taxonID	type	description	language	source
8F1D4A79D244487694DDFC17A20FA32E.taxon	type_taxon	Type species: Ancillaria lanceolata Martens, 1901	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D244487694DDFC17A20FA32E.taxon	description	Description. Shell small to large for the family, 12 – 50 mm in length, elongate oval. Protoconch dome shaped, later part covered with callus, rendering whorl count usually impossible. Spire high, conical; primary callus medium thick (Fig. 1 C — pc) and covering all whorls except initial part of protoconch, slightly impressed at sutures, glossy, microshagreened; secondary callus (Fig. 1 C – E — sc) from thin to thick, extending to right-ventral side of last whorl and continuing up as tongue-shaped pad, reaching suture of antepenultimate whorl. It is overlaying primary callus except ventral side of last whorl. Body whorl cloak glossy, covered by very weak and indistinct spiral striation. Olivoid groove very shallow and indistinct, terminating in very short rounded-triangular extension on outer lip to obsolete. Rear edge of anterior band from very indistinct to sharp. Anterior band usually strongly different from shell cloak in colour, rarely similar to body whorl cloak, sometimes distinctly separated into upper and lower bands. Columella smooth or with weak spiral cords. Aperture narrow oval, acute posteriorly. Shell differently coloured, from milky-white to different shades of orange and brown, often with bluish tint. Operculum either small subtriangular, or large, narrow lanceolate. Nucleus terminal. Foregut anatomy. Proboscis not long, cylindrical or conical. Salivary glands seemingly fused, compact, formed by ramified tubes, embracing the nerve ring. Gland of Leiblein large, broadly tubular and folded anteriorly, anteriormost part creamy, corresponding to dorsal glandular folds of oesophagus, main part brownish. Accessory salivary gland present, single, situated on right side of rhynchodaeum, massive, tubular. Gland narrowing at half-length, attached to rhynchodaeum and following backward along the oesophagus. Valve of Leiblein distinct [examined in T. alcocki and T. monachalis]. Radula short, spanning less than half of proboscis length. Radula (Fig. 4) of about 35 – 90 rows of teeth, normally 55 – 75. Rachidians with very deep notch on the anterior side and long tongue-shaped projection of the base of posterior edge, corresponding in shape and engaging the notch of the neighbouring tooth, with 3 long and narrow cusps, central one being the largest. Lateral teeth unicuspid, subtriangular.	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D244487694DDFC17A20FA32E.taxon	discussion	Remarks. Although the shell shape is rather distinctive in Turrancilla, it bears clear resemblance to Amalda, particularly T. reboriae. Turrancilla can be distinguished from Amalda in having a generally much weaker (or absent) olivoid groove; correspondingly, the olivoid band is not or much less differentiated from the body whorl cloak. The genera can be readily distinguished by very different operculums: in Amalda it is elongate-oval with subcentral nucleus, while in Turrancilla the operculum strongly narrows in the lower part and the nucleus is terminal. The radula of Amalda is also very different from Turrancilla in having a nearly straight anterior edge of the rachidian teeth (Fig. 4 E, after Kantor et al. 2020: fig. 7) and lacking projection of the posterior edge. On the contrary, similar radular morphology with deeply notched rachidian tooth is found in the conchologically very different genus Entomoliva Bouchet & Kilburn, 1991 (Fig. 4 F)	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D247487A94DDFE2AA56EA3C2.taxon	description	Figs 3, 4 A – B, 5	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D247487A94DDFE2AA56EA3C2.taxon	materials_examined	Type locality. East Africa, Zanzibar Channel, 5 ° 27 ' S, 39 ° 18 ' E, depth 463 m; Valdivia stn. 245. Syntypes: ZMB Mol 61070 (studied syntype SL 22.1 mm). Sequenced material. SOLOMON ISLANDS • 2 lv; NW Santa Isabel I.; 7 ° 39 ' S, 157 ° 43 ' E; depth 495 – 650 m; SALOMON 2, stn CP 2213; MNHN-IM- 2007 - 33299, MNHN-IM- 2007 - 33300 • 1 lv; NW Santa Isabel I.; 7 ° 42 ' S, 157 ° 44 ' E; depth 550 – 682 m; SALOMON 2, stn CP 2214; MNHN-IM- 2007 - 33306 • 1 lv; Vella Gulf, 7 ° 48 ' S, 156 ° 52 ' E; depth 590 – 600 m; SALOMON 2, stn CP 2267; MNHN-IM- 2007 - 33310 • 1 lv; Guadalcanal I.; 9 ° 25 ' S, 160 ° 31 ' E; depth 456 – 551 m; SALOMONBOA 3, stn CP 272; MNHN-IM- 2007 - 36211 • 1 lv; Savo; 9 ° 12 ' S, 160 ° 55 ' E; depth 706 – 722 m; SALOMONBOA 3, stn CP 2777; MNHN-IM- 2007 - 36185 • 3 lv; Sealark channel; 9 ° 37 ' S, 160 ° 47 ' E; depth 502 – 621 m; SALOMONBOA 3, stn CP 2850; MNHN-IM- 2007 - 36187, MNHN-IM- 2007 - 36188, MNHN-IM- 2007 - 36208 • 2 lv; Kaoka Bay; 9 ° 40 ' S, 160 ° 45 ' E; depth 650 – 725 m; SALOMONBOA 3, stn CP 2858; MNHN-IM- 2007 - 36204, MNHN-IM- 2007 - 36214 PAPUA NEW GUINEA • 3 lv; S of Lae, Huon Gulf; 6 ° 56 ' S, 147 ° 8 ' E; depth 700 – 740 m; BIOPAPUA stn. CP 3632; MNHN-IM- 2009 - 25006, MNHN-IM- 2019 - 22747, MNHN-IM- 2009 - 25007 • 1 lv; Huon Gulf; 6 ° 47 ' S, 147 ° 14 ' E; depth 650 m; PAPUA NIUGINI, stn CP 4000; MNHN-IM- 2013 - 19870 • 1 lv; NW Manam I.; 4 ° 00 ' S, 144 ° 54 ' E; depth 800 – 860 m; PAPUA NIUGINI, stn CP 4078; MNHN-IM- 2013 - 19425 • 1 lv; Mambare Bay; 7 ° 52 ' S, 148 ° 4 ' E; depth 575 – 655 m; BIOPAPUA stn. CP 3729; MNHN-IM- 2019 - 22751 • 1 lv; Mambare Bay; 7 ° 50 ' S, 148 ° 3 ' E; depth 895 – 1150 m; BIOPAPUA stn. CP 3731; MNHN-IM- 2019 - 22749 SOLOMON SEA • 1 lv; Dampier Strait; 5 ° 35 ' S, 148 ° 13 ' E; depth 630 – 870 m; PAPUA NIUGINI, stn CP 4014; MNHN-IM- 2013 - 19990 • 1 lv; Induna I.; 4 ° 35 ' S, 152 ° 25 ' E; depth 575 – 616 m; PAPUA NIUGINI, stn CP 4266; MNHN-IM- 2013 - 45474 • 1 lv; New Britain, Ainto Bay; 6 ° 10 ' S, 149 ° 18 ' E; depth 510 – 743 m; MADEEP, stn CP 4339; MNHN-IM- 2013 - 46439 PHILIPPINES • 1 lv; Bohol Sea; 9 ° 31 ' N, 124 ° 01 ' E; depth 602 – 738 m; PANGLAO 2005, stn CP 2350; MNHN-IM- 2007 - 31962 • 1 lv; Bohol Sea, off Balicasag I.; 9 ° 33 ' S, 124 ° 41 ' E; depth 713 – 731 m; PANGLAO 2005, stn CP 2398; MNHN-IM- 2007 - 31967 • 1 lv; Luzon I.; 15 ° 20 ' N, 121 ° 37 ' E; depth 593 – 600 m; AURORA 2007; MNHN-IM- 2007 - 34773. MADAGASCAR • 1 lv; off Majunga and Cap Saint-André; 15 ° 31 ' S, 45 ° 42 ' E; depth 760 – 777 m; MIRIKY, stn CP 3270; MNHN-IM- 2007 - 36849 • 1 lv; off Majunga and Cap Saint-André; 15 ° 21 ' S, 45 ° 57 ' E; depth 780 – 1020 m; MIRIKY, stn CP 3279; MNHN-IM- 2007 - 36713. MOZAMBIQUE CHANNEL • 1 lv; Zambeze transect; 19 ° 36 ' S, 36 ° 47 ' E; depth 636 m; MAINBAZA, stn CC 3154; MNHN-IM- 2007 - 38771. Non sequenced material. SOLOMON ISLANDS • 1 dd; NW Santa Isabel I.; 7 ° 42 ' S, 157 ° 44 ' E; depth 550 – 682 m; SALOMON 2, stn CP 2214; MNHN-IM- 2007 - 33305. MOZAMBIQUE CHANNEL • 2 dd (badly worn); Zambeze transect; 19 ° 35 ' S, 36 ° 46 '; depth 518 – 524 m; MAINBAZA CC 3153 (SL to 28 mm, apex broken off).	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D247487A94DDFE2AA56EA3C2.taxon	description	Description. Shell of small size for the genus (up to 22 mm), oval bullet-shaped, slender. Protoconch low dome shaped, smooth, from light to distinctly brownish (Fig. 3 I – J). First whorl with distinctly impressed suture, further suture overlaid by callus. Protoconch borders indistinct, rendering whorl count impossible. Spire medium to high, weakly convex in profile; primary callus moderately thick and covering all whorls except protoconch, slightly impressed at sutures; surface of callus glossy, microshagreened; secondary callus thin, extending to right-ventral side of last whorl and continuing up as fairly thin, tongue-shaped pad, reaching suture of antepenultimate whorl. Surface of secondary callus very similar to that of primary one, borders indistinct. Body whorl cloak glossy, covered by very weak and indistinct spiral striation. Olivoid groove shallow, sometimes indistinct, better seen on ending part of last whorl, terminating in very short rounded-triangular extension on outer lip. Rear edge of anterior band weak, detectable by slight elevation of shell surface due to thickening. Anterior band slightly different from shell cloak in colour, not separated into upper and lower bands. Plication plate with deep and broad groove in upper third, separating adapical largest spiral ridge, followed by three-four spiral ridges progressively weakening abapically. Columella smooth. Aperture narrow oval, acute posteriorly, widest medially; outer lip slightly thickened, slightly opisthocline, with slight, blunt labral denticle; siphonal notch moderately deep, asymmetrical. Shell from uniformly milky-white, to creamy on body whorl cloak, sometimes with slightly darker irregularly spaced axial bands, corresponding to growth lines (Fig. 3 G). Operculum narrow-lanceolate, thick, spanning about 2 / 3 of AL, dark-brown, with terminal nucleus and prominent growth lines. Radula (Fig. 4 A – B), studied in two specimens (MNHN-IM- 2019 - 22749, SL 17.7 mm, MNHN-IM- 2007 - 38771, SL 10.5 mm), of about 55 rows of teeth, 12 - 14 nascent, typical for the genus. Alary processes well developed, long, spanning about 40 % of total radular membrane length. Radula about 1 mm long (11 % of AL), radula membrane about 65 – 95 µm in width (1.2 – 1.1 % of AL correspondingly). Diagnostic cox- 1 sites: ' A' in the site 73, ' A' in the site 169, ' G' in the site 196.	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D247487A94DDFE2AA56EA3C2.taxon	discussion	Remarks. The species exhibits some degree of variation in the shell outline as well as in the number of ridges on plication plate that can occasionally be five. In some specimens not only protoconch, but also uppermost teleoconch whorls can be orange, or the suture area can be highlighted by light orange spiral band. Most of the specimens retain the protoconch, but in some it can be decollated together with upper teleoconch whorls. Within the species three subclades are distinguished and recognized by ASAP as separate PSHs. Each subclade is restricted to a single geographic area. Since specimens belonging to all three subclades are morphologically very close, we consider this structuring as geographically determined. The largest clade includes the specimens collected off Solomons and Papua New Guinea. The second clade includes three specimens from the Philippines, and finally, the third one three specimens collected off Madagascar and Mozambique. The species differs from other congeners in having milky-white or creamy small shell (not reaching 30 mm) without clear distinction in the cloak, olivoid band and with poorly differentiated anterior band.	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D247487A94DDFE2AA56EA3C2.taxon	distribution	Distribution. From east Africa to Papua New Guinea and Philippines (Fig. 5).	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D24A487C94DDFF2FA304A00E.taxon	description	Figs 4 C – D, 6 A – F, 7 A Turrancilla sp. 2 — Kantor et al. 2017: figs. 6 T (shell), 12 H (radula).	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D24A487C94DDFF2FA304A00E.taxon	materials_examined	Type material. Holotype: presently in Conchology, Inc., will be donated to a Philippine Institution (Poppe et al. 2017). Type locality. Philippines, Mactan I., Punta Engaño, 250 – 300 m. Sequenced material. PHILIPPINES • 2 lv; Bohol Sea, off Balicasag I., 9 ° 29 ' N, 123 ° 44 ' E, 271 – 318 m, PANGLAO 2005, stn CP 2340 (MNHN-IM- 2007 - 35726, MNHN-IM- 2007 - 35727) • 1 lv; Bohol Sea, off Pamilacan I., 9 ° 25 ' N, 123 ° 50 ' E, 544 – 712 m, PANGLAO 2005, stn CP 2341 (MNHN-IM- 2007 - 31947) • 1 lv; Bohol / Sulu Sea sill, 8 ° 49 ' N, 123 ° 38 ' E, PANGLAO 2005, stn CP 2360 (MNHN-IM- 2007 - 31954) • 2 lv; Bohol Sea, Maribojoc Bay, 9 ° 39 ' N, 123 ° 43 ' E, 481 – 505 m, PANGLAO 2005, stn CP 2404 (MNHN-IM- 2007 - 31952, MNHN-IM- 2007 - 31953). Non sequenced material. PHILIPPINES • 6 dd; Bohol Sea, off Balicasag I., 9 ° 29 ' N, 123 ° 44 ' E, 271 – 318 m, PANGLAO 2005, stn CP 2340 • 3 dd; Bohol Sea, Maribojoc Bay; 9 ° 35 ' N, 123 ° 42 ' E; depth 642 – 669 m; PANGLAO 2005, stn CP 2397.	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D24A487C94DDFF2FA304A00E.taxon	description	Description. Shell of small size for the genus (up to 15 mm), ovate. Protoconch low dome shaped, smooth, light yellowish-brown. First whorl with distinct impressed suture, further whorls’ suture overlaid by callus. Protoconch borders indistinct, rendering whorl count impossible. Spire low, conical; primary callus moderately thick and covering all whorls except protoconch, slightly impressed at sutures; especially on penultimate whorl, surface of callus glossy, distinctly microshagreened; secondary callus thin, extending to right-ventral side of last whorl and continuing up as fairly thin, tongue-shaped pad, reaching suture of antepenultimate whorl. Surface of secondary callus very similar to that of primary one, borders indistinct. Body whorl cloak glossy, covered by very weak and indistinct thin spiral striation. Olivoid groove very shallow but distinct, terminating in very short and narrow rounded extension on outer lip. Olivoid band usually slightly lighter than cloak. Rear edge of anterior band rather distinct, in a shape of sharp step. Anterior band darker than cloak, brown, microshagreened, separated into upper and about three times narrower lower band by low but distinct ridge. Both upper and lower parts of anterior band weakly convex. Plication plate well separated by broad and deep groove, with well pronounced 4 spiral ridges progressively weakening abapically. Columella smooth. Aperture narrow oval, acute posteriorly; outer lip slightly thickened, slightly opisthocline, with slight, blunt labral denticle; siphonal notch moderately deep, asymmetrical. Shell of different shades of brown, with a light broad band in suture area. Plication plate from creamy to light purplish. Radula (Fig. 4 C – D) studied in one specimen (MNHN-IM- 2007 - 31947, SL 9.9 mm), typical for the genus. Radula membrane narrow, about 40 µm in width (0.7 % of AL). Diagnostic cox- 1 sites. ' C' in the site 64, ' C' in the site 70.	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D24A487C94DDFF2FA304A00E.taxon	discussion	Remarks. This very distinct small species (reaching 14.3 mm in SL) has some similarity to the species of Ancilla and was originally described in this genus. Nevertheless, the radula and molecular data undoubtedly place it in Turrancilla.	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D24A487C94DDFF2FA304A00E.taxon	distribution	Distribution. Philippines (Fig. 7 A).	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D24D487D94DDFD0AA00FA5B2.taxon	description	Figs 8, 9 A, 10, 12 A – B	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D24D487D94DDFD0AA00FA5B2.taxon	materials_examined	Type material. Holotype: Zoological Survey of India, ZSI M 918 / 1, SL 33 mm. Type locality. Off Andaman Is., 13 ° 17 ' 15 " N, 93 ° 10 ' E, 338 m. Sequenced material. SOLOMON ISLANDS • 1 lv; E Santa Isabel I.; 8 ° 17 ' S, 160 ° 00 ' E; depth 464 – 523 m; SALOMON 2, stn CP 2184; MNHN-IM- 2007 - 33304 • 1 lv; E Santa Isabel I.; 8 ° 17 ' S, 160 ° 00 ' E; depth 487 – 541 m; SALOMON 2, stn CP 2186 MNHN-IM- 2007 - 33282 • 2 lv; NW Santa Isabel I.; 7 ° 38 ' S, 157 ° 42 ' E; depth 400 – 475 m; SALOMON 2, stn CP 2212, MNHN-IM- 2007 - 33316, MNHN-IM- 2007 - 33317 • 1 lv; NW Santa Isabel I.; 7 ° 39 ' S, 157 ° 43 ' E; depth 495 – 650 m; SALOMON 2, stn CP 2213; MNHN-IM- 2007 - 33323 • 2 lv; Choiseul I.; 6 ° 39 ' S, 156 ° 14 ' E; depth 490 – 520 m; SALOMON 2, stn CP 2226; MNHN-IM- 2007 - 33295, MNHN-IM- 2007 - 33296 • 2 lv; Choiseul I.; 6 ° 37 ' S, 156 ° 13 ' E; depth 508 – 522 m; SALOMON 2, stn CP 2227; MNHN-IM- 2007 - 33319, MNHN-IM- 2007 - 33320 • 1 lv; Choiseul I.; 6 ° 35 ' S, 156 ° 11 ' E; depth 609 – 625 m; SALOMON 2, stn CP 2228; MNHN-IM- 2007 - 33318 • 2 lv; Vella Gulf; 7 ° 56 ' S, 156 ° 51 ' E; depth 460 – 487 m; SALOMON 2, stn CP 2262; MNHN-IM- 2007 - 33326, MNHN-IM- 2007 - 33327 • 2 lv; E Guadalcanal I.; 9 ° 19 ' S, 160 ° 06 ' E; depth 416 – 425 m; SOLOMONBOA 3, stn CP 2767; MNHN-IM- 2009 - 11939, MNHN-IM- 2009 - 11940 • 1 lv; E Guadalcanal I.; 9 ° 36 ' S, 160 ° 46 ' E; depth 448 – 523 m; SOLOMONBOA 3, stn CP 2849; MNHN-IM- 2007 - 36205. PAPUA NEW GUINEA • 1 lv; New Ireland; 2 ° 24 ' S, 150 ° 34 ' E; depth 388 – 572 m; KAVIENG 2014, stn DW 4424; MNHN-IM- 2013 - 58149 • 1 lv; New Ireland; 2 ° 23 ' S, 150 ° 35 ' E; depth 534 – 650 m; KAVIENG 2014, stn DW 4439; MNHN-IM- 2013 - 58320. Non-sequenced material. INDONESIA, Tanimbar Is. • 4 lv, 5 dd, 2 lv; 08 ° 20 ' S, 132 ° 11 ' E; depth 399 – 405 m; KARUBAR, stn CP 59; MNHN-IM- 2023 - 452 • 2 lv, 4 dd; 08 ° 42 ' S, 131 ° 53 ' E; depth 356 – 368 m; KARUBAR, stn CP 69; MNHN-IM- 2023 - 453 • 3 dd; 08 ° 50 ' S, 131 ° 33 ' E; depth 400 – 401 m; KARUBAR, stn CP 76; MNHN-IM- 2023 - 454 • 2 lv, 2 dd; 08 ° 57 ' S, 131 ° 27 ' E; depth 346 – 352 m; KARUBAR, stn CP 77; MNHN-IM- 2023 - 447, MNHN-IM- 2023 - 448 • 2 lv; 09 ° 23 ' S, 131 ° 00 ' E; depth 285 – 297 m; KARUBAR, stn CP 83; MNHN-IM- 2023 - 451. PHILIPPINES • 8 lv, 5 dd; Luzon I.; 15 ° 58 ' N, 121 ° 49 ' E; depth 422 – 431 m; AURORA 2007, stn CP 2658; MNHN uncatalogued • 1 lv, 1 dd; 16 ° 00 ' N, 121 ° 51 ' E; 431 – 432 m, AURORA 2007, stn CP 2735; MNHN-IM- 2023 - 454; SOLOMON ISLANDS • 5 lv; N Guadalcanal I.; 09 ° 15 ' S, 160 ° 09 ' E; depth 477 – 505 m; SALOMON 1, stn CP 1792; MNHN-IM- 2023 - 454 • 1 lv; N Guadalcanal I.; 09 ° 19 ' S, 160 ° 25 ' E; depth 469 – 481 m; SALOMON 1, stn CP 1796; MNHN uncatalogued • 2 lv; NW San Cristobal I.; 10 ° 10 ' S, 161 ° 22 ' E; depth 439 – 486 m; SALOMON 1, stn CP 1836; MNHN uncatalogued • 6 dd; SE Santa Isabel I.; 8 ° 17 ' S, 160 ° 00 ' E; depth 464 – 523 m; SALOMON 2, stn CP 2184; MNHN uncatalogued • 1 lv; SE Santa Isabel I.; 8 ° 17 ' S, 160 ° 00 ' E; depth 487 – 541 m; SALOMON 2, stn CP 2186; MNHN uncatalogued • 1 lv, 1 dd; SE Santa Isabel I.; 8 ° 18 ' S, 160 ° 01 ' E; depth 495 – 677 m; SALOMON 2, stn CP 2188; MNHN uncatalogued • 4 lv; SE Santa Isabel I.; 8 ° 17 ' S, 160 ° 00 ' E; depth 482 – 604 m; SALOMON 2, stn CP 2187; MNHN uncatalogued • 2 lv; NW Santa Isabel I.; 7 ° 38 ' S, 157 ° 42 ' E; depth 400 – 475 m; SALOMON 2, stn CP 2212; MNHN-IM- 2023 - 457 (radula prepared), MNHN-IM- 2023 - 458 • 1 lv; NW Santa Isabel I.; 7 ° 42 ' S, 157 ° 44 ' E; depth 550 – 682 m; SALOMON 2, stn CP 2214; MNHN uncatalogued • 1 lv; Vella Gulf; 7 ° 56 ' S, 156 ° 51 ' E; depth 460 – 487 m; SALOMON 2, stn CP 2262; MNHN uncatalogued. TAIWAN • 3 lv, 1 dd; NE coast off Taschi; 24 ° 48 ' N, 122 ° 08 ' E; 326 – 331 m; TAIWAN 2001, stn CP 102; MNHN uncatalogued • 1 lv; NE coast off Taschi; 24 ° 49 ' N, 122 ° 06 ' E; 367 – 424 m; TAIWAN 2001, stn CP 103; MNHN uncatalogued.	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D24D487D94DDFD0AA00FA5B2.taxon	description	Description. Shell of large size for the genus (up to 48 mm), elongate-oval. Protoconch and uppermost teleoconch whorls decollated in all available specimens, including smallest ones with shell length of only 9 mm. Spire moderately high, conical; primary callus rather thin, covering all remaining whorls; surface of callus weakly glossy, strongly microshagreened. Spiral striation visible through primary callus on apertural side of shell. Secondary callus thick, forming tongue-shaped pad extending to right-ventral side of last whorl and continuing up as thick pad reaching suture of antepenultimate whorl. Surface of secondary callus strongly microshagreened and similar to that of primary callus. Body whorl cloak glossy, covered by distinct very narrow, wavy, and closely spaced spiral striation; distinct lighter band in lower part adjoining anterior band. Olivoid groove not pronounced. Rear edge of anterior band distinct in a shape of sharp step. Anterior band usually distinctly darker than cloak, but occasionally of same colour. Upper anterior band usually very weakly convex to nearly flat, separated from lower anterior band by additional distinct ridge. Lower anterior band significantly narrower, nearly flat to weakly convex and separated from plication plate by deep and broad groove. Plication plate with 5 – 7, exceptionally 8 distinct narrow spiral ridges. Columella with indistinct spiral cords. Aperture narrow oval, acute posteriorly, widest medially; outer lip thickened, slightly opisthocline, sometimes with weak extension corresponding to labral denticle; siphonal notch of medium depth, asymmetrical. Columella crossed by variably distinct in colour brown spiral band corresponding to lower part of anterior band, sometimes brownish in area corresponding to upper anterior band. Shell colour variable from very pale creamy to medium dark of different shades of orange and brown, sometimes part of shell cloak with bluish tint crossed by irregular interrupting and very narrow brown lines. Primary callus usually slightly different from cloak, sometimes slightly darker. Secondary callus rose-creamy on ventral part near aperture and of same shade as primary callus on dorsal side. Upper edge of secondary callus usually highlighted by narrow darker band. Anterior band distinctly darker than cloak, brownish with still darker lower part. Plication plate white to creamy, columella intersected by brown line corresponding to lower plication plate. Operculum large, narrow lanceolate, with terminal nucleus. Radula (Fig. 9 A) studied in one specimen (MNHN-IM- 2023 - 457 – SL 35.1 mm), typical for the genus, of about 60 transverse rows of teeth, 5 nascent. Radula about 1.65 mm in length (0.7 % of AL), membrane about 130 µm in width (0.55 % of AL). Diagnostic cox- 1 sites. ' T' in the site 322, ' G' in the site 328, ' C' in the site 343.	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D24D487D94DDFD0AA00FA5B2.taxon	discussion	Remarks. The species was originally described from the Andaman Islands, whereas all our sequenced specimens were collected from the Solomons and Papua New Guinea, which are considerably distant from the type locality. In addition to the sequenced specimens, we have extensive material from the Tanimbar Islands (Indonesia) that is morphologically indistinguishable (see, e. g., Fig. 8 J). The species was also recently reported off Java (Indonesia), by Tan & Islami (2021). Morphologically, our specimens, as well as those illustrated by Tan & Islami, are very similar. Although the holotype is a worn and faded specimen (Fig. 8 A – B), it cannot be reliably distinguished from the specimens examined and illustrated herein based on the main characters. Therefore, until conspecificity with the specimens from the type locality or nearby regions can be confidently rejected through molecular data, we apply the name T. glans to this molecularly defined species. Turrancilla glans is quite similar to T. monachalis (Ninomiya, 1988) and T. heraldei sp. nov., both of which are molecularly distinct species. It differs from T. monachalis in attaining a larger size (the largest T. glans specimen available to us has a SL of 47.8 mm – MNHN-IM- 2007 - 33317, while the largest T. monachalis specimen has a SL of 37 mm) and having a narrower lower part of the anterior band (average ratio lab / uab 0.29 vs. 0.43). Additionally, the species exhibit markedly different operculum morphology: in T. glans, it is typically large, spanning nearly the entire aperture, and narrow lanceolate, whereas in T. monachalis, it is small and broadly triangular. For differences with T. heraldei sp. nov., see the description of the latter species.	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D24D487D94DDFD0AA00FA5B2.taxon	distribution	Distribution. From Andaman Islands to Philippines, Taiwan, Tanimbar Islands in Indonesia estward to Solomon Islands (Fig. 10).	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D251486494DDFB6BA302A00E.taxon	description	Figs 7 B, 9 B – C, 11, 12 C – D, 13	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D251486494DDFB6BA302A00E.taxon	materials_examined	Type material. Holotype: MNHN-IM- 2007 - 31959. Type locality. PHILIPPINES; Bohol Sea, off Balicasag I.; 9 ° 32 ' N, 123 ° 39 ' E; depth 757 – 760 m; PANGLAO 2005, stn CP 2336. Sequenced material. PHILIPPINES • 2 lv; Bohol Sea, Maribojoc Bay; 9 ° 38 ' N, 123 ° 43 ' E; depth 584 – 596 m; PANGLAO 2005, stn CP 2333; MNHN-IM- 2007 - 31949, MNHN-IM- 2007 - 31951 • 1 lv; Bohol Sea, off Balicasag I.; 9 ° 32 ' N, 123 ° 39 ' E; depth 757 – 760 m; PANGLAO 2005, stn CP 2336; MNHN-IM- 2007 - 31959 • 2 lv; Bohol Sea, off Pamilacan I.; 9 ° 25 ' N, 123 ° 50 ' E; depth 544 – 712 m; PANGLAO 2005, stn CP 2341; MNHN-IM- 2007 - 31960, MNHN-IM- 2007 - 31961 • 2 lv; Bohol Sea, Maribojoc Bay; 9 ° 36 ' N, 123 ° 44 ' E; depth 382 – 434 m; PANGLAO 2005, stn CP 2395; MNHN-IM- 2007 - 31965, MNHN-IM- 2007 - 31966. Non-sequenced material. PHILIPPINES • 2 dd; Bohol Sea, Maribojoc Bay; 9 ° 38 ' N, 123 ° 43 ' E; depth 584 – 596 m; PANGLAO 2005, stn CP 2333 • 2 lv; Bohol Sea; 9 ° 34 ' N, 123 ° 38 ' E; depth 729 – 773 m; PANGLAO 2005, stn CP 2335; MNHN-IM- 2023 - 449 (radula voucher), MNHN-IM- 2023 - 450 • 17 dd; Bohol Sea, off Balicasag I.; 9 ° 32 ' N, 123 ° 39 ' E; depth 757 – 760 m; PANGLAO 2005, stn CP 2336 • 2 dd; Bohol Sea; 9 ° 24 ' N, 123 ° 53 ' E; depth 1240 – 1258 m; PANGLAO 2005, CP 2342 • 2 lv, 15 dd; Bohol Sea; 9 ° 31 ' N, 124 ° 03 ' E; depth 810 – 812 m; PANGLAO 2005, stn CP 2351 • 26 dd; Bohol Sea; 9 ° 27 ' N, 124 ° 03 ' E; depth 923 – 1260 m; PANGLAO 2005, stn CP 2352 • 1 lv, 2 dd; Bohol Sea; 9 ° 24 ' N, 124 ° 11 ' E; depth 1764 – 1775 m; PANGLAO 2005, stn CP 2355 • 4 dd; Bohol / Sulu Sea sill; 8 ° 52 ' N, 123 ° 37 ' E; depth 569 – 583 m; PANGLAO 2005, stn CP 2358 Bohol / Sulu Sea sill; 8 ° 45 ' N, 123 ° 19 ' E; depth 338 – 351 m; PANGLAO 2005, stn CP 2383 • 13 dd; Bohol Sea; 9 ° 27 ' N, 123 ° 35 ' E; depth 762 – 786 m; PANGLAO 2005, stn CP 2388 • 25 dd; Bohol Sea; 9 ° 27 ' N, 123 ° 43 ' E; depth 672 – 645 m; PANGLAO 2005, stn CP 2389 • 5 dd; Bohol Sea; 9 ° 28 ' N, 123 ° 38 ' E; depth 784 – 786 m; PANGLAO 2005, stn CP 2390 • 1 dd; Bohol Sea, off Balicasag I.; 9 ° 29 ' N, 123 ° 40 ' E; depth 470 – 566 m; PANGLAO 2005, CP 2394 • 6 dd; Bohol Sea, Maribojoc Bay; 9 ° 35 ' N, 123 ° 42 ' E; depth 642 – 669 m; PANGLAO 2005, stn CP 2397 • 3 dd; Bohol Sea, Maribojoc Bay; 9 ° 39 ' N, 123 ° 43 ' E; depth 481 – 505 m; PANGLAO 2005, stn CP 2404 • 2 lv, 1 dd; Bohol Sea, Maribojoc Bay; 9 ° 39 ' N, 123 ° 46 ' E; depth 387 – 453 m; PANGLAO 2005, stn CP 2405 • 1 dd; N Panay I.; 11 ° 58 ' N, 122 ° 02 ' E; depth 486 – 551 m; MUSORSTOM 3, stn CP 135 • 1 dd, attached to shell of Xenophora pallidula (Reeve, 1842); Palawan (Fig. 13).	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D251486494DDFB6BA302A00E.taxon	description	Description (holotype). Shell of medium size for the genus (SL 39.0 mm), elongate oval, slender. Protoconch low dome shaped, smooth, creamy coloured. About one whorl with distinct shallow suture, further suture overlaid by callus. Protoconch borders indistinct, rendering whorl count impossible. Spire moderately high, conical; with weakly convex sides. Primary callus rather thin, covering all remaining whorls; surface of callus weakly glossy, strongly microshagreened. Spiral striation visible through primary callus on apertural side of shell. Secondary callus moderately thick, forming tongue-shaped pad extending to right-ventral side of last whorl and continuing up as pad reaching suture of antepenultimate whorl. Surface of secondary callus strongly microshagreened and similar to that of primary callus. Body whorl cloak glossy, covered by distinct very narrow, wavy, and closely spaced spiral striation; distinct but very narrow lighter band in lower part adjoining anterior band. Olivoid groove not pronounced. Rear edge of anterior band distinct in a shape of sharp step, obscured in ending part of last whorl adjacent to aperture lip. Anterior band with darker narrow brown band in lower part of upper anterior band followed by a lighter band adjacent to border between upper and lower parts of anterior band. Lower anterior band separated by distinct elevated ridge, darker orange-brown with irregularly spaced still darker transverse lines. Upper anterior band nearly flat, very weakly concave medially, lower anterior band nearly flat. Lower anterior band narrower, separated from plication plate by deep and broad groove. Plication plate with 7 equally developed ridges. Columella with few weak spiral cords in lower part, crossed by two distinct brown spiral bands, the upper narrower and corresponding to darker spiral band of the upper anterior band, the lower broader band corresponding to abapical part of lower anterior band, adjacent to plication plate. Aperture narrow oval, acute posteriorly, widest medially; outer lip sharp, nearly orthocline. Siphonal notch moderately deep, asymmetrical. Body whorl cloak orange-brownish, with bluish tint on dorsal side and irregularly spaced axial darker brown lines of varying width, especially distinct on dorsal side. Primary and secondary calluses of more intense orange, plication plate creamy. Radula (Fig. 9 B – C) studied in one specimen (MNHN-IM- 2023 - 449 – SL 31.8 mm), typical for the genus, of about 75 transverse rows of teeth, 5 nascent. Radula about 1.92 mm in length (10.1 % of AL), membrane about 175 µm in width (0.92 % of AL). Diagnostic cox- 1 sites. ' G' in the site 4, ' A' in the site 175.	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D251486494DDFB6BA302A00E.taxon	discussion	Remarks. The species is moderately variable in shell slenderness and colouration. At least one specimen is very light (Fig. 11 I – K) with light yellowish primary and secondary calluses and light-creamy body whorl cloak. Generally, the new species strongly resembles T. glans, differing, in several characters. First, protoconch is retained in all examined specimens of T. heraldei, even dead-collected ones, while it is always missing and upper whorls are strongly eroded in T. glans. Secondly, most of the specimens of T. heraldei sp. nov. have the two distinct and narrow spiral brown bands crossing the columella (Fig. 12 C – D), while only one distinctly does it in T. glans (Fig. 12 A – B). It should be mentioned, that the second, upper band is fading in dead collected shells and can be absent in some lightly-coloured live specimens (e. g.: Fig. 11 I – K). Finally, T. glans can attain a larger size, being up to 48 mm in SL vs 39 in T. heraldei sp. nov. One specimen was found attached to the shell of Xenophora pallidula (Reeve, 1842), collected in the Philippines (Fig. 13).	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D251486494DDFB6BA302A00E.taxon	etymology	Etymology. The species is named after Prof. Francisco M. Heralde III, from the University of Philippines, Manila with whom (at that time a PhD student) we processed the specimens of the new species during the PANGLAO 2005 expedition, in recognition of his participation in the “ bar-code team ” of MNHN expeditions to the Philippines.	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D251486494DDFB6BA302A00E.taxon	distribution	Distribution. Philippines (Fig. 7 B).	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D255486794DDFD0AA02FA78A.taxon	description	(Figs 9 D – F, 14, 15)	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D255486794DDFD0AA02FA78A.taxon	description	Turrancilla alcocki – Tan, Islami, 2021: 321, fig. 7 E.	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D255486794DDFD0AA02FA78A.taxon	materials_examined	Type material. Holotype – Zoological Survey of India, SL 38 mm (after Smith 1906). Type locality. Andaman Is., 11 ° 26 ' 30 " N, 92 ° 43 ' 45 " E, 378 fms (691 m), Investigator, stn 322, Sequenced specimens. PHILIPPINES • 1 lv; Luzon I.; 15 ° 58 ' N, 121 ° 49 ' E; depth 422 – 431 m; AURORA 2007, stn CP 2658; MNHN-IM- 2009 - 23361 (radula voucher, SL 32.1 mm) • 1 lv; Luzon I.; 15 ° 45 ' N, 121 ° 45 ' E; depth 562 m; AURORA 2007, stn CP 2663; MNHN-IM- 2009 - 11921 PAPUA NEW GUINEA • 1 lv; Mambare Bay; 7 ° 52 ' S, 148 ° 1 ' E; depth 498 – 501 m; BIOPAPUA, stn CP 3728; MNHN-IM- 2009 - 22260 SOLOMON SEA • 1 lv; New Britain, Ainto Bay; 6 ° 7 ' S, 149 ° 12 ' E; depth 260 m; MADEEP, stn CP 4331; MNHN-IM- 2013 - 46027 • 1 lv; New Britain, Ainto Bay; 6 ° 8 ' S, 149 ° 17 ' E; depth 410 – 614 m; MADEEP, stn CP 4338; MNHN-IM- 2013 - 46396 • 1 lv; New Britain, Ainto Bay; 6 ° 10 ' S, 149 ° 18 ' E; depth 510 – 743 m; MADEEP, stn CP 4339; MNHN-IM- 2013 - 46429 JAPAN • 1 lv; appr. 32 ° 34 ′ N, 132 ° 22 ′ E; from fishermen; MNHN-IM- 2019 - 620 Non-sequenced specimens. INDONESIA • 4 lv, 5 dd; Tanimbar Is.; 08 ° 41 ' S, 131 ° 47 ' E; depth 410 – 413 m; KARUBAR, stn CP 70; MNHN-IM- 2023 - 442, MNHN-IM- 2023 - 441 • 3 lv, 5 dd; Tanimbar Is.; 08 ° 36 ' S, 131 ° 33 ' E; depth 676 – 699 m; KARUBAR, stn CP 72 • 4 lv, 3 dd; Tanimbar Is.; 08 ° 46 ' S, 131 ° 36 ' E; depth 451 – 452 m; KARUBAR, stn CP 75; MNHN-IM- 2023 - 443, MNHN-IM- 2023 - 444 • 1 lv; Tanimbar Is.; 08 ° 50 ' S, 131 ° 33 ' E; depth 400 – 401 m; KARUBAR, stn CP 76; MNHN-IM- 2023 - 440 • 1 lv; Makassar; 00 37 ’ S, 119 33 ’ E; depth 675 m; CORINDON stn CH 240; MNHN-IM- 2023 - 445. SOLOMON ISLANDS • 3 lv, 3 dd; N Guadalcanal I.; 09 ° 21 ' S, 159 ° 56 ' E; depth 582 – 594 m; SALOMON 1, stn CP 1749 • 1 lv; N Guadalcanal I.; 09 ° 16 ' S, 159 ° 55 ' E; depth 693 – 696 m; SALOMON 1, stn CP 1750 • 1 lv; NE Guadalcanal I.; 09 ° 35 ' S, 160 ° 43 ' E; depth 367 – 500 m; SALOMON 1, stn CP 1805 • 1 lv; SW Santa Isabel I.; 08 ° 25 ' S, 159 ° 27 ' E; depth 440 – 521 m; SALOMON 2, stn CP 2194 • 1 lv, 1 dd; SW Santa Isabel I.; 08 ° 26 ' S, 159 ° 26 ' E; depth 543 – 593 m; SALOMON 2, stn CP 2195 • 1 lv; SW Santa Isabel I.; 08 ° 26 ' S, 159 ° 26 ' E; depth 724 – 765 m; SALOMON 2, stn CP 2196 • 1 lv; Vella Gulf; 07 ° 52 ' S, 156 ° 51 ' E; depth 515 – 520 m; SALOMON 2 stn CP 2264 PHILIPPINES • 6 dd; Luzon / Mindoro Is.; 13 ° 40 ' N, 120 ° 43 ' E; MUSORSTOM 2 stn CP 25; MNHN-IM- 2023 - 446 • 6 dd; Luzon / Mindoro Is.; 13 ° 50 ' N, 120 ° 29 ' E; depth 441 – 550 m; MUSORSTOM 2 stn CP 78 • 1 dd; between Ambil I. and Luzon I.; 13 ° 48 ' N, 120 ° 31 ' E; depth 640 – 688 m; MUSORSTOM 3, stn CP 106 • 1 dd; between Buad I. & Panay I.; 11 ° 50 ' N, 121 ° 42 ' E; depth 815 – 821 m; MUSORSTOM 3 stn CP 128 • 1 dd; • 2 dd; Bohol Sea; 9 ° 27 ' N, 124 ° 03 ' E; depth 923 – 1260 m; PANGLAO 2005, stn CP 2352 • 1 dd; Bohol Sea; 9 ° 24 ' N, 124 ° 11 ' E; depth 1764 – 1775 m; PANGLAO 2005, stn CP 2355 • 1 dd; Bohol, Sulu Sea sill; 08 ° 49 ' N, 123 ° 02 ' E; depth 2120 – 2149 m; PANGLAO 2005, stn CP 2386	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D255486794DDFD0AA02FA78A.taxon	description	Description. Shell of large size for the genus (up to 50 mm with corroded apex), ovate. Protoconch and upper teleoconch whorls strongly eroded in all available specimens. spire medium low (based on least eroded specimens Fig. 14 E). Primary callus thick and covering all remaining whorls, slightly impressed at sutures; surface of callus weakly glossy, strongly microshagreened. Secondary callus thick, forming tongue-shaped pad extending to right-ventral side of last whorl and continuing up as fairly thin, reaching suture of antepenultimate whorl and following parallel to suture. Surface of secondary callus strongly microshagreened. Body whorl cloak glossy, covered by distinct spiral striation. Olivoid groove not pronounced; some specimens with very shallow depression corresponding to position of groove, usually visible only at oblique light. Rear edge of anterior band distinct, in shape of low but sharp step. Anterior band distinctly different from shell cloak in colour, darker, of different shades of brown-orange, usually with lighter narrow band adapically. Upper anterior band weakly convex, sometimes nearly flat, separated from lower anterior band by additional distinct ridge. Lower anterior band narrower, nearly flat to weakly convex and separated from plication plate by deep and broad groove. Plication plate of different morphology, with 7 – 8 low but distinct narrow spiral ridges (Fig. 15 A – B) in specimens from Philippines and Japan, or 3 – 4 low indistinct rounded ridges in specimens from Papua New Guinea and Indonesia (Fig. 15 C – D). Columella either smooth (in specimens with thicker columellar callus) or with very low indistinct spiral cords on columella and parietal part of last whorl. Aperture narrow oval, acute posteriorly, widest medially; outer lip sharp, slightly opisthocline, with slight rounded angulation corresponding to position of labral denticle; siphonal notch moderately deep, asymmetrical. Shell colour variable. Last whorl cloak from light to dark orange-brown, with part of the shell (usually dorsal side of last whorl) with bluish tint. Primary callus of the same shade as cloak, sometimes slightly darker. Secondary callus usually creamy on ventral part and of same shade as primary callus on dorsal side. Anterior band distinctly darker than cloak, brownish with lighter median spiral band. Plication plate white to creamy. Operculum typical for the genus, large, dark-brown, spanning nearly entire aperture, lanceolate, thick, with terminal often broken off nucleus. Right edge narrowly thickened and separated by distinct groove running entire operculum length, broadening with operculum growth. Radula (Fig. 9 D – F) studied in two specimens (MNHN-IM- 2009 - 23361 – SL 32.3 mm, MNHN-IM- 2023 - 445 – SL 36.6 mm), typical for the genus, of about 75 – 90 transverse rows of teeth, 10 – 15 nascent. Radula about 2.12 – 2.76 mm in length (10.4 – 11.5 % of AL), membrane about 175 – 260 µm in width (0.86 – 1.08 % of AL). Diagnostic cox- 1 sites. ' G' in the site 181, ' G' in the site 310.	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D255486794DDFD0AA02FA78A.taxon	discussion	Remarks. Radulae of studied specimens differ in the shape of the rachidian teeth. In MNHN-IM- 2023 - 445 the cusps are shorter and broader (Fig. 9 F) than in MNHN-IM- 2009 - 23361 (Fig. 9 D – F). The radula of the latter specimens showed heavy wear of the rachidians of 14 – 15 anteriormost rows of teeth (Fig. 9 E), while lateral teeth have only slight abrasion of the tips. The species is represented in our material by a few sequenced specimens: two from the Philippines, one from Japan, and four from Papua New Guinea. Additionally, many specimens were collected in eastern Indonesia. There is significant variability in shell shape, which is largely determined by the degree of erosion of the upper whorl. Sometimes shell completely truncated with nearly flat apex and even upper part of secondary callus on last whorl can be affected by erosion. This variability produces shell outlines ranging from short and truncate to moderately elongated. Furthermore, there is a notable difference in the morphology of the plication plate, forming two morphologically distinct groupings: one from the Philippines and Japan with numerous, sharp, distinct spiral ridges, and another from Papua New Guinea and Indonesia with fewer, indistinct, rounded ridges. However, the specimens from the Philippines and Japan do not constitute a molecular clade, which prevents their separation into two species. Among the non-sequenced material from the Philippines, we found a live-collected specimen (MNHN-IM- 2023 - 446) and several dead ones with plication plate morphology intermediate between the two forms. The specimens from Papua New Guinea and Indonesia closely resemble the holotype (not examined but illustrated by Annandale & Stewart, 1909), differing only in their slightly more slender shells. Therefore, we apply the name T. alcocki to the molecularly defined species, at least until material suitable for sequencing can be obtained from the type locality or nearby regions. The species was also recently recorded off Java, Indonesia, by Tan & Islami (2021). Occasionally, very pale specimens are collected in both Indonesia and the Philippines. These specimens display a light cream cloak with even lighter primary and secondary calluses, as well as a slightly darker anterior spiral line in the lower anterior band (e. g., MNHN-IM- 2023 - 445 – Fig. 14 L). The specimen from Japan (Fig. 14 E) differs from all others in having a more elongated shell shape and less erosion of the upper whorls. Nevertheless, it shares the more numerous spiral ridges on the plication plate with the specimens from the Philippines, while the latter exhibit closer similarity to those from Papua New Guinea.	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D255486794DDFD0AA02FA78A.taxon	distribution	Distribution. From Andaman Islands to Philippines, Tanimbar Islands (Indonesia), Papua New Guinea and Solomons (Fig. 16).	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D256486B94DDF98EA371A0C6.taxon	description	(Figs 16, 17, 18, 19 A, 20 A – B)	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D256486B94DDF98EA371A0C6.taxon	materials_examined	Type material. Holotype NSMT 644460, paratype 2, NSMT 644461. Type locality. Off Southwest Taiwan, 40 fms deep. Studied material (sequenced). TAIWAN • 2 lv; off Liuqiu I.; 22 ° 13 ' N, 120 ° 25 ' E; depth 286 – 347 m; TAIWAN 2013, stn CP 4085; MNHN-IM- 2013 - 41618, MNHN-IM- 2013 - 41629 • 5 lv; off Liuqiu I.; 22 ° 12 ' N, 120 ° 24 ' E; depth 343 – 393 m; TAIWAN 2013, stn CP 4086; MNHN-IM- 2013 - 41613, MNHN-IM- 2013 - 41614, MNHN-IM- 2013 - 41615, MNHN-IM- 2013 - 41616, MNHN-IM- 2013 - 41617 • 2 lv; off Liuqiu I.; 22 ° 11 ' N, 120 ° 22 ' E; depth 387 – 510 m; TAIWAN 2013, stn CP 4087; MNHN-IM- 2013 - 41630, MNHN-IM- 2013 - 41631 • 5 lv; off Liuqiu I.; 22 ° 11 ' N, 120 ° 23 ' E; depth 372 – 436 m; TAIWAN 2013, stn CP 4088; MNHN-IM- 2013 - 41608, MNHN-IM- 2013 - 41609, MNHN-IM- 2013 - 41610, MNHN-IM- 2013 - 41611, MNHN-IM- 2013 - 41612 • 4 lv; off Liuqiu I.; 22 ° 13 ' N, 120 ° 25 ' E; depth 310 – 330 m; TAIWAN 2013, stn CP 4089; MNHN-IM- 2013 - 41619, MNHN-IM- 2013 - 41620, MNHN-IM- 2013 - 41621, MNHN-IM- 2013 - 41622 • 4 lv; off Liuqiu I.; 22 ° 13 ' N, 120 ° 23 ' E; depth 329 – 371 m; TAIWAN 2013, stn CP 4090; MNHN-IM- 2013 - 41623, MNHN-IM- 2013 - 41624, MNHN-IM- 2013 - 41625, MNHN-IM- 2013 - 41626, MNHN-IM- 2013 - 41627 • 5 lv; off Liuqiu I.; 22 ° 13 ' N, 120 ° 24 ' E; depth 310 – 346 m; NanHai 2014, stn CP 4099; MNHN-IM- 2013 - 44003, MNHN-IM- 2013 - 44005, MNHN-IM- 2013 - 44006, MNHN-IM- 2013 - 44007, MNHN-IM- 2013 - 44008 • 4 lv; off Liuqiu I.; 22 ° 13 ' N, 120 ° 32 ' E; depth 327 – 372 m; DongSha 2014, stn CP 4120; MNHN-IM- 2013 - 50002, MNHN-IM- 2013 - 50003, MNHN-IM- 2013 - 50004, MNHN-IM- 2013 - 50007.	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D256486B94DDF98EA371A0C6.taxon	description	Description. Shell of medium size for the genus (up to 37 mm), elongate oval. Protoconch low dome shaped, smooth, light yellow-brownish, of about one whorl with distinct impressed suture, further whorls’ sutures overlaid by callus. Protoconch borders indistinct, rendering whorl count impossible. Spire low, conical; primary callus moderately thick. Primary callus thick and covering all whorls except initial part of protoconch; surface of callus weakly glossy, strongly microshagreened. Secondary callus very thick, forming tongue-shaped pad extending to right-ventral side of last whorl and continuing up as thick pad, covering most of spire whorls. Surface of secondary callus strongly microshagreened and similar to that of primary callus. Body whorl cloak glossy, covered by fairly distinct spiral striation, often cloak with distinct lighter band in lower part adjoining anterior band. Olivoid groove not pronounced, in some specimens shallow depression in anterior part of last whorl close to aperture, corresponding to position of groove (e. g. MNHN-IM- 2013 - 41616 – Fig. 17 P) and terminating in obtuse and very short, rounded extension on outer lip. Rear edge of anterior band distinct in a shape of sharp step. Anterior band different from shell cloak in colour, darker, of different shades of brown-orange. Upper anterior band weakly convex, sometimes nearly flat, separated from lower anterior band by additional distinct ridge. Upper anterior band with submedian darker spiral band. Lower anterior band narrower, nearly flat to weakly convex and separated from plication plate by deep and broad groove. Plication plate with 5 – 6 (5 in holotype) distinct narrow spiral ridges. Columella smooth. Aperture narrow oval, acute posteriorly, widest medially; outer lip sharp, slightly opisthocline, occasionally with weak extension corresponding to labral denticle; siphonal notch medium deep, asymmetrical. Columella crossed by blurred brown spiral band corresponding to lower part of anterior band, rarely with another spiral band corresponding to upper anterior band. Shell colour variable from very pale creamy to different shades of orange and brown, sometimes part of shell with bluish tint. Primary callus of the same shade as cloak, sometimes slightly darker. Secondary callus creamy on ventral part near aperture and of same shade as primary callus on dorsal side. Anterior band distinctly darker than cloak, brownish with darker median spiral band in upper part. Plication plate white to creamy. Operculum vestigial, spanning only about 1 / 5 of aperture (Fig. 17 K), broadly triangular, very thin (Fig. 17 L), situated on dorsal part of the foot in live animal and difficult to find in preserved specimen (partially or completely concealed with contracted parapodia: Fig. 18). Radula (Fig. 20 A – B) studied in one specimen (MNHN-IM- 2013 - 44003, SL 30.7 mm), typical for the genus, of about 70 transverse rows of teeth, 10 – 15 nascent. Radula about 1.6 mm in length (7.5 % of AL), membrane about 135 µm in width (0.6 % of AL). Diagnostic cox- 1 sites. ' C' in the site 35, ' A' in the site 55.	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D256486B94DDF98EA371A0C6.taxon	discussion	Remarks. The species exhibits moderate variability in shell outline and colouration. It was originally described without a precise locality, from a depth of approximately 75 m off southwestern Taiwan. However, all our specimens were collected much deeper, from 286 to 387 m. The species closely resembles T. glans but differs in its smaller size (the largest specimen of T. monachalis in our collection measures 37 mm in SL, while T. glans reaches 47.8 mm). Another distinguishing feature is the relative width of the anterior and posterior parts of the anterior band. In T. monachalis, the average width of the lower part constitutes 43 % of the width of the anterior part (range: 37 – 47 %), whereas in T. glans, it is 29 % of the width of the anterior part (range: 26 – 30 %) (Fig. 19). While these conchological differences are subtle, the species have entirely different operculum morphology. Furthermore, molecular data clearly confirm the existence of two distinct species. Particular attention should be given to the opercula of the type specimens. In the original description, Ninomiya (1988) illustrated the holotype in a partially lateral position, hindering detailed examination of the operculum. Examination of the holotype and paratype 2 clearly demonstrated that they possess the operculum of Amalda, which is oval, thin, and has a subcentral nucleus shifted towards the lower edge (Fig. 17 D). This is entirely different from the opercula of our specimens. Ninomiya did not specify the source of the material, which most likely came from dealers. It is possible that the opercula were intentionally or unintentionally switched with those of an Amalda species, especially since the vestigial opercula of T. monachalis are challenging to observe and retrieve.	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D256486B94DDF98EA371A0C6.taxon	distribution	Distribution. Taiwan (Fig. 16).	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D25D486D94DDFF2FA012A076.taxon	description	(Figs 6 G – I, 20 C – D)	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D25D486D94DDFF2FA012A076.taxon	materials_examined	Type material. Holotype MNHN-IM- 2000 - 1459, SL 12.9 mm, and paratype MNHN, SL 10.3 mm. Type locality. Off Mauritania, 20 ° 30 ' N, 18 ° 34 ' W, 1855 m, R / V Atalante, EUMELI 2, stn CP 05, 06 February 1991.	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D25D486D94DDFF2FA012A076.taxon	description	Description. Shell of small size for the genus (up to 13 mm), ovate. Protoconch low dome shaped, smooth, glossy, white, of about one whorl with distinct impressed suture, further suture overlaid by callus. Spire low, conical; primary callus medium thick and covering all whorls except protoconch, slightly impressed at sutures; surface of callus glossy, microshagreened; secondary callus medium thick, extending to right-ventral side of last whorl and continuing up as fairly thin, tongue-shaped pad, reaching suture of antepenultimate whorl. Surface of secondary callus very similar to that of primary one, borders indistinct. Body whorl cloak glossy, covered by very weak and indistinct thin spiral striation. Olivoid groove from very shallow and indistinct to rather distinct. Rear edge of anterior band distinct, in a shape of low but sharp step. Anterior band is of same colour as cloak, microshagreened, not separated into upper and lower parts. Plication plate well separated by broad and deep groove, with poorly pronounced 3 rounded spiral ridges (paratype) or lacking ridges completely (holotype). Columella smooth. Aperture oval, rounded posteriorly; outer lip sharp, in side view slightly opisthocline, with slight, angulation, not corresponding to olivoid groove; siphonal notch moderately deep, asymmetrical. Colour white with slightly yellowish sutural band, overlaid by secondary callus. Operculum spanning 2 / 3 of aperture (but partly worn), subtriangular, very thin and transparent, yellow, nucleus corroded, but probably terminal (Kantor & Bouchet 1999: fig. 2 C). Distinct, narrow, longitudinal, slightly curved rib on its inner surface. Columellar muscle attachment zone long, ovate. Radula (Fig. 20 C – D) studied in holotype, typical for the genus, of only about 35 transverse rows of teeth. Radula about 0.6 mm in length (8.9 % of AL), membrane about 150 µm in width (2.2 % of AL).	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D25D486D94DDFF2FA012A076.taxon	discussion	Remarks. This species is known from two specimens only, reaching 12.9 mm in SL. It is the only representative of the genus in the Atlantic Ocean known so far. Judging from the small size of the penis in the holotype, it may not be fully mature.	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D25C486E94DDFDD2A318A296.taxon	description	(Fig. 21 A – E)	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D25C486E94DDFDD2A318A296.taxon	materials_examined	Type material. Holotype of Turrancilla apicalis Kira, 1959 – Osaka Museum of Natural History, 8082. Holotype of Ancillus apicalis Ninomiya, 1988 – NSMT 64458, paratype 1 NSMT 64459. Type localities: Turrancilla apicalis – Tosa; Ancillus apicalis – Off Daiosaki Point, Shima Peninsula, Japan, 200 – 250 m in depth.	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D25C486E94DDFDD2A318A296.taxon	discussion	Remarks. The name was first introduced by T. Kira in the first edition of his Coloured Illustrations of the Shells of Japan (Kira 1954) without any description. Therefore, the name was long considered a nomen nudum. However, in the second edition of the book, Kira (1959) provided a brief description in Japanese, thus validating the name (Bieler & Petit 1990). In subsequent printings of the book, Kira referred to the species as Turrancilla suavis (Yokoyama, 1926) (a fossil from the Cenozoic: Hijikata Formation). This nomenclature was later adopted by various authors (e. g., Higo et al. 1999). Ninomiya (1988) appears to have overlooked this validation and described the species again under the genus Ancillus, referencing Kira. Consequently, Ninomiya’s name is a junior subjective synonym and a secondary homonym of Turrancilla apicalis Kira, 1959. Comparison of the holotypes of Turrancilla apicalis (Osaka Museum of Natural History; Fig. 21 A – C) and Ancillus apicalis (Fig. 21 D – E) leaves little doubt that the specimens are conspecific. Ninomiya (1988) also compared the species to Ancilla suavis Yokoyama, 1926, and concluded that it represents a distinct species, differing “ in having a remarkably deep depression on the subsutural band, a slender shape, and a larger size ” (p. 147). The holotype of Ancilla suavis was deposited in UMUT CM 23364. However, according to the museum’s website, it is currently missing (https: // umdb. um. u-tokyo. ac. jp / DKoseibu / en / Collection / detail. php? umutNo = 23364). The existing illustration of the type does not allow for resolution of the synonymy between suavis and apicalis. Thus, we follow Ninomiya’s (1988) opinion and consider them as two distinct species. T. apicalis is recognized as the only recorded Recent species of the genus in Japan (Okutani 2017). The single specimen we sequenced from Japan falls within T. alcocki but differs from the types of T. apicalis by having a broader shell and a higher aperture. Nevertheless, it is important to emphasize that we are working with very limited material, and interspecific variation remains unstudied. The species is very similar to T. glans, if distinguishable at all. A minor difference is that, in T. glans, the colour band crossing the columella (corresponding to the lower anterior band) is distinct and narrow. In contrast, in T. apicalis, it is either absent (as in the holotypes of A. apicalis and T. apicalis) or represented by a broad band with blurred edges (as seen in the paratype of A. apicalis). Due to the limited material from Japan and the lack of molecular data, the variability of the species could not be examined. Considering the large geographic distance between specimens confidently identified as T. apicalis and our specimens of T. glans, we take a conservative approach and regard T. apicalis as a separate species, pending additional molecular data from Japan.	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D25E486F94DDFF67A1ADA187.taxon	description	(Fig. 21 F – H)	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D25E486F94DDFF67A1ADA187.taxon	materials_examined	Type material. Holotype USNM 784573. Type locality. Off Punto Fijo, Gulf of Venezuela, Venezuela, 35 m.	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
8F1D4A79D25E486F94DDFF67A1ADA187.taxon	discussion	Remarks. Petuch (1987) attributed the species to Turrancilla without any discussion. The species is known only from the empty shell of the holotype, and its taxonomic position remains unclear. However, it differs from other Turrancilla species in several aspects. First, it was collected at much shallower depths (35 m), while the shallowest occurrence of Turrancilla spp. is around 200 m, typically below 350 m, and as deep as 1855 m. Second, in other Turrancilla species, the suture is overlaid by a thick secondary callus, rendering it invisible from the exterior; its position can only be inferred from a shallow depression. In contrast, in “ T. ” williamsoni, the suture, though overlaid by a relatively thinner callus, is clearly visible. Third, the olivoid groove is very distinct, while the olivoid band is relatively much narrower than in Turrancilla species that exhibit a very weak groove (T. akontistes and T. reboriae). Finally, the secondary callus forms a distinct keel-like thickening just above the suture, which is absent in studied Turrancilla species. Conversely, features of “ T. ” williamsoni are found in various Amalda species (see, e. g., Kantor et al. 2020), which, in the Western Atlantic, are known to inhabit a similar depth range (Malacolog 4.1.1 – Rosenberg 2009). Thus, in the absence of data on the operculum and radula, we conditionally reassign the species to the genus Amalda.	en	Kantor, Yuri, Strano, Giorgio, Vervaet, Fred, Weddingen, Mélanie, Puillandre, Nicolas (2025): Revision of the genus Turrancilla (Neogastropoda: Olivoidea: Ancillariidae) in an integrative taxonomy framework. Zootaxa 5647 (5): 401-434, DOI: 10.11646/zootaxa.5647.5.1, URL: https://doi.org/10.11646/zootaxa.5647.5.1
