ -25.378448, -49.282760,\ 10.II.2018, L.Graf ”, ♂, “ DZUP 547050 ”, “ Brasil, Paraná, Curitiba,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.28276&amp;materialsCitation.latitude=-25.378448" title="Search Plazi for locations around (long -49.28276/lat -25.378448)">Parque Tanguá</a>,\ -25.378448, -49.282760,\ 10.II.2018, L.Graf ”. Brazil, SC, DZUP. ♀, “ DZUP 029307 ” “Brasil, Santa Catarina, Bom\ Jardim da Serra, Alt. 1355 m,\ 28 22 84S., 49 38 32W\ 5.xi.2006, A.J.C. Aguiar, A. \ Martins, L.R.R. Faria Jr.”. Brazil, SP, DZUP. ♀, “ DZUP 540124 ”,” Brasil, SP, 10 km a SW\ de S. J. Barreiro, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.642223&amp;materialsCitation.latitude=-22.708" title="Search Plazi for locations around (long -49.642223/lat -22.708)">Serra\ da Bocaina</a>, 1690m,\ 22.708ºS 44.635ºW,\ 23.x.2011, G. Melo ”, ♂, “ DZUP 540125 ”, “ Brasil, SP, 10 km a SW\ de S. J. Barreiro, 1585m,\ 22º43’32’’S 44º37’33’’W,\ 29.iii.2010, Melo &amp; Luz ”. Brazil, RS, MCT. ♀, “ S. F. de Paula, RS, Brasil\ Pró-Mata 7.10.1995 \ leg. Birgit Harter \\ 17649 H: 701\ F: 85 12:00\\ Caenohalictus \ obscuros sp. n.\ Moure det. 1999\\ Caenohalictus \ cyanolucens\ Gonçalves det.”, ♀, “ São Francisco de Paula \ RS, Brasil \ Pró-Mata 5.10.1996 \ leg. Birgit Harter \\ 11583 H: 701\ F: 85 13:00 13:30\\ Caenohalictus \ obscuros sp. n.\ Moure det. 1999”, ♀, “ São Francisco de Paula \ RS, Brasil \ Pró-Mata 4.10.1996 \ leg. Birgit Harter \\ 11478 H: 701\ F: 85 10:00 11:00\\ Caenohalictus \ obscuros sp. n.\ Moure det. 1999”, ♀, “ São Francisco de Paula \ RS, Brasil \ Pró-Mata 4.10.1996 \ leg. Birgit Harter \\ 11479 H: 701\ F: 85 10:00 11:00\\ Caenohalictus \ obscuros sp. n.\ Moure det. 2000”, ♀, “ São Francisco de Paula \ RS, Brasil \ Pró-Mata 4.10.1996 \ leg. Birgit Harter \\ 11448 H: 728\ F: 86 12 80 ♂\\ Caenohalictus \ obscuros sp. n.\ Moure det. 2001”, ♀, “ São Francisco de\ Paula, Pró-Mata, RS,\ Brasil 12.09.1997 \ leg. Birgit Harter \\ 13857 H: 701\ F: 85 13 20 141\ 21ºC\\ Caenohalictus \ obscuros sp. n.\ Moure det. 1999”, ♀, “ S. F. de Paula, RS, Brasil \ Pró-Mata 7.10.1995 \ leg. Birgit Harter \\ 12898 H: 701\ F: 85 12 00 13 00\\ Caenohalictus \ obscuros sp. n.\ Moure det. 1999”, ♀, “ São Francisco de\ Paula, Pró-Mata, RS,\ Brasil 12.09.1997 \ leg. Birgit Harter \\ 13813 H: 701\ F: 85 12:00-\ 12:30\\ Caenohalictus \ obscuros sp. n.\ Moure det. 1999”, ♀, “ São Francisco de\ Paula, Pró-Mata, RS,\ Brasil 12.09.1997 \ leg. Birgit Harter \\ 17650 H: 701\ F: 85 12:00 25ºC\\ Caenohalictus \ obscuros sp. n.\ Moure det. 1999”, ♀, “ São Francisco de\ Paula, Pró-Mata, RS,\ Brasil 27.09.1997 \ leg. Birgit Harter \\ 14139 H: 701\ F: 85 11:00 19ºC\\ Caenohalictus \ obscuros sp. n.\ Moure det. 1999”, ♀, “ São Francisco de\ Paula, Pró-Mata, RS,\ Brasil 12.09.1997 \ leg. Birgit Harter \\ 14122 H: 701\ F: 85 II 13 20 14 00\\ Caenohalictus \ obscuros sp. n.\ Moure det. 1999”, ♀, “ São Francisco de\ Paula, Pró-Mata, RS,\ Brasil 12.09.1997 \ leg. Birgit Harter \\ 13871 H: 701\ F: 85 12-12:30 25ºC\\ Caenohalictus \ obscuros sp. n.\ Moure det. 1999”, ♂, “ São Francisco de\ Paula, Pró-Mata, RS,\ Brasil 26.2.1997 \ leg. Birgit Harter \\ 14780 H: 538\ F: 85 12:30-13:00\\ Caenohalictus \ obscuros sp. n.\ Moure det. 1999”, ♀, “ São Francisco de\ Paula, Pró-Mata, RS,\ Brasil 12.10.1998 \ leg. Birgit Harter \\ 16734 H: 701\ F: 85 11-11:30”, ♀, “ São Francisco de\ Paula, Pró-Mata, RS,\ Brasil 12/10/1998 \ leg. Birgit Harter \\ 16866 H: 701\ F: 85 11-11:30”, ♀, “ São Francisco de\ Paula, Pró-Mata, RS,\ Brasil 12.10.1998 \ leg. Birgit Harter \\ 16904 H: 701\ F: 85 10-10:30”, ♀, “ São Francisco de\ Paula, Pró-Mata, RS,\ Brasil 12.10.1998 \ leg. Birgit Harter \\ 16671 H: 701\ F: 85 11-11:30”, ♀, “ São Francisco de\ Paula, Pró-Mata, RS,\ Brasil 12.10.1998 \ leg. Birgit Harter \\ 16765 H: 701\ F: 85 11-11:30”, ♀, “ São Francisco de\ Paula, Pró-Mata, RS,\ Brasil 12.10.1998 \ leg. Birgit Harter \\ 16578 H: 701\ F: 85 11-11:30”, ♀, “ São Francisco de\ Paula, Pró-Mata, RS,\ Brasil 12.10.1998 \ leg. Birgit Harter \\ 16733 H: 701\ F: 85 11-11:30”, ♀, “ São Francisco de\ Paula, Pró-Mata, RS,\ Brasil 12.10.1998 \ leg. Birgit Harter \\ 16903 H: 701\ F: 85 10-10:30”, ♀, “ São Francisco de\ Paula, Pró-Mata, RS,\ Brasil 12/10/1998 \ leg. Birgit Harter \\ 16757 H: 701\ F: 85 11-11:30”, ♀, “ São Francisco de\ Paula, Pró-Mata, RS,\ Brasil 12.10.1998 \ leg. Birgit Harter \\ 16768 H: 701\ F: 85 11-11:30”, ♀, “ São Francisco de\ Paula, Pró-Mata, RS,\ Brasil 12/10/1998 \ leg. Birgit Harter \\ 16639 H: 701\ F: 85 11-11:30”, ♀, “ São Francisco de\ Paula, Pró-Mata, RS,\ Brasil 12/10/1998 \ leg. Birgit Harter \\ 16863 H: 701\ F: 85 11-11:30”, ♀, “ São Francisco de\ Paula, Pró-Mata, RS,\ Brasil 12/10/1998 \ leg. Birgit Harter \\ 16751 H: 701\ F: 85 11-11:30”, ♀, “ São Francisco de\ Paula, Pró-Mata, RS,\ Brasil 12/101998\ leg. Birgit Harter \\ 16755 H: 701\ F: 85 11-11:30”, ♂, “ São Francisco de\ Paula, Pró-Mata, RS,\ Brasil 12.04.1998 \ leg. Birgit Harter \\ 17433 H: 901\ F: 86 10:30-11:00♂\\ Caenohalictus \ obscuros\ sp. nova.\ Moure det. 1999”, ♂, “ São Francisco de\ Paula, Pró-Mata, RS,\ Brasil 12.04.1998 \ leg. Birgit Harter \\ 16022 H: 250\ F: 86 11:30-\12:00\\ Caenohalictus \ obscuros sp. n.♂\ Moure det. 1999”.</p></div>	https://treatment.plazi.org/id/8C7BE1229E1FFFE976F019412AA5F883	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cappola, Victor;Gonçalves, Rodrigo B.	Cappola, Victor, Gonçalves, Rodrigo B. (2026): A review of Caenohalictus species from southern Brazil (Apoidea, Caenohalictini). Zootaxa 5782 (3): 401-448, DOI: 10.11646/zootaxa.5782.3.1, URL: https://doi.org/10.11646/zootaxa.5782.3.1
8C7BE1229E1AFFF676F01B2D2D50F9D6.text	8C7BE1229E1AFFF676F01B2D2D50F9D6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caenohalictus implexus Moure 1950	<div><p>4. Caenohalictus implexus Moure, 1950</p><p>(Figs. 1A, 10–11, 28D–F, 30C)</p><p>Caenohalictus implexus Moure, 1950 . Type data: holotype ♀ Type locality: Paraguay, San Pedro, San Estanislao. Museo Argentino de Ciencias Naturales. Not examined.</p><p>Diagnosis. Small species (5–6 mm) similar to C. tesselatus and C. leticiae sp. nov. by the rugulose metapostnotum and olivaceus green body. It can be differentiated from C. tesselatus by black scape on both sexes (Fig. 10B, 11B), the lack of dense, yellow branched setae on mesoscutum of females (Fig. 10C) and S1 with few metallic reflections. Females can be separated from C. leticiae sp. nov. by the scutellum areolate and sparsely punctate (Fig. 10C), tarsi light brown to yellow and head as long as wide (Fig. 10B); mesepisternum and metanotum rugulose. Males differ in having flagellomeres 2–11 homogeneously covered by intermixed trichodea and placodea sensillae (Fig. 1A), mesoscutum entirely very sparsely punctate and propodeum lateral surface lineolate. The male is described here for the first time.</p><p>Redescription</p><p>Female: Body length x̄ =6.09 (N=5). ITD x̄ =1.19 (N=5). UID x̄ =0.93 (N=5). LID x̄ =0.88 (N=5). CLL x̄ =0.32 (N=5). HL x̄ =1.44 (N=5). HW x̄ =1.56 (N=5). Head. Metallic green, with golden reflections. Labrum: black; apical elevation present, but small; base rugose. Mandible: mostly black, with its apex dark amber and with a yellowish median stripe; preapical teeth present. Maxillary palpomeres length: 0.5 times MOD or less. Clypeus: punctate to sparsely punctate (I=PD; I&gt;PD); areolate; short, thin and sparse setae in the middle (1.5 MOD), longer on the apex, with a comb of very short setae; apical dark band occupying less than 1/2 its length, with a golden reflection edge. Epistomal lobe: angle obtuse and not projected onto the clypeus. Supraclypeal area: sparsely punctated (3–4 pd); areolate. Parocular area: yellowish setae, plumose and long (1.5–2.5 MOD). Antennae: scape black; flagellomeres black, yellowish anteriorly. Frons: punctuate to densely punctate in a few areas; areolate. Eye setae: short (0.5–0.7 MOD). Gena: metallic green; thin and sparse long plumose setae (2–4 MOD); areolate; sparsely punctate to punctate. Mesosoma. Mesoscutum: sparsely punctated (I&gt;PD); areolate. Scutellum: areolate; sparsely punctated to punctate. Mesepisternum: rugose on the preepisternum and on the hypoepimeral area, areolate below the scrobal sulcus. Metanotum: rugulose-punctulate. Metepisternum: rugose. Legs: coxa to femur with metallic green reflections, tibia and tarsus light brown or yellow; yellowish plumose setae. Metapostnotum: medium depression occupying most of the dorsal area; longitudinally sculptured, areolate-rugose, irregularly or branched. Propodeum: sparse and thin long plumose setae; pit weakly carinate, not enclosed above. Metasoma. Terga: mostly metallic green or goldish green, sometimes with a black area in the disc of T1, T2, T3 and T4; marginal area brown or greenish brown. Sterna: S1 brownish with green metallic reflections; S2–S6 brownish, with green metallic reflections; thin, long and yellowish plumose setae.</p><p>Description</p><p>Male: Body length x̄ =5.64 (N=5). ITD x̄ =0.96 (N=5). UID x̄ =0.84 (N=5). LID x̄ =0.53 (N=5). CLL x̄ =0.28 (N=5). HL x̄ =1.30 (N=5). HW x̄ =1.35 (N=5). Head. Cuprous or metallic green. Labrum: yellow. Mandible: mostly yellow, only a small anterior portion being black, with amber apical spot occupying around 1/4 of its length; lacking preapical teeth. Maxillary palpomeres length: 0.5–1.0x MOD. Clypeus: punctate to densely puncate; finely coriaceous; yellow spot occupying around 1/3 of the clypeal area, with a dark edge separating it from the metallic area. Epistomal lobe: angle obtuse and not projected onto the clypeus. Supraclypeal area: coriaceous; punctate. Parocular area: dense, short, thick and white plumose setae, on the clypeus and supraclypeal area with thinner and yellow setae. Antennae: scape black with weak dark green reflections; flagellomeres black to yellow anteriorly; crenulate, F2 longer than each F3–F10. Frons: areolate-rugulose; inconspicuous punctate, blending with the sculpture. Eye setae: small (0.5–0.7x MOD). Gena: same color of the integument; coriaceous to rugulose; punctate to sparsely punctate; long and thick white plumose setae, gradually becoming thin and short above. Mesosoma. Integument color: cuprous or metallic green. Mesoscutum: sparsely punctate to densely punctate on the lateral and posterior edges; coriaceous to polished, areolate on a few areas. Scutellum: polished to coriaceous on the edges; confused punctate. Mesepisternum: confused rugulose. Metanotum: rugulose; punctate to densely punctate. Metepisternum: confused rugulose. Legs: fore coxa metallic green, middle and hind coxa dark brown, hind coxa with green reflections, legs dark brown, with green reflections on trocanter and femur, fore tibia yellow anteriorly, middle and hind tibia with an apical yellow spot, basitarsus and tarsi yellow. Metapostnotum: median depression occupying most of the dorsal area; areolate-rugose; micro lineolate, rugulose anteriorly on the lateral. Propodeum: sparse, short and thin plumose setae; pit weakly carinate, not enclosed above. Metasoma. Terga: metallic green anteriorly; black area starting around in the middle of the terga on T1–T3, T4 onwards occupying almost all of the area; marginal area dark brown. Sterna: S1 dark brown with green reflections, dark brown elsewhere. Genitalia. Gonobase: wider than long, as wide as gonocoxite length. Volsella: enlarged, as long as gonapophysis, wider than long. Gonapophysis: ventral prong present; small concave dorsal flanges, with the length a little smaller than gonapophysis; dorsal depressions and dorsal ridge present; dorsal crest present. Gonocoxite: longitudinally striate. Gonostylus: VA short and with an oblong lobe with small setae on its apex. VP absent. MGL sigmoid. IGP with a dense and thick basal setose plate. OGP with a few small setae on apex and a dorsal setose oblong lobe.</p><p>Comments. The female paratype deposited in DZUP was examined.</p><p>Variation. Females and males can vary in color due the presence and extension of cupreous reflexes.</p><p>Distribution. Brazil: Goiás, Minas Gerais, Paraná, São Paulo. Paraguay.</p><p>Examined material. Brazil, MG, DZUP. ♀, “ DZUP 540265 ”, ”PASSOS - MG Brasil\ 5-10-XI-1961 \ C. Elias leg”, ♀, “ DZUP 540266 ”, ”PASSOS - MG Brasil \ 5-10-XI-1961 \ C. Elias leg”, ♀, “ DZUP 540267 ”, ”PASSOS - MG Brasil \ 5-10-XI-1961 \ C. Elias leg”, ♂, “ DZUP 540268 ”, ”PASSOS - MG Brasil\ 5-10-XI-1961 \ C. Elias leg”, ♂, “ DZUP 540269 ”, ” IBIÁ - M. Gerais \ BRASIL - 17/12/65\ C.Elias leg”, ♂, “ DZUP 540270 ”, ”PASSOS - MG\ BR 02-07 - VIII - 62\ Claudionor Elias”, ♀, “ DZUP 540271 ”, ” POÇOS CALDAS \ MG - Brasil XI 61\ Claudionor Elias”, ♂, “ DZUP 540272 ”, ” POÇOS CALDAS \ MG - Brasil XI 61\ Claudionor Elias”, ♂, “ DZUP 540273 ”, ”IBIÁ - BRASIL \ MG- 20/10/1965 \ C.T. &amp; C. Elias ”, ♂, “ DZUP 540274 ”, ”IBIÁ - M. Gerais \ BRASIL - 17/12/65\ C.Elias leg”, ♂, “ DZUP 540275 ”, ”IBIÁ - BRASIL \ MG- 10/12/1965 \ C. Elias leg”, ♂, “ DZUP 540276 ”, ”IBIÁ - BRASIL \ MG- 10/12/1965 \ C. Elias leg”, ♂, “ DZUP 540277 ”, ” PATOS de MINAS - MG \ BRASIL - 23/11/965\ Claudionor Elias”, ♂, “ DZUP 540278 ”, ” PATOS de MINAS - MG \ BRASIL - 23/11/965\ Claudionor Elias” . Brasil, PR, DZUP. ♀, “ DZUP 539961 ”, “FOZ DO IGUAÇU\ Parana Brasil \ 7.XII.1966 \ Exc.Dept. ZOO”, ♀, “ DZUP 539962 ”,” FOZ do IGUAÇU-PR \ Brasil 15.XII.1965 \ V. Graf - L. Azevedo ”, ♀, “ DZUP 539963 ”, “ FOZ DO IGUAÇU \ Parana Brasil \ 7.XII.1966 \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-51.47541&amp;materialsCitation.latitude=-24.53696" title="Search Plazi for locations around (long -51.47541/lat -24.53696)">Exc.Dept.</a> ZOO”, ♀, “ DZUP 539964 ”,” FOZ DO IGUAÇU \ Parana Brasil \ 7.XII.1966 \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-51.47541&amp;materialsCitation.latitude=-24.53696" title="Search Plazi for locations around (long -51.47541/lat -24.53696)">Exc.Dept.</a> ZOO”, ♀, “ DZUP 539965 ”, “ FOZ DO IGUAÇU \ Parana Brasil \ 7.XII.1966 \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-51.47541&amp;materialsCitation.latitude=-24.53696" title="Search Plazi for locations around (long -51.47541/lat -24.53696)">Exc.Dept.</a> ZOO”, ♀, “ DZUP 539966 ”, “ FOZ do IGUAÇU-PR \ Brasil 15.XII.1965 \ V. Graf - L. Azevedo ”, ♀, “ DZUP 539967 ”, “ Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-51.47541&amp;materialsCitation.latitude=-24.53696" title="Search Plazi for locations around (long -51.47541/lat -24.53696)">Manoel Ribas</a>,\ P1 GI, 24.53696ºS,\ 51.475409ºW, 01.III.2022,\ L.V.Graf, rede entomológica”, ♂, “ DZUP 539969 ”, “ Brasil, PR, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-51.47541&amp;materialsCitation.latitude=-24.53696" title="Search Plazi for locations around (long -51.47541/lat -24.53696)">Imbituva</a>,\ Ponto P1-GVI, 837m,\ 25.1453ºS 50.5075ºW,\ 22.xi.2020, A. Martins \ &amp; D. Muniz, Busca ativa”, ♂, “ DZUP 539970 ”, “ Brasil, Paraná, Castro,\ -24.7594525, -50.0965344,\ 28.V.2021, L.Graf ”, ♂, “ DZUP 539971 ”, “ Brasil, Paraná, São\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-51.47541&amp;materialsCitation.latitude=-24.53696" title="Search Plazi for locations around (long -51.47541/lat -24.53696)">João do Triunfo</a>, P2 GIII,\ 25.6375ºS 50.3109ºW,\ 27.V.2022, L.V.Graf,\ rede entomológica”, ♂, “ DZUP 539972 ”, “ Brasil, Paraná, São\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-51.47541&amp;materialsCitation.latitude=-24.53696" title="Search Plazi for locations around (long -51.47541/lat -24.53696)">Mateus do Sul</a>, SIX, CV \ -25.866367, -50.409012,\ 09.III.2021, L.Graf ”, ♂, “ DZUP 539973 ”, “ Brasil, Paraná, São\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-51.47541&amp;materialsCitation.latitude=-24.53696" title="Search Plazi for locations around (long -51.47541/lat -24.53696)">Mateus do Sul</a>, SIX, BRA \ -25.848244, -50.416959,\ 11.III.2021, L.Graf ”, ♂, “ DZUP 539974 ”, “ Brasil, PR, São Mateus\ do Sul, UN-SIX/Petrobras\ 20.III.2012 \ Rafael Kamke leg.\\ Ponto amostral\ MATA DO TÚNEL \\ CJS 11712 ”, ♂, “ DZUP 539975 ”, “ Brasil, PR, São Mateus\ do Sul, UN-SIX/ Petrobras\ 16.IV.2013 \ Rafael Kamke leg.\\ Ponto amostral\ VACÓDROMO\\ CJS 13494”, ♂, “ DZUP 606936 ”, “ Brasil, Paraná, Parque\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-51.47541&amp;materialsCitation.latitude=-24.53696" title="Search Plazi for locations around (long -51.47541/lat -24.53696)">Estadual de Vila Velha</a>,\ Área I, 25º13’53’’S,\ 49º59’35’’W, 1.V.2024,\ Gonçalves, Graf, Pundek” . Paraguay, DZUP. ♂, “ DZUP 540178 ”, “Colony, Carlos\ Pfannl, Paraguay\ II-51\ F. H. Shade ” . Brasil, SP, DZUP. ♀, “ DZUP 540243 ”, ” Brasil, São Paulo, Cajuru,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15&amp;materialsCitation.latitude=-21.2" title="Search Plazi for locations around (long -49.15/lat -21.2)">Fazenda Rio Grande</a>,\ 21°12’S 49°09’W, 25.iii.2000 \ G.A.R. Melo”, 20 ♀, DZUP 540245 - DZUP 540264, “Cajuru-SP\ 30 / XII / 85\ Moure &amp; Camargo col.” .</p></div>	https://treatment.plazi.org/id/8C7BE1229E1AFFF676F01B2D2D50F9D6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cappola, Victor;Gonçalves, Rodrigo B.	Cappola, Victor, Gonçalves, Rodrigo B. (2026): A review of Caenohalictus species from southern Brazil (Apoidea, Caenohalictini). Zootaxa 5782 (3): 401-448, DOI: 10.11646/zootaxa.5782.3.1, URL: https://doi.org/10.11646/zootaxa.5782.3.1
8C7BE1229E05FFF176F01AD82B04FD76.text	8C7BE1229E05FFF176F01AD82B04FD76.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caenohalictus incertus (Schrottky 1902)	<div><p>5. Caenohalictus incertus (Schrottky, 1902)</p><p>(Figs. 12–13, 27D–F, 30A)</p><p>Augochlora (Augochloropsis) incerta Schrottky, 1902 . Type data: holotype ♀ Type locality: Brazil, São Paulo, Botucatu, Vitoriana. Museu de Zoologia, Universidade de São Paulo. Examined through photographs.</p><p>Halictus vallensis Vachal, 1904 . Type data: holotype ♀ Type locality: Peru, Cuzco, Callanga. Muséum National d’Histoire Naturelle, Paris. Examined. Junior synonym (Moure &amp; Hurd 1987).</p><p>Halictus (Paragapostemon) cicindulus Vachal, 1903 . Type data: holotype ♂ Type locality: Peru, Cuzco, Callanga. Muséum National d’Histoire Naturelle, Paris. Examined. New synonym.</p><p>Diagnosis. This species is similar to C. arcuatus sp. nov. by the transverse striation on metapostnotum, but in C. incertus the striae are parallel to weakly curved. It can be distinguished by both sexes being generally larger; integument color metallic green; terga with metallic area occupying the entire disc, and with green or golden reflections (Fig. 12D, 13D) and clypeus and supraclypeal area polished to slightly coriaceous. Females are distinct by having the clypeus with a longitudinal depression on the apex, and apical dark band occupying around one half of clypeus length (Fig. 12B). Males have the head as long as wide (Fig. 13B).</p><p>Redescription</p><p>Female: Body length x̄ =7.20 (N=5). ITD x̄ =1.44 (N=5). UID x̄ =1.16 (N=5). LID x̄ =1.06 (N=5). CLL x̄ =0.41 (N=5). HL x̄ =1.75 (N=5). HW x̄ =1.91 (N=5). Head. Metallic green. Labrum: dark brown; apical elevation presente; base rugose. Mandible: mostly black, with its apex dark amber, sometimes with a yellowish median depression; preapical teeth present. Maxillary palpomeres length: 0.5 times MOD or less. Clypeus: punctate to densely punctate (I=PD; i&lt;pd); with a longitudinal depression at the apex; coriaceous irregularly on the edges and more inconspicuous or polished between the points; thin and long setae (2.5–3 MOD approximately), on its apex with shorter, denser and little thicker setae (2 MOD), mixed with a comb of very short setae; apical dark band occupying a little less than one half its length. Epistomal lobe: angle obtuse and not projected onto the clypeus. Supraclypeal area: punctated (I=PD); polished in the middle, coriaceous elsewhere. Parocular area: whitish setae, plumose and short in the forehead, plumose and long at the lower parocular area. Antennae: scape dark brown; flagellomeres dark brown to light brown, yellowish anteriorly. Frons: slightly punctate, blending with the lineolate to areolate sculpture. Eye setae: very short (less than 0.5 MOD). Gena: metallic green; plumose setae, short to long (0.5–4x MOD); lineolate to polished, with sparse punctation (I&gt;PD). Mesosoma. Mesoscutum: sparsely punctated (I&gt;PD); lineolate, more inconspicuous or sometimes polished on the disc. Scutellum: polished between punctation; punctulated to sparsely punctated. Mesepisternum: lineolate. Metanotum: lineolate; few conflected punctures at the disc, sparsely punctated elsewhere; few plumose setae laterally. Metepisternum: lineolate. Legs: coxa to femur with metallic green reflections, tibia brown and tarsus yellow; yellowish plumose setae. Metapostnotum: medium depression occupying most of the dorsal area; transversally striae lineations, sometimes weakly curved. Propodeum: sparse and thick long plumose setae; pit not carinate. Metasoma. Terga: mostly metallic green or goldish green, sometimes with a black area on the disc of T1, T2, T3 and T4; marginal area brown or greenish brown. Sterna: S1 brownish with green metallic reflections; S2–S6 yellowish, brownish, dark brown or black with green metallic reflections; plumose setae, with longer and denser setae on S4 and S5.</p><p>Male: Body length x̄ =6.11 (N=5). ITD x̄ =1.09 (N=5). UID x̄ =0.91 (N=5). LID x̄ =0.64 (N=5). CLL x̄ =0.34 (N=5). HL x̄ =1.42 (N=5). HW x̄ =1.49 (N=5). Head. Goldish green, metallic green above the frons. Labrum: yellow. Mandible: black; amber apical spot occupying around 1/5 of its length, in the middle with a strong yellow spot occupying a little more than 1/3 of the mandible’s length; lacking preapical teeth. Maxillary palpomeres length: 0.5–1.0x MOD. Clypeus: punctate to densely puncate; finely coriaceous; yellow band occupying only the apex, around 1/6 of the clypeal area. Epistomal lobe: angle obtuse and not projected onto the clypeus. Supraclypeal area: finely coriaceous; punctate. Parocular area: dense, thick, white plumose setae, yellow and thinner on the clypeus and supraclypeal area. Antennae: scape black with green reflections; flagellomeres black to light brown or yellow anteriorly, with a strong yellow spot on the anterior portion of F1, sometimes extending to more flagellomeres; crenulate, F2 as long as each F3–F10. Frons: coriaceous-rugulose with inconspicuous punctation, blending with the sculpture.Eye setae: 0.7–1.0x MOD. Gena: metallic green; coriaceous; sparsely punctate; dense, long and thick white plumose setae. Mesosoma. Integument color: metallic green. Mesoscutum: sparsely punctate to densely punctate on the lateral and posterior edges; polished. Scutellum: polished; confused punctate, sparse on the disc and denser at the edges. Mesepisternum: coriaceous to polished ventrally. Metanotum: polished, with a few finely coriaceous spots; slightly and sparsely punctate. Metepisternum: coriaceous. Legs: fore coxa metallic green, middle and hind coxa dark brown, hind coxa with green reflections, legs dark brown, with green reflections on trochanter and femur, fore tibia yellow anteriorly, middle and hind tibia with an apical yellow spot, basitarsus and tarsi yellow. Metapostnotum: median depression occupying most of the dorsal area; transversally rugose lineations, semi circle shaped anteriorly only; polished; densely punctate. Propodeum: dense, short and thick plumose setae on the sides, weakly near the metapostnotum; pit weakly carinate, not enclosed above. Metasoma. Terga: dark cuprous anteriorly; black area with gold reflections; marginal area dark brown or amber, with green reflections on the lateral of the terga, and on T1 anteriorly. Sterna: black, with green or gold reflections, sometimes very weakly. Genitalia. Gonobase: globose, as long as wide, width 2/3 of the gonocoxite length. Volsella: small, length 1/3 of the gonapophysis, as long as wide, with a small setose plate near gonocoxite. Gonapophysis: ventral prong and dorsal flanges absent; dorsal crest present. Gonocoxite: non striated, smooth surface. Gonostylus: VA absent. VP absent. MGL reniform to arcuated, concave. IGP with a dense and thin basal setose lobule. OGP with three long branched setae on apex.</p><p>Variation. The terga from the females of this species can vary from metallic green or goldish green, and the sterna color is very variable, ranging from yellow to black.</p><p>Comments. RBG examined the types of Halictus vallensis and Halictus cicindulus (MHNP) and the size and sculpturing pattern strongly suggests that they are conspecific. Both specimens are within the delineation of Caenohalictus incertus, so we follow the synonym proposed by Moure &amp; Hurd (1987) for H. vallensis and proposed a new synonym with H. cicindulus .</p><p>Distribution. Brazil: Bahia, Espírito Santo, Mato Grosso, Mato Grosso do Sul, Minas Gerais, Paraná, Rio Grande do Sul, Rio de Janeiro, Santa Catarina, and São Paulo. Paraguay. Peru.</p><p>Examined material. Brazil, BA, DZUP. 6♀, DZUP 540173 DZUP 540358, ”MARACÁS - Bahia \ BRASIL - 23/11/1964 \ Claudionor Elias”,, ”CACULE - BA\ Brasil - VII - 61\ S. LAROCA leg”, 4♀, “ DZUP 540363 - DZUP 540367, ”MARACÁS - Bahia \ BRASIL - 23/11/1964 \ Claudionor Elias ”,, ♀, “ DZUP 540368 ”, ”ITAPETINGA - BAHIA \ BRASIL 23 -27/VII/67\ C. &amp; C. T. Elias leg.”, ♀, “ DZUP 540369 ”, ”MARACÁS - Bahia \ BRASIL - 23/11/1964 \ Claudionor Elias ”, ♀, “ DZUP 540370 ”, ”IBIASSUCÊ\ CACULE - BA\ Brasil 21-VII-61\ Salgado, Laroca”, ♀, “DZUP 540371”, ”MARACÁS - BA\ Brasil 18-VIII-64\ C. ELIAS, LEG”, 9♀, DZUP 540372 - DZUP 540380, ”MARACÁS - Bahia \ BRASIL - 23/11/1964 \ Claudionor Elias”,, ♀, “DZUP 540381”, ”CACULE - BA\ Brasil 21-VII-61\ Salgado, Laroca”, ♀, “DZUP 540382”, ”MARACÁS - Bahia \ BRASIL - 23/11/1964 \ Claudionor Elias”, 27♀, DZUP 540383DZUP 540411, ”MARACÁS - BA\ Brasil 18-VIII-64\ C. ELIAS, LEG”,, ♀, “DZUP 540412”, ”MARACÁS - BA\ Brasil 18-VIII-64\ C. ELIAS, LEG”, ♀, “DZUP 540413”, ”ITAPETINGA - BAHIA \ BRASIL 23-27/VII/67\ C. &amp; C. T. Elias leg.”, ♀, “DZUP 540414”, ”CACULE - BA\ Brasil - VII 61\ S. LAROCA leg”, ♀, “DZUP 540415”, ”CACULE - BA\ Brasil 21 - VII 61\ Salgado Laroca”, ♀, “DZUP 540416”, ”MARACÁS - Bahia \ BRASIL - 23/11/1964 \ Claudionor Elias”, ♀, “DZUP 540417”, ”MARACÁS - BA\ Brasil 18-VIII-64\ C. ELIAS, LEG”, ♀, “DZUP 540418”, ”VITORIA da CONQUISTA\ B A H I A Brasil\ 21/ 25-V-1961 \ F. M. Oliveira”, ♂, “DZUP 540419”, ”MARACÁS - Bahia \ BRASIL - 23/11/1964 \ Claudionor Elias”, ♂, “DZUP 540420”, ”MARACÁS - Bahia \ BRASIL - 23/11/1964 \ Claudionor Elias”, ♀, “DZUP 540421”, ”CACULE BA\ Brasil 21- VII 61\ Salgado, Laroca”, ♀, “DZUP 540422”, ”CACULE BA\ Brasil 21-VII 61\ Salgado, Laroca”. Brasil, ES, DZUP. ♂, “DZUP 540170”, ”Sta. TERESA - ES\ Brasil 19-II-64\ C. ELIAS leg”, ♀, “DZUP 540452”, ”S.J.Petrópolis - ES\ Brasil 12 - II - 64\ C. ELIAS, - LEG”, ♀, “DZUP 540453”, ”S.J.Petrópolis - ES\ Brasil 12 - II - 64\ C. ELIAS, - LEG”, ♀, “DZUP 540454”, ”Sta. TERESA-ES\ Brasil I - II - 64\ C. ELIAS leg”, ♂, “DZUP 540455”, ”Sta. TERESA-ES\ Brasil 27 - II - 64\ C. ELIAS leg”, ♀, “DZUP 540456”, ”F U N D A O - E S\ Brasil 20 - V - 64\ C. ELIAS, - LEG”, ♀, “DZUP 540457”, ”CARIACICA - BRASIL \ ES - 10 a 15/4/1967 / Claudionor Elias”, ♀, “DZUP 540458”, ”VIANA - BRASIL \ ES - 5/7/1966 \ C.T. &amp; C. Elias”. Brasil, MS, DZUP. ♀, “DZUP 540423”, ”Naviraí - MT\ 21-I-1977 \ Pereira, M.A. leg”, ♀, “DZUP 540426”, ”SISBIOTA - CNPq/FAPESP\ BR: MS: Aquidauana\ 20°25’59”S, 55°39’20.8”W\ 11-26.x.2011 Malaise 08\ Lamas, Nihei &amp; eq. Col.”. Brasil, MT, DZUP. ♂, “DZUP 540171”, ”Chapada de Guimarâes\ MT - Brasil - 30 - I - 65\ Sebastião Laroca leg”, ♀, “DZUP 540424”, ”Chap. Guimarães-MT\ 21.XI.1983 \ Exc. Dep. Zool-UFPR\ (Polonoroeste)”, ♀, “DZUP 540425”, ”Chap. Guimarães-MT\ 21.XI.1983 \ Exc. Dep. Zool-UFPR\ (Polonoroeste)”. Brasil, MG, DZUP. ♂, “DZUP 540169”, ”PASSOS - MG\ Brasil 17-VIII-62\ Claudionor Elias”, ♀, “DZUP 540360”, ”BRAZOPOLIS - MG\ Brasil XII - 61\ Claudionor Elias”, ♀, “DZUP 540361”, ”BRAZOPOLIS - MG\ Brasil XII - 61\ Claudionor Elias”, ♀, “DZUP 540362”, ”BRAZOPOLIS - MG\ Brasil XII - 61\ Claudionor Elias”, ♀, “DZUP 540366”, ”BRAZOPOLIS - MG\ Brasil XII - 61\ Claudionor Elias”, ♀, “DZUP 540404”, ”Sta. JULIANA\ MG - BRASIL \ 4/VI/1965 \ C. Elias leg”, ♀, “DZUP 540405”, ”Sta. JULIANA\ MG - BRASIL \ 4/VI/1965 \ C. Elias leg”, ♀, “DZUP 540427”, ”Brasil, Minas Gerais,\ Viçosa, UFV, mata da\ caixa da água,\ 2.vii.1992, G. Melo,\ em flor de Guapira sp”, ♀, “DZUP 540428”, ” Brasil, Minas Gerais,\ Viçosa, UFV, mata da\ caixa da água,\ 2.vii.1992, G. Melo,\ em flor de Guapira sp”, ♀, “DZUP 540429”, ”Brasil, Minas Gerais,\ Viçosa, UFV, mata da\ caixa da água,\ 2.vii.1992, G. Melo,\ em flor de Guapira sp”, ♂, “DZUP 540430”, ” Brasil, Minas Gerais,\ Viçosa, UFV, mata da\ caixa da água,\ 2.vii.1992, G. Melo,\ em flor de Guapira sp”, ♂, “DZUP 540431”, ”Brasil, Minas Gerais,\ Viçosa, UFV, mata da\ caixa da água,\ 2.vii.1992, G. Melo,\ em flor de Guapira sp”, ♂, “DZUP 540432”, ” Brasil, Minas Gerais,\ Viçosa, UFV, mata da\ caixa da água,\ 2.vii.1992, G. Melo,\ em flor de Guapira sp”, ♂, “DZUP 540433”, ”Brasil, Minas Gerais,\ Viçosa, UFV, mata da\ caixa da água,\ 2.vii.1992, G. Melo,\ em flor de Guapira sp”, ♂, “DZUP 540434”, ” Brasil, Minas Gerais,\ Ouro Preto, 1400m \ 20°22’S 43°30’W\ 8-9.i.2010, K. Ramos\ &amp; V. Kanamura”, ♂, “DZUP 540435”, ”Brasil, Minas Gerais,\ Viçosa, UFV, mata da\ caixa da água,\ 2.vii.1992, G. Melo,\ em flor de Guapira sp”, ♀, “DZUP 540436”, ”Viçosa - MG\ BRASIL 07/10/92\ G. A. R. MELO”, ♀, “DZUP 540437”, ”BRAZOPOLIS - MG\ Brasil XII - 61\ Claudionor Elias”, ♀, “DZUP 540438”, ”PATOS de MINAS - MG\ BRASIL - 23/11/965\ Claudionor Elias”, ♀, “DZUP 540439”, ”PATOS de MINAS - MG\ BRASIL - 23/11/965\ Claudionor Elias”, ♂, “DZUP 540440”, ”TAPIRA-MG\ Brasil 27-V 65\ C. ELIAS leg”, ♂, “DZUP 540441”, ”TAPIRA-MG\ Brasil 27-V 65\ C. ELIAS leg”, ♀, “DZUP 540442”, ”Sta. JULIANA\ MG - BRASIL \ 29/VII/1965 \ C. Elias leg”, ♂, “DZUP 540443”, ”Sta. JULIANA\ MG - BRASIL \ 4/VI/1965 \ C. Elias leg”, ♀, “DZUP 540444”, ”Brasil, Minas Gerais,\ 18km W de Capitólio,\ Rio Turvo, 800m \ 20°38’S 46°13’W\ 15.v.1999. GAR Melo”, ♀, “DZUP 540445”, ”POÇOS CALDAS -MG\ Brasil 23-XI-62\ Claudionor Elias”, ♀, “DZUP 540446”, ”PASSOS MG\ BR - 16-21 - XII - 63\ Claudionor Elias”, ♂, “DZUP 540447”, ”PASSOS MG\ BR - 16-21 - XII - 63\ Claudionor Elias”, ♀, “DZUP 540448”, ”SÃO GOTARDO\ MG- BRASIL \ 11/VI/1965 \ C. Elias leg”, ♀, “DZUP 540449”, ”SÃO GOTARDO\ MG- BRASIL \ 11/VI/1965 \ C. Elias leg”, ♀, “DZUP 540450”, ”JACUI - MG\ Brasil 27-XI-63\ C. ELIAS, LEG”, ♂, “DZUP 540451”, ”JACUI - MG\ Brasil 27-XI-63\ C. ELIAS, LEG”. Brazil, PR, DZUP. ♀, “DZUP 539925”, ”Brasil, Paraná, Campo\ Largo, Rio Verde, 900m,\ 25º25’S 49º27’W,\ 09.x.2005, G. Melo”, ♂, “DZUP 539926”, “ Brasil, Paraná,\ Morretes, Est. Exp.\ IAPAR, 04.xii.2004,\ G.A.R. Melo”, ♂, “DZUP 539927”, “Brasil, Paraná,\ Morretes, Est. Exp.\ IAPAR, 04.xii.2004,\ G.A.R. Melo”, ♂, “DZUP 539928”, “ Brasil, Paraná,\ Morretes, Est. Exp.\ IAPAR, 04.xii.2004,\ G.A.R. Melo”, ♂, “DZUP 539929”, “Brasil, Paraná,\ Morretes, IAPAR,\ 06.xii.2004, 9:20-11:30\ K.S. Ramos”, ♂, “DZUP 539930”, “ Brasil, Paraná,\ Morretes, IAPAR,\ 06.xii.2004, 15:30-16:30\ K.S. Ramos”, ♀, “DZUP 539931”, ”Morretes-PR Brasil\ (IAPAR)\ 04-11.II.1985 \ C.I.I.F. (Malaise)”, ♀, “DZUP 539932”, ”GUAIRA - PR \ Brasil 11-XII-1965 \ V. Graf - L. Azevedo”, ♀, “DZUP 539933”, ”GUAIRA - PR \ Brasil 11-XII-1965 \ V. Graf - L. Azevedo”, ♀, “DZUP 539934”, ”FOZ DO IGUAÇU\ Parana Brasil \ 7.XII.1966 \ Exc.Dept. ZOO”, ♀, “DZUP 539935”, ”PARANAVAI - PR \ Brail 15 - V - 61\ Laroca e Marston”, ♂, “DZUP 539936”, “Brasil, Paraná, Imbituva,\ P1 GVI, 25.13366ºS,\ 50.52290ºW, 27.II.2022,\ L.V.Graf, rede entomológica”, ♂, “DZUP 539937”, “ Brasil, Paraná, Ibati,\ Ponto 5LT, 03.xii.2021,\ 23.7283ºS 50.2961ºW, 650m, Muniz&amp;Barbosa, Busca ativa”, ♂, “DZUP 539938”, “Brasil, Paraná, Manoel Ribas,\ -24.5220645, -51.4769617,\ 26.V.2021, L.Graf”, ♀, “DZUP 539944”,”H. M. Taura\ 18.12.85\ Carqueja\ UEM.MGA. PR. ”. Brazil, RJ, DZUP. ♂, “DZUP 539953”, “Morro Da. MARTA\ Rio Janeiro Guanabara\ Brasil X.1957 \ M. Alvarenga leg.”, 53♂, DZUP 540279 - DZUP 540332, “SILVESTRE\ Guanabara Brasil \ 4-IV-1960 \ M. Alvarenga leg”, ♀, “DZUP 540333”, “S. Bento D. Caxias\ Est. Rio Janeiro\ BRASIL VI-1953 \ P.A. Telles”, ♀, “DZUP 540334”, “S. Bento\ Duque de Caxias\ Est. do Rio BRASIL \ VI-1954 P.A. Teles”, ♀, “DZUP 540335”, “S. Bento D. Caxias\ Est. Rio Janeiro\ BRASIL VI-1953 \ P.A. Telles”, ♀, “DZUP 540336”, “S. Bento D. Caxias\ Est. Rio Janeiro\ BRASIL VI-1953 \ P.A. Telles”, ♀, “DZUP 540337”, “S. Bento D. Caxias\ Est. Rio Janeiro\ BRASIL VI-1953 \ P.A. Telles”, ♂, “DZUP 540338”, “Retiro Pedro do Rio\ Est do Rio Brasil\ III-1954 \ C.A.C. Seabra Coll.”, ♂, “DZUP 540339”, “Retiro Pedro do Rio\ Est do Rio Brasil\ III-1954 \ C.A.C. Seabra Coll.”, ♀, “DZUP 540340”, “S. Bento D. Caxias\ Est. Rio Janeiro\ BRASIL VI-1953 \ P.A. Telles”, ♀, “DZUP 540341”, “S. Bento D. Caxias\ Est. Rio Janeiro\ BRASIL VI-1953 \ P.A. Telles”, ♀, “DZUP 540342”, “S. Bento\ Duque de Caxias\ Est. do Rio BRASIL \ 30-V-1954 P.A. Teles”, ♀, “DZUP 540343”, “Ilha do Governador\ D. Federal BRASIL \ 25-VII-954\ M Alvarenga Coll.”, ♀, “DZUP 540344”, “Ilha do Governador\ D. Federal BRASIL \ 29-VII-1954 \ M Alvarenga Coll.”, ♀, “DZUP 540345”, “Ilha do Governador\ D. Federal BRASIL \ 25-VII-954\ M Alvarenga Coll.”, ♀, “DZUP 540346”, “Ilha do Governador\ D. Federal BRASIL \ 25-VII-954\ M. Alvarenga Coll.”, ♀, “DZUP 540347”, “Ilha do Governador\ D. Federal BRASIL \ 25-VII-954\ M Alvarenga Coll.”, ♀, “DZUP 540348”, “Ilha do Governador\ D. Federal BRASIL \ 25-VII-954\ M Alvarenga Coll.”, ♀, “DZUP 540349”, “Ilha do Governador\ D. Federal BRASIL \ 25-VII-954\ M Alvarenga Coll.”, ♀, “DZUP 540350”, “Ilha do Governador\ D. Federal BRASIL \ 25-VII-954\ M. Alvarenga Coll.”, ♀, “DZUP 540351”, “S. Bento D. Caxias\ Est. Rio Janeiro\ BRASIL VI-1953 \ P.A. Telles”, ♀, “DZUP 540352”, “Rep. RIO GRANDE\ GUANABARA 3/65\ BRASIL \ F. M. Oliveira l”, ♀, “DZUP 540353”, “Rep. RIO GRANDE\ GUANABARA 3/65\ BRASIL\ F. M. Oliveira l”. Brazil, RS, DZUP. ♀, “DZUP 539939”,”N. PETROPOLIS, RS\ Joaneta, 8.1.93,\ leg. Isabel A. Santos\\ 9234 ninhos\ f:22 no paredão”, ♀, “DZUP 539940”,”N. PETROPOLIS, RS\ Joaneta, 8.1.93,\ leg. Isabel A. Santos\\ 9236 ninhos\ f:22 no paredão”, ♀, “DZUP 539941”,”N. PETROPOLIS, RS\ Joaneta, 8.1.93,\ leg. Isabel A. Santos\\ 9238 ninhos\ f:22 no paredão”. Brazil, SC, DZUP. ♀, “DZUP 539942”, “Córrego Grande\ Florianópolis, SC, Brazil \ 04.III.2006 \ Josefina Steiner leg.\\ Caenohalictus incertus \ R. B. Gonçalves det\\ Schinus terebinthifolius\ Anacardiaceae \ J. Steiner det. 2006\ Lâmina 43\\ CJS 3494”, ♀, “DZUP 539943”,“Córrego Grande\ Florianópolis, SC, Brazil \ 04.III.2006 \ Josefina Steiner leg.\\ Caenohalictus incertus \ Rafael Kamke det. 2008\\ Schinus terebinthifolius\ Anacardiaceae \ J. Steiner det. 2006\ Lâmina 42\\ CJS 3495”. Brazil, SP, DZUP. ♀, “DZUP 539945”,”Pedregulho - SP\ Brasil 8-XI-62\ Claudionor Elias”, ♀, “DZUP 539946”,”BRASIL-SÃO PAULO\ Batatais XII.1943 \ Pe. Pereira col.\\ Paragapostemon ♀ \ incertus (5chr.)\ P. Moure det. 1944”, 7♂, “DZUP 539947- DZUP 540172 “Pedregulho - SP\ Brasil 8-XI-62\ Claudionor Elias”. Paraguay, DZUP. ♀, “DZUP 540174”, ” Paraguay, Amambay,\ Parque Nacional Cerro\ Corá, 280 m, 12.ii.2007 \ 22°39’50”S 55°59’18”W\ M.G.Hermes coll”, ♀, “DZUP 540175”, ” Paraguay, Amambay,\ Pq. Nac. Cerro Corá,\ campo-cerrado en la\ zona del Cerro Muralla,\ 22°39’S 55°59’W, 280 m,\ 12.ii.2007, B. Garcete”, ♂, “DZUP 540176”, “Colony, Carlos\ Pfannl, Paraguay \ III-51\ F. H. Shade”, ♂, “DZUP 540177”, “Colony, Carlos\ Pfannl, Paraguay \ III-51\ F. H. Shade”.</p></div>	https://treatment.plazi.org/id/8C7BE1229E05FFF176F01AD82B04FD76	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cappola, Victor;Gonçalves, Rodrigo B.	Cappola, Victor, Gonçalves, Rodrigo B. (2026): A review of Caenohalictus species from southern Brazil (Apoidea, Caenohalictini). Zootaxa 5782 (3): 401-448, DOI: 10.11646/zootaxa.5782.3.1, URL: https://doi.org/10.11646/zootaxa.5782.3.1
8C7BE1229E02FFFF76F01F392B79FBAF.text	8C7BE1229E02FFFF76F01F392B79FBAF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caenohalictus leticiae Cappola & Gonçalves 2026	<div><p>6. Caenohalictus leticiae sp nov.</p><p>(Figs. 3B, 14–15, 28G–I, 30C)</p><p>Diagnosis. Small species (5–6 mm) similar to C. tesselatus and C. implexus by the rugulose metapostnotum and olivaceus green body. It can be differentiated from C. tesselatus by black scape on both sexes (Fig. 14B), the lack of dense, yellow branched setae on mesoscutum of females (Fig. 14C) and S1 with few metallic reflections. Females are separated from C. implexus by scutellum weakly coriaceous and sparsely to densely punctate (Fig. 14C); tarsi dark brown; head wider than long; mesepisternum and metanotum rugose. Males differ from C. implexus in having flagellomeres 2–11 with anterior large patches of placodea sensillae (Fig. 3B), mesoscutum punctate at sides of parapsidal lines and posterior margin, propodeum lateral surface mostly polished and punctate (Fig. 15C).</p><p>Description</p><p>Female: Body length x̄ =6.57 (N=5). ITD x̄ =1.26 (N=5). UID x̄ =1.00 (N=5). LID x̄ =0.94 (N=5). CLL x̄ =0.33 (N=5). HL x̄ =1.49 (N=5). HW x̄ =1.65 (N=5). Head. Metallic green, with golden reflections. Labrum: black; apical elevation presente; base rugose. Mandible: dark brown, with apical amber spot occupying 1/2 of its length, sometimes a little more; preapical teeth present. Maxillary palpomeres length: 0.7 the size of MOD or less. Clypeus: punctate to sparsely punctate (I=PD; I&gt;PD), with a longitudinal depression at the apex; coriaceous, at the black apical stripe more inconspicuously or polished; thin plumose setae (2–2.5 MOD approximately); apical dark band occupying less than 1/2 its length, sometimes with a purple reflection edge. Epistomal lobe: angle obtuse and not projected onto the clypeus. Supraclypeal area: sparsely punctated to densely punctated; coriaceous. Parocular area: yellowish, very thin and dense plumose setae (1.5–2 MOD). Antennae: scape black; flagellomeres black, light brown to yellowish anteriorly, with a strong yellow spot in F1, F2 or F3. Frons: punctuate to densely punctate in a few areas, more inconspicuously on the coriaceous area. Eye setae: about 1 MOD. Gena: metallic green, sometimes with golden reflections; thin and sparse long plumose setae (2–4 MOD); slightly coriaceous; punctate. Mesosoma. Mesoscutum: sparsely punctated to densely punctated on the edges; areolate to micro lineolate. Scutellum: coriaceous; sparsely punctulate anteriorly and on the disc, punctate to densely punctate posteriorly. Mesepisternum: rugose to lineolate ventrally. Metanotum: rugulose-sparsely punctulate to punctate. Metepisternum: rugose. Legs: coxa to femur brown with metallic green reflections, tibia to tarsi brown or light brown; whitish/yellowish plumose setae. Metapostnotum: medium depression occupying most of the dorsal area; longitudinally sculptured, areolate-rugose, irregularly or branched; coriaceous. Propodeum: dense and thin long plumose setae; pit not carinate, not enclosed above. Metasoma. Terga: goldish green; dark violaceous area on the disc of T1–T3, sometimes small on T1 and even smaller on T3; marginal area brown with light goldish green reflections. Sterna: S1 dark brown or black with green metallic reflections. S2–S6 dark brown or black, with green metallic reflections; dense and thin plumose setae on the post gradular area, glabrous on the pre gradular area.</p><p>Male: Body length x̄ =5.58 (N=5). ITD x̄ =1.04 (N=5). UID x̄ =0.90 (N=5). LID x̄ =0.56 (N=5). CLL x̄ =0.31 (N=5). HL x̄ =1.34 (N=5). HW x̄ =1.42 (N=5). Head. Metallic green, goldish green on the parocular areas and below the antennal socket. Labrum: yellow. Mandible: black; amber apical spot occupying around 1/3 of its length, in the middle with a strong yellow spot occupying a little more than 1/3 of its length; lacking preapical teeth. Maxillary palpomeres length: 0.5–0.7x MOD. Clypeus: punctate; coriaceous to polished below; yellow spot occupying 1/2 to 1/3 its length, with a dark edge separating it from the metallic area. Epistomal lobe: angle obtuse and not projected onto the clypeus. Supraclypeal area: polished to slightly coriaceous; densely punctate. Paraocular area: pubescence white, thin and sparse plumose setae. Antennae: scape black, with weak green reflections; flagellomeres black to dark brown anteriorly with yellow spots, flagellomeres 2–11 with anterior large patches of sensilla placodea; crenulate, F2 longer than each F3– F10. Frons: coriaceous-rugulose with inconspicuous punctation, blending with the sculpture. Eye setae: 0.7–1.0x MOD. Gena: metallic green to goldish green; coriaceous-rugulose; punctate to sparsely punctate; long and thin yellow plumose setae. Mesosoma. Integument color: metallic green. Mesoscutum: sparsely punctate to densely punctate on the lateral and posterior edges; coriaceous. Scutellum: polished to coriaceous posteriorly; confused punctate on the disc to densely punctate on the edges. Mesepisternum: confused rugulose. Metanotum: rugulose; punctate to densely punctate. Metepisternum: confused rugulose. Legs: coxa metallic green, except the black midcoxa, trochanter and femur black with green reflections, fore leg tibia dark brown to yellow, yellow elsewhere, mid leg tibia dark brown with yellow apex, yellow to brown elsewhere, hind leg tibia black, dark brown to light brown elsewhere; white to yellow plumose setae. Metapostnotum: median depression occupying most of the dorsal area; areolate-rugose; coriaceous-rugulose to coriaceous. Propodeum: dense, short and thin plumose setae; pit carinate, not enclosed above. Metasoma. Terga: dark blue anteriorly; black on the disc; dark brown on the marginal area. Sterna: black to dark brown. Genitalia. Gonobase: wider than long, was wide as1/2 the gonocoxite length. Volsella: enlarged, almost as long as the gonapophysis, as long as wide. Gonapophysis: with ventral prong; small dorsal flanges; dorsal depressions and dorsal ridge; dorsal crest. Gonocoxite: longitudinally striate. Gonostylus: VA absent. VP absent. MGL reniform. IGP slightly expanded. OGP with a few small setae on apex.</p><p>Etymology. Named in honor of Letícia Vanessa Graf, a melittologist who contributed with specimens from Paraná State and collaborated on urban bee ecology projects with the senior author.</p><p>Distribution. Brazil: Paraná.</p><p>Type Material. Holotype. Brazil, PR, DZUP. ♀, “ DZUP 540093 ”, “BRASIL, PARANÁ,\ COLOMBO, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15935&amp;materialsCitation.latitude=-25.321852" title="Search Plazi for locations around (long -49.15935/lat -25.321852)">EMBRAPA</a> \ 25º19’18.66’’S 49º9’33.66’’W \ 8.I.2020 935m \ MIKICH, S.B. leg. E5479” . Paratype. Brazil, PR, DZUP. ♀, “ DZUP 540094 ”, ” BRASIL, PARANÁ,\ COLOMBO, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15935&amp;materialsCitation.latitude=-25.321852" title="Search Plazi for locations around (long -49.15935/lat -25.321852)">EMBRAPA</a> \ 25º19’18.66’’S 49º9’33.66’’W \ 8.I.2020 935m \ MIKICH, S.B. leg. E5480”, ♀, “ DZUP 540095 ”,” Brasil, PR, São Mateus\ do Sul, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15935&amp;materialsCitation.latitude=-25.321852" title="Search Plazi for locations around (long -49.15935/lat -25.321852)">UN-SIX/Petrobras</a> \ 24.I.2013 \ Rafael Kamke leg.\\ Ponto amostral\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15935&amp;materialsCitation.latitude=-25.321852" title="Search Plazi for locations around (long -49.15935/lat -25.321852)">VACÓDROMO</a> \\ CJS 12640”, ♀, “ DZUP 540096 ”,” Brasil, PR, São Mateus\ do Sul, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15935&amp;materialsCitation.latitude=-25.321852" title="Search Plazi for locations around (long -49.15935/lat -25.321852)">UN-SIX/Petrobras</a> \ 27.II.2013 \ Rafael Kamke leg.\\ Ponto amostral\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15935&amp;materialsCitation.latitude=-25.321852" title="Search Plazi for locations around (long -49.15935/lat -25.321852)">MATA DO TÚNEL</a> \\ CJS 12829”, ♀, “ DZUP 540097 ”,” Brasil, PR, São Mateus\ do Sul, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15935&amp;materialsCitation.latitude=-25.321852" title="Search Plazi for locations around (long -49.15935/lat -25.321852)">UN-SIX/Petrobras</a> \ 28.II.2013 \ Rafael Kamke leg.\\ Ponto amostral\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15935&amp;materialsCitation.latitude=-25.321852" title="Search Plazi for locations around (long -49.15935/lat -25.321852)">VACÓDROMO</a> \\ CJS 13017”, ♂, “ DZUP 540098 ”, “ Brasil, PR, São Mateus\ do Sul, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15935&amp;materialsCitation.latitude=-25.321852" title="Search Plazi for locations around (long -49.15935/lat -25.321852)">UN-SIX/Petrobras</a> \ 27.II.2013 \ Rafael Kamke leg.\\ Ponto amostral\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15935&amp;materialsCitation.latitude=-25.321852" title="Search Plazi for locations around (long -49.15935/lat -25.321852)">JAGUATIRICA</a> \\ CJS 12944”, ♂, “ DZUP 540099 ”, “ Brasil, PR, São Mateus\ do Sul, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15935&amp;materialsCitation.latitude=-25.321852" title="Search Plazi for locations around (long -49.15935/lat -25.321852)">UN-SIX/Petrobras</a> \ 21.I.2011 \ Rafael Kamke leg.\\ Ponto amostral\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15935&amp;materialsCitation.latitude=-25.321852" title="Search Plazi for locations around (long -49.15935/lat -25.321852)">MATA DO TÚNEL</a> \\ CJS 9683”, ♂, “ DZUP 540100 ”, “ Brasil, PR, São Mateus\ do Sul, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15935&amp;materialsCitation.latitude=-25.321852" title="Search Plazi for locations around (long -49.15935/lat -25.321852)">UN-SIX/Petrobras</a> \ 27.II.2013 \ Rafael Kamke leg.\\ Ponto amostral\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15935&amp;materialsCitation.latitude=-25.321852" title="Search Plazi for locations around (long -49.15935/lat -25.321852)">JAGUATIRICA</a> \\ CJS 12947”, ♂, “ DZUP 540101 ”, “ Brasil, Paraná, Ipiranga,\ -24.9361383, -50.5479950,\ 29.V.2021, L.Graf ”, ♂, “ DZUP 540102 ”, “ Brasil, Paraná, Colombo, P3,\ 25.363624ºS, 49.145306ºW,\ 28.IV.2022, L. V. Graf, rede\ entomológica ”, ♀, “ DZUP 606115 ”,” Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15935&amp;materialsCitation.latitude=-25.321852" title="Search Plazi for locations around (long -49.15935/lat -25.321852)">Parque\ Estadual de Vila Velha</a>,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15935&amp;materialsCitation.latitude=-25.321852" title="Search Plazi for locations around (long -49.15935/lat -25.321852)">Área I</a>, 25º13’53’’S,\ 49º59’35’’W, 09.I.2024,\ Gonçalves, Graf, Pundek ”, ♂, “ DZUP 606935 ”, “ Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15935&amp;materialsCitation.latitude=-25.321852" title="Search Plazi for locations around (long -49.15935/lat -25.321852)">Parque\ Estadual de Vila Velha</a>,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15935&amp;materialsCitation.latitude=-25.321852" title="Search Plazi for locations around (long -49.15935/lat -25.321852)">Área I</a>, 25º13’53’’S,\ 49º59’35’’W, 1.V.2024,\ Gonçalves, Graf, Pundek ”, ♂, “ DZUP 547668 ”, “ Brasil, Paraná, Curitiba,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15935&amp;materialsCitation.latitude=-25.321852" title="Search Plazi for locations around (long -49.15935/lat -25.321852)">Cemit. Parque Iguaçu</a>,\ -25.419348, -49.310204,\ 05.III.2018, L.Graf ”, ♂, “ DZUP 547403 ”, “ Brasil, Paraná, Curitiba,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15935&amp;materialsCitation.latitude=-25.321852" title="Search Plazi for locations around (long -49.15935/lat -25.321852)">Quartel PQ Reg. Man. 5</a>,\ - 25.396512, -49.226759,\ 26.I.2018, L.Graf ”, ♂, “ DZUP 548149 ”, “ Brasil, Paraná, Curitiba,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15935&amp;materialsCitation.latitude=-25.321852" title="Search Plazi for locations around (long -49.15935/lat -25.321852)">Quartel PQ Reg. Man. 5</a>,\ -25.396512, -49.226759,\ 03.IV.2018, L.Graf ”, ♂, “ DZUP 546813 ”, “ Brasil, Paraná, Curitiba,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15935&amp;materialsCitation.latitude=-25.321852" title="Search Plazi for locations around (long -49.15935/lat -25.321852)">Cemit. Parque Iguaçu</a>,\ -25.419348, -49.310204,\ 06.II.2018, L.Graf ” .</p></div>	https://treatment.plazi.org/id/8C7BE1229E02FFFF76F01F392B79FBAF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cappola, Victor;Gonçalves, Rodrigo B.	Cappola, Victor, Gonçalves, Rodrigo B. (2026): A review of Caenohalictus species from southern Brazil (Apoidea, Caenohalictini). Zootaxa 5782 (3): 401-448, DOI: 10.11646/zootaxa.5782.3.1, URL: https://doi.org/10.11646/zootaxa.5782.3.1
8C7BE1229E0DFFFC76F01CEC2ACAFAD2.text	8C7BE1229E0DFFFC76F01CEC2ACAFAD2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caenohalictus longipalpis Cappola & Gonçalves 2026	<div><p>7. Caenohalictus longipalpis sp. nov.</p><p>(Figs. 2C, 16–17, 26B, 29G–I, 31A)</p><p>Diagnosis. Easily separated from other Caenohalictus from southern Brazil by having long maxillary palpomeres, with each palpomere measuring about 2 to 3 times the MOD in females (Fig. 16A), and about 2.5 times the MOD in males. Females can also be distinguished by the propodeal pit enclosed by strong carinae, carinae arched outwards (Fig. 2C). The elongate palpomeres and propodeal pit carina also distinguish it from C. oresicoetes . From the C. sesquipalpis sp. nov., females can be separated also by longer head, male S6 having a slight invagination (Fig. 26B), and longer eye setae, measuring around 1.0 times the MOD. The elongate palpomeres and head also distinguish these species from C. aerugineus .</p><p>Description</p><p>Female: Body length x̄ =9.95 (N=5). ITD x̄ =1.69 (N=5). UID x̄ =1.25 (N=5). LID x̄ =1.17 (N=5). CLL x̄ =0.52 (N=5). HL x̄ =2.33 (N=5). HW x̄ =2.27 (N=5). Head. Bluish green or goldish green. Labrum: black; apical elevation present, but small; base rugose. Mandible: black; apical amber spot occupying 1/2 of its length; preapical teeth present. Maxillary palpomeres length: 1.0–2.5 the size of MOD. Clypeus: punctate (I=PD); areolate; long, thin and dense plumose setae; apical dark band occupying around 1/2 of its length, median part more produced. Epistomal lobe: angle approximately orthogonal, projected onto the clypeus. Supraclypeal area: punctate to sparsely punctate; coriaceous. Parocular area: yellowish setae, long, thin and dense, a little shorter on the frons. Antennae: scape dark brown; flagellomeres dark brown; antennal socket located in a strongly concave depression. Frons: slightly punctate; coriaceous to rugulose near the ocellus. Eye setae:approximately 1 MOD. Gena: very thin and long plumose setae (2.5–4x MOD); micro lineolate with inconspicuous punctation; dark green or goldish green. Mesosoma. Mesoscutum: punctate to sparsely punctate anteriorly and on the middle of the disc, densely punctate elsewhere; areolate to granulate. Scutellum: polished between punctation; densely punctated. Mesepisternum: areolate to lineolate. Metanotum: polished on the middle, areolate on the sides; densely punctated. Metepisternum: areolate. Legs: dark brown, with fore leg tarsi light brown to yellow; yellow plumose setae. Metapostnotum: medium depression occupying only the median area, a little more expanded anteriorly, weakly striate to rugose; granulate. Propodeum: few long and thin plumose setae; pit carinate, with the carina forming two semi circles from both sides of the pit anteriorly, not enclosed above. Metasoma. Terga: brown on the disc and marginal area, sometimes with blue or purple reflections; green or greenish blue anteriorly; on T4 green or goldish green. Sterna: S1 brown with green, purple or greenish blue reflections; S2–S6 dark brown; sparse and thin white plumose setae on the post gradular area, glabrous on the pre gradular area.</p><p>Male: Body length x̄ =8.84 (N=2). ITD x̄ =1.47 (N=3). UID x̄ =1.17 (N=3). LID x̄ =0.88 (N=3). CLL x̄ =0.49 (N=3). HL x̄ =2.00 (N=3). HW x̄ =2.06 (N=3). Head. Metallic blue on its entirety or metallic green, with faint blue tones, goldish green on the vertex. Labrum: yellow basaly, dark brown on apex. Mandible: black; amber apical spot occupying nearly 1/2 of its length; lacking preapical teeth. Maxillary palpomeres length: 1.5–2.5x MOD. Clypeus: punctate to densely puncate; coriaceous; yellow spot occupying around 1/2 of the clypeal area, with a dark edge separating it from the metallic area. Epistomal lobe: angle approximately orthogonal, projected onto the clypeus. Supraclypeal area: coriaceous to areolate; inconspicuously punctate. Parocular area: yellow, short and plumose setae, thicker on the lower parocular area and around the antennal socket, thinner and brown on the clypeus and lower portion of the supraclypeal area. Antennae: scape and flagellomeres dark brown, lighter anteriorly and with yellow spots on the flagellomeres; crenulate, F2 as long as each F3–F10. Frons: areolate; inconspicuously punctate, blending with the sculpture. Eye setae: 0.7–1.0x MOD. Gena: greenish blue; lineolate to areolate; punctate to densely punctate; long and yellow plumose setae. Mesosoma. Integument color: blue on its entirety or with pronotum metallic blue, dark green on the pronotum lobe, dark green elsewhere, with a few goldish green spots and some bluish tones on the edges. Mesoscutum: densely punctate to punctate, sparsely punctate on the middle of the disc and anteriorly; areolate to granulate. Scutellum: weakly lineolate on the edges, almost polished on the disc; densely punctate. Mesepisternum: areolate to lineolate. Metanotum: coriaceous, weakly on the disc; punctate to densely punctate. Metepisternum: areolate. Legs: black or dark brown, with fore coxa with dark green reflections, tibia to tarsi light brown to yellow anteriorly. Metapostnotum: median depression occupying only the median area, a little more expanded anteriorly; striae longitudinally rugose, branched or areolate, sometimes very weakly; granulate. Propodeum: dense, short and thick plumose setae on the sides, weakly near the metapostnotum; pit carinate, enclosed, parallel. Metasoma. Terga: blue or green anteriorly; black on the disc, sometimes with some purple reflections; dark brown on the marginal area. Sterna: S1 metallic blue or green, dark brown elsewhere. Genitalia. Gonobase: wider than long, as wide as the gonocoxite length. Volsella: small, length a little less than 1/3 of the gonapophysis, wider than long. Gonapophysis: small ventral prong; small dorsal flanges, starting almost at middle of gonapophysis, with same width as the gonapophysis; dorsal crest present; with a few dorsal setae in the edge, near the flanges; with a few ventral setae on the middle. Gonocoxite: longitudinally striate. Gonostylus: VA membranous large lobe, with dense and small setae at the apex, and longer setae in the lower edges; MGL reniform to arcuated, concave, with a lower lobule; IGP expanded and with a large setose lobe; OGP with a few long and branched setae.</p><p>Variation. The males of this species can vary in color from entirely blue to dark green, with some metallic green, bluish and goldish spots.</p><p>Etymology. The specific epithet is compound from the Latin comparative longior, longius (“longer”), abbreviated here as longi- in reference to the elongated condition of the structure, and from the Latin noun palpus (genitive palpi), meaning “palp.” The name refers to the proportionally elongated maxillary palpus observed in this species. The epithet is treated as a noun in apposition.</p><p>Distribution. Brazil: Minas Gerais, Paraná, Rio Grande do Sul, and Santa Catarina.</p><p>Type material. Holotype. Brasil, SC, DZUP. ♀, “ DZUP 540114 ”, “ Florianópolis, SC\ Brasil, 02.IV.2001 \ T. M. Guerra col.\\ 8 14:10”. Paratype. Brasil, Paraná, DZUP. ♀, “ DZUP 540107 ”,” Brasil, Paraná, São\ José dos Pinhais, M2,\ 25.3820ºS, 49.0224ºW,\ 20.VI.2023, L.V.Graf,\ prato verde \\ Coord. corretas M2:\ 25º38’26.11’’S \ 49º02’16.80’’W ”, ♀, “ DZUP 540108 ”,” S.José Pinhais - PR \ Brasil ( Br277-Km54)\ 05-12.XI.1984 \ C.I.I.P. (Malaise)”, ♀, “ DZUP 540109 ”,” S.José Pinhais - PR \ Brasil ( Br277-Km54)\ 10-17.XII.1984 \ C.I.I.P. (Malaise)”, ♀, “ DZUP 540110 ”,” Brasil, Paraná, São\ José dos Pinhais, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.032726&amp;materialsCitation.latitude=-25.658588" title="Search Plazi for locations around (long -49.032726/lat -25.658588)">S2</a>,\ 25.3949ºS, 49.0143ºW,\ 15.VI.2022, L.V.Graf,\ rede entomológica \\ Coord. corretas S2:\ 25º39’30.92’’S \ 49º01’57.81’’W ”, ♀, “ DZUP 540111 ”,” Brasil, Paraná, Pi raquara,\ Mananciais da Serra;\ 9-20.ii.2003; malaise;\ AJCAguiar ”, ♀, “ DZUP 540112 ”,” Brasil, Paraná, Castro,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.0925&amp;materialsCitation.latitude=-24.7587" title="Search Plazi for locations around (long -50.0925/lat -24.7587)">Ponto P1-GVII</a>, 1020m,\ 24.7587ºS, 50.0925ºW,\ 21.xi.2020, A. Martins \ &amp; D. Muniz, Busca ativa ”, ♀, “ DZUP 540113 ”,” Brasil, Paraná, Castro,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.5221&amp;materialsCitation.latitude=-24.4537" title="Search Plazi for locations around (long -49.5221/lat -24.4537)">Cargill</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.5221&amp;materialsCitation.latitude=-24.4537" title="Search Plazi for locations around (long -49.5221/lat -24.4537)">M1</a>, 24.4537ºS,\ 49.5221ºW, 02.V.2023,\ L.V.Graf, prato azul ”, ♂, “ DZUP 540115 ”, “ Brasil, PR, Piraquara,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.9839&amp;materialsCitation.latitude=-25.4967" title="Search Plazi for locations around (long -48.9839/lat -25.4967)">Mananciais da Serra</a>,\ 25.4967ºS 48.9839ºW,\ 1010m, 16.ii-11.iii.2019,\ Melo &amp; Martins, Malaise ”, ♂, “ DZUP 540116 ”, “ Brasil, PR, Piraquara,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.9839&amp;materialsCitation.latitude=-25.4967" title="Search Plazi for locations around (long -48.9839/lat -25.4967)">Mananciais da Serra</a>,\ 25.4967ºS 48.9839ºW,\ 1010m, 16.ii-11.iii.2019,\ Melo &amp; Martins, Malaise ”. Brasil, SC, DZUP. ♂, “ DZUP 540117 ”, “ Florianópolis, SC\ Brasil, 03.IV.2001 \ T. M. Guerra col.\\ 9 10:04” .</p></div>	https://treatment.plazi.org/id/8C7BE1229E0DFFFC76F01CEC2ACAFAD2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cappola, Victor;Gonçalves, Rodrigo B.	Cappola, Victor, Gonçalves, Rodrigo B. (2026): A review of Caenohalictus species from southern Brazil (Apoidea, Caenohalictini). Zootaxa 5782 (3): 401-448, DOI: 10.11646/zootaxa.5782.3.1, URL: https://doi.org/10.11646/zootaxa.5782.3.1
8C7BE1229E0FFFF976F019DD291BFA8A.text	8C7BE1229E0FFFF976F019DD291BFA8A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caenohalictus palumbes (Vachal 1903)	<div><p>8. Caenohalictus palumbes (Vachal, 1903)</p><p>(Figs. 18, 19, 29A–C, 31D)</p><p>Halictus (Paragapostemon) palumbes Vachal, 1903 . Type data: lectotype ♀ Type locality: Brazil, Rio de Janeiro, Nova Friburgo. Naturhistorisches Museum in Wien. Examined through photographs.</p><p>Diagnosis. Caenohalictus palumbes is a large species, measuring from 8 to 10mm, with maxillary palpomeres measuring about 1x the MOD and eyes not enlarged as in C. longipalpis sp. nov. and C. sesquipalpis sp. nov. It resembles C. cyanolucens sp. nov. and C. parviformis sp. nov., but can be separated by the larger size in both sexes, females with scutellum densely punctate and polished, propodeal pit enclosed, weak carinae almost parallel and S1 with strong metallic reflections. From males of C. cyanolucens sp. nov., it can be easily separated by flagellomeres 2–11 homogeneously covered by intermixed trichodea and placodea sensillae and mandible with preapical tooth. The male preapical tooth is found only in C. palumbes among examined species. The male is described for the first time.</p><p>Redescription</p><p>Female: Body length x̄ =9.37 (N=4). ITD x̄ =1.69 (N=4). UID x̄ =1.32 (N=4). LID x̄ =1.30 (N=4). CLL x̄ =0.52 (N=4). HL x̄ =2.09 (N=4). HW x̄ =2.29 (N=4). Head. Metallic green or cuprous, sometimes with golden reflections. Labrum: black; apical elevation absent; base rugose. Mandible: black; apical amber spot occupying 1/3 of its length; preapical teeth present. Maxillary palpomeres length:0.7–1.0 the size of MOD. Clypeus:punctate to sparsely punctate (I=PD; I&gt;PD); areolate; thin and long setae (2.5–3 MOD approximately), on its apex with shorter, denser and little thicker setae (2 MOD), intermixed with a comb of very short setae; apical dark band occupying a little less than 1/2 its length. Epistomal lobe: angle approximately orthogonal, projected onto the clypeus. Supraclypeal area: sparsely punctated (I&gt;PD); areolate. Parocular area: brown setae, plumose and long (1,5– 2 MOD). Antennae: scape black; flagellomeres black to dark brown. Frons: slightly punctate, blending with the lineolate to areolate sculpture. Eye setae: approximately 1 MOD. Gena: thin and sparse long plumose setae (2–4 MOD); coriaceous; sparsely punctate; metallic green, turquoise or goldish green. Mesosoma. Mesoscutum: sparsely punctated to punctated in a few areas; granulate. Scutellum: coriaceous posteriorly, only on the apical edge; densely punctated (&lt;1pd). Mesepisternum: areolate to rugulose, coriaceous below. Metanotum: rugulose-punctulate to coriaceous. Metepisternum: areolate. Legs: fore coxa and femur dark brown with metallic dark green reflections, dark brown elsewhere; fore leg with white plumose setae on trochanter and femur, light brown elsewhere, median and hind leg with dark brown plumose setae. Metapostnotum: medium depression occupying only the median area, a little more expanded anteriorly; weakly striate, irregular or branched shaped, rugose; granulate. Propodeum: dense and thin long plumose setae; pit weakly carinate, not enclosed above. Metasoma. Terga: green or golden anteriorly; black on the disc; marginal area dark brown; on T3 with a blue, green or goldish green reflection on the middle, between the black area and marginal area; T4 without the black area, also with the same reflections on the middle before the marginal area. Sterna: S1 bright metallic green; S2–S6 black to dark brown; dense and thin plumose setae on the post gradular area, glabrous on the pre gradular area.</p><p>Description</p><p>Male: Body length x̄ =9.5 (N=5). ITD x̄ =1.53 (N=5). UID x̄ =1.35 (N=5). LID x̄ =1.02 (N=5). CLL x̄ =0.51 (N=5). HL x̄ =2.02 (N=5). HW x̄ =2.17 (N=5). Head. Metallic green, goldish green or dark green, above de frons black, dark blue, turquoise or metallic green. Labrum: yellow. Mandible: black; amber apical spot occupying around 1/3 of its length; preapical teeth present. Maxillary palpomeres length: 0.5–1.0x MOD. Clypeus: sparsely punctate to densely punctate below; areolate to micro lineolate; yellow spot occupying around 1/2 of the clypeal area, with a dark edge separating it from the green area. Epistomal lobe: angle obtuse and not projected onto the clypeus. Supraclypeal area: areolate; sparsely punctate. Parocular area: yellow and thick plumose setae, dense around the antennal socket, with brown and thin plumose setae on the supraclypeal area, lower parocular area and clypeus. Antennae: scape black, sometimes with weak cuprous reflections; flagellomeres black to yellow anteriorly; crenulate, F2 longer than each F3–F10. Frons: areolate-rugulose; inconspicuous punctation, blending with the sculpture. Eye setae: long (1.0– 1.2x MOD). Gena: black, turquoise, metallic green or goldish green; coriaceous-rugulose; slightly punctate; long and thick yellow plumose setae, gradually becoming thin and short above the middle. Mesosoma. Integument color: black, goldish green, turquoise or metallic green with a few bluish spots. Mesoscutum: inconspicually and sparsely punctate; granulate. Scutellum: lineolate; punctate to densely punctate. Mesepisternum: areolate. Metanotum: areolate; slightly punctate. Metepisternum: areolate. Legs: dark brown, with dark blue reflections on the coxa, except on the middle coxa or brown to light brown, with metallic green reflections from coxa to femur, except on the middle coxa. Metapostnotum: median depression occupying only the median area, a little more expanded anteriorly; striae areolate-rugose, sometimes weakly; granulate. Propodeum: dense, short and thick plumose setae on the sides, weakly near the metapostnotum; pit weakly carinate, not enclosed above, sometimes weakly. Metasoma. Terga: dark green, blue or metallic green with blue reflections, with a small black area before the dark brown marginal area. Sterna: S1 same color of the integument; brown or dark brown to black elsewhere. Genitalia. Gonobase: wider than long, width a little smaller than the length of the gonocoxite. Volsella: length a little more than 1/2 of gonapophysis, wider than long. Gonapophysis: ventral prong and dorsal flanges absent; dorsal crest absent; with small dorsal setae on the outer edges in the middle. Gonocoxite: longitudinally striate. Gonostylus: VA vestigial; VP absent; MGL reniform; IGP dorsal process as an oblong lobe, with dense, small and very thick setae, lower apex with the shape of a hook; OGP with 3 long setae.</p><p>Variation. This species can vary greatly in size, from 8 to 10 mm and also exhibits a great range of body colors, especially on metasoma: opaque green, metallic green, metallic blue, cupreous, black or turquoise. Vachal (1903:99) already recognized the color variation for this species.</p><p>Distribution. Brazil: Minas Gerais, Paraná, Rio de Janeiro, Rio Grande do Sul, São Paulo.</p><p>Examined material. Brazil, MG, DZUP. ♀, “540163”, “ P. de CALDAS-MG \ Brasil XI-61\ Claudionor Elias l. ” . Brasil, PR, DZUP. ♀, ♂, “ DZUP 539976 ”, “ Flôres\ de\ Arruda \\ Curityba-Paraná\ 21-IX-1942 \\ Caenohalictus \ palumbes\ P. Moure (Vach)\ 1952”, ♀, “ DZUP 539977 ”, “Curityba-Paraná\ 13-IX-1942 ”, ♀, “ DZUP 539978 ”, “Curityba-Paraná\ IX-1941 \\ palumbes\ Vach\ det. J.S. Moure 19”, ♀, “ DZUP 539979 ”, “CURITIBA\ Paraná BRASIL \ XI.1955. MOURE \ LANÇE, MICHENER ”, ♀, “ DZUP 539980 ”, “Curityba-Paraná\ XII.39\\ palumbes\ Vach\ det. J.S. Moure 19”, ♂, “ DZUP 539981 ”, “ Brasil, Paraná, 10Km N de\ Bocaiúva do Sul, Santana,\ 15º06’S, 49º06’W, 900m \ 12.x.2002, GMelo, LZanette &amp;\ AAguiar ”, ♂, “ DZUP 539982 ”, “ CURITIBA \ Paraná BRASIL \ I - 1961 ”, ♂, “ DZUP 539984 ”, “ Brasil, Paraná, Campina\ Grande do Sul, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.1&amp;materialsCitation.latitude=-15.1" title="Search Plazi for locations around (long -49.1/lat -15.1)">Pico</a> \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.1&amp;materialsCitation.latitude=-15.1" title="Search Plazi for locations around (long -49.1/lat -15.1)">Caratuva</a>, 100-1480m,\ 25º14’S 48º50’W,\ 17.ii.2007, G.A.R. Melo,\ F. vivallo ”, ♀, “ DZUP 540147 ”, “ BRASIL, PARANÁ,\ COLOMBO, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.1&amp;materialsCitation.latitude=-15.1" title="Search Plazi for locations around (long -49.1/lat -15.1)">EMBRAPA</a> \ 25º19’18.66’’S 49º9’33.66’’W \ 26.VIII.2019 935m \ MIKICH, S.B. leg. E2988”, ♀, “ DZUP 540148 ”, “ BRASIL, PARANÁ,\ COLOMBO, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.1&amp;materialsCitation.latitude=-15.1" title="Search Plazi for locations around (long -49.1/lat -15.1)">EMBRAPA</a> \ 25º19’18.66’’S 49º9’33.66’’W \ 9.IX.2019 935m \ MIKICH, S.B. leg. E3079”, ♀, “ DZUP 540149 ”, “ BRASIL, PARANÁ,\ COLOMBO, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.1&amp;materialsCitation.latitude=-15.1" title="Search Plazi for locations around (long -49.1/lat -15.1)">EMBRAPA</a> \ 25º19’18.66’’S 49º9’33.66’’W \ 3.X.2019 935m \ MIKICH, S.B. leg. E3572”, ♀, “ DZUP 600472 ”, “ Curitiba \ X.943\ R.Huke \\ Paragapostemon \ palumbes\ (Vachal) ♀ ”, ♂, “ DZUP 600474 ”, “ Curitiba - PR \ 7 / II / 1965 \ R.Lange leg.”, ♀, “ DZUP 169070 ”, “ Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.1&amp;materialsCitation.latitude=-15.1" title="Search Plazi for locations around (long -49.1/lat -15.1)">Tunas do\ Paraná, Parque Estadual\ de Campinhos</a>, 25º02’S \ 49º05’W, Alt. 860m,\ 11.ix.2008, G. Weiss ” . Brasil, RJ, DZUP. ♀, “ DZUP 540128 ”, “ Brasil, Rio de Janeiro,\ Nova Friburgo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-42.483334&amp;materialsCitation.latitude=-22.366667" title="Search Plazi for locations around (long -42.483334/lat -22.366667)">Macaé\ de Cima</a>, 1500m,\ 22º22’S, 42º29’W,\ 15.xi.2005, P. Grossi ” . Brasil, RS, DZUP. ♂, “ DZUP 539983 ”, “ CANELA, RS \ Brasil 26.VIII.1985 \ M. Hoffmann col.\\ col. MCN 91336\\ Caenohalictus ♂\ hoffmann” . Brasil, SP, DZUP. ♀, “ DZUP 540126 ”, “ SÃO PAULO \ IX-1953 \ C. Jordão ”, ♀, “ DZUP 540127 ”, “ C. do Jordão-SP \ Brasil II 1958 \ K.Lenko leg”. MCT. ♂, “ São Francisco de\ Paula, Pró-Mata, RS,\ Brasil 09.03.1997 \ leg. Birgit Harter \\ 12811 H: 239\ F: 85 11:30 ♀\\ Caenohalictus palumbes \ Vachal, 1903\ B. Harter det. 1999”, ♂, “ São Francisco de\ Paula, Pró-Mata, RS,\ Brasil 22.2.1997 \ leg. Birgit Harter \\ 14691 H: 228\ F: 86 10:30\\ Caenohalictus palumbes \ Vachal, 1903\ B. Harter det. 1999” .</p></div>	https://treatment.plazi.org/id/8C7BE1229E0FFFF976F019DD291BFA8A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cappola, Victor;Gonçalves, Rodrigo B.	Cappola, Victor, Gonçalves, Rodrigo B. (2026): A review of Caenohalictus species from southern Brazil (Apoidea, Caenohalictini). Zootaxa 5782 (3): 401-448, DOI: 10.11646/zootaxa.5782.3.1, URL: https://doi.org/10.11646/zootaxa.5782.3.1
8C7BE1229E0AFFF876F019252959FC06.text	8C7BE1229E0AFFF876F019252959FC06.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caenohalictus parviformis Cappola & Gonçalves 2026	<div><p>9. Caenohalictus parviformis sp. nov.</p><p>(Figs. 1C–D, 2A, 20, 31D)</p><p>Diagnosis. This species has the epistomal lobe projected over the clypeus and maxillary palpomeres about 1 MOD. Females can be distinguished from C. palumbes by having smaller size, approximately 7.5mm, elongated head (Fig. 20B), scutellum sparsely punctate(Fig. 20C); propodeal fossa not enclosed above (Fig. 2B), S1 with weak metallic reflections; terga olive-green. From C. cyanolucens sp. nov. by the elongate head, conspicuous eye setae; gena with the same color as the integument, olive-green, opaque. From C. shulthessi, it can be separated also by the bright color. Males are not known.</p><p>Description</p><p>Female: Body length x̄ =7.40 (N=2). ITD x̄ =1.46 (N=2). UID x̄ =1.10 (N=2). LID x̄ =1.00 (N=2). CLL x̄ =0.45 (N=2). HL x̄ =1.81 (N=2). HW x̄ =1.85 (N=2). Head. Metallic green, with a few bluish reflections on the edge of the parocular fovea, sometimes inconspicuous. Labrum: black; apical elevation presente; base rugose. Mandible: mostly black, with a small apical amber spot; preapical teeth present. Maxillary palpomeres length: 0.7–1.0 the size of MOD. Clypeus: punctate to sparsely punctulate; areolate; thin plumose setae (2–2,5 MOD approximately); middle of the apical dark band going to almost half of the clypeus, triangular shaped, with the sides slightly declining towards the edges. Epistomal lobe: angle approximately orthogonal, projected onto the clypeus. Supraclypeal area: sparsely punctated (I&gt;PD); areolate. Parocular area: whitish setae, plumose and long (2–3 MOD), thick on the parocular area. Antennae: Scape black; flagellomeres black to light brown or yellowish anteriorly. Frons: slightly punctate, blending with the areolate sculpture, finely on the frons. Eye setae: approximately 1 MOD. Gena: thick plumose setae (1.5– 4x MOD); micro lineolate to finely areolate on a few areas; punctulate to sparse punctate; metallic green. Mesosoma. Mesoscutum: sparsely punctated to very sparsely punctated; granulate. Scutellum: coriaceous; sparsely punctated (&gt;1pd). Mesepisternum: areolate to micro canalictulated ventrally. Metanotum: rugulose to areolate. Metepisternum: areolate. Legs: fore coxa and femur dark brown with metallic dark green reflections, with fore tibia to tarsi light brown with a few yellow areas, dark brown elsewhere; yellowish plumose setae. Metapostnotum: medium depression occupying only the median area, a little more expanded anteriorly; weakly striate, irregular or branched shaped, rugose; granulate. Propodeum: thick and long plumose setae, thinner and shorter near the propodeal pit; pit not carinate, enclosed, parallel. Metasoma. Terga: dark green; marginal area dark brown. Sterna: S1 dark brown. S2–S6 dark brown; dense and thin plumose setae on the post gradular area, glabrous on the pre gradular area.</p><p>Male: Not known.</p><p>Etymology. A compound name from Latin word parvus (“small”) and the suffix -formis (“in the form of, shaped as”), referring to the peculiar condition of this species, which belongs to a group characterized by being comparatively large in size, but with a body size resembling that of the group with smaller size.</p><p>Distribution. Brazil: Minas Gerais and Paraná.</p><p>Type material. Holotype. Brasil, PR, DZUP. ♀, “ DZUP 606585 ”, “ Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.993057&amp;materialsCitation.latitude=-25.23139" title="Search Plazi for locations around (long -49.993057/lat -25.23139)">Parque\ Estadual de Vila Velha</a>,\ Área I, 25º13’53’’S,\ 49º59’35’’W, 03.III.2024,\ Gonçalves, Graf, Pundek ” . Paratype. Brasil, MG, DZUP. ♀, “ DZUP 540103 ”, “ Brasil, Minas Gerais,\ Ouro Preto, 1400m \ 20º22’ S 43º30’ W \ 8-9.i.2010, K. Ramos \ &amp; V. Kanamura ” .</p></div>	https://treatment.plazi.org/id/8C7BE1229E0AFFF876F019252959FC06	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cappola, Victor;Gonçalves, Rodrigo B.	Cappola, Victor, Gonçalves, Rodrigo B. (2026): A review of Caenohalictus species from southern Brazil (Apoidea, Caenohalictini). Zootaxa 5782 (3): 401-448, DOI: 10.11646/zootaxa.5782.3.1, URL: https://doi.org/10.11646/zootaxa.5782.3.1
8C7BE1229E34FFC676F01CEC2D64FEC6.text	8C7BE1229E34FFC676F01CEC2D64FEC6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caenohalictus schulthessi (Vachal 1903)	<div><p>10. Caenohalictus schulthessi (Vachal, 1903)</p><p>(Fig. 21, 31B)</p><p>Halictus (Paragapostemon) Schulthessi Vachal, 1903 . Type data: holotype MNHP &lt;F&gt;. Type locality: Brazil, Santa Catarina. Examined .</p><p>Diagnosis. This species has the epistomal lobe projected over the clypeus and maxillary palpomeres about 1 MOD. The combination of body mostly black, with few metallic reflections; forecoxa brown; antennae brown; hind tibia and tarsi with dark setae; propodeal pit strongly ridged distinguished from C. palumbes, C. parviformis sp. nov. and C. cyaneolucens sp. nov.</p><p>Redescription (from notes taken by RBG in MNHP).</p><p>Female: Body length: 8 mm. Body dark with dark blue highlights elsewhere, flagellomeres dark, fore coxa dark, tergal marginal zones brown, S1 dark. Hind tibia and tarsus with dark setae, remaining body setae yellowish. Head with triangular aspect. Epistomal lobe forming straight angle. Maxillary palpomeres length = MOD. Occipital ridge rounded. Gena shorter than eye. Pronotal anterior angle obtuse. Mesoscutum rounded anteriorly, alveolate to coriaceous at the disc. Scutellum and metanotum weakly convex. Metapostnotum with a triangular median portion, weakly concave, rugulose, rugulose at sides of this region, remaining metapostnotum microalveolate. Propodeal pit carinae parallel to weakly divergent above. Tibial spur with three subapical teeth. Terga mostly lineolate, a little coriaceous.</p><p>Male: Not known.</p><p>Variation. We examined specimens from São Paulo and Minas Gerais that agree well with C. schulthessi type, but differ in having the head with opaque green reflections rather than dark blue as the holotype. Color variation is known for other species, as C. palumbes and for males of C. longipalpis sp. nov.</p><p>Distribution. Brazil: Minas Gerais, Santa Catarina, and São Paulo.</p><p>Examined material. Brazil, MG, DZUP. ♀, “ DZUP 540145 ”, “ Brasil, Minas Gerais,\ Itamonte, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.69389&amp;materialsCitation.latitude=-22.349167" title="Search Plazi for locations around (long -44.69389/lat -22.349167)">Serra Negra</a> \ 1- 3.xi.2007 D.M.Takiya ”, ♀,“ DZUP 540146 ”, “ Brasil, Minas Gerais,\ Itamonte, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.69389&amp;materialsCitation.latitude=-22.349167" title="Search Plazi for locations around (long -44.69389/lat -22.349167)">Serra Negra</a>,\ 22º20’57’’S 44º41’38’’W \ 1-3.xi.2007, K.S.Ramos ”. 12♀, DZUP 540132 - DZUP 540143, “ SÃO PAULO \ s.p - 16/x/70\ MOURE &amp; MIELKE ”,, ♀, “ DZUP 540144 ”, “ C. do Jordão-SP \ Brasil II 1958 \ K.Lenko leg”, ♀, “ DZUP 540161 ”, “ SÃO PAULO \ s.p - 16/ x/70\ MOURE &amp; MIELKE ”, ♀, “ DZUP 540162 ”, “ SÃO PAULO \ s.p - 16/x/70\ MOURE &amp; MIELKE ” .</p></div>	https://treatment.plazi.org/id/8C7BE1229E34FFC676F01CEC2D64FEC6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cappola, Victor;Gonçalves, Rodrigo B.	Cappola, Victor, Gonçalves, Rodrigo B. (2026): A review of Caenohalictus species from southern Brazil (Apoidea, Caenohalictini). Zootaxa 5782 (3): 401-448, DOI: 10.11646/zootaxa.5782.3.1, URL: https://doi.org/10.11646/zootaxa.5782.3.1
8C7BE1229E35FFC576F01DF12CA4FB96.text	8C7BE1229E35FFC576F01DF12CA4FB96.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caenohalictus sesquipalpis Cappola & Gonçalves 2026	<div><p>11. Caenohalictus sesquipalpis sp. nov.</p><p>(Figs. 2D, 3C, 22–23, 26A, 29J–L, 31C)</p><p>Diagnosis. It has some similarities to C. longipalpis sp. nov., such as having longer maxillary palpomeres than the other species and enlarged eyes. It can be separated by palpomeres measuring approximately 0.7 to 1.5 times the MOD in both sexes; females having a linear propodeal fossa, without opening anteriorly and without adjacent carinae (Fig. 2D), and the males with dark head and mesosoma (Fig. 23) and with a strong invagination surrounded by a membrane on S6 (Fig. 26A). Females and males are separated from C. aerugineus and C. oresicoetes by the enclosed propodeal pit, more opened on C. aerugineus and deeper on C. oresicoetes .</p><p>Description</p><p>Female: Body length x̄ =9.78 (N=3). ITD x̄ =1.61 (N=3). UID x̄ =1.33 (N=3). LID x̄ =1.13 (N=3). CLL x̄ =0.49 (N=3). HL x̄ =2.12 (N=3). HW x̄ =2.31 (N=3). Head. Goldish green or dark green, sometimes with gold reflections. Labrum: black; apical elevation present, but small; base rugose. Mandible: black; apical amber spot occupying 1/3 of its length; preapical teeth present. Maxillary palpomeres length: 1.0–1.5 times MOD. Clypeus: punctate to sparsely punctulate; areolate; long, thin and sparse setae in the middle (3 MOD), and denser and thicker setae on apex; apical dark band occupying almost 2/3 of its length, more in middle. Epistomal lobe: angle approximately orthogonal, projected onto the clypeus. Supraclypeal area: punctate to sparsely punctate; coriaceous to areolate. Parocular area: whitish setae, plumose and long (2–3 MOD), thicker on the parocular area. Antennae: scape and flagellomeres black; antennal socket located in a weakly concave depression. Frons: slightly punctate, blending with the areolate sculpture. Eye setae: approximately 1 MOD. Gena: thin and sparse long plumose setae (2–4 MOD); coriaceous; sparsely punctate; dark green. Mesosoma. Mesoscutum: very sparsely punctated anteriorly to densely punctated posteriorly, with a blend of both on the disc; areolate to granulate. Scutellum: polished between punctation; densely punctated (&lt;1pd). Mesepisternum: areolate to coriaceous. Metanotum: polished surface; densely punctated; inconspicuously areolate on the sides. Metepisternum: areolate. Legs: dark brown, with fore leg tarsi light brown to yellow; coxa with white setae, legs with yellowish setae, median and hind basitarsus with white to brown setae. Metapostnotum: medium depression occupying only the median area, a little more expanded anteriorly; weakly striated, irregular or branched shaped, rugose; granulate. Propodeum: sparse and thin long plumose setae; pit not carinate, enclosed, parallel. Metasoma. Terga: green anteriorly; on the disc dark brown with purple or blue reflections, weakly on T4; marginal area light brown, with green reflections on T3 and T4. Sterna: S1 dark brown; S2–S6 dark brown; dense and thin plumose setae on the post gradular area, glabrous on the pre gradular area.</p><p>Male: Body length x̄ =9.44 (N=1). ITD x̄ =1.48 (N=2). UID x̄ =1.22 (N=2). LID x̄ =0.86 (N=2). CLL x̄ =0.48 (N=2). HL x̄ =2.02 (N=2). HW x̄ =2.13 (N=2). Head. Dark green, almost black, with a few lighter green areas on the lower parocular area and on the inner orbit of the eye, with weak green reflections on the clypeus and on the supraclypeal area. Labrum: yellow. Mandible: black; amber apical spot occupying nearly 1/2 of its length; lacking preapical teeth. Maxillary palpomeres length: 0.7–1.5x MOD. Clypeus: punctate; coriaceous; yellow spot occupying around 1/2 of its length, hill-shaped, with a dark band before green base. Epistomal lobe: angle approximately orthogonal, projected onto the clypeus. Supraclypeal area: areolate to rugulose; with inconspicuous punctation. Parocular area: yellow to white plumose, thick and long setae, on the clypeus thinner and brown. Antennae: scape and flagellomeres black, yellow anteriorly on the flagellomeres; crenulate, F2 as long as each F3–F10. Frons: areolate; inconspicuous punctation, blending with the sculpture. Eye setae: 0.7–1.0x MOD. Gena: dark metallic blue; lineolate to rugulose; slightly punctate; long plumose setae. Mesosoma. Integument color: black to dark green on the mesoscutum and scutellum; goldish green on the pronotum lobe, metanotum, metapostnotum and propodeum; dark blue on the sides and ventrally of mesosoma. Mesoscutum: punctate in a few areas, sparsely to very sparsely punctate elsewhere. Scutellum: weakly lineolate on the edges, almost polished on the disc; densely punctate. Mesepisternum: areolate. Metanotum: coriaceous, weakly on the disc; punctate to densely punctate. Metepisternum: areolate. Legs: brown, on the fore coxa with green reflections, tibia to tarsi lighter brown, with a few yellow areas on the tibia and basitarsus; white to yellow setae. Metapostnotum: median depression occupying only the median area, a little more expanded anteriorly; longitudinal striae rugose, branched or areolate, sometimes very weakly; areolate. Propodeum: dense and thin plumose setae on the sides, more sparse near the metapostnotum; pit not carinate, enclosed, parallel. Metasoma. Terga: green anteriorly; brown on the disc and on the marginal area, on the marginal area of T3 and T4 with green reflections. Sterna: S1 dark brown, light brown elsewhere, almost yellow, with the apex of S2 and S3 yellow. Volsella: length 3/4 the gonapophysis, longer than wide. Gonapophysis: with small ventral prong; small dorsal flanges starting almost at the middle of gonapophysis, with same width as the gonapophysis; dorsal crest with a few dorsal setae on the outer edge, near the flanges, and in the middle; with a few ventral setae in the middle of the gonapophysis; gonapophysis wider subapically. Gonocoxite: longitudinally striate. Gonostylus: VA membranous and enlarged, with dense and short setae in the apex to the middle, and 6 long setae in the lower edges; MGL reniform to arcuated, concave; IGP expanded and with a large and concave lobe, with dense and small setae on the concave edge; OGP with a few long and branched setae.</p><p>Etymology. A compound name from Latin prefix sesqui- (“one and a half times, unusually long”) and palpis (“palpus, palp”), in reference to the long maxillary palpomeres, measuring about one and a half times the median ocellus diameter. The epithet is treated as a noun in apposition.</p><p>Distribution. Brazil: Paraná and São Paulo.</p><p>Type material. Holotype. Brazil, PR, DZUP. ♀, “ DZUP 024897 ”, “ Brasil, Paraná,\ São José dos Pinhais,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.183334&amp;materialsCitation.latitude=-25.516666" title="Search Plazi for locations around (long -49.183334/lat -25.516666)">Aeroporto Afonso Pena</a>,\ KF, 25º31’S 49º11’W,\ 14.v.2005, A.Martins \ R.Gonçalves.\\ SJP 2676” . Paratype. Brazil, PR, DZUP. ♀, “ DZUP 540118 ”, “ 25-11-2000 \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15935&amp;materialsCitation.latitude=-25.321852" title="Search Plazi for locations around (long -49.15935/lat -25.321852)">TIROL DAS TORRES</a>, UBE-\ RABA, CURITIBA, PARANÁ \ 900m, O. MIELKE LEG.”, ♀, “ DZUP 540119 ”, “ BRASIL, PARANÁ,\ COLOMBO, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15935&amp;materialsCitation.latitude=-25.321852" title="Search Plazi for locations around (long -49.15935/lat -25.321852)">EMBRAPA</a> \ 25º19’18.66’’S 49º9’33.66’’W \ 9.IX.2019 935m \ MIKICH, S.B. leg. E3080”, ♂, “ DZUP 540120 ”, “ Curityba-Paraná \ X-1944 ”, ♂, “ DZUP 540121 ”, “ Brasil, Paraná,\ Mangueirinha,\ 25º47’19’’S, 52º07’02’’O \ 10.IX.2024, D.A.Roher,\ rede entomol. ”, ♂, “ DZUP 540159 ”, “ Brasil, Paraná, Honório \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.15935&amp;materialsCitation.latitude=-25.321852" title="Search Plazi for locations around (long -49.15935/lat -25.321852)">Serpa</a>, 26º07’56’’S,\ 52º31’09’’W, 23.V.2025,\ Graf L. V., prato colorido ”, ♂, “ DZUP 540160 ”, “ Curityba-Paraná \ 12-ix-1944 \\ Caenohalictus \ palumbes\\ UMNG-ins\ 001448”. Brasil, SP, DZUP. ♀, “ DZUP 540164 ”, “ C. do Jordão-SP \ Brasil II 1958 \ K.Lenko leg” .</p></div>	https://treatment.plazi.org/id/8C7BE1229E35FFC576F01DF12CA4FB96	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cappola, Victor;Gonçalves, Rodrigo B.	Cappola, Victor, Gonçalves, Rodrigo B. (2026): A review of Caenohalictus species from southern Brazil (Apoidea, Caenohalictini). Zootaxa 5782 (3): 401-448, DOI: 10.11646/zootaxa.5782.3.1, URL: https://doi.org/10.11646/zootaxa.5782.3.1
8C7BE1229E37FFCF76F0188329E3F9CC.text	8C7BE1229E37FFCF76F0188329E3F9CC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caenohalictus tesselatus (Moure 1940)	<div><p>12. Caenohalictus tesselatus (Moure, 1940)</p><p>(Figs. 24–25, 28A–C, 30D)</p><p>Paragapostemon tesselatus Moure, 1940 . Type data: holotype ♀ Type locality: Brazil, Paraná, Curitiba. Departamento de Zoologia, Universidade Federal do Paraná. Examined .</p><p>Diagnosis. Small species (5–6 mm) similar to C. implexus and C. leticiae sp. nov. by the rugulose metapostnotum and olivaceus green body. Both sexes can be differentiated from these species by the scape with metallic reflections (Fig. 24B), the short apical dark band on clypeus. Females of C. tesselatus have dense, yellow long branched setae, especially on the head and mesoscutum (Fig. 24C), and the propodeal lateral surface granulate-areolate to granulate-lineolate. Males have S1 entirely bright green while in the other species the metallic reflections are reduced.</p><p>Redescription</p><p>Female: Body length x̄ =6.44 (N=5). ITD x̄ =1.18 (N=5). UID x̄ =1.03 (N=5). LID x̄ =0.92 (N=5). CLL x̄ =0.32 (N=5). HL x̄ =1.37 (N=5). HW x̄ =1.64 (N=5). Head. Metallic green or cuprous, sometimes with golden reflections. Labrum: dark brown; apical elevation present, but small; base rugose. Mandible: black with metallic green reflections basally, cuprous elsewhere; apical dark amber spot occupying 1/4 of its length; preapical teeth present. Maxillary palpomeres length: 0.7 the size of MOD or less. Clypeus: sparsely punctate (I&gt;PD); coriaceous, polished near and at the black apical stripe; thin and long setae (2,5–3 MOD approximately); apical dark band occupying less than 1/3 its length. Epistomal lobe: angle obtuse and not projected onto the clypeus. Supraclypeal area: sparsely punctated (I&gt;PD); coriaceous. Parocular area: yellowish setae, plumose and long (2–3 MOD), thicker in the upper parocular area and thinner in the lower parocular area. Antennae: scape black with greenish or golden reflections; flagellomeres black to brown, lighter and with yellow spots anteriorly, stronger in F1 and F2, smoother elsewhere. Frons: sparsely punctate, more inconspicuous at the coriaceous to areolate sculpture. Eye setae: approximately 1 MOD. Gena: thin and long plumose setae (2–5 MOD); coriaceous; sparsely punctate; metallic green, sometimes with golden reflections. Mesosoma. Mesoscutum: sparsely punctated (I&gt;PD); coriaceous to areolate. Scutellum: coriaceous; sparsely punctated on the disc, punctate on the edges. Mesepisternum: rugulose to coriaceous. Metanotum: coriaceous; inconspicuous punctation. Metepisternum: rugulose. Legs: coxa to femur dark brown, with green metallic reflections, tibia to tarsi light brown posteriorly to yellow anteriorly; whitish plumose setae. Metapostnotum: medium depression occupying most of the dorsal area; longitudinally striate, confused-rugose, carinate or branched; coriaceous. Propodeum: thin and short plumose setae, pit carinate, not enclosed above. Metasoma. Terga: green or cuprous anteriorly; black on the disc; marginal area light brown or cuprous, with green or goldish green reflections. Sterna: S1 dark brown or black with green metallic reflections; S2–S6 dark brown or black, with green metallic reflections; dense and thin plumose setae on the post gradular area, glabrous on the pre gradular area.</p><p>Male: Body length x̄ =6.20 (N=5). ITD x̄ =1.05 (N=5). UID x̄ =0.98 (N=5). LID x̄ =0.67 (N=5). CLL x̄ =0.30 (N=5). HL x̄ =1.36 (N=5). HW x̄ =1.51 (N=5). Head. Cuprous to goldish green. Labrum: yellow. Mandible: mostly yellow, only a small anterior portion being black; amber apical spot occupying around 1/4 of its length; lacking preapical teeth. Maxillary palpomeres length: 0.5–0.7x MOD. Clypeus: punctate to sparsely punctate; coriaceous; yellow spot occupying only the apex, around 1/4 to 1/5 its length. Epistomal lobe: angle obtuse and not projected onto the clypeus. Supraclypeal area: coriaceous; punctate to sparsely punctate. Paraocular area: pubescence white, thick and dense plumose setae, on the supraclypeal area and on the clypeus thinner and yellow plumose setae. Antennae: scape metallic green or cuprous, with black apex; flagellomeres dark brown to yellow anteriorly; flagellomeres 2–11 with anterior large patches of sensilla placodea; crenulate, F2 longer than each F3– F10. Frons: coriaceous-rugulose; inconspicuous punctation, blending with the sculpture. Eye setae: long (1.0–1.2x MOD). Gena: metallic green to goldish green; micro lineolate-rugulose; slightly punctate to sparsely punctate; long and thick white plumose setae. Mesosoma. Integument color: cuprous to goldish green. Mesoscutum: sparsely punctate to very sparsely punctate; coriaceous. Scutellum: polished to coriaceous on the edges; very sparsely punctated to sparsely punctate to punctate. Mesepisternum: coriaceous-rugulose to coriaceous to areolate. Metanotum: coriaceous-rugulose; punctate to sparsely punctate. Metepisternum: areolate-rugulose to coriaceous-rugulose. Legs: coxa, trocanter and femur metallic green, except the middle leg coxa, that is black, fore leg tibia dark brown to yellow, yellow elsewhere, middle leg tibia dark brown with yellow apex, yellow elsewhere, hind leg tibia dark brown, yellow elsewhere; with white to yellow plumose setae. Metapostnotum: median depression occupying most of the dorsal area; striae areolate-rugose, coriaceous-rugulose to coriaceous. Propodeum: dense and thick plumose setae on the sides, thinner and more sparse near the metapostnotum; pit carinate, not enclosed above. Metasoma. Terga: dark green or cuprous anteriorly; black or dark brown on the disc; brown or dark brown marginal area. Sterna: S1 dark brown to metallic green, dark brown to light brown elsewhere, with green reflections. Genitalia. Gonobase: wider than long, was wide as 1/2 the gonocoxite. Volsella: as long as 1/2 gonapophysis, wider than long. Gonapophysis: with small ventral prong; dorsal flanges; dorsal depressions; dorsal ridge; dorsal crest. Gonocoxite: longitudinally striate. Gonostylus: VA long and narrowly oblong lobe with very small setae on its apex, inconspicuous; VP absent; MGL reniform to sigmoid; IGP with a dense and thick basal setose plate; OGP with dense and small setae.</p><p>Variation. Some specimens have a goldish green coloration, but most are green in color.</p><p>Distribution. Brazil: Goiás, Minas Gerais, Paraná, Rio Grande do Sul, Santa Catarina and São Paulo.</p><p>Examined material. Brazil, GO, DZUP. ♀, “ DZUP 540189 ”, “Faz. Nova Orlandia/ Jataí, Go - Brasil/ I. 964 - Martins, Morgante &amp; Silva”, ♀, “ DZUP 540190 ”, “Cabeceiras\ (Lagôa Formosa)\ Goiás 24-27. X. 1964 \ Exp. Dep. Zool ”, ♀, “ DZUP 540191 ”, “Faz. Nova Orlandia/ Jataí, Go - Brasil/ I. 964 - Martins, Morgante &amp; Silva” . Brasil, MG, DZUP. 55♂, DZUP 540187 - DZUP 540242, “ POÇOS CALDAS \ MG - Brasil XI 61\ Claudionor Elias” . Brazil, PR, DZUP. ♀, “ DZUP 539968 ”, “Brasil, Paraná,\ Guarapuava, P2, Am.16,\ -25.4305, -51.7651,\ 06.I.2022, L. S. Passos,\ rede entomológica”, ♀, “ DZUP 539991 ”, “CURITIBA-- PR \ Br. 18-X-1967 \ S. LAROCA LEG \\ sb 3”, ♀, “ DZUP 539992 ”, “CURITIBA--PR\ Br. 18-X-1967 \ S. LAROCA LEG \\ 3 sb” , ♂, “ DZUP 539993 ”, “CURITIBA- -PR\ Br. 11-XI-1967 \ S. LAROCA LEG \\ 1 sb” , ♀, “ DZUP 539994 ”, “C U R I T I B A\ Paraná BRASIL \ 22-IX-1960 ” , ♀, “ DZUP 539995 ”, “CURITIBA-PARANÁ\ BRASIL- 23/9/1955 \ Moure &amp; Michener”, ♀, “ DZUP 539996 ”, “ CURITIBA - PR \ Brasil-IX.61\ S. L A R O C A”, ♀, “ DZUP 539997 ”, “ CURITIBA 950m \ Paraná BRASIL \ 27- IX-1955 Moure,\ Michener e Lange”, ♂, “ DZUP 539998 ”, “Curitiba\ 27.9.55”, ♀, “ DZUP 539999 ”, “C U R I T I B A\ Paraná BRASIL \ 22-IX-1960 ”, ♂, “ DZUP 540000 ”, “ CURITIBA, Paraná \ B r a s i l\ III-1954 \ Pe.J.S.Moure leg.”, ♂, “ DZUP 540001 ”, “CURITIBA-PARANÁ\ BRASIL- 23/9/1955 \ Moure &amp; Michener”, ♂, “ DZUP 540002 ”, “CURITIBA-PARANÁ\ BRASIL- 23/9/1955 \ Moure &amp; Michener”, ♀, ♂, “ DZUP 540003 ”, “ CURITIBA, Paraná \ B r a s i l\ III-1954 \ Pe.J.S.Moure leg.”, ♀, “ DZUP 540004 ”, “ Brasil, Paraná, Curitiba,\ Centro Politécnico,\ 5.xi.2005, G. Melo ” , ♂, “ DZUP 540005 ”, “ Est. Curitiba-Paranaguá \ Km 8 Paraná BRASIL \ 14-IX-1955 Lange,\ Michener e Moure” , ♀, “ DZUP 540006 ”, “ Est. Curitiba-Paranaguá \ Km 8 Paraná BRASIL \ 14-IX-1955 Lange,\ Michener e Moure” , ♀, “ DZUP 540007 ”, “ Est. Curitiba-Paranaguá \ Km 8 Paraná BRASIL \ 14-IX-1955 Lange,\ Michener e Moure” , ♀, “ DZUP 540008 ”, “ Est. Curitiba-Paranaguá \ Km 27 Paraná BRASIL \ 14-IX-1955 Lange,\ Michener e Moure” , ♂, “ DZUP 540009 ”, “ Est. Curitiba-Paranaguá \ Km 27 Paraná BRASIL \ 14-IX-1955 Lange,\ Michener e Moure” , ♂, “ DZUP 540010 ”, “ Est. Curitiba-Paranaguá \ Km 27 Paraná BRASIL \ 14-IX-1955 Lange,\ Michener e Moure” , ♂, “ DZUP 540011 ”, “ Est. Curitiba-Paranaguá \ Km 27 Paraná BRASIL \ 14-IX-1955 Lange,\ Michener e Moure” , ♂, “ DZUP 540012 ”, “ Est. Curitiba-Paranaguá \ Km 8 Paraná BRASIL \ 14-IX-1955 Lange,\ Michener e Moure” , ♀, “ DZUP 540013 ”, “C A S T R O PR \ Brasil 8 IX 1966 \ Marinoni, Fonseca”, ♂, “ DZUP 540014 ”, “CASTRO-PR\ Brasil I - 61\ A. SALGADO”, ♀, “ DZUP 540015 ”, “CASTRO-PR\ Brasil-XII 64\ F. Giacomel”, ♀, “ DZUP 540016 ”, “C A S T R O PR \ Brasil 8 IX 1966 \ Marinoni, Fonseca”, ♀, “ DZUP 540017 ”, “GUARAPUAVA 1120m \ Paraná BRASIL \ 7-IX-1955 \ Michener e Moure”, ♂, “ DZUP 540018 ”, “GUARAPUAVA 1120m \ Paraná BRASIL \ 7-IX-1955 \ Michener e Moure”, ♂, “ DZUP 540019 ”, “GUARAPUAVA 1120m \ Paraná BRASIL \ 7-IX-1955 \ Michener e Moure”, ♀, “ DZUP 540020 ”, “GUARAPUAVA 1120m \ Paraná BRASIL \ 7-IX-1955 \ Michener e Moure”, ♂, “ DZUP 540021 ”, “GUARAPUAVA 1120m \ Paraná BRASIL \ 7-IX-1955 \ Michener e Moure”, ♂, “ DZUP 540022 ”, “GUARAPUAVA 1120m \ Paraná BRASIL \ 7-IX-1955 \ Michener e Moure”, ♀, “ DZUP 540023 ”, “ Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.5229&amp;materialsCitation.latitude=-25.13366" title="Search Plazi for locations around (long -50.5229/lat -25.13366)">Imbituva</a>,\ P1 GVI, 25.13366ºS,\ 50.52290ºW, 27.II.2022,\ L. V.Graf, rede entomológica” , ♀, “ DZUP 540024 ”, “ Brasil, PR, São Mateus \ do Sul, UN-SIX\ 27.II.2009 \ Rafael Kamke leg.\\ Ponto amostral\ Mata do túnel\ 12:00-13:00\\ CJS 6383” , ♀, “ DZUP 540025 ”, “ Brasil, PR, São Mateus \ do Sul, UN-SIX\ 26.I.2009 \ Rafael Kamke leg.\\ Ponto amostral\ Bracatingal\ 11:00-12:00\\ CJS 5612” , ♀, “ DZUP 540026 ”, “ Brasil, PR, São Mateus \ do Sul, Fazenda Durgo \ 25.I.2009 \ Rafael Kamke leg.\\ Ponto amostral\ Fazenda Durgo\ 15:00 - 16:00\\ CJS 5922” , ♀, “ DZUP 540027 ”, “ Brasil, PR, São Mateus \ do Sul, UN-SIX/ Petrobras \ 23.I.2011 \ Rafael Kamke leg.\\Ponto amostral\CAB(antigo MDI)\\ CJS9883 ” , ♂,“ DZUP540028 ”,“TIJ.SUL(VOSSOROCA)\ PR - BRASIL 14/9/75\ Pe.Moure &amp; Camargo”, ♀, “ DZUP 540029 ”, “ Brasil, Paraná, Campo \ Magro, P1, -25.348060,\ -49.439137, 19.XI.2021,\ L. V.Graf, rede entomo.” , ♀, “ DZUP 540030 ”, “ Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.14134&amp;materialsCitation.latitude=-25.35695" title="Search Plazi for locations around (long -49.14134/lat -25.35695)">Colombo</a>,\ P3, -25.35695, -49.14134,\ 23.VIII.2021, L. V. Graf,\ rede entomológica” , ♀, “ DZUP 540031 ”, “ Brasil, Paraná, São \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.6571&amp;materialsCitation.latitude=-26.0159" title="Search Plazi for locations around (long -50.6571/lat -26.0159)">Mateus do Sul</a>, P2 GV,\ 26.0159ºS, 50.6571ºW \ 01.IX.2022, L. V. Graf,\ rede entomológica” , ♀, “ DZUP 540032 ”, “ Brasil, Paraná, União da\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-51.06281&amp;materialsCitation.latitude=-26.14459" title="Search Plazi for locations around (long -51.06281/lat -26.14459)">Vitória</a>, P1 GV, 26.14459ºS,\ 51.06281ºW, 23.II.2022,\ L. V.Graf, rede entomológica” , ♂, “ DZUP 540035 ”, “COLOMBO 950 m \ Paraná BRASIL \ 18-VIII-1955 Lange,\ Michener e Moure”, ♀, “ DZUP 540037 ”, “ Brasil, Paraná, São Mateus \ do <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.65715&amp;materialsCitation.latitude=-26.01587" title="Search Plazi for locations around (long -50.65715/lat -26.01587)">Sul</a>, P2 GV, 26.01587ºS,\ 50.65715ºW 24.II.2022,\ L. V.Graf, rede entomológica” , ♀, “ DZUP 540038 ”, “ Brasil, Paraná, União da\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-51.06281&amp;materialsCitation.latitude=-26.14459" title="Search Plazi for locations around (long -51.06281/lat -26.14459)">Vitória</a>, P1 GV, 26.14459ºS,\ 51.06281ºW, 23.II.2022,\ L. V.Graf, rede entomológica” , ♀, “ DZUP 540039 ”, “ Brasil, Paraná, União da\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-51.06281&amp;materialsCitation.latitude=-26.14459" title="Search Plazi for locations around (long -51.06281/lat -26.14459)">Vitória</a>, P1 GV, 26.14459ºS,\ 51.06281ºW, 23.II.2022,\ L. V.Graf, rede entomológica” , ♀, “ DZUP 540040 ”, “ Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.5229&amp;materialsCitation.latitude=-25.1337" title="Search Plazi for locations around (long -50.5229/lat -25.1337)">Imbituva</a>,\ P1 GVI, 25.1337ºS,\ 50.5229ºW, 28. V.2022,\ L. V.Graf, rede\ entomológica” , ♀, “ DZUP 540041 ”, “ Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.5229&amp;materialsCitation.latitude=-25.1337" title="Search Plazi for locations around (long -50.5229/lat -25.1337)">Imbituva</a>,\ P1 GVI, 25.1337ºS,\ 50.5229ºW, 28. V.2022,\ L. V.Graf, rede\ entomológica” , ♂, “ DZUP 540042 ”, “COLOMBO 950 m \ Paraná BRASIL \ 18-VIII-1955 Lange,\ Michener e Moure”, ♀, “ DZUP 540044 ”, “ Brasil, Paraná,\ Guarapuava, 26º GAC,\ 25º23’S 51º28’N,\ 1100 m, 24.x.2003,\ G. Melo &amp; S. Bazilio ” , ♂, “ DZUP 540045 ”, “ Brasil, Paraná, Campo \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.366665&amp;materialsCitation.latitude=-25.35" title="Search Plazi for locations around (long -49.366665/lat -25.35)">Magro</a>, 1070m,\ 25º21’S 49º22’W,\ 5.iv.2008, G. Melo ” , ♂, “ DZUP 540046 ”, “ Brasil, Paraná, Campo \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.366665&amp;materialsCitation.latitude=-25.35" title="Search Plazi for locations around (long -49.366665/lat -25.35)">Magro</a>, 1070m,\ 25º21’S 49º22’W,\ 5.iv.2008, G. Melo ” , ♂, “ DZUP 540047 ”, “ Brasil, Paraná, Campo \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.366665&amp;materialsCitation.latitude=-25.35" title="Search Plazi for locations around (long -49.366665/lat -25.35)">Magro</a>, 1070m,\ 25º21’S 49º22’W,\ 5.iv.2008, G. Melo ” , ♀, “ DZUP 540048 ”, “ Brasil, Paraná, Tibagi,\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.3642&amp;materialsCitation.latitude=-24.8364" title="Search Plazi for locations around (long -50.3642/lat -24.8364)">Ponto</a> 2LT, 25.xi.2021,\ 24.8364ºS 50.3642ºW,\ 775m, Muniz&amp;Barbosa,\ Busca ativa” , ♀, “ DZUP 540049 ”, “ Est. Paraná \ Campinha\ 15- XI-51\ M. Paranaense \\ Museu Paranaense\ Nº 9760” , ♀, “ DZUP 540050 ”, “ Est. Paraná \ Campininha\ 13-XI-51\ M. Paranaense \\ Museu Paranaense\ Nº 9761” , ♀, “ DZUP 540051 ”, “ Brasil, Paraná, São \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.3109&amp;materialsCitation.latitude=-25.6375" title="Search Plazi for locations around (long -50.3109/lat -25.6375)">João do Triunfo</a>, P2 GIII,\ 25.6375ºS 50.3109ºW,\ 27. V.2022, L. V.Graf,\ rede entomológica” , ♀, “ DZUP 540052 ”, “ Brasil, Paraná, São \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.6571&amp;materialsCitation.latitude=-26.0159" title="Search Plazi for locations around (long -50.6571/lat -26.0159)">Mateus do Sul</a>, P2 GV,\ 26.0159ºS, 50.6571ºW \ 01.IX.2022, L. V. Graf,\ rede entomológica” , ♀, “ DZUP 540053 ”, “ Brasil, Paraná, São \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.6571&amp;materialsCitation.latitude=-26.0159" title="Search Plazi for locations around (long -50.6571/lat -26.0159)">Mateus do Sul</a>, P2 GV,\ 26.0159ºS, 50.6571ºW \ 01.IX.2022, L. V. Graf,\ rede entomológica” , ♀, “ DZUP 540054 ”, “ Brasil, Paraná, São \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.6571&amp;materialsCitation.latitude=-26.0159" title="Search Plazi for locations around (long -50.6571/lat -26.0159)">Mateus do Sul</a>, P2 GV,\ 26.0159ºS, 50.6571ºW \ 01.IX.2022, L. V. Graf,\ rede entomológica” , ♀, “ DZUP 540055 ”, “ Brasil, Paraná, São \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.6571&amp;materialsCitation.latitude=-26.0159" title="Search Plazi for locations around (long -50.6571/lat -26.0159)">Mateus do Sul</a>, P2 GV,\ 26.0159ºS, 50.6571ºW \ 01.IX.2022, L. V. Graf,\ rede entomológica” , ♀, “ DZUP 540056 ”, “ Brasil, Paraná, São \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.6571&amp;materialsCitation.latitude=-26.0159" title="Search Plazi for locations around (long -50.6571/lat -26.0159)">Mateus do Sul</a>, P2 GV,\ 26.0159ºS, 50.6571ºW \ 01.IX.2022, L. V. Graf,\ rede entomológica” , ♀, “ DZUP 540057 ”, “ Brasil, Paraná, São \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.6571&amp;materialsCitation.latitude=-26.0159" title="Search Plazi for locations around (long -50.6571/lat -26.0159)">Mateus do Sul</a>, P2 GV,\ 26.0159ºS, 50.6571ºW \ 01.IX.2022, L. V. Graf,\ rede entomológica” , ♀, “ DZUP 540058 ”, “ Brasil, Paraná, Ponta \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.27548&amp;materialsCitation.latitude=-25.1393" title="Search Plazi for locations around (long -50.27548/lat -25.1393)">Grossa</a>, P1 GIII,\ 25.1393ºS, 50.27548ºW \ 26.VIII.2022, L. V.Graf,\ rede entomológica” , ♀, “ DZUP 540059 ”, “ Brasil, Paraná, São José \ dos <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=49.175278&amp;materialsCitation.latitude=-25.595278" title="Search Plazi for locations around (long 49.175278/lat -25.595278)">Pinhais</a> 25º35’43’’S,\ 49º10’31’’O, 27.VII.2024 \ Graf L. V., rede entomol.” , ♀, “ DZUP 540060 ”, “ Brasil, Paraná, São \ José dos <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.0224&amp;materialsCitation.latitude=-25.382" title="Search Plazi for locations around (long -49.0224/lat -25.382)">Pinhais</a>, M2,\ 25.3820ºS, 49.0224ºW,\ 20. VI.2023, L. V.Graf,\ rede entomológica\\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.0224&amp;materialsCitation.latitude=-25.382" title="Search Plazi for locations around (long -49.0224/lat -25.382)">Coord.</a> corretas M2:\ 25º38’26.11’’S \ 49º02’16.80’’W ” , ♀, “ DZUP 540061 ”, “ Brasil, Paraná, São \ José dos <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.0143&amp;materialsCitation.latitude=-25.3949" title="Search Plazi for locations around (long -49.0143/lat -25.3949)">Pinhais</a>, S2,\ 25.3949ºS, 49.0143ºW,\ 20. VI.2023, L. V.Graf,\ rede entomológica\\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.0143&amp;materialsCitation.latitude=-25.3949" title="Search Plazi for locations around (long -49.0143/lat -25.3949)">Coord.</a> corretas S2:\ 25º39’30.92’’S \ 49º01’57.81’’W ” , ♀, “ DZUP 540062 ”, “ Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.2697&amp;materialsCitation.latitude=-25.1477" title="Search Plazi for locations around (long -50.2697/lat -25.1477)">Ponta</a> \ Grossa, P1- GIII, 781m,\ 25.1477ºS, 50.2697ºW,\ 20.xi.2020, A. Martins \ &amp; D. Muniz, Busca ativa” , ♂, “ DZUP 540063 ”, “ Brasil, Paraná, São José \ dos <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.090664&amp;materialsCitation.latitude=-25.562145" title="Search Plazi for locations around (long -49.090664/lat -25.562145)">Pinhais</a>, P4,\ -25.562145, -49.090665,\ 10.XII.2021, L. V.Graf, rede” , ♂, “ DZUP 540064 ”, “ Brasil, Paraná, Ponta \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.2697&amp;materialsCitation.latitude=-25.1477" title="Search Plazi for locations around (long -50.2697/lat -25.1477)">Grossa</a>, P1- GIII, 781m,\ 25.1477ºS, 50.2697ºW,\ 30.xi.2021, A.L. Martins,\ Busca ativa” , ♂, “ DZUP 540065 ”, “ Brasil, Paraná, Ponta \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.2697&amp;materialsCitation.latitude=-25.1477" title="Search Plazi for locations around (long -50.2697/lat -25.1477)">Grossa</a>, P1- GIII, 781m,\ 25.1477ºS, 50.2697ºW,\ 30.xi.2021, A.L. Martins,\ Busca ativa” , ♂, “ DZUP 540066 ”, “ Brasil, PR, São Mateus \ do Sul, UN-SIX/ Petrobras \ 10.I.2012 \ Rafael Kamke leg.\\ Ponto amostral\ HP\\ CJS 10963” , ♂, “ DZUP 540067 ”, “ Brasil, PR, São Mateus \ do Sul, UN-SIX/ Petrobras \ 11.I.2012 \ Rafael Kamke leg.\\ Ponto amostral\ CAB\\ CJS 11010” , ♂, “ DZUP 540068 ”, “ Brasil, PR, São Mateus \ do Sul, UN-SIX/ Petrobras \ 11.I.2012 \ Rafael Kamke leg.\\ Ponto amostral\ CAB\\ CJS 11015” , ♂, “ DZUP 540069 ”, “ Brasil, PR, São Mateus \ do Sul, UN-SIX/ Petrobras \ 11.I.2012 \ Rafael Kamke leg.\\ Ponto amostral\ CAB\\ CJS 11014” , ♂, “ DZUP 540070 ”, “ Brasil, PR, São Mateus \ do Sul, UN-SIX/ Petrobras \ 11.I.2012 \ Rafael Kamke leg.\\ Ponto amostral\ CAB\\ CJS 11013” , ♂, “ DZUP 540071 ”, “ Brasil, PR, São Mateus \ do Sul, UN-SIX/ Petrobras \ 11.I.2012 \ Rafael Kamke leg.\\ Ponto amostral\ CAB\\ CJS 11012” , ♂, “ DZUP 540072 ”, “ Brasil, PR, São Mateus \ do Sul, UN-SIX/ Petrobras \ 11.I.2012 \ Rafael Kamke leg.\\ Ponto amostral\ CAB\\ CJS 11011” , ♂, “ DZUP 540073 ”, “ Brasil, PR, São Mateus \ do Sul, UN-SIX/ Petrobras \ 23.I.2011 \ Rafael Kamke leg.\\ Ponto amostral\ CAB (antigo MDI)\\ CJS 9893 ” , ♂, “ DZUP 540074 ”, “ Brasil, PR, São Mateus \ do Sul, UN-SIX/ Petrobras \ 23.I.2011 \ Rafael Kamke leg.\\ Ponto amostral\ CAB (antigo MDI)\\ CJS 9892 ” , ♂, “ DZUP 540075 ”, “ Brasil, PR, São Mateus \ do Sul, UN-SIX/ Petrobras \ 20.I.2011 \ Rafael Kamke leg.\\ Ponto amostral\ MATA DO TÚNEL \\ CJS 9560 ” , ♂, “ DZUP 540076 ”, “ Brasil, PR, São Mateus \ do Sul, UN-SIX/ Petrobras \ 23.I.2011 \ Rafael Kamke leg.\\ Ponto amostral\ CAB (antigo MDI)\\ CJS 9891 ” , ♂, “ DZUP 540077 ”, “ Brasil, PR, São Mateus \ do Sul, UN-SIX/ Petrobras \ 20.I.2011 \ Rafael Kamke leg.\\ Ponto amostral\ CAVA\\ CJS 9639” , ♂, “ DZUP 540078 ”, “ Brasil, PR, São Mateus \ do Sul, UN-SIX/ Petrobras \ 21.I.2011 \ Rafael Kamke leg.\\ Ponto amostral\ MATA DO TÚNEL \\ CJS 9719 ” , ♂, “ DZUP 540079 ”, “ Brasil, PR, São Mateus \ do Sul, UN-SIX/ Petrobras \ 21.I.2011 \ Rafael Kamke leg.\\ Ponto amostral\ MATA DO TÚNEL \\ CJS 9681 ” , ♂, “ DZUP 540080 ”, “ Brasil, Paraná, União da\ Vitória, P2 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-51.17856&amp;materialsCitation.latitude=-26.07979" title="Search Plazi for locations around (long -51.17856/lat -26.07979)">Controle</a>,\ 26.07979ºS, 51.17856ºW,\ 22.II.2022, L. V.Graf, rede” , ♂, “ DZUP 540081 ”, “ Brasil, Paraná, União da\ Vitória, P2 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-51.17856&amp;materialsCitation.latitude=-26.07979" title="Search Plazi for locations around (long -51.17856/lat -26.07979)">Controle</a>,\ 26.07979ºS, 51.17856ºW,\ 22.II.2022, L. V.Graf, rede” , ♂, “ DZUP 540082 ”, “ Brasil, Paraná, União da\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-51.0628&amp;materialsCitation.latitude=-26.1446" title="Search Plazi for locations around (long -51.0628/lat -26.1446)">Vitória</a>, P1 GV,\ 26.1446ºS, 51.0628ºW,\ 25. V.2022, L. V.Graf,\ rede entomológica” , ♂, “ DZUP 540083 ”, “ Brasil, Paraná, União da\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-51.0628&amp;materialsCitation.latitude=-26.1446" title="Search Plazi for locations around (long -51.0628/lat -26.1446)">Vitória</a>, P1 GV,\ 26.1446ºS, 51.0628ºW,\ 25. V.2022, L. V.Graf,\ rede entomológica” , ♂, “ DZUP 540084 ”, “ Brasil, Paraná, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.5229&amp;materialsCitation.latitude=-25.1337" title="Search Plazi for locations around (long -50.5229/lat -25.1337)">Imbituva</a>,\ P1 GVI, 25.1337ºS,\ 50.5229ºW, 28. V.2022,\ L. V.Graf, rede\ entomológica” , ♂, “ DZUP 540085 ”, “ Brasil, Paraná, Ponta \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.27548&amp;materialsCitation.latitude=-25.1393" title="Search Plazi for locations around (long -50.27548/lat -25.1393)">Grossa</a>, P1 GIII,\ 25.1393ºS, 50.27548ºW \ 26.VIII.2022, L. V.Graf,\ rede entomológica” , ♂, “ DZUP 540086 ”, “ Brasil, Paraná, São José \ dos Pinhais, M3,\ 25.3823ºS, 49.0131ºW,\ 19. VI.2022, L. V.Graf,\ rede entomológica\\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.0131&amp;materialsCitation.latitude=-25.3823" title="Search Plazi for locations around (long -49.0131/lat -25.3823)">Coord.</a> corretas M3:\ 25º38’23.44’’S \ 49º01’29.34’’W ” , ♂, “ DZUP 540087 ”, “ Brasil, Paraná, São José \ dos <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=49.175278&amp;materialsCitation.latitude=-25.595278" title="Search Plazi for locations around (long 49.175278/lat -25.595278)">Pinhais</a>, 25º35’43’’S,\ 49º10’31’’O, 27.VII.2024,\ Graf L. V., rede entomol.” , ♂, “ DZUP 540088 ”, “ Brasil, Paraná, São \ José dos <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.0143&amp;materialsCitation.latitude=-25.3949" title="Search Plazi for locations around (long -49.0143/lat -25.3949)">Pinhais</a>, S2,\ 25.3949ºS, 49.0143ºW,\ 13.III.2023, L. V.Graf,\ rede entomológica\\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.0143&amp;materialsCitation.latitude=-25.3949" title="Search Plazi for locations around (long -49.0143/lat -25.3949)">Coord.</a> corretas S2:\ 25º39’30.92’’S \ 49º01’57.81’’W ” , ♂, “ DZUP 540089 ”, “ Brasil, Paraná, São José \ dos Pinhais, M3,\ 25.3823ºS, 49.0131ºW,\ 19. VI.2022, L. V.Graf,\ rede entomológica\\ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.0131&amp;materialsCitation.latitude=-25.3823" title="Search Plazi for locations around (long -49.0131/lat -25.3823)">Coord.</a> corretas M3:\ 25º38’23.44’’S \ 49º01’29.34’’W ” , ♂, “ DZUP 540090 ”, “ Brasil, Paraná, São \ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.6571&amp;materialsCitation.latitude=-26.0159" title="Search Plazi for locations around (long -50.6571/lat -26.0159)">Mateus do Sul</a>, P2 GV,\ 26.0159ºS, 50.6571ºW \ 01.IX.2022, L. V. Graf,\ rede entomológica” , ♂, “ DZUP 540091 ”, “ Brasil, Paraná, São \ José dos <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.0128&amp;materialsCitation.latitude=-25.62375" title="Search Plazi for locations around (long -49.0128/lat -25.62375)">Pinhais</a>, M1,\ 25.3724ºS, 49.0128ºW,\ 14.III.2023, L. V. Graf,\ rede entomológica\\ Coord. corretas M1:\ 25º37’25.50’’S\ 49º01’28.99’’W”, ♀, “ DZUP 540092 ”, “Brasil, Paraná, Ponta\ Grossa, P1- GIII, 781m,\ 25.1477ºS, 50.2697ºW,\ 30.xi.2021, A.L. Martins,\ Busca ativa”, ♀, “ DZUP 540150 ”, “ BRASIL, PARANÁ,\ COLOMBO, EMBRAPA\ 25º19’18.66’’S 49º9’33.66’’W \ 22.XI.2019 935m \ MIKICH, S.B. leg. E4847”, ♀, “ DZUP 540151 ”, “ BRASIL, PARANÁ,\ COLOMBO, EMBRAPA\ 25º19’18.66’’S 49º9’33.66’’W \ 22.XI.2019 935m \ MIKICH, S.B. leg. E4855”, ♂, “ DZUP 540152 ”, “ BRASIL, PARANÁ,\ COLOMBO, EMBRAPA\ 25º19’18.66’’S 49º9’33.66’’W \ 22.XI.2019 935m \ MIKICH, S.B. leg.E4865”, ♂, “ DZUP540153 ”, “ BRASIL, PARANÁ,\ COLOMBO, EMBRAPA\ 25º19’18.66’’S 49º9’33.66’’W \ 8.I.2020 935m \ MIKICH, S.B. leg. E5482”, ♂, “ DZUP 540154 ”, “ BRASIL, PARANÁ,\ COLOMBO, EMBRAPA\ 25º19’18.66’’S 49º9’33.66’’W \ 14.I.2020 935m \ MIKICH, S.B. leg. E5731”, ♀, “ DZUP 540155 ”, “ BRASIL, PARANÁ,\ COLOMBO, EMBRAPA\ 25º19’18.66’’S 49º9’33.66’’W \ 6.XII.2019 935m \ MIKICH, S.B. leg. E5252”, ♀, “ DZUP 540156 ”, “ BRASIL, PARANÁ,\ COLOMBO, EMBRAPA\ 25º19’18.66’’S 49º9’33.66’’W \ 15.I.2020 935m \ MIKICH, S.B. leg. E5833”, ♀, “ DZUP 600476 ”, “Curitiba - PR \ 19 / IX / 1964 \ R. Lange leg.”, ♀, “ DZUP 600477 ”, “Curitiba - PR \ 7 / XII / 1964 \ R. Lange leg.\\ R.Lange\ No.500”, ♂, “ DZUP 540180 ”, “CURITYBA-PARANÁ\ (Parolim)\ 24. VI.38\ Coll Claretiano”, ♂, “ DZUP 540181 ”, “ CURITIBA \ Paraná BRASIL \ 22-IX-1960 ”, ♂, “ DZUP 540182 ”, “GUARAPUAVA 1120m \ Paraná BRASIL \ 7- IX-1955 \ Michener e Moure”, ♂, “ DZUP 540183 ”, “ CURITIBA \ Paraná BRASIL \ 1-XI- I955\ Michener e Lange”, ♂, “ DZUP 540184 ”, “ CURITIBA \ Paraná BRASIL \ 12-IX-1960 ”, ♂, “ DZUP 540185 ”, “COLOMBO 950 m \ Paraná BRASIL \ 18-VIII-1955 Lange,\ Michener e Moure”, ♂, “ DZUP 540188 ”, “ R.dos Papagaios\ PR Brasil - I - X 59\ P. D. HURD leg”, ♀, “ DZUP 540202 ”, “S. L. PURUNÃ-GO\ Brasil 15 - X - 65\ Fioravante e Graf”. Brasil, RS, DZUP. ♀, “ DZUP 540033 ”, “ CANELA, RS \ Brasil 10.XI.1984 \ M. Hoffmann col.\\ col. MCN 66.739”, ♀, “ DZUP 540034 ”, “ CANELA, RS \ Brasil 9.X.1989 \ M. Hoffmann col.\\ col. MCN 95.640”, ♂, “ DZUP 540043 ”, “B. de COTEGIPE\ R.Gr.S.- BRASIL \ 15/I/1967 \ F. Giacomel lg”. RS, MCT. ♂, “ MCP123567 ”, “ BR - RS - POA\ S. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.02472&amp;materialsCitation.latitude=-25.1477" title="Search Plazi for locations around (long -49.02472/lat -25.1477)">Fco de Paula</a>, Pró-\ Mata 09.02.2000 \ leg.: A. Köhler \\ 8986\ 168\\ MCP \ 123567\\ Caenohalictus \ tesselatus\ Gonçalves det. ”, ♂, “ MCP123568 ”, “ BR - RS - POA\ S. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.02472&amp;materialsCitation.latitude=-25.1477" title="Search Plazi for locations around (long -49.02472/lat -25.1477)">Fco de Paula</a>, Pró-\ Mata 09.02.2000 \ leg.: A. Köhler \\ 8987\ 168\\ MCP \ 123568”, ♂, “ MCP123569 ”, “ BR - RS - POA\ S. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.02472&amp;materialsCitation.latitude=-25.1477" title="Search Plazi for locations around (long -49.02472/lat -25.1477)">Fco de Paula</a>, Pró-\ Mata 09.02.2000 \ leg.: A. Köhler \\ 9019\ 168\\ MCP \ 123569”, ♂, “ MCP123570 ”, “ BR - RS - POA\ S. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.02472&amp;materialsCitation.latitude=-25.1477" title="Search Plazi for locations around (long -49.02472/lat -25.1477)">Fco de Paula</a>, Pró-\ Mata 09.02.2000 \ leg.: A. Köhler \\ 8994\ 168\\ MCP \ 123570”, ♂, “ MCP123582 ”, “ BR - RS - POA\ S. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.02472&amp;materialsCitation.latitude=-25.1477" title="Search Plazi for locations around (long -49.02472/lat -25.1477)">Fco de Paula</a>, Pró-\ Mata 14.01.2000 \ leg.: A. Köhler \\ 8592\ 228\\ MCP \ 123582”, ♂, “ MCP123583 ”, “ BR - RS - POA\ S. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.02472&amp;materialsCitation.latitude=-25.1477" title="Search Plazi for locations around (long -49.02472/lat -25.1477)">Fco de Paula</a>, Pró-\ Mata 08.02.2000 \ leg.: A. Köhler \\ 8862\ 168\\ MCP \ 123583”, ♂, “ MCP123584 ”, “ BR - RS - POA\ S. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.02472&amp;materialsCitation.latitude=-25.1477" title="Search Plazi for locations around (long -49.02472/lat -25.1477)">Fco de Paula</a>, Pró-\ Mata 08.02.2000 \ leg.: A. Köhler \\ 8866\ 168\\ MCP \ 123584”, ♂, “ MCP123585 ”, “ BR - RS - POA\ S. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.02472&amp;materialsCitation.latitude=-25.1477" title="Search Plazi for locations around (long -49.02472/lat -25.1477)">Fco de Paula</a>, Pró-\ Mata 08.02.2000 \ leg.: A. Köhler \\ 8760\ 168\\ MCP \ 123585”, ♂, “ MCP123588 ”, “ BR - RS - POA\ S. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.02472&amp;materialsCitation.latitude=-25.1477" title="Search Plazi for locations around (long -49.02472/lat -25.1477)">Fco de Paula</a>, Pró-\ Mata 08.02.2000 \ leg.: A. Köhler \\ 8863\ 168\\ MCP \ 123588”, ♂, “ MCP123575 ”, “ BR - RS - POA\ S. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.02472&amp;materialsCitation.latitude=-25.1477" title="Search Plazi for locations around (long -49.02472/lat -25.1477)">Fco de Paula</a>, Pró-\ Mata 09.02.2000 \ leg.: A. Köhler \\ 9011\ 168\\ MCP \ 123575”, ♂, “ MCP123594 ”, “ BR - RS - POA\ S. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.02472&amp;materialsCitation.latitude=-25.1477" title="Search Plazi for locations around (long -49.02472/lat -25.1477)">Fco de Paula</a>, Pró-\ Mata 08.02.2000 \ leg.: A. Köhler \\ 8851\ 168\\ MCP \ 123594”, ♂, “ MCP123576 ”, “ BR - RS - POA\ S. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.02472&amp;materialsCitation.latitude=-25.1477" title="Search Plazi for locations around (long -49.02472/lat -25.1477)">Fco de Paula</a>, Pró-\ Mata 09.02.2000 \ leg.: A. Köhler \\ 9023\ 168\\ MCP \ 123576”, ♂, “ MCP123573 ”, “ BR - RS - POA\ S. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.02472&amp;materialsCitation.latitude=-25.1477" title="Search Plazi for locations around (long -49.02472/lat -25.1477)">Fco de Paula</a>, Pró-\ Mata 09.02.2000 \ leg.: A. Köhler \\ 8992\ 168\\ MCP \ 123573”, ♂, “ MCP123586 ”, “ BR - RS - POA\ S. Fco de Paula, Pró-\ Mata 08.02.2000 \ leg.: A. Köhler \\ 8860\ 168\\ MCP \ 123586”, ♂, “ MCP123596 ”, “ BR - RS - POA\ S. Fco de Paula, Pró-\ Mata 08.02.2000 \ leg.: A. Köhler \\ 8861\ 168\\ MCP \ 123596”. Brasil, SC, DZUP. ♀, “ DZUP 540036 ”, “ BRASIL \ STA.CATARINA\ Caçador\ Col.Lorchzatto ”. Brasil, SP, DZUP. ♀, “ DZUP 540179 ”, “ Brasil, SP, 10 km ao SW\ de S. J. Barreiro, 1585m,\ 22°43’32”S 44°37’33”W,\ 29.iii.2010, Melo &amp; Luz”, ♂, “ DZUP 540186 ”, “BOTUCATU\ S. Paulo BRASIL \ II-1954 \ W. Bokermann ” .</p></div>	https://treatment.plazi.org/id/8C7BE1229E37FFCF76F0188329E3F9CC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cappola, Victor;Gonçalves, Rodrigo B.	Cappola, Victor, Gonçalves, Rodrigo B. (2026): A review of Caenohalictus species from southern Brazil (Apoidea, Caenohalictini). Zootaxa 5782 (3): 401-448, DOI: 10.11646/zootaxa.5782.3.1, URL: https://doi.org/10.11646/zootaxa.5782.3.1
