identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
C62094596522568D9C1F4BB86E22A615.text	C62094596522568D9C1F4BB86E22A615.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pupiformidae Lee & Kim 2026	<div><p>Pupiformidae fam. nov.</p><p>Diagnosis.</p><p>As for the type genus.</p><p>Type genus.</p><p>Pupiformus gen. nov.</p></div>	https://treatment.plazi.org/id/C62094596522568D9C1F4BB86E22A615	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lee, Jimin;Kim, Il-Hoi	Lee, Jimin, Kim, Il-Hoi (2026): A new family of siphonostomatoid Copepoda (Pupiformidae fam. nov.) from the Korea Strait. ZooKeys 1284: 217-230, DOI: 10.3897/zookeys.1284.194182
81DBED1A73E1529A88968F9CEA372381.text	81DBED1A73E1529A88968F9CEA372381.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pupiformus apodus Lee & Kim 2026	<div><p>Pupiformus apodus sp. nov.</p><p>Figs 1, 2</p><p>Type locality.</p><p>Korea Strait, east of Jeju Island (33°30'55"N, 127°46'49"E), depth 116–123 m.</p><p>Type material.</p><p>Holotype (intact ♀, MABIK CR 00261359) and paratypes (6 intact ♀♀, MABIK CR 00261360; 2 dissected ♀♀) from washings of benthic animals dredged from the <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.78028&amp;materialsCitation.latitude=33.515278" title="Search Plazi for locations around (long 127.78028/lat 33.515278)">Korea Strait</a>, collected by J. Lee, 20 August 2025 . Intact type specimens have been deposited in the Marine Biodiversity Institute of Korea (MABIK), Seocheon, Korea. Dissected paratypes are kept in the collection of I. - H. Kim.</p><p>Etymology.</p><p>The name is derived from the Greek words a (= without) and pod (= a foot), referring to the absence of legs in the new species.</p><p>Description.</p><p>Female. Body (Fig. 1 A – C) pupiform, cylindrical, appearing immovable, 6 - segmented, consisting of cephalothorax, metasome, and genitoabdomen, with thick, hard integument. Body length 850–970 μm, based on 7 specimens. Body length of figured and described specimen 895 μm. Maximum width 367 μm across cephalothorax. Cephalothorax 324 μm long, wider than long. Cephalothoracic shield clearly divided into 3 parts (dorsal tergite and 2 lateral pleurites) by 2 longitudinal, unsclerotized grooves (Fig. 1 C, E). Metasome 4 - segmented, comprising second to fifth pedigerous somites; sizes of these somites 145 × 356, 116 × 360, 102 × 313, and 90 × 273 μm, respectively, from anterior to posterior. Fourth metasomite with tubercle on each dorsolateral side. Genitoabdomen (Fig. 2 B) with tubercle on each dorsolateral side (Fig. 2 D), distinct anal operculum (Fig. 2 B – D); genital apertures (Fig. 2 B) positioned at proximal region of ventral surface. Caudal rami (Fig. 2 C) as small rugose distal process on genitoabdomen; each caudal ramus with 3 indistinct, minute setae.</p><p>Rostrum (Figs 1 D, 1 E, 2 A) well developed, with bluntly tapering distal apex. Antennule (Fig. 2 E) short, 53 μm long, 4 - segmented; armature formula 1, 2, 6, and 7 + aesthetasc; three setae on first and second segments plumose, other setae naked; terminal segment consisting of broader proximal third and narrowed distal two-thirds, its aesthetasc pointed distally, as long as segment. Antenna (Fig. 2 F) stout, 4 - segmented, consisting of coxa, basis, and 2 - segmented endopod; first to third segments unarmed; last segment (second endopodal segment; Fig. 2 G) short, wider than long, with terminal claw bearing 3 teeth on concave margin, 1 blunt distal spine, and 3 setae (2 distal and 1 proximal).</p><p>Oral cone (Figs 1 D, 1 E, 2 H) short, but distinct, 102 × 75 μm in anterior view. Mandible (Fig. 2 I) consisting of 1 - segmented palp, 38 × 16 μm, bearing 2 thick, distal setae of 41 and 34 μm long, respectively, and 1 small subdistal, naked seta of 11 μm long; mandibular stylet absent. Maxillule (Fig. 2 J) 2 - segmented; proximal segment unarmed, not articulated from ventral surface of cephalothorax; distal segment 24 × 12 μm, bearing 3 setae (1 plumose proximal seta, and 1 plumose and 1 naked distal setae). Maxilla absent, but its vestige presents as sclerotized spot positioned lateral to base of maxilliped (Fig. 1 D). Maxilliped (Fig. 2 K) 3 - segmented, consisting of coxobasis and 2 - segmented endopod; second endopodal segment slender, 30 μm long, twice longer than first endopodal segment, distally bearing 1 small seta and 1 claw; latter claw 15 μm long, bearing 2 denticles subdistally.</p><p>Leg 1–4 vestigial, represented by large, sclerotized pads (Fig. 1 B, C). Legs 5 and 6 absent.</p><p>Male. Unknown.</p><p>Remarks.</p><p>The division of the cephalothoracic shield of P. apodus sp. nov. into the dorsal tergite and two lateral pleurites is clearly defined and consistent in all examined specimens. This characteristic also appears in P. rugosus sp. nov. described below.</p></div>	https://treatment.plazi.org/id/81DBED1A73E1529A88968F9CEA372381	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lee, Jimin;Kim, Il-Hoi	Lee, Jimin, Kim, Il-Hoi (2026): A new family of siphonostomatoid Copepoda (Pupiformidae fam. nov.) from the Korea Strait. ZooKeys 1284: 217-230, DOI: 10.3897/zookeys.1284.194182
6BC5E53C6B1E55B0B0C698B7D7A9274A.text	6BC5E53C6B1E55B0B0C698B7D7A9274A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pupiformus Lee & Kim 2026	<div><p>Pupiformus gen. nov.</p><p>Diagnosis (female).</p><p>Body pupiform, cylindrical, 6 - segmented, consisting of cephalothorax, 4 - segmented metasome, and genitoabdomen. Cephalothoracic shield clearly divided into dorsal tergite and 2 lateral pleurites. Metasome comprising second to fifth pedigerous somites. Genitoabdomen lacking any trace of segmentation; genital apertures positioned ventrally. Caudal rami absent or rudimentary. Rostrum developed. Antennule short, 4 - or 5 - segmented; terminal segment bearing 1 aesthetasc on proximal anterior margin. Antenna 4 - segmented, consisting of coxa, basis, and 2 - segmented endopod; exopod absent; second endopodal segment bearing small claw distally. Oral cone short, but distinct, comprising anterior and posterior lips (labrum and labium). Mandible represented by 1 - segmented palp; stylet absent. Maxillule 2 - segmented; proximal segment (precoxa or pedetal) unarmed; distal segment (palp or fusion of inner and outer lobes) with 3 setae (2 apical and 1 proximal). Maxilla absent. Maxilliped 3 - or 4 - segmented, consisting of coxa, basis, and 2 - segmented endopod, or coxobasis and 2 - segmented endopod; second endopodal segment bearing claw at tip. Legs 1–4 vestigial, represented by large sclerotized pads. Legs 5 and 6 absent.</p><p>Etymology.</p><p>The generic name refers to the pupiform body shape of the species within the genus. Gender masculine.</p><p>Type species.</p><p>Pupiformus apodus sp. nov. (original designation).</p><p>Other included species.</p><p>Pupiformus rugosus sp. nov.</p></div>	https://treatment.plazi.org/id/6BC5E53C6B1E55B0B0C698B7D7A9274A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lee, Jimin;Kim, Il-Hoi	Lee, Jimin, Kim, Il-Hoi (2026): A new family of siphonostomatoid Copepoda (Pupiformidae fam. nov.) from the Korea Strait. ZooKeys 1284: 217-230, DOI: 10.3897/zookeys.1284.194182
8EB438B2C52D5EBF8B023E5B7F3835A1.text	8EB438B2C52D5EBF8B023E5B7F3835A1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pupiformus rugosus Lee & Kim 2026	<div><p>Pupiformus rugosus sp. nov.</p><p>Figs 3, 4, 5</p><p>Type locality.</p><p>Korea Strait, east of Jeju Island (33°30'55"N, 127°46'49"E), depth 116–123 m.</p><p>Type material.</p><p>Holotype (intact ♀, MABIK CR 00261361) and paratypes (11 intact ♀♀, MABIK CR 00261362; 2 dissected ♀♀) collected with Pupiformus apodus sp. nov. from washings of benthic animals dredged from the same locality as the former species . Intact type specimens have been deposited in the Marine Biodiversity Institute of Korea (MABIK), Seocheon, Korea. Dissected paratypes are kept in the collection of I. - H. Kim.</p><p>Etymology.</p><p>The name is an adjective (Latin: rugosus) and refers to the rugose dorsal tubercles of the new species.</p><p>Description.</p><p>Female. Body (Figs 3 A, 3 B, 5) shaped and segmented as in P. apodus gen. et sp. nov. Body length of figured and described specimen 501 μm. Mean body length 502 μm, based on 7 specimens. Maximum width 242 μm. Cephalothorax 180 × 242 μm, gradually broadened posteriorly, consisting of clearly divided dorsal tergite and lateral pleurites (Fig. 3 C, D); tergite bearing 5 tubercles (arranged as 2, 1, and 2) on anterior half and large posterolateral tubercles each bearing about 10 irregularly arranged, wart-like processes (Fig. 3 A, C). Ventral surface of cephalothorax bearing 3 pairs of sclerotized areas: one on lateral side of maxillule, another between oral cone and maxilliped, and the other lateral to maxilliped. Pleurites with uneven lateral and ventral margins (Fig. 3 A, C). Metasomites each with pair of tubercles at dorsal or dorsolateral tubercles bearing about 10 or more globular processes. Fourth metasomite with sclerotized region on both sides of ventral surface (Fig. 3 B). Genitoabdomen 105 × 160 μm in ventral view, with several processes on each side of posterior margin, small, excavated anal region bearing pair of small lobes (Fig. 3 A, B); genital apertures positioned proximal region of ventral surface; anal operculum absent. Caudal rami absent.</p><p>Rostrum (Fig. 3 B) roughly trapezoid in anterior view (Fig. 3 C), well separated from tergite. Antennule (Fig. 4 A) small, 42 μm long, 5 - segmented; armature formula 1, 5, 3, 3, and 10 + aesthetasc; seta of first segment and first seta of second segment plumose, other setae naked; aesthetasc on last segment extending slightly over distal end of terminal segment; terminal segment bearing produced, nipple-like anterodistal corner. Antenna (Fig. 4 B) 4 - segmented, consisting of coxa, basis, and 2 - segmented endopod; lengths of segments 11, 33, 30, and 7 μm, respectively, from proximal to distal; terminal segment (second endopodal segment) short, bearing 1 claw bearing 3 denticles, 1 small spine, and 2 small setae.</p><p>Oral cone (Fig. 3 C) distinct, bearing membranous apex. Mandible (Fig. 4 C) represented by 1 - segmented palp bearing 2 naked distal setae (58 and 48 μm long, respectively). Maxillule (Fig. 4 D) 2 - segmented; proximal segment unarmed; distal segment with 3 setae (plumose proximal seta and 2 naked, unequal distal setae). Maxilla (Fig. 3 B) absent. Maxilliped (Fig. 4 E) 4 - segmented; proximal 3 segments (syncoxa, basis, and first endopodal segments) unarmed; basis and 2 endopodal segments 42, 12, and 26 μm long, respectively; slender second endopodal segment distally bearing small claw (10 μm long) and 1 small seta.</p><p>Legs 1–4 vestigial, represented by large, sclerotized pads (Fig. 3 B); each pad bearing 1–3 hair-like setules on its lateral and posterior margins. Legs 5 and 6 absent.</p><p>Male. Unknown.</p><p>Remarks.</p><p>Pupiformus rugosus sp. nov. and P. apodus sp. nov. are very similar, sharing key diagnostic features of the genus. However, P. rugosus sp. nov. differs significantly from P. apodus sp. nov. in several female morphological details: (1) the body is smaller, about 500 μm long, compared to roughly 900 μm in P. apodus sp. nov.; (2) the cephalothorax and metasomites have rugose dorsal tubercles, whereas in P. apodus sp. nov., these structures are either absent or smooth; (3) caudal rami are absent in P. rugosus sp. nov., but are present as rudimentary processes in P. apodus sp. nov.; (4) the antennule has five segments, vs four in P. apodus sp. nov.; (5) the mandibular palp has two long, naked setae at its tip, whereas P. apodus sp. nov. has two stout, spiniform distal setae and one small, naked, subdistal seta; and (6) the maxilliped is 4 - segmented, vs three segments in P. apodus sp. nov.</p></div>	https://treatment.plazi.org/id/8EB438B2C52D5EBF8B023E5B7F3835A1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lee, Jimin;Kim, Il-Hoi	Lee, Jimin, Kim, Il-Hoi (2026): A new family of siphonostomatoid Copepoda (Pupiformidae fam. nov.) from the Korea Strait. ZooKeys 1284: 217-230, DOI: 10.3897/zookeys.1284.194182
