taxonID	type	description	language	source
8625131CFFB2FFA48DF362E3B5C89A91.taxon	description	(Figs 1 – 11)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFB2FFA48DF362E3B5C89A91.taxon	diagnosis	Diagnosis. Male. Head. Median clypeal lobe trapezoidal or ill-defined (Fig. 1 A, B); mandible with two to three apical teeth; Metasoma. Tergal process present as a pair of median subnavel-shaped tubercles mesad, located on each side of an elliptical or subcircular depression, the surrounding area may be setose; hypopygium with wide or narrow spiculum, posterior margin straight or medially incurved (Fig. 2 A, B). Genitalia. Harpe with apical margin wide and mesad (Fig. 3 GH) or narrow and slightly posterad (Fig. 3 A, B), outer margin rounded or angulated, subapical margin angled or rounded; outer lobe of the dorsal aedeagal valve usually strongly ornamented; basal half of aedeagal ventral valve can be ornamented by a basal process and apical half ventral margin with teeth ventrad; basal cover plate subtrapezoidal, however can be absent in some species. Female. Unknown.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFB2FFA48DF362E3B5C89A91.taxon	description	Description. Male. Head. Median clypeal lobe trapezoidal or ill-defined (Fig. 1 A, B), with one angulated or rounded median tooth. Median clypeal carina simple, complete or incomplete, with only exception in D. bicarinatus, which is double. Mandible with two to three apical teeth. Metasoma. Metasomal tergal process as a pair of median subnavel-shaped tubercles, located on each side of elliptical or subcircular depression, located on median third of tergal width (Fig. 1 C), tubercles strongly or weakly mesad, often accompanied by a strongly deep fovea in former case and a shallower in latter, both biconcave in dorsal view, surrounding area of depression often setose, with one tuft of setae on each side. Hypopygium with wide or narrow spiculum, posterior margin straight or medially incurved (Fig. 2 A, B). Genitalia. Some distinct patterns can be found in several species (Fig. 2 C, D) related to the previous definitions of stellatus and vallensis species-groups sensu Azevedo (1999 a). Harpe with apical margin wide and mesad (Fig. 3 GH) or narrow and slightly posterad (Fig. 3 A, B), outer margin rounded or angulated, subapical margin angled or rounded. Cuspis simple, rounded and without serration. Digitus usually short, about 2 × as long as wide, sometimes distinctly elongated, about 4 × as long as wide (Fig. 3 E, F). Ventral aedeagal valve can have serrated or spinose areas. Many species have a basal ventral process, taking the shape of filamentous (sinuous or rounded) (Figs 3 G, 6 M), lamellar (serrated or smooth) (Figs 4 A, 5 M) or denticulate projection (multidentate or unidentate) (Figs 3 E, 9 L), usually in the most basal or median portion of the ventral aedeagal valve, with the exception of D. torem, which is in the entire basal portion (Fig. 10). Dorsal aedeagal valve much more ornamented, its outer lobe with serrated, filamentous and spinose projections, its inner lobe membranous and slightly setose, rarely with projections, with D. eliasi sp. nov. and D. mortarioni sp. nov. being exceptions (Figs 5 H, 7 L), dorsal margin of both outer or inner lobes usually with arched or straight anterad tooth in its basal half; aedeagal dorsal valve apex usually narrow and long, except in D. basidentatus and D. cerutus (Figs 5 J, 6 B), with gibbous and short valve. Basal cover plate subtrapezoidal, posterior margin usually strongly incurved medially, anterior margin outcurved medially, basal cover plate absent in some species (Figs 5 B, 9 F). Apodema usually does not extend anterior to basal plate. Female. Unknown. Species included. D. basidentatus Azevedo, 1999; D. bicarinatus Azevedo, 1999; D. bifurcatus Azevedo, 1999; D. bisserratus Azevedo, 1999; D. cerutus Azevedo, 1999; D. digitatus Azevedo, 1999; D. dilatatus Azevedo, 1999; D. extrarramis Azevedo, 1999; D. largidentatus Azevedo, 1999; D. planus Azevedo, 1999; D. polidentatus Azevedo, 1999; D. rosangelae Colombo & Azevedo, 2016; D. rotundus Azevedo, 1999; D. scavatus Azevedo, 1999; D. stellatus Azevedo, 1999; D. strictus Azevedo, 1999; D. torem Colombo & Azevedo, 2018; D. tortuosus Azevedo, 1999; D. triangularis Azevedo, 1999, and D. vallensis Evans, 1979.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFB2FFA48DF362E3B5C89A91.taxon	distribution	Distribution. Neotropical region: Argentina (recorded for the first time); Bolivia (new records); Brazil (several states with new records); Colombia; Costa Rica; Dominican Republic; Ecuador (new records); French Guiana (recorded for the first time); Mexico; Panama; Paraguay; Peru (new records); Suriname (new records); Trinidad & Tobago; Venezuela.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFB6FFA58DF36020B2CF9CC7.taxon	description	(Fig. 3 A – D) Description. Male. Head. Mandible with three distal teeth; median clypeal lobe ill-defined, with one angulate tooth, median clypeal carina high and straight in profile or nearly so, complete or nearly so; frons smooth, punctures small, and sparse; vertex crest straight or nearly so in dorsal view. Mesosoma. Dorsal pronotal area with anterior margin coriaceous. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad; depression deep with surrounding area glabrous; posterior hypopygeal margin straight or nearly so. Genitalia. Harpe with apical margin narrow and slightly posterad, outer margin rounded, inner margin angled subapically; apex of digitus short, about 2 × as long as wide; ventral aedeagal valve with apex anterior to apex of dorsal aedeagal valve, wide, progressively narrowing apicad, outer margin outcurved, not diverging apicad, apex acute and not bifid, inner margin incurved, with distinct concavity, outer margin smooth, ventral margin smooth and without teeth, basal ventral process absent; dorsal aedeagal valve narrow and long with two pairs of apical lobes, outer lobe with outer margin smooth, inner margin with teeth on apical portion in dorsal view, apical portion with two pairs of teeth, and one smooth ramified filament, ventro-posterad, basal portion ventral margin smooth, dorsal margin with a straight tooth anterad, inner lobe membranous, gibbous, rounded, larger than outer lobe and its ventral area not covered by outer lobe when seen in lateral view, dorsal margin smooth, ventral margin smooth; basal cover plate, trapezoidal, and not divided, posterior margin strongly incurved medially, anterior margin slightly outcurved medially; apodema without apical membrane, not extending anterior to basal plate. Female. Unknown.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFB6FFA58DF36020B2CF9CC7.taxon	diagnosis	Differential diagnosis. This species is similar to D. lorgari sp. nov. by having the ventral aedeagal valves wide, narrowing progressively apicad, outer margin entirely rounded, inner apical margin with concavity, and the outer lobe of the dorsal aedeagal valves with filament ventro-posterad. However, D. angroni sp. nov. has two pairs of apical teeth in the outer lobe of the dorsal aedeagal valves, along with another pair of apical teeth on the inner margin of the outer lobe apical portion in dorsal view, whereas D. lorgari sp. nov. has no teeth on the outer lobe of the dorsal aedeagal valves, instead having a ventro-posterad filament.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFB6FFA58DF36020B2CF9CC7.taxon	materials_examined	Material examined. Holotype, ♂, BRAZIL, Acre, Bujari, FES, Antimary, 09 º 20 ’ 01 ’’ S 68 º 19 ’ 17 ’’ W, 18 – 31. xii. 2017, arm. Malaise grande, Z. F. M. Silva & J. A. Rafael col. (INPA). Paratypes: BRAZIL, Acre, 1 ♂, Bujari, FES, Antimary, 09 º 20 ’ 01 ’’ S 68 º 19 ’ 17 ’’ W, 22. ix – 06. x. 2016, arm. Malaise grande, Z. F. M. Silva & J. A. Rafael col. (INPA *); Rondônia, 1 ♂, Guajará Mirim, Rio Pacaás Novo, 11 º 11 ’ 13 ’’ S 64 º 51 ’ 21 ’’ W, 07 – 11. IX. 1999, J. F. Vidal col. (INPA); 1 ♂, Itapuã do Oeste, FLONA Jamari, 09 º 15 ’ 36 ’’ S 62 º 54 ’ 96 ’’ W, iii. 2017, arm. Malaise, Z. F. M. Silva & J. A. Rafael col. (INPA); 1 ♂, Porto Velho, ESEC Morro Três Irmãos, v. 2017, 09 º 00 ’ 09 ’’ S 64 º 32 ’ 40 ’’ W, arm. Malaise, V. S. Silva & J. A. Rafael col. (INPA).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFB6FFA58DF36020B2CF9CC7.taxon	etymology	Etymology. The specific epithet refers to the Primarch Angron, from the Warhammer 40 k media.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFB6FFA58DF36020B2CF9CC7.taxon	distribution	Distribution. Brazil: Acre and Rondônia.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFB7FFA58DF36593B37599F1.taxon	description	(Fig. 3 E, F)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFB7FFA58DF36593B37599F1.taxon	diagnosis	Diagnosis. Male. Head. Mandible with two distal teeth; clypeus with one angulate median tooth, median carina incomplete apically. Metassoma. Tergal process with tubercles fully mesad, depression with surrounding area setose; posterior hypopygeal margin slightly incurved medially. Genitalia. Harpe wide, apical margin narrow, inner margin angled subapically, outer margin entirely rounded; apex of digitus elongated, about 4 × as long as wide; basivolsela ventral margin sinuous; ventral aedeagal valve with apex aligned with apex of dorsal aedeagal valve, wide, progressively narrowing apicad, apex acute and not bifid, basal ventral process dentate, with three distinct teeth; dorsal aedeagal valve short and gibbous. Female. Unknown.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFB7FFA58DF36593B37599F1.taxon	materials_examined	Material revised. Holotype, ♂, BRAZIL, Pará, Tucuruí, Rio Tocantins, interception trap, 7. vii. 1986, N. Degalier col. (MPEG). Paratype: BRAZIL, Pará, 1 ♂, Tucuruí, Rio Tocantins, Arm. Inter. 13 a 21. III. 1985, Pará, N. Degallier (UFES nº 12111). New material. ECUADOR, Otongachi, 1 ♂, La Unión del Toachi, V. 2003, leg Giovanni Onore, Museo Genova, coll. G. Pagliano, Acquisto 2012 (MCSN); BRAZIL, Pará, 1 ♂, Juruti, Mauari, [02 ° 09 ’ 12 ” S 56 ° 05 ’ 14 ” W], 01. XII. 2008, arm. Malaise, O. T. Silveira & equipe col. (MPEG).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFB7FFA58DF36593B37599F1.taxon	distribution	Distribution. This species was known from Brazil: Amapá, Pará (Azevedo 1999 a) and Mato Grosso (Azevedo 2003). Now, it is recorded for the first time from Ecuador.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFB7FFAB8DF36080B2E89DE7.taxon	description	(Fig. 3 G – J)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFB7FFAB8DF36080B2E89DE7.taxon	diagnosis	Diagnosis. Male. Head. Mandible with three distal teeth; clypeus with two median carina. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad, depression with surrounding area setose; posterior hypopygeal margin straight or nearly so. Genitalia. Harpe with apical margin wide and mesad, outer margin rounded, inner margin rounded subapically; apex of digitus short, about 2 × as long as wide; basivolsela ventral margin rounded; ventral aedeagal valve with apex posterior to apex of dorsal aedeagal valve, entirely narrow, outer margin sinuous, diverging apicad, apex acute and not bifid, inner margin smooth, outer margin smooth, ventral margin smooth and with two pairs of teeth, basal ventral process filamentous, and sinuous, originating laterally; aedeagal dorsal valve narrow and long, with two pairs of apical lobes, outer lobe, with outer margin smooth, inner margin smooth in dorsal view, apical portion with one pair of teeth, dorsad, basal portion ventral margin smooth, dorsal margin smooth, inner lobe membranous and gibbous, dorsal margin with arched tooth anterad, ventral margin smooth; basal cover plate, trapezoidal, anterior margin strongly incurved medially, posterior margin strongly incurved medially, and not divided; apodema extending anterior to basal plate. Female. Unknown.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFB7FFAB8DF36080B2E89DE7.taxon	materials_examined	Material revised. Holotype, ♂, BRAZIL, Pará, Tucuruí, Rio Tocantins, right margin, 26. iii. 1984 (MPEG).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFB7FFAB8DF36080B2E89DE7.taxon	distribution	Distribution. Brazil: Pará (Azevedo 1999 a).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFB9FFA98DF364B3B1589DAF.taxon	description	(Fig. 3 K – M)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFB9FFA98DF364B3B1589DAF.taxon	diagnosis	Diagnosis. Male. Head. Mandible with three distal teeth; clypeus with one median carina. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad, depression with surrounding area setose; posterior hypopygeal margin straight or nearly so. Genitalia. Harpe with apical margin narrow and slightly posterad, outer margin rounded, inner margin rounded subapically; apex of digitus short, about 2 × as long as wide; basivolsela ventral margin sinuous; ventral aedeagal valve with apex aligned with apex of dorsal aedeagal valve, entirely narrow, outer margin sinuous, not diverging apicad, apex acute and not bifid, inner margin smooth, outer margin smooth, ventral margin without teeth, basal ventral process absent; aedeagal dorsal valve narrow and long, with two pairs of apical lobes, outer lobe medially divided longitudinally, with outer margin smooth, inner margin smooth in dorsal view, apical portion with two pairs of teeth, ventrad and overlapping each other, basal portion ventral margin smooth, dorsal margin smooth, inner lobe membranous and gibbous, dorsal margin smooth, ventral margin smooth; basal cover plate, trapezoidal, posterior margin strongly incurved medially; apodema not extending anterior to basal plate. Female. Head. Head length 1.36 × its width; punctures large and dense; mandible with four distal teeth; first four antennal segments in a ratio of 16: 2: 1: 2, eleventh segment length 2.1 × its width; eye subcircular, with 7 facets distant from the mandible by 1.4 × its length; clypeal median lobe trapezoidal, clypeal carina high in profile; vertex crest straight; occipital carina length 0.43 × that of the head, visible in dorsal view. Mesosoma. Coriaceus; dorsum with small, deep and dense punctures; pronotal collar microstriate transversally, length 0.38 × that of the dorsal pronotal area; dorsal pronotal area length 1.14 × its width; mesonotum length 0.38 × that of the dorsal pronotal area, posterior margin convex and slightly embraced by the metapectal-propodeal complex; propodeal declivity weakly coriaceus and microstriate transversally; legs weakly coriaceus, midtibia without spurs, forefemur length 2.13 × its width; pleurosternum posterior margin slightly produced anterad medially; metasternum elongated with lateral margins elevated. Metasoma. Petiole long and coriaceus; segments nearly polished, first one with transverse microstriae, tranversal section subcircular.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFB9FFA98DF364B3B1589DAF.taxon	materials_examined	Material revised. Holotype, ♂, BRAZIL, Pará, Tucuruí, Rio Tocantins, base 4, 17. xii. 1986, interception trap, N. Degallier col. (MPEG). Paratypes: BRAZIL, Pará, Tucuruí, 1 ♂, Base 4, 17. VI. 1986, Interception, N. Degallier col. (UFES nº 12157); 1 ♂, 22. II. 1986, Interception, N. Degallier col. (UFES nº 12155); 4 ♂, Interception, 9. XI. 1985, N. Degallier col. (UFES nº 12151 – 12154); 1 ♂, Rio Tocantins, V. 1986, interception, N. Degallier col. (UFES nº 12160); 1 ♂, Fruta, Carniça, Fezes, 15. X. 1987, N. Degallier col. (UFES nº 12158). Vouchers: BRAZIL, Alagoas, 1 ♂, Quebrângulo, Res. Biol. Pedra Talhada, 08 – 11. IX. 2002 – Malaise B 2, Penteado-Dias & equipe col., BIOTA-FAPESP (UFES nº 12168); 1 ♂, Malaise B 3, Penteado-Dias & equipe col. (UFES nº 12169); 1 ♂, Malaise B 4, Penteado-Dias & equipe col. (UFES nº 12167); 1 ♂, Molecular B nº 00342 (UFES nº 12166); 2 ♂, Malaise T 2, Penteado-Dias & equipe col. (UFES nº 12164, 12165); 1 ♂, Malaise T 3, Penteado-Dias & equipe col. (UFES nº 12171); 1 ♂, Molecular B nº 00455 (UFES nº 12172); 1 ♂, Malaise T 4, Penteado-Dias & equipe col., Molecular B nº 00456 (UFES nº 12170); 1 ♂, PB, João Pessoa, Mata do Buraquinho, 07 º 08 ’ 25 ’’ S 34 º 51 ’ 38 ’’ W, arm. Malaise T 5, 28 – 31. VII. 2002, STP. Amarante & eq. col., BIOTA-FAPESP (UFES nº 12163); 1 ♂, SE, Sta Luzia do Itanhy, Crasto, 11 º 22 ’ 37,4 ’’ S 37 º 24 ’ 50,3 ’’ W, 01 – 04. VIII. 2001, Malaise B 2, M. T. Tavares & Eq. col., BIOTA-FAPESP (UFES nº 12174); 1 ♂, Malaise T 1, M. T. Tavares & Eq. col., BIOTA-FAPESP (UFES nº 12173); 1 ♂, Molecular DOC 166, Molecular B nº 00166 (UFES nº 12178); 1 ♂, Malaise T 3, M. T. Tavares & Eq. col., BIOTA-FAPESP (UFES nº 12179); 1 ♂, Malaise T 4, M. T. Tavares & Eq. col., BIOTA-FAPESP (UFES nº 12177); 1 ♂, 1. VIII. 2001, varredura 23, M. T. Tavares & Eq. col., BIOTA-FAPESP (UFES nº 12175); 1 ♂, 11 º 22 ’ 37,4 ’’ S 7 º 24 ’ 50,3 ’’ W, 29. VII – 01. VIII. 2001, Malaise B 2, M. T. Tavares & Eq. col., BIOTA-FAPESP (UFES nº 12176). New material. BOLIVIA, Cochabamba, 8 ♂, 67.5 km E Villa Tunari, Est. Biol. Valle del Saita, 300 m, 17 º 6 ’ 19 ’’ S 64 ° 46 ’ 57 ’’ W, 7 – 9. II. 1999, FIT, rain forest, F. Génier col. (CNCI *); 2 ♂, 9 – 13. II. 1999, low land rain forest, FIT, F. Génier col. (CNCI); 4 ♂, Univ. San Simón, 300 m, 17 º 6 ’ 19 ’’ S 64 º 46 ’ 67 ’’ W, 9 – 13. II. 1999, FIT, low land rain for., F. Génier col. (CNCI); 1 ♂, 117 Km E Cochabamba on, Villa Tunari road, 1040 m, 17 º 6.53 ’ S 65 º 41.20 ’ W, FIT, 1 – 6. II. 1999, R. S. Hanley col. (CNCI); 1 ♂, Est. Biol. Valle del Saita, Univ. San Simón, 67.5 km, NE Villa Tunari, 300 m, 17 º 6.52 ’ S 64 º 47.87 ’ W — FIT, 9 – 13. II. 1999, R. S. Hanley col. (CNCI); 1 ♂, 17 ° 6 ’ 5.25 ” S 64 ° 47 ’ 8.72 ” W, 9 – 13. II. 1999, FIT # 077, R. S. Hanley col. (CNCI); 1 ♂, Est. Biol. Villa Tunari, 17 ° 6.52 ’ S 64 ° 47.87 ’ W, 5 – 7. II. 1999, FIT, R. S. Hanley col. (CNCI); 1 ♂ Villa Tunari, 16 ° 54 ’ 55 ” S 65 ° 22 ’ 06 ” W, 6. X. 2001, trop. for., MT, H. Heider (CNCI); 14 ♂, La Paz, Noryungas, Coroico, El Bagante, 1500 m, [16 ° 11 ’ S 67 ° 44 ’ W], 18. IV. 1997, cloud forest, B – 03, L. Masner s. s col. (CNCI *); BRAZIL, Acre, 2 ♂, Bujari, FES, Antimary, 09 º 20 ’ 01 ’’ S 68 º 19 ’ 17 ’’ W, 18 – 31. xii. 2017, arm. Malaise grande, J. A. Rafael & E. F. Morato col. (INPA); 2 ♂, 19. xi – 3. xii. 2016, arm., E. F. Morato & J. A. Rafael col., Rede BIA (INPA); 2 ♂, 22. ix – 06. x. 2016, arm., Z. F. M. Silva & J. A. Rafael col. (INPA); 2 ♂, Manoel Urbano, RESEX, Cazumbá – Iracema, Santo Antônio, 9 º 08 ’ 31.1 ’’ S 68 º 56 ’ 44.0 ’’ W, iii. 2018, arm. Malaise, E. F. Morato col. (INPA); Manoel Urbano, RESEX, Castanheira, 9 º 08 ’ 31.1 ’’ S 68 º 56 ’ 44.0 ’’ W, 5 ♂ xi. 2017, 2 ♂ iv. 2018, arm. Malaise, E. F. Morato col. (INPA *); 1 ♂, Porto Acre, Res. Humaitá, mata primária, S 9 ° 45 [‘ S] 6 w 7 ° 40 ’ [W] (2000 ha), 10 – 11. IV. 1998 — Arm. Malaise, E. F. Morato col. (UFAC); 3 ♂, Sen. Guiomard, FES Catuaba, 10 º 04 ’ 28 ’’ S 67 º 37 ’ 00 ’’ W, 14 – 31. i. 2017, arm. Malaise, E. F. Morato & J. A. Rafael col. (INPA *); Amazonas, 1 ♂, Ipixuna, Rio Gregório, Com. Lago grande, 07 º 10 ’ 11.7 ’’ S 70 º 49 ’ 10.3 ’’ W, 22. V. 2011, 04: 00 – 06: 30 h, arm luminosa, dossel, J. A. Rafael, J. T. Câmara, R. F. Silva, A. Somavilla, C. Gonçalves, A. Agudelo, leg. (INPA); 1 ♂ Novo Airão, PARNA Jaú, Rio Papagaio, Igarapé, terra firme, mata, 02 ° 00 ’ 03 ” S 62 ° 43 ’ 54 ” W, VI. 2001, arm. Malaise, A. L. Henriques & equipe col. (INPA); 1 ♂, Novo Aripuanã, Igarapé Arauazinho, 06 ° 17 ’ 44 ” S 60 ° 21 ’ 30 ” W, IV. 2005, arm. Malaise, F. F. Xavier, F. Godoi & eq. col. (INPA); Bahia, 1 ♂, Barra do Rocha, Faz. Iacina, 14 º 11 ’ S 39 º 34 ’ W, 19. XI. 2002, arm. Malaise (CPDC); Buerarema, Faz. Sempre Viva, 15 º 02 ’ S 39 º 14 ’ W, 2 ♂ 25. VIII. 2002, 5 ♂ 28. XI. 2002, arm. Malaise (CPDC *); 2 ♂, Firmino Alves, Faz. Bela Vista, 14 º 59 ’ S 39 º 55 ’ W, 09. IV. 2003, arm. Malaise (CPDC); 1 ♂, Itacaré, Faz. Muchirão, 14 º 20 ’ S 39 º 18 ’ W, 17. VIII. 2002, arm. Malaise (CPDC); 3 ♂, Itajuípe, Faz. Trindade, [14 ° 40 ’ S 39 ° 22 ’ W], 27. XI. 2002, arm. Malaise (CPDC *); 1 ♂, Jitaúna, Faz. Boa Paz, [14 ° 01 ’ S 39 ° 53 ’ W], 15. VIII. 2002, arm. Malaise (CPDC); 1 ♂, Santa Luzia, Faz. Conceição, 15 º 20 ’ S 39 º 26 ’ W, 14. III. 2001, arm. Malaise, J. A. Cardoso & J. R. Maia (CPDC); S. J. da Vitória, Faz. São Roque, 15 º 04 ’ S 39 º 18 ’ W, 2 ♂ 12. I. 2001, 1 ♂ 12. III. 2001; arm. Malaise, J. A. Cardoso & J. R. Maia (CPDC); 1 ♂, Ubaitaba, Faz. Casarão de Pedra, 14 º 18 ’ S 39 º 19 ’ W, 16. VIII. 2002, arm. Malaise (CPDC); 1 ♂, Una, Faz. Piruma, cabruca, 15 º 15 ’ S 39 º 07 ’ W, 08. III. 2001, arm. Malaise, J. A. Cardoso & J. R. Maia (CPDC); Goiás, 1 ♂, Alto Paraíso, Pq. Nac. Chapada dos Veadeiros, 14 º 00 ’ S 47 º 41 ’ W, 15 – 25. IX. 2005, arm. Malaise 20, Aguiar & eq. col. (MZSP); Mato Grosso, 1 ♂, Sinop, 12 º 31 ’ S 55 º 37 ’ W, 350 m, X. 1974, M. Alvarenga (CNCI); 1 ♂, Sinop, [11 ° 52 ’ S 55 ° 30 ’ W], II. 1976, M. T., O. Roppa (CNCI); Minas Gerais {= Mato Grosso}, 1 ♂, Sinop, [11 ° 52 ’ S 55 ° 30 ’ W], XI – 1975, M. Alvarenga (PMAE); Pará, 2 ♂, Juruti, Estrada Mutum, [02 ° 09 ’ 12 ” S 56 ° 05 ’ 14 ” W], 01. XII. 2008, arm. Malaise, O. T. Silveira & equipe col. (MPEG *); 1 ♂, Juruti, Estrada Mutum, 28. II – 04. III. 2008, arm. Malaise, O. T. Silveira & equipe col. (MPEG); 1 ♂, Juruti, Estrada Pacoval, 28. II – 04. III. 2008, arm. Malaise, O. T. Silveira & equipe col. (MPEG); 1 ♂, Juruti, Galileia, 10. V. 2009, arm. Malaise, O. T. Silveira & equipe col. (MPEG); 1 ♂, Juruti, Galileia, Ramal Mauari, Galileia, 26 – 29. V. 2008, arm. Malaise, J. N. Santos & L. A. Quaresma col. (MPEG); 1 ♂, Juruti, Mineração Alcoa, Capiranga, 03. XI. 2007, arm. Malaise, O. T. Silveira & equipe col. (MPEG); 1 ♂, Juruti, Mineração Alcoa, Capiranga, 30. V – 03. VI. 2008, arm. Malaise, J. N. Santos & L. A. Quaresma col. (MPEG); Melgaço, Floresta Nacional Caxiuanã, Trilha Est. Cie. Ferreira Pena, [01 ° 42 ’ 30 ” S 51 ° 31 ’ 45 ” W], 1 ♂, 14. XI – 19. XI. 2003, Arm. Malaise 2, A. P. Aguiar & J. D. Dias, Ponto P 05058 (MPEG); 2 ♂, 16. XI – 22. XI. 2003, arm. Malaise 3, A. P. Aguiar & J. Dias, Ponto P 05105 a (MPEG); 2 ♂, 19. XI – 25. XI. 2003, arm. Malaise 2, A. P. Aguiar & J. Dias, Ponto P 05141 (MPEG *); 1 ♂, 19. XI – 25. XI. 2003, arm. Malaise 1, A. P. Aguiar & J. Dias, Ponto P 05140 (MPEG); 1 ♂, 22. XI – 25. XI. 2003, arm. Malaise 19, A. P Aguiar & J. Dias, Ponto P 05146 (MPEG); 1 ♂, 22. XI – 25. XI. 2003, arm. Moerike, A. P Aguiar & J. Dias, Ponto P 05151 (MPEG); 1 ♂, Trilha Igarapé Curua, 23. XI – 26. XI. 2003, arm. Malaise 8, A. P Aguiar & J. Dias (MPEG); 1 ♂, Óbidos, Fazenda Pajurá, 01 º 37 ’ 21 ” S, 55 º 23 ’ 14 ” W, 05 – 11. IX. 2001, arm. Malaise, J. A. Rafael & J. F. Vidal col. (INPA); Rondônia, 1 ♂, Candeias do Jamari, Batalhão da Polícia Ambiental, BR 364, Km 2, 102 m, 08 º 47 ’ S 63 º 42 ’ W, 28. I – 04. II. 2006, Malaise, D. F. Mugrabi col. (UFES nº 126890); 1 ♂, Fazenda Rancho Grande, 62 km s. Ariquemes, 10 ’ 18 ’ S 62 ’ 53 ’’ W, 8 – 20. IV. 1997, 275 m, A. Rehn & C. Alexander, hilltop (UCDC); FRENCH GUIANA, Saül, 1 ♂, 7 km N 63 km SE of Eaux Claires, Mt. Fuméc, 490 m, [03 ° 36 ’ N 53 ° 35 ’ W], 1 – 8. VI. 1997, FIT, J. Ashe, R. Brooks (CNCI); PERU, Madre de Dios, 4 ♂, Cocha Salvador, 310 m, [11 ° 59 ’ 53.2 ” S 71 ° 13 ’ 50.1 ” W], 20 – 21. X. 2000, FIT, R. Brooks (CNCI *); 1 ♂, Pantiocolla Lodge, 400 m, [12 ° 39 ’ S 71 ° 13 ’ W], 25 – 26. X. 2000, FIT, R. Brooks (CNCI); SURINAME, Marowijne, 1 ♂, Nassau Montain, 500 m, 4 ° 4836 ” ’ N 54 ° 31 ’ 16 ” W, 2 – 4. VI. 1999, FIT, Z. H. Falin & B. DeDijn (CNCI).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFB9FFA98DF364B3B1589DAF.taxon	distribution	Distribution. This species was known from Brazil: Pará (Azevedo 1999 a), Alagoas, Paraíba, Sergipe (Redighieri & Azevedo 2006), Piauí and Maranhão (Colombo et al. 2018). Now, it is recorded for the first time from Bolivia, French Guiana, Peru, Suriname, and the Brazilian states of Acre, Amazonas, Bahia, Goiás, Mato Grosso, and Rondônia.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFBBFFAF8DF3657BB1949D03.taxon	description	(Fig. 4 A – D)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFBBFFAF8DF3657BB1949D03.taxon	diagnosis	Diagnosis. Male. Head. Mandible with two distal teeth; clypeus with one median carina. Metasoma. Metasomal segment II with median tergal process tubercles strongly mesad, depression with surrounding area setose; posterior hypopygeal margin slightly incurved medially. Genitalia. Harpe with apical margin narrow and slightly posterad, outer margin rounded, inner margin angled subapically; apex of digitus short, about 2 × as long as wide; basivolsela ventral margin rounded; ventral aedeagal valve with apex aligned with apex of dorsal aedeagal valve, entirely narrow, outer margin straight or nearly so, not diverging apicad, apex acute, inner margin smooth, outer margin smooth, ventral margin smooth and without teeth, basal ventral process lamellar and serrated, occupying median third of ventral aedeagal valve basal half, originating medially; aedeagal dorsal valve narrow and long, with two pairs of apical lobes, outer lobe, with outer margin smooth, inner margin with teeth on median portion in dorsal view, apical portion with two pairs of teeth, ventrad, with ventral and dorsal margin serrated, basal portion ventral margin smooth, dorsal margin smooth, inner lobe membranous, gibbous, about as large as outer lobe and its ventral area not covered by outer lobe when seen in lateral view, dorsal margin with arched tooth anterad, ventral margin smooth; basal cover plate, trapezoidal, anterior margin slightly outcurved medially, posterior margin strongly incurved medially, and not divided; apodema not extending anterior to basal plate. Female. Head. Head length 1.44 × its width; punctates large and dense; mandible with four distal teeth; median clypeal lobe trapezoidal, clypeal carina high in profile; first four antennal segments in a ratio of 9: 2: 1: 1, eleventh segment length 2 × its width; eye subcircular, with 7 facets, distant from the mandible by its length; vertex crest straight or nearly so; occipital carina length 0.41 × that of the head, visible in dorsal view. Mesosoma. Coriaceus; pronotal collar length 0.33 × that of the dorsal pronotal area, lateral area microstriate; dorsal pronotal area length 1.29 × its width, lateral area microstriate; mesonotum length 0.33 × that of the dorsal pronotal area, posterior margin convex and slightly embraced by the metapectal-propodeal complex; metapectal-propodeal complex elongated, lateral area microstriate; pleurosternum basal portion microstriate; epicnemium with transversal striae; metasternum diamond shaped; forefemur length 2.22 × its width, midtibia without spurs. Metasoma. Elongated and stout, length 2 × that of the mesosoma, petiole long and coriaceus-striate, segments polished, tranversal section circular.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFBBFFAF8DF3657BB1949D03.taxon	discussion	Remarks. Previous revisions did not mention or illustrate the presence of a lamellar and serrated basal ventral process.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFBBFFAF8DF3657BB1949D03.taxon	materials_examined	Material revised. Holotype, ♂, BRAZIL, Distrito Federal, Brasília, Ecological Reserve of IBGE, 15 ° 5 ’ 58 ” S 47 ° 51 ’ 02 ” W, forest ecotone, 18. iii. 1982, window trap, J. Dalmácio col. (IBGE). Paratypes: BRAZIL, Distrito Federal, Brasília, 1 ♂, Reserva Ecológica do IBGE, 15 º 55 ’ 58 ’’ S 47 º 51 ’ 02 ’’ W, Data: 9 – 16. XII. 1982, Janela, Beira da Mata (UFES nº 12207); 1 ♂, Data: 20 – 27. I. 1983, Janela, Beira da Mata (UFES nº 12205); 1 ♂, Data: 21 – 30. XII. 1982, Janela, Beira da Mata (UFES nº 12204). New material. BRAZIL, Pará, 1 ♂, Melgaço, Floresta Nacional Caxiuanã, Trilha Igarapé Curua, [01 ° 42 ’ 30 ” S 51 ° 31 ’ 45 ” W], 17. XI – 23. XI. 2003, arm. Malaise 7, A. P. Aguiar & J. D. Dias, Ponto P 05105 b (MPEG); 1 ♂, Serra Norte, Itacaiunas, [06 ° 00 ’ 01 ” S 50 ° 17 ’ 26 ” W], xi – xii. 1982, CDC light trap, eq. J. Arias (OSUC); 1 ♂, Vitória do Xingu, Rio Xingu, Igarapé Di Maria, [02 ° 53 ’ 02 ” S 52 ° 0 ’ 16 ” W], 12 – 16. VIII. 2008, arm. Malaise, O. T. Silveira & equipe col. (MPEG); São Paulo, 1 ♂, Teodora Sampaio, [22 ° 31 ’ 58 ” S 52 ° 10 ’ 04 ” W], XII – 1977, M. Alvarenga (PMAE); 1 ♂, Descalvado, Faz. Itaúnas, Cerrado, 21 º 54 ’ S 47 º 37 ’ W, 01. XII. 2005, arm. Malaise, N. W. Perioto & equipe col. (UFES nº 79594).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFBBFFAF8DF3657BB1949D03.taxon	distribution	Distribution. Brazil: Distrito Federal, São Paulo (Azevedo 1999 a). Now, it is recorded for the first time from the Brazilian state of Pará.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFBDFFAF8DF364D7B2C19A2D.taxon	description	(Fig. 4 E – G)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFBDFFAF8DF364D7B2C19A2D.taxon	diagnosis	Diagnosis. Male. Head. Mandible with three distal teeth; clypeus with one angulate median tooth, median carina incomplete apically. Mesosoma. Mesopectus with one pair of spines in posterior margin. Metassoma. Tergal process with tubercle partially mesad, depression with surrounding area setose; posterior hypopygeal margin straight or nearly so. Genitalia. Harpe narrow, apical margin narrow, inner margin angled subapically, outer margin straight, angulated medially; apex of digitus elongated, about 4 × as long as wide; basivolsela ventral margin sinuous; ventral aedeagal valve with apex anterior to apex of dorsal aedeagal valve, wide, progressively narrowing apicad, apex acute and not bifid, basal ventral process dentate, with seven small teeth; dorsal aedeagal valve short and gibbous. Female. Unknown.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFBDFFAF8DF364D7B2C19A2D.taxon	materials_examined	Material revised. Holotype, ♂, ECUADOR, Queredo, Pichilingue, v. 1978, V. Mendoza col. (USNM). Paratypes: ECUADOR, Pichincha, Tinalandia, 2 ♂, 14 – VI – 1976, S. + J. Peck (UFES nº 12291, 12292). New material. ECUADOR, Pichincha, 1 ♂, 16 km SE Sto Domingo, Tinalandia, 500 m, [0 ° 16 ’ 3 ” S 79 ° 3 ’ 39 ” W], 4 – 14. VI. 1976, S + J, Peck (CNCI); 1 ♂, 47 km S Sto Domingo, Rio Palenque, 200 m, [0 ° 39 ’ S 79 ° 18 ’ W], 18 – 30. V. 1975, S. Peck (CNCI); 1 ♂, 49 km S Sto. Domingo, [0 ° 40 ’ S 79 ° 17 ’ W], M. T., J. Belwood (CNCI).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFBDFFAF8DF364D7B2C19A2D.taxon	distribution	Distribution. Ecuador (Azevedo 1999 a).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFBDFFAC8DF363C4B2929D93.taxon	description	(Fig. 4 H – K)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFBDFFAC8DF363C4B2929D93.taxon	description	Description. Male. Head. Mandible with three distal teeth; median clypeal lobe ill-defined, with one angulate tooth, median clypeal carina high and straight in profile or nearly so, complete or nearly so; frons smooth, punctures small, and sparse; vertex crest straight or nearly so in dorsal view. Mesosoma. Dorsal pronotal area with anterior margin coriaceous. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad; depression deep with surrounding area setose; posterior hypopygeal margin straight or nearly so. Genitalia. Harpe with apical margin narrow and slightly posterad, outer margin rounded, inner margin angled subapically; apex of digitus short, about 2 × as long as wide; ventral aedeagal valve with apex anterior to apex of dorsal aedeagal valve, entirely narrow, outer margin sinuous, diverging apicad, apex acute and not bifid, inner margin straight, without concavity, outer margin smooth, ventral margin of ventral aedeagal valve smooth and with one pair of subapical teeth, basal ventral process lamellar, occupying only base of ventral aedeagal valve, originating medially; dorsal aedeagal valve narrow and long with two pairs of apical lobes, outer lobe with outer margin smooth, inner margin smooth in dorsal view, apical portion with two pairs of teeth, both ventrad, with subapical one extremely long, basal portion ventral margin with projection ventro-anterad serrated apically, dorsal margin with a straight tooth anterad, inner lobe membranous, gibbous, rounded, larger than outer lobe with its ventral area not covered by outer lobe when seen in lateral view, dorsal margin smooth, ventral margin smooth; basal cover plate absent; apodema without apical membrane, not extending anterior to basal plate. Female. Unknown. Variations. Vertex outcurved; posterior hypopigeal margin medially incurved.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFBDFFAC8DF363C4B2929D93.taxon	diagnosis	Differential diagnosis. This species differs from all other species of vallensis species-group by having an extremely long pair of subapical teeth ventrad on the outer lobe of the dorsal aedeagal valves.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFBDFFAC8DF363C4B2929D93.taxon	materials_examined	Material examined. Holotype, ♂, BRAZIL, Bahia, Ilhéus, Faz. São José, 14 º 42 ’ S 39 º 11 ’ W, 26. IV. 2002, arm. Malaise (CPDC). Paratypes: BRAZIL, Bahia, 1 ♂, Encruzilhada, 15 º 32 ’ 25 ’’ S 40 º 50 ’ 12 ’’ W, 800 m, 10 – 12. xii. 2007, arm. luminosa, J. A. Rafael et al. col (INPA); 1 ♂ Ibicuí, Faz. Maravilha, 14 º 52 ’ S 40 º 00 ’ W, 26. XI. 2001, arm. Malaise (CPDC); 2 ♂, Iguaí, Faz. Veneza, [14 ° 45 ’ S 40 ° 05 ’ W], 28. XI. 2002, arm. Malaise (CPDC *); 2 ♂, Itororó, Faz. Guanabara, 15 º 03 ’ S 039 º 06 ’ W, 25. XI. 2002, arm. Malaise (CPDC *); 1 ♂, Itororó, Faz. Santa Cruz, 15 º 01 ’ S 040 º 02 ’ W, 24. XI. 2002, arm. Malaise (CPDC); 1 ♂, Santa Luzia, Faz. Conceição, 15 º 20 ’ S 39 º 26 ’ W, 14. III. 2001, arm. Malaise, J. A. Cardoso & J. R. Maia (CPDC); Pernambuco, Caruaru, [08 ° 17 ’ S 35 ° 59 ’ W], IV – 1972, 1 ♂ M. Alvarenga (UFES nº 13301), previously determined as D. planus, M. T., J. Lima (CNCI).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFBDFFAC8DF363C4B2929D93.taxon	etymology	Etymology. The specific epithet refers to the Primarch Corvus, from the Warhammer 40 k media.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFBDFFAC8DF363C4B2929D93.taxon	distribution	Distribution. Brazil: Bahia and Pernambuco.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFBEFFAD8DF36527B0869CE3.taxon	description	(Fig. 4 L – N)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFBEFFAD8DF36527B0869CE3.taxon	diagnosis	Diagnosis. Male. Head. Mandible with three distal teeth; clypeus with one median carina. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad, depression with surrounding area setose; posterior hypopygeal margin slightly incurved medially. Genitalia. Harpe with apical margin narrow and slightly posterad, outer margin rounded, inner margin angled subapically; apex of digitus short, about 2 × as long as wide; basivolsela ventral margin rounded; ventral aedeagal valve with apex posterior to apex of dorsal aedeagal valve, wide, abruptly narrowing apicad, outer margin outcurved, diverging apicad, apex slightly serrated, acute and not bifid, outer margin smooth, ventral margin without teeth, basal ventral process absent; aedeagal dorsal valve narrow and long with two pairs of apical lobes, outer lobe, with outer margin smooth, inner margin smooth in dorsal view, apical portion with one pair of teeth, ventrad, basal portion ventral margin smooth, dorsal margin smooth, inner lobe membranous, gibbous, larger than outer lobe and its ventral area not covered by outer lobe when seen in lateral view, dorsal margin smooth; basal cover plate, trapezoidal, anterior margin strongly outcurved medially, posterior margin straight or nearly so, and not divided; apodema not extending anterior to basal plate. Female. Unknown.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFBEFFAD8DF36527B0869CE3.taxon	discussion	Remarks. Previous revisions did not mention or illustrate the presence of a pair of apical teeth ventrad on the outer lobe of dorsal aedeagal valve, a character useful for this species identification.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFBEFFAD8DF36527B0869CE3.taxon	materials_examined	Material revised. Holotype, ♂, TRINIDAD & TOBAGO, Tobago, Saint John Prov., Hermitage River, Bridge Charlotteville; 12 – 21. iii. 1979 (USNM). Paratypes: ECUADOR, Napo, 1 ♂, Huahua sumaco, Km. 44, on Hollin-Loreto Rd, XII. 14.89, mal. trap, M. J. Wasbauer, H. Real (UFES nº 12649). Vouchers: BRAZIL, Acre, 1 ♂, Pq. Nac. Serra, Divisor, Sul – 6, 18 – 99. iii. 1997, arm. Malaise, E. F. Morato col. (UFES nº 12653); 1 ♂, 9, 23 – 24. iii. 1997, arm. Malaise, E. F. Morato col. (UFES nº 12652); ECUADOR, Sucumbios, 1 ♂, 0.5 ºS 76.5 ºW, 290 m, Sacha Lodge, 3 – 16. viii. 1994, P. Hibbs, Malaise Trap (UFES nº 12641); 1 ♂, 13 – 23. iv. 1994, P. Hibbs, Malaise Trap (UFES nº 12646); 1 ♂, 13 – 25. vii. 1994, P. Hibbs, Malaise Trap (UFES nº 12643); 1 ♂, 14 – 24. v. 1994, P. Hibbs, Malaise Trap (UFES nº 12642); 1 ♂, Sacha Lodge, 0.5 ºS 76.5 ºW, 14 – 24. iii. 1994, P. Hibbs, MT. 290 m (UFES nº 12645). New material. BOLIVIA, Cochabamba, 2 ♂, 109 km E Cochabamba, 17 º 8 ’ 52 ’’ S 65 º 42 ’ 54 ’’ W, 1400 m, cloud forest, FIT, 1 – 6. II. 1999 – F. Génier col. (CNCI *); 1 ♂, Villa Tunari road, 1480 m, 17 º 8 ’ 84 ’’ S 65 º 42 ’ 49 ’’ W, FIT, 1 – 6. II. 1999, R. S. Hanley col. (CNCI); 1 ♂, 117 km E Cochabamba at Lagunita, 1000 m, 17 ° 6 ’ 22 ” S 65 ° 40 ’ 57 ” W, 5 – 8. II. 1999, FIT, mountain evergreen forest, F. Génier col. (CNCI); 1 ♂, 1 – 6. II. 1999, FIT, mountain evergreen for., F. Génier col. (CNCI); 2 ♂, 117 km E Cochabamba on Villa Tunari road, 1040 m, 17 ° 6.53 ’ S 65 ° 41.20 ’ W, FIT, 1 – 6. II. 1999, R. S. Hanley col. (CNCI); 1 ♂, 6 – 8. II. 1999, R. S. Hanley col. (CNCI); 1 ♂, 117 km E Cochabamba Villa Tunari road, 1040 m, 17 ° 6.53 ’’ S 65 ° 41.203 ’’ W, 10 – 12. II. 1999 – FIT # 73, R. S. Hanley col. (CNCI); BRAZIL, Acre, 3 ♂, Manuel, Urbano, RESEX Cazumbá-Iracema, Morumbá, 9 º 07 ’ 53.8 ’’ S 68 º 53 ’ 17.3 ’’ W, xi. 2017, arm. Suspensa, E. F. Morato col (INPA *); 1 ♂, Sen. Guiomard FES, Catuaba, ix. 2016, 10 º 04 ’ 30 ’’ S 67 º 37 ’ 00 ’’ W, arm. Malaise, E. F. Morato col. (INPA); 1 ♂, Roraima, Amajari, Tepequém, 15 – 31. iii. 2016, arm. Malaise, R. Boldrini & J. A. Rafel col. (INPA); ECUADOR, Napo, 5 ♂, Limnococha, 250 m, [0 ° 24 ’ S 76 ° 37 ’ W], 15 – 28 – IV. 1976, S. & J. Peck (CNCI *); 1 ♂, Yuturi Lodge, [0 ° 32 ’ 54 ” S 76 ° 2 ’ 29 ” W], 20 – 21. III. 1999, FIT, R. Brooks (CNCI); 3 ♂, Rio Napo, Yuturi Lodge, 270 m, 0 º 32 ’ 54 ’’ S 76 º 2 ’ 18 ’’ W, 20 – 21. III. 1999, FIT, R. Brooks, D. Brzoska (CNCI *); Sucumbios, 1 ♂, Rio Napo, Sacha Lodge, 0 º 30 ’’ S 76 º 30 ’ W, 225 m, 3 – 13. viii. 1994, MT, P. Hibbs col. (CNCI); 1 ♂, Sacha Lodge, 270 m, 0 ° 28 ’ 14 ” S, 76 ° 27 ’ 35 ” W, 21 – 24. III. 1999, FIT, R. Brooks (CNCI); PERU, Pasco, 3 ♂, Vila Rica road, 1475 m, 10 ° 47 ’ S 75 ° 18 ’ W, 15 – 18. X. 1999, FIT, R. Brooks (CNCI *); [TRINIDAD & TOBAGO], Trinidad, 1 ♂, 8 Km N. Arima, Simla Res. Sta., lower montane rain for., 260 m, [10 ° 40 ’ N 61 ° 18 ’ W], 14 – 24. VI. 1993, FIT, S. & J. Peck (CNCI); 1 ♂, MT, S. & J. Pack, 93 – 47 (CNCI); 1 ♂, 24. VI – 8. VII. 1993, FIT, S. & J Peck, 93 – 57 (CNCI); 1 ♂, 8 Km Arima, Simla Res. Sta., lower montane rain for., 260 m, 24. VI – 8. VII. 1993, MT, S. & J. Peck, 93 – 56 (CNCI); 1 ♂, 8 Km N Arima, Simla Res. Sta. trop. for., 240 m, 6 – 10. VI. 1993, FIT, S. & J. Peck (CNCI); 2 ♂, 260 m, 6 – 14. VI. 1993, FIT, S. & J. Peck (CNCI *); 1 ♂, 13 Km S. Arima, 2 Km N. Talparo, Quesnell Farm, rain for., [10 ° 31 ’ 31 ” N 61 ° 16 ’ 16 ” W], 50 m, 12 – 22. VI. 1993, S. & J. Peck (CNCI); 1 ♂, 22. VI – 8. VII. 1993, S. & J. Peck (CNCI); 1 ♂, 22. VI – 8. VII. 1993, MT, S. & J. Peck, 93 – 45 (CNCI); VENEZUELA, Aragua, 2 ♂, Maracay, Pozo del Diablo, [10 ° 18 ’ N 67 ° 36 ’ W], 11. I. 1996, MT + FIT, M. J. Sharkey (CNCI).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFBEFFAD8DF36527B0869CE3.taxon	distribution	Distribution. This species was known from Brazil: Rondônia; Ecuador; Trinidad & Tobago, Venezuela (Azevedo 1999 a); Brazil: Acre, Ecuador (Azevedo 2003). Now, it is recorded for the first time from Bolivia and Peru.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFBFFFB28DF365B7B0AF9B57.taxon	description	(Fig. 5 A – D)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFBFFFB28DF365B7B0AF9B57.taxon	diagnosis	Diagnosis. Male. Head. Mandible with three distal teeth; clypeus with one median carina. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad, depression with surrounding area setose; posterior hypopygeal margin slightly incurved medially. Genitalia. Harpe with apical margin narrow and slightly posterad, outer margin rounded, inner margin angled subapically; apex of digitus short, about 2 × as long as wide; basivolsela ventral margin rounded; ventral aedeagal valve with apex posterior to apex of dorsal aedeagal valve, wide, progressively narrowing apicad, outer margin outcurved, not diverging apicad, apex dilated, acute and not bifid, inner margin smooth, outer margin smooth, ventral margin smooth and without teeth, basal ventral process absent; aedeagal dorsal valve narrow and long, with two pairs of apical lobes, outer lobe, with outer margin smooth, inner margin smooth in dorsal view, apical portion with two pairs of teeth, ventro-posterad and ventro-anterad, both narrow, basal portion ventral margin smooth, dorsal margin smooth, inner lobe membranous, gibbous, larger than outer lobe and its ventral area not covered by outer lobe when seen in lateral view, dorsal margin smooth, ventral margin smooth; basal cover plate, trapezoidal, anterior margin straight or nearly so, posterior margin strongly incurved medially, and not divided; apodema not extending anterior to basal plate. Female. Unknown. Variations. Genitalia can vary greatly in size.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFBFFFB28DF365B7B0AF9B57.taxon	discussion	Remarks. Previous revisions did not mention or illustrate the presence of two pairs of apical teeth ventro-posterad and ventro-anterad on the outer lobe of dorsal aedeagal valve, a character useful for this species identification.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFBFFFB28DF365B7B0AF9B57.taxon	materials_examined	Material revised. Holotype, ♂, COSTA RICA, Puntarenas, Monteverde, 1400 m, 10 ° 43 ’ N 84 ° 02 ’ W, 12 – 15. vi. 1974, Malaise trap, W. A. Harding & J. P. Donahue col. (LACM). Paratypes: COSTA RICA, Alajuela, 1 ♂, 28 – 11 – 1970, O. Peck (UFES nº 12657); 1 ♂, Guanacaste, 3 km SE R. Naranjo, 13 – 31 Jul. 1993, F. D. Parker (UFES nº 12658); 1 ♂, Heredia, Est. Biol. La Selva, 10.43 ºN 84.02 ºW, 26. vi. – 1. vii. 93 TRAP 1, B. V. Brown & D. H. Feener (UFES nº 12656); 1 ♂, S. J., C. Nara. NE Quenos, 16 Jul. 1975, W. J. Hanson (UFES nº 12663); MEXICO, Vera Cruz, 1 ♂, Cordoba, 4 sept. 1974, W. Hanson, G. Bohart (UFES nº 12655); PANAMA, Canal Zone, 1 ♂, Pan. Barro Colorado I. V. 6.1981, R. B. & L. S. Kimsey (UFES nº 12698). Vouchers: PANAMA, Bocas del Toro, Wekso, Teribe, PILA, 17 – 24 oct. 1999, Tr Malaise 50 m. s. n. m, A. Santos, 23 ♂ (UFES nº 12664 – 12686); 1 ♂, Colón, Pr. Parq. Nac. Portobello, San Antonio, 16 – 29 feb. 2000, tr. Malaise, P. González, A. Santos, 0795 (UFES nº 12687); CZ, 1 ♂, B. Colorado Is, 15 – 31. VII. 1994, FIT, D. Banks (CNCI); 5 ♂, Fortuna, Div. Cont. Chiriqui-, B. Toro, 8 – 11 Abr 1999, 1070 – 1090 m, Malaise, R. Cambra, A. Santos (UFES nº 12693 – 12697); 1 ♂, Pr. Chilibre, Parque Nacional Chegres, Campo Chagres, Tr. Malaise, 24 – 28 enero. 2000, A. Santos, P. Gonzalez, 0783 (UFES nº 12688); 2 ♂, Panama Pr., P. Nac. Altos de Campana, 10 – 14 ago. 1999, Tr. Malaise, A. Santos, P. González (UFES nº 12691, 12692); 2 ♂, Panama, Pr. P. Nac. Sob. Camino Plantación Malaise, 24 ago – 2 sep. 1999 m A. Santos, P. Gonzalez (UFES nº 12689, 12690). New material. COSTA RICA, Alajuela, 1 ♂, Est. Biol. Sam Ramon, areas conc. arenal, 700 m, [10 ° 13 ’ N 84 ° 37 ’ W], 1 – 5. X. 1995, PT. G. Carballo (CNCI); Guanacaste, 1 ♂, Arenales, W side, Volcán Cacao, 900 m, [10 º 55 ’ 43 ” N 85 º 28 ’ 10 ” W], 1988 – 1989 (MUCR); 1 ♂, Est. Pitilla, 9 km S, Santa Cecilia, 700 m, [10 ° 58 ’ 35 ” N 85 ° 25 ’ 42 ” W], V 1989, P. Hanson (MUCR); Heredia, 1 ♂, La Selva, [10 ° 25 ’ 53 ” N 84 ° 00 ’ 18 ” W] 10 – 31. XII. 1996, J. Pitts (CNCI); Limón, 1 ♂, P. N. Tortuguero, Est. 4 – esquinas, Om, [10 ° 23 ’ 46 ” N 83 ° 29 ’ 08 ” W], IX – X 1989, J. Solano (MUCR); 1 ♂, sur de Iroquois, 300 m, [10 ° 11 ’ 9.29 ” N 83 ° 37 ’ 20.03 ” W], 23 / V / 1987, Col. Paul Hanson (MUCR); 1 ♂, Pen. de Osa, Est. Fund. Neot. Agua Buena, 7 km W Rincon de Osa, 80 m, rain for., [08 ° 34 ’ N 83 ° 30 ’ W], 21 – 25. VI. 1997, FIT, S. & J. Peck (CNCI); Puntarenas, San Vito, Jardin Bot. Las Cruces, 1200 m, [08 ° 47 ’ 08 ” N 82 ° 57 ’ 35 ” W], 1 ♂ VII – VIII / 88, 1 ♂ 05 / VI / 1988, Col. P. Hanson (MUCR); San José, 2 ♂, P. N. Braulio Carrillo, 8 Km NE Tunel, 1100 m, [10 ° 09 ’ 35 ” N 83 ° 58 ’ 28 ” W], 15 / V / 88, Col. Hanson (MUCR *); 1 ♂ San Gerardo under C. Chirripo, 1400 m, CR 15, [09 ° 29 ’ N 83 ° 29 ’ W], 1 – 1 X – 1986, L. Masner S. S. (MUCR); ECUADOR, Pichincha, 1 ♂, 47 km S Sto Domingo, Rio Palenque, 200 m, [0 ° 39 ’ S 79 ° 18 ’ W], 18 – 30. V. 1975, S Peck (CNCI); PANAMA, Chiriquí, 2 ♂, 15 km NW Hato del Volcano, [08 ° 46 ’ N 82 ° 38 ’ W], 1200 m, 24 – 31. V. 1997, Peck & Howden (CNCI *); Div. Cont. Chiriqui-B. Toro, Fortuna, [08 ° 46 ’ 16.5 ” N 82 ° 11 ’ 27.3 ” W], 1 ♂ 8 – 11. abr. 1999, 1 ♂ 9 – 11. abril. 1999 Tr. Amar., Malaise, R. Cambra, A. Santos (MIUP); Coclé, 1 ♂, El Cope, 19. XI. 1994, Windson & Edwards (CNCI); 1 ♂, Par. Nac. Omar Torrijos H., [08 º 41 ’ 56 ” N 80 º 39 ’ 17 ” W], 26 – 30. sep. 1999, Tr Malaise, 570 – 670, m. s. n. m., A. Santos, P. González (MIUP); Darién, 25 ♂, PN Darién, Pirre, Est. Rancho Frio, 80 m, [08 ° 00 ’ N 77 ° 45 ’ W], 16 nov 00 – 17 ene 2001, Malaise, R. Cambra, A. Santos (MIUP *).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFBFFFB28DF365B7B0AF9B57.taxon	distribution	Distribution. Costa Rica, Mexico, and Panama (Azevedo 1999 a). Now, it is recorded for the first time from Ecuador.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA0FFB08DF362E3B1A09E23.taxon	description	(Fig. 5 E – H) Description. Male. Head. Mandible with three distal teeth; median clypeal lobe trapezoidal, with one rounded tooth, median clypeal carina high and outcurved in profile, incomplete; frons coriaceous, punctures small, and dense; vertex crest straight or nearly so in dorsal view. Mesosoma. Dorsal pronotal area with anterior margin coriaceous. Metasoma. Metasomal segment II with median tergal process tubercles strongly mesad, depression very deep with surrounding area setose; posterior hypopygeal margin straight or nearly so. Genitalia. Harpe with apical margin wide and mesad, outer margin rounded, inner margin angled subapically; apex of digitus short, about 2 × as long as wide; ventral aedeagal valve with apex aligned with apex of dorsal aedeagal valve, entirely narrow, outer margin sinuous, not diverging apicad, apex acute and not bifid, inner margin straight, without concavity, outer margin smooth, ventral margin smooth and with two pairs of subapical teeth, basal ventral process filamentous, and sinuous, originating laterally; dorsal aedeagal valve narrow and long with two pairs of apical lobes, outer lobe with outer margin spinose, inner margin smooth in dorsal view, apical portion with two pairs of teeth, dorsad and dorso-posterad, basal portion ventral margin smooth, dorsal margin smooth, inner lobe membranous, gibbous, rounded, about as large as outer lobe and almost covered by outer lobe when seen in lateral view, dorsal margin with arched tooth anterad, and posterad tooth, ventral margin smooth; basal cover plate, trapezoidal, and not divided, posterior margin strongly incurved medially, anterior margin slightly outcurved medially; apodema with distinct apical membrane, extending anterior to basal plate. Female. Unknown. Variations. Vertex crest outcurved; tergal process tubercles very weakly mesad.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA0FFB08DF362E3B1A09E23.taxon	diagnosis	Differential diagnosis. This species is similar to D. bicarinatus by having the ventral aedeagal valves with two pairs of teeth on its ventral margin. However, D. eliasi sp. nov. has two pairs of apical teeth on the outer lobe of the dorsal aedeagal valves and a clypeus with one clypeal carina, whereas D. bicarinatus has one pair of apical teeth on the outer lobe of the dorsal aedeagal valves and two carinas in its clypeus.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA0FFB08DF362E3B1A09E23.taxon	materials_examined	Material examined. Holotype, ♂, BRAZIL, Pará, Altamira, Rio Xingu, Ilha Grande, [04 ° 26 ’ 45 ” S 53 ° 50 ’ 32 ” W], 06 – 10. XI. 2007, arm. Malaise, O. T. Silveira & equipe col. (MPEG). Paratypes: BRAZIL, Pará, 2 ♂, same data as holotype (MPEG *); 1 ♂, same data as holotype except 26 – 30. I. 2008 (MPEG).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA0FFB08DF362E3B1A09E23.taxon	etymology	Etymology. The specific epithet refers to the first author’s paternal grandfather, Elias, in memoriam.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA0FFB08DF362E3B1A09E23.taxon	distribution	Distribution. Brazil: Pará.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA2FFB08DF367F7B39899B9.taxon	description	(Fig. 5 I – L)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA2FFB08DF367F7B39899B9.taxon	diagnosis	Diagnosis. Male. Head. Mandible with three distal teeth; clypeus with one median carina. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad, depression with surrounding area setose; posterior hypopygeal margin outcurved. Genitalia. Harpe with apical margin narrow and slightly posterad, outer margin rounded, inner margin angled subapically; apex of digitus short, about 2 × as long as wide; basivolsela ventral margin rounded; ventral aedeagal valve with apex anterior to apex of dorsal aedeagal valve, entirely narrow, outer margin sinuous, not diverging apicad, apex acute and not bifid, inner margin smooth, outer margin smooth, ventral margin without teeth, basal ventral process absent; aedeagal dorsal valve narrow and long, with two pairs of apical lobes, outer lobe, with outer margin smooth, inner margin smooth in dorsal view, apical portion with three pairs of teeth, ventrad, subapical projection weakly serrated, sub rectangular, basal portion ventral margin smooth, dorsal margin smooth, inner lobe membranous, gibbous, about as large as outer lobe and almost covered by outer lobe when seen in lateral view, dorsal margin smooth, ventral margin smooth; basal cover plate, trapezoidal, anterior margin straight or nearly so, posterior margin slightly incurved medially, and not divided; apodema not extending anterior to basal plate. Female. Unknown.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA2FFB08DF367F7B39899B9.taxon	discussion	Remarks. Previous revisions did not mention or illustrate the presence of a third ventrad pair of teeth in the outer lobe of dorsal aedeagal valve, behind the subquadrangular lamellar serrated projection, a character useful for this species identification.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA2FFB08DF367F7B39899B9.taxon	materials_examined	Material revised. Holotype, ♂, BRAZIL, São Paulo, São José do Barreiro, Serra da Bocaina; 4. xi. 1965, F. M. Oliveira col. (DZUP). Paratypes: BRAZIL, Espírito Santo, Santa Teresa, Estação Biológica Santa Lúcia, 1 ♂, 638 m, 19 º 58 ’ 18,5 ’’ S 40 º 32 ’ 07,6 ’’ W, 9 – 12. IV. 2001, Malaise B 5, C. O. Azevedo & equipe col. (UFES nº 12747), 1 ♂, 15 – 19. X. 2010, Malaise, Trilha 2, M. T. Tavares & equipe col. (UFES nº 126439); Paraná, 2 ♂, Morretes, Parque Est. do Pau Oco, 25 º 34 ’ 27,5 ’’ S 48 º 53 ’ 33 ’’ W, 10 – 13. IV. 2002, Malaise, M. T. Tavares & Eq. col. (UFES nº 12753, 12754). New material. BRAZIL, Espírito Santo, 1 ♂, Atílio Vivácqua, Serra das Torres, 20 º 59 ’ S 41 º 12 ’ W, 14 – 19. iv. 2007, arm. Malaise, C. Waichert & equipe col. (UFES nº 172356); 1 ♂, Sta Leopoldina, Bragança, 770 m, 20 º 10 ’ 14 ’’ S 40 º 34 ’ 56 ’’ W, 8 – 15. x. 2012, Malaise, C. O. Azevedo & eq. col. (UFES nº 131400); Santa Maria de Jetibá-Faz. Clarindo Kruger, 20 º 04 ’ 27.9 ’’ S 40 º 44 ’ 51.3 ’’ O, 2 ♂ 6 – 13. xii. 2002, 1 ♂ 29. xi – 06. xii. 2002, Armadilha Malaise, M. T. Tavares, C. O. Azevedo e eq. col. (UFES nº 14355, 14358, 14364); 2 ♂, Sta. Leopoldina, Bragança, 20 º 10 ’ 14 ’’ S 40 º 34 ’ 56 ’’ W, 770 m, 8 – 15. x. 2012, Malaise, C. O. Azevedo & eq. col. (UFES nº 129138, UFES nº 131353); Rio de Janeiro, 8 ♂, Nova Friburgo, 17 km E ou BR, 22 º 23 ’ 04 ’’ S 42 º 33 ’ 30 ’’ W, old secondary for., FIT, 29.12000, F. Génier (CNCI *); 1 ♂, Teresópolis, Sitio Davi, 22 º 26 ’ S 42 º 56 ’ W, v. 2005, arm. Möricke, Vert. A. Peronti & A. L. B. G. (UFES nº 172357); São Paulo, 1 ♂, São Carlos, Faz. Canchim, [21 ° 58 ’ S 47 ° 50 ’ W], 4 – 6. I. 1996, M. Sharkey (CNCI); 1 ♂, Mogi-Guaçu, [22 ° 22 ’ S 46 ° 57 ’ W], 3 – 8. I. 1970, J. M. & B. A. Campbell (CNCI).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA2FFB08DF367F7B39899B9.taxon	distribution	Distribution. Brazil: Rio de Janeiro, São Paulo (Azevedo 1999 a), Espírito Santo (Azevedo 2003), Bahia, Espírito Santo, Paraná, and Rio de Janeiro (Redighieri & Azevedo 2006).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA2FFB18DF36148B1969B1F.taxon	description	(Fig. 5 M – P)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA2FFB18DF36148B1969B1F.taxon	description	Description. Male. Head. Mandible with three distal teeth; median clypeal lobe subtrapezoidal, with one angulate tooth, median clypeal carina high and straight in profile or nearly so, complete or nearly so; frons coriaceous, punctures small, and dense; vertex crest straight or nearly so in dorsal view. Mesosoma. Dorsal pronotal area with anterior margin coriaceous. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad, depression deep with surrounding area glabrous; posterior hypopygeal margin straight or nearly so. Genitalia. Harpe with apical margin wide and mesad, outer margin slightly angulate, inner margin angled subapically; apex of digitus short, about 2 × as long as wide; ventral aedeagal valve with apex anterior to apex of dorsal aedeagal valve, entirely narrow, outer margin sinuous, diverging apicad, apex acute and bifid, inner margin straight, without concavity, outer margin spinose, ventral margin smooth and without teeth, basal ventral process lamellar, occupying only base of ventral aedeagal valve, originating laterally; dorsal aedeagal valve narrow and long with two pairs of apical lobes, outer lobe with outer margin spinose, inner margin with teeth on median portion, apical portion with three pairs of teeth, two ventrad, and one wide dorsad, basal portion ventral margin with projection ventro-posterad fully smooth, dorsal margin smooth, inner lobe membranous, gibbous, rounded, larger than outer lobe and its ventral area not covered by outer lobe when seen in lateral view, dorsal margin with arched tooth anterad, ventral margin smooth; basal cover plate, trapezoidal, and not divided, posterior margin strongly incurved medially, anterior margin straight or nearly so; apodema without apical membrane, not extending anterior to basal plate. Female. Unknown Variations. Hypopygeal posterior margin medially incurved.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA2FFB18DF36148B1969B1F.taxon	diagnosis	Differential diagnosis. This species is similar to D. roboutei sp. nov. by having at least two pairs of apical teeth ventrad in the outer lobe of the dorsal aedeagal valves. However, D. firminoi sp. nov. has two pairs of apical teeth ventrad and a lamellar basal ventral process, whereas D. roboutei sp. nov. has three pairs of apical teeth on its outer lobe and a filamentous basal ventral process.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA2FFB18DF36148B1969B1F.taxon	materials_examined	Material examined. Holotype, ♂, BOLIVIA, Cochabamba, Vila Tunari, 325 m, 6 ° 59 ’ 642 ” S 65 ° 26 ’ 103 ” W, 4 – 6. XI. 2000, MT, M. Hauser (CNCI). Paratypes: BOLIVIA, Cochabamba, 1 ♂, 67.5 km E Villa Tunari, Est. Biol. Valle del Saita, 300 m, 17 ° 6 ’ 19 ” S 64 ° 46 ’ 57 ” W, 7 – 9. II. 1999, FIT, rain forest, F. Génier col. (CNCI); Santa Cruz, 1 ♂, Rio Mamore 2 km. N. mouth of Rio Chapare, VII – 31 – 1965, J. K. Bouseman Collector (UFES nº 14199), previously determined as D. stellatus.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA2FFB18DF36148B1969B1F.taxon	etymology	Etymology. The specific epithet refers to the first author’s maternal grandfather, Firmino, in memoriam.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA2FFB18DF36148B1969B1F.taxon	distribution	Distribution. Bolivia.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA3FFB68DF362ABB3709DCB.taxon	description	(Fig. 6 A – D) Description. Male. Head. Mandible with three distal teeth; median clypeal lobe ill-defined, with one angulate tooth, median clypeal carina high and straight in profile or nearly so, complete or nearly so; frons smooth, punctures small, and sparse; vertex crest straight or nearly so in dorsal view. Mesosoma. Dorsal pronotal area with anterior margin coriaceous. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad, depression shallow with surrounding area glabrous; posterior hypopygeal margin straight or nearly so. Genitalia. Harpe with apical margin narrow and slightly posterad, outer margin rounded, inner margin rounded subapically; apex of digitus short, about 2 × as long as wide; ventral aedeagal valve with apex anterior to apex of dorsal aedeagal valve, wide, progressively narrowing apicad, outer margin outcurved, not diverging apicad, apex acute and not bifid, inner margin incurved, with distinct concavity, outer margin smooth, ventral margin smooth and without teeth, basal ventral process absent; dorsal aedeagal valve narrow and long with two pairs of apical lobes, outer lobe with outer margin with small apical filament, inner margin with teeth on median portion in dorsal view, apical portion entirely ventrad, its dorsal margin entirely spinose, basal portion ventral margin smooth, dorsal margin with a straight tooth anterad, inner lobe membranous, gibbous, rounded, larger than outer lobe and its ventral area not covered by outer lobe when seen in lateral view, dorsal margin with arched tooth anterad, ventral margin smooth; basal cover plate, trapezoidal, and not divided, posterior margin strongly incurved medially, anterior margin slightly outcurved medially; apodema without apical membrane, not extending anterior to basal plate. Female. Unknown Variations. Tergal process with setae tufts on the lateral sides of depression.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA3FFB68DF362ABB3709DCB.taxon	diagnosis	Differential diagnosis. This species differs from all other species of vallensis species-group by having an entirely ventrad outer lobe of the dorsal aedeagal valves, with its dorsal margin entirely spinose.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA3FFB68DF362ABB3709DCB.taxon	materials_examined	Material examined. Holotype, ♂, BRAZIL, Acre, Sen. Guiomard, FES Catuaba, 10 º 04 ’ 28 ’’ S 67 º 37 ’ 00 ’’ W, 14 – 31. i. 2017, arm. Malaise, E. F. Morato & J. A. Rafael col., Rede BIA (INPA). Paratypes: BRAZIL, Amazonas, 1 ♂, Benjamin Constant, Igarapé Crajarizinho, 04 ° 26 ’ 35 ” S 69 ° 59 ’’ 58 ” W 9. IX. 2005, arm. Malaise, J. A. Rafael & F. F. Xavier Fº col. (INPA); 1 ♂, Lábrea, Ramal Apaeral, Km 09, Sitio São Raimundo, 07 ° 19 ’ 10 ” S 64 ° 40 ’ 07 ” W, VI. 2005, arm Malaise, F. F. Xavier Fº col. (INPA); 1 ♂, PEF. Palmeiras do Javari, 5 º 8.16 ’’ S 72 º 48.72 ’’ W, 1 – 15. iv. 2016, arm. Malaise, M. L. Oliveira & F. F. Xavier F. col. (INPA); Rondônia, 1 ♂, Fazenda Rancho Grande, 62 km. S. Ariquemes, 165 m, S 10.32 W 62.48, 12 – 22 November 1991, E. M. Fisher collector (UCDC); Itapuã do Oeste, FLONA Jamari, 9 º 15 ’ 36 ’’ S 62 º 54 ’ 96 ’’ W, 1 ♂ iii. 2017, 3 ♂ ix. 2016, arm. Malaise, Z. F. M. Silva & J. A. Rafael col (INPA *); 1 ♂, Nova Mamoré, PES, Guarajá-Mirim, 10 º 18 ’ 48 ’’ S 64 º 33 ’ 10 ’’ W, xii. 2016, arm. Malaise, A. A. Silva col. (INPA); Porto Velho, 1 ♂, ESEC, Morro Três Irmãos, i. 2017, arm. Malaise, V. S. Silva & J. A. Rafael col. (INPA); 1 ♂, viii. 2016, 9 º 00 ’ 09 ’’ S 64 º 32 ’ 40 ’’ W, arm. Malaise, V. S. Silva & J. A. Rafael col. Rede BIA (INPA); 1 ♂, Ramal Maria da Gorda, Comunidade São Sebastião, 8 º 42 ’ 30 ’’ S 64 º 02 ’ 51 ’’ W, 6 – 14. xi. 2021, arm. Malaise, M. B. Zani & J. L. Calabazas col. (INPA).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA3FFB68DF362ABB3709DCB.taxon	etymology	Etymology. The specific epithet refers to the Primarch Fulgrim, from the Warhammer 40 k media.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA3FFB68DF362ABB3709DCB.taxon	distribution	Distribution. Brazil: Acre, Amazonas, and Rondônia.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA4FFB78DF3649FB2369FB7.taxon	description	(Fig. 6 E – H)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA4FFB78DF3649FB2369FB7.taxon	description	Description. Male. Head. Mandible with three distal teeth; median clypeal lobe ill-defined, with one angulate tooth, median clypeal carina high and straight in profile or nearly so, complete or nearly so; frons coriaceous, punctures small, and sparse; vertex crest straight or nearly so in dorsal view. Mesosoma. Dorsal pronotal area with anterior margin coriaceous. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad, depression shallow with surrounding area glabrous; posterior hypopygeal margin outcurved. Genitalia. Harpe with apical margin narrow and slightly posterad, outer margin rounded, inner margin rounded subapically; apex of digitus short, about 2 × as long as wide; ventral aedeagal valve with apex anterior to apex of dorsal aedeagal valve, wide, abruptly narrowing apicad, outer margin outcurved, diverging apicad, apex acute and not bifid, inner margin straight, without concavity, outer margin smooth, ventral margin basal half slightly spinose and without teeth, basal ventral process absent; dorsal aedeagal valve narrow and long with two pairs of apical lobes, outer lobe with outer margin smooth, inner margin smooth in dorsal view, apical portion with, subapical projection strongly serrated, filamentous, ventro-anterad, and one apical smooth filament, ventro-posterad, dorsal margin smooth, inner lobe membranous, gibbous, rounded, about as large as outer lobe and almost covered by outer lobe when seen in lateral view, dorsal margin with arched tooth anterad, ventral margin smooth; basal cover plate, trapezoidal, and not divided, posterior margin straight or nearly so, anterior margin strongly outcurved medially; apodema without apical membrane, extending anterior to basal plate. Female. Unknown. Variations. Mandibles bidentate.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA4FFB78DF3649FB2369FB7.taxon	diagnosis	Differential diagnosis. This species is similar to D. rosangelae by having one pair of smooth ventro-anterad filamentous projections on the outer lobe of the dorsal aedeagus valves. However, D. horusi sp. nov. has no basal ventral process nor teeth on its outer lobe, whereas D. rosangelae has two pairs of apical teeth ventrad on the outer lobe of the dorsal aedeagal valves and a unidentate basal ventral process.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA4FFB78DF3649FB2369FB7.taxon	discussion	Remarks. This species is composed entirely from specimens previosly assigned as D. rosangelae paratypes.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA4FFB78DF3649FB2369FB7.taxon	materials_examined	Material examined. Holotype, ♂, BRAZIL, Espírito Santo, Santa Maria de Jetibá, Faz. Paulo Seick, 20 º 02 ’ 31,1 ’’ S 40 º 41 ’ 51,3 ’’ W, 06 – 13. xii. 2002, Armadilha Malaise B 2, M. Tavares, C. Azevedo e eq. col. (UFES nº 14372). Paratypes: BRAZIL, Espírito Santo, 1 ♂, João Neiva, Alto Bérgamo, mata, 478 m, 19 º 44 ’ 46 ’’ S 40 º 26 ’ 43 ’’ W, 27. X – 3. XI. 2008, arm. Malaise, C. O. Azevedo & equipe col. (UFES nº 23018); 15 ♂, Santa Maria de Jetibá, Fazenda Clarindo Krüger, 20 º 04 ’ 27,9 ’’ S 40 º 44 ’ 51,3 ’’ W, 29. XI – 06. XII. 2002, 06 – 13. XII. 2002, Armadilha Malaise, Tavares, Azevedo e eq. col. (UFES nº 14798,14820, 14823, 14847, 14878, 14880, 14900, 14964, 15025, 15244, 15296, 15343, 15344, 15346, 15349); 1 ♂, Santa Teresa, Est. Biol. Sta. Lúcia, [19 º 58 ’ S 40 º 32 ’ W], 23. II. 2001, varredura, Azevedo & Kawada col. (UFES nº 15540).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA4FFB78DF3649FB2369FB7.taxon	etymology	Etymology. The specific epithet refers to the Primarch Horus, from the Warhammer 40 k media.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA4FFB78DF3649FB2369FB7.taxon	distribution	Distribution. Brazil: Espírito Santo.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA5FFB78DF3677EB22E9BD8.taxon	description	(Fig. 6 I – L) Description. Male. Head. Mandible with three distal teeth; median clypeal lobe trapezoidal, with one angulate tooth, median clypeal carina high and outcurved in profile, incomplete; frons weakly coriaceous, punctures small, and sparse; vertex crest straight or nearly so in dorsal view. Mesosoma. Dorsal pronotal area with anterior margin coriaceous. Metasoma. Metasomal segment II with median tergal process tubercles strongly mesad, depression very deep with surrounding area setose. Genitalia. Harpe with apical margin wide and mesad, outer margin rounded, inner margin angled subapically; apex of digitus short, about 2 × as long as wide; ventral aedeagal valve with apex anterior to apex of dorsal aedeagal valve, entirely narrow, outer margin outcurved, not diverging apicad, apex rounded and not bifid, inner margin straight, without concavity, outer margin smooth, ventral margin smooth, and with one pair of subapical teeth, basal ventral process filamentous, and rounded, originating laterally; dorsal aedeagal valve narrow and long with two pairs of apical lobes, outer lobe with outer margin spinose, inner margin smooth in dorsal view, apical portion with two pairs of teeth, dorsad and dorso-posterad, basal portion ventral margin smooth, dorsal margin smooth, inner lobe membranous, gibbous, rounded, larger than outer lobe and its dorsal area not covered by outer lobe when seen in lateral view, dorsal margin with arched tooth anterad, ventral margin smooth; basal cover plate, trapezoidal, and not divided, posterior margin strongly incurved medially, anterior margin slightly outcurved medially; apodema with distinct apical membrane, extending anterior to basal plate. Female. Unknown.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA5FFB78DF3677EB22E9BD8.taxon	diagnosis	Differential diagnosis. This species differs from all other species of vallensis species-group by having the ventral aedeagal valves with rounded apex and with one pair of teeth on the ventral margin.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA5FFB78DF3677EB22E9BD8.taxon	materials_examined	Material examined. Holotype, ♂, BRAZIL, Amapá, Serra do Navio, 0 º 52 ’ 28.1 ’’ N 51 º 57 ’ 45.4 ’’ W, 20. xi. 2014, varredura, J. A. Rafael & F. F. Xavier F. col (INPA). Paratypes: BRAZIL, Amapá, 1 ♂, Serra do Navio, 0 º 52 ’ 28.1 ’’ N 51 º 57 ’ 45.4 ’’ W, 20. xi. 2014, varredura, J. A. Rafael & F. F. Xavier F. col (INPA); Pará, 1 ♂, Rio Nhamundá, 01 º 35 ’ 11 ’’ S 57 º 37 ’ 32 ’’ W, 25 m, 17 – 20. V. 2008, Malaise, J. A. Rafael et al. leg (INPA).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA5FFB78DF3677EB22E9BD8.taxon	etymology	Etymology. The specific epithet refers to the Primarch Jaghatai, from the Warhammer 40 k media.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA5FFB78DF3677EB22E9BD8.taxon	distribution	Distribution. Brazil: Amapá and Pará.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA5FFB58DF3636EB17E9ECF.taxon	description	(Fig. 6 M – O)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA5FFB58DF3636EB17E9ECF.taxon	diagnosis	Diagnosis. Male. Head. Mandible with three distal teeth; clypeus with one median carina. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad, depression with surrounding area setose; posterior hypopygeal margin straight or nearly so. Genitalia. Harpe with apical margin wide and mesad, outer margin rounded, inner margin rounded subapically; apex of digitus short, about 2 × as long as wide; basivolsela ventral margin rounded; ventral aedeagal valve with apex posterior to apex of dorsal aedeagal valve, entirely narrow, outer margin sinuous, diverging apicad, apex acute and not bifid, inner margin serrated, outer margin smooth, ventral margin with one pair of subapical teeth, basal ventral process filamentous, and rounded, originating laterally; aedeagal dorsal valve narrow and long, with two pairs of apical lobes, outer lobe, with outer margin smooth, inner margin smooth, apical portion with two pairs of teeth, ventro-anterad, basal portion ventral margin with projection ventro-anterad, dorsal margin smooth, inner lobe membranous, gibbous, about as large as outer lobe and almost covered by outer lobe when seen in lateral view, dorsal margin with arched tooth anterad, ventral margin smooth; basal cover plate, trapezoidal, anterior margin straight or nearly so, posterior margin strongly incurved medially, and not divided; apodema not extending anterior to basal plate. Female. Unknown.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA5FFB58DF3636EB17E9ECF.taxon	materials_examined	Material revised. Holotype, ♂, BRAZIL, Amazonas, Estirão do Equador, Rio Javari; x. 1979; M. Alvarenga col. (PMAE). New material. BRAZIL, Amazonas, 2 ♂, Benjamin Constant, Igarapé Crajarizinho, 04 ° 26 ’ 35 ” S 69 ° 59 ’ 58 ” W, 9. IX. 2005, arm Malaise, J. A. Rafael & F. F. Xavier Fº col. (INPA *); 1 ♂, PEP, Palmeiras do Javari, 5 º 8.16 ’’ S 72 º 48.72 ’’ W, 1 – 15. iv. 2016, arm. Malaise, M. L. Oliveira & F. F. Xavier F. col. (INPA); Rondônia, 1 ♂, Porto Velho, ESEC, Morro Três Irmãos, viii. 2016, 9 º 00 ’ 09 ’’ S 64 º 32 ’ 40 ’’ W, arm. Malaise, V. S. Silva & J. A. Rafael col., Rede BIA (INPA).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA5FFB58DF3636EB17E9ECF.taxon	distribution	Distribution. Brazil: Amazonas (Azevedo 1999 a). Now, it is recorded for the first time from the Brazilian state of Rondônia.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA7FFB58DF3679BB1A0994F.taxon	description	(Fig. 7 A – D) Description. Male. Head. Mandible with three distal teeth; median clypeal lobe ill-defined, with one angulate tooth, median clypeal carina high and straight in profile or nearly so, complete or nearly so; frons smooth, punctures small, and sparse; vertex crest straight or nearly so in dorsal view. Mesosoma. Dorsal pronotal area with anterior margin carinate. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad, depression deep with surrounding area glabrous; posterior hypopygeal margin straight or nearly so. Genitalia. Harpe with apical margin narrow and slightly posterad, outer margin rounded, inner margin rounded subapically; apex of digitus short, about 2 × as long as wide; ventral aedeagal valve with apex anterior to apex of dorsal aedeagal valve, wide, progressively narrowing apicad, outer margin sinuous, not diverging apicad, apex acute and not bifid, inner margin incurved, with distinct concavity, outer margin smooth, ventral margin smooth and without teeth, basal ventral process absent; dorsal aedeagal valve narrow and long with two pairs of apical lobes, outer lobe with outer margin smooth, inner margin with apical filament in dorsal view, apical portion with, one smooth filament, ventro-posterad, basal portion ventral margin smooth, dorsal margin smooth, inner lobe membranous, gibbous, rounded, larger than outer lobe and its ventral area not covered by outer lobe when seen in lateral view, dorsal margin with arched tooth anterad, ventral margin smooth; basal cover plate, trapezoidal, and not divided, posterior margin strongly incurved medially, anterior margin slightly outcurved medially; apodema without apical membrane, not extending anterior to basal plate. Female. Unknown.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA7FFB58DF3679BB1A0994F.taxon	diagnosis	Differential diagnosis. This species is similar to D. angroni sp. nov. by having a similar ventral aedeagal valves: wide, narrowing progressively apicad, outer margin entirely rounded, inner apical margin with concavity and with a ventro-posterad filament on the outer lobe of the dorsal aedeagal valves. However, D. lorgari sp. nov. has no teeth on the outer lobe of the dorsal aedeagal valves nor in the inner margin of the outer lobe in dorsal view, whereas D. angroni sp. nov. has apical teeth both in the outer lobe and in its inner margin in dorsal view.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA7FFB58DF3679BB1A0994F.taxon	materials_examined	Material examined. Holotype, ♂, BRAZIL, Pará, Melgaço, Floresta Nacional Caxiuanã, Trilha Igarapé Curua, 17. XI – 23. XI. 2003, arm. Malaise 7, A. P. Aguiar & J. D. Dias, Ponto P 05105 b (MPEG). Paratype: BRAZIL, Pará, 1 ♂, Melgaço, Floresta Nacional Caxiuanã, Trilha Est. Cie. Ferreira Pena, [01 ° 42 ’ 30 ” S 51 ° 31 ’ 45 ” W] 17. XI – 22. XI. 2003, arm. Malaise 5, A. P. Aguiar & J. Dias, Ponto P 05107 b (MPEG).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA7FFB58DF3679BB1A0994F.taxon	etymology	Etymology. The specific epithet refers to the Primarch Lorgar, from the Warhammer 40 k media.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA7FFB58DF3679BB1A0994F.taxon	distribution	Distribution. Brazil: Pará.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA7FFBA8DF3601BB1A09C77.taxon	description	(Fig. 7 E – H) Description. Male. Head. Mandible with three distal teeth; median clypeal lobe ill-defined, with one angulate tooth, median clypeal carina high and straight in profile or nearly so, complete or nearly so; frons smooth, punctures small, and sparse; vertex crest straight or nearly so in dorsal view. Mesosoma. Dorsal pronotal area with anterior margin carinate. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad, depression deep with surrounding area glabrous; posterior hypopygeal margin straight or nearly so. Genitalia. Harpe with apical margin narrow and slightly posterad, outer margin rounded, inner margin angled subapically; apex of digitus short, about 2 × as long as wide; ventral aedeagal valve with apex anterior to apex of dorsal aedeagal valve, wide, progressively narrowing apicad, outer margin outcurved, not diverging apicad, apex acute and not bifid, inner margin incurved, with distinct concavity, outer margin smooth, ventral margin smooth and without teeth, basal ventral process absent; dorsal aedeagal valve narrow and long with two pairs of apical lobes, outer lobe with outer margin smooth, inner margin with teeth on median portion in dorsal view, apical portion with two pairs of teeth, posterad, basal portion ventral margin smooth, dorsal margin with a straight tooth anterad, inner lobe membranous, gibbous, rounded, larger than outer lobe and its ventro-posterial area not covered by outer lobe when seen in lateral view, dorsal margin smooth, ventral margin smooth; basal cover plate, trapezoidal, and not divided, posterior margin strongly incurved medially, anterior margin slightly incurved medially; apodema without apical membrane, not extending anterior to basal plate. Female. Unknown.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA7FFBA8DF3601BB1A09C77.taxon	diagnosis	Differential diagnosis. This species is similar to D. angroni sp. nov. by having a similar ventral aedeagal valves: wide, narrowing progressively apicad, outer margin entirely rounded, inner apical margin with concavity and outer lobe of the dorsal aedeagus valves with apical teeth. However, this species has the apical teeth in the outer lobe of the dorsal aedeagal valve posterad and without filamentous projections, whereas D. angroni sp. nov. has the outer lobe with apical teeth ventrad and a filament ventro-posterad.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA7FFBA8DF3601BB1A09C77.taxon	materials_examined	Material examined. Holotype, ♂, BRAZIL, Pará, Juruti, Estrada Mutum, [02 ° 09 ’ 12 ” S 56 ° 05 ’ 14 ” W], 07. XII. 2008, arm. Malaise, O. T. Silveira & equipe col. (MPEG).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA7FFBA8DF3601BB1A09C77.taxon	etymology	Etymology. The specific epithet refers to the Primarch Magnus, from the Warhammer 40 k media.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA7FFBA8DF3601BB1A09C77.taxon	distribution	Distribution. Brazil: Pará.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA8FFBB8DF3653EB2DF9FD3.taxon	description	(Fig. 7 I – L)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA8FFBB8DF3653EB2DF9FD3.taxon	description	Description. Male. Head. Mandible with three distal teeth; median clypeal lobe ill-defined, with one angulate tooth, median clypeal carina high and straight in profile or nearly so, complete or nearly so; frons smooth, punctures small, and sparse; vertex crest straight or nearly so in dorsal view. Mesosoma. Dorsal pronotal area with anterior margin coriaceous. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad, depression deep with surrounding area glabrous; posterior hypopygeal margin slightly incurved medially. Genitalia. Harpe with apical margin narrow and slightly posterad, outer margin rounded, inner margin rounded subapically; apex of digitus short, about 2 × as long as wide; ventral aedeagal valve with apex anterior to apex of dorsal aedeagal valve, wide, progressively narrowing apicad, outer margin outcurved, diverging apicad, apex strongly serrated, rounded and not bifid, inner margin straight, without concavity, outer margin strongly serrated, ventral margin smooth and without teeth, basal ventral process absent; dorsal aedeagal valve narrow and long with two pairs of apical lobes, outer lobe with outer margin spinose, inner margin smooth in dorsal view, apical portion with one pair of teeth ventrad, subapical projection strongly serrated and subcircular, basal portion ventral margin smooth, dorsal margin smooth, inner lobe membranous, flat ventrad, larger than outer lobe and its ventral area not covered by outer lobe when seen in lateral view, dorsal margin smooth, ventral margin with one filament ventrad; basal cover plate, trapezoidal, and medially divided, posterior margin strongly incurved medially, anterior margin slightly outcurved medially; apodema without apical membrane, not extending anterior to basal plate. Female. Unknown.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA8FFBB8DF3653EB2DF9FD3.taxon	diagnosis	Differential diagnosis. This species is similar to D. rotundus by having a subapical serrated on the outer lobe of the dorsal aedeagus valves. However, D. mortarioni sp. nov. has a distinct strongly serrated apex of ventral aedeagal valve, no basal ventral process and inner lobe of the dorsal aedeagus valves with filamentous projection ventrad, whereas, D. rotundus has a smooth apex of ventral aedeagal valve, unidentate basal ventral process and inner lobe of the dorsal aedeagus valves without projections.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA8FFBB8DF3653EB2DF9FD3.taxon	materials_examined	Material examined. Holotype, ♂, ECUADOR, Dureno, (Orient), [0 ° 3 ’ S 76 ° 42 ’ W], 150 – 200 m, 23 – 28. IX. 1977 (CNCI). Paratypes: BRAZIL, Acre, 1 ♂, Pq. Nac, Serra, Divisor, Sul – 9, 23 – 24. iii. 1997, arm. Malaise, E. F. Morato col. (UFES nº 12651) previously determined as D. digitatus; ECUADOR, Sucumbios, 1 ♂, 0.5 ºS 76.5 ºW, 290 m, Sacha Lodge, Malaise, 22. ii – 4. iii. 1994, P. Hibbs, col. (UFES nº 12644) previously determined as D. digitatus; Rio Napo, Sacha Lodge, 0 ° 30 ’ S 76 ° 30 ’ W, 225 m, 1 ♂ 3 – 13. VIII. 1994, 1 ♂ 12 – 22. VI. 1995, 2 ♂ 23. IV – 4. V. 1994, 1 ♂ 26. VII – 3. VIII. 1994, MT, P. Hibbs (CNCI *).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA8FFBB8DF3653EB2DF9FD3.taxon	etymology	Etymology. The specific epithet refers to the Primarch Mortarion, from the Warhammer 40 k media.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFA8FFBB8DF3653EB2DF9FD3.taxon	distribution	Distribution. Ecuador and Brazil: Acre.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFAAFFB88DF366F2B27B9B8B.taxon	description	(Fig. 7 M – P) Description. Male. Head. Mandible with three distal teeth; median clypeal lobe ill-defined, with one angulate tooth, median clypeal carina high and straight in profile or nearly so, complete or nearly so; frons smooth, punctures small, and dense; vertex crest straight or nearly so in dorsal view. Mesosoma. Dorsal pronotal area with anterior margin coriaceous. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad, depression deep with surrounding area glabrous. Genitalia. Harpe with apical margin narrow and slightly posterad, outer margin rounded, inner margin angled subapically; apex of digitus short, about 2 × as long as wide; ventral aedeagal valve with apex anterior to apex of dorsal aedeagal valve, wide, progressively narrowing apicad, outer margin outcurved, not diverging apicad, apex rounded and not bifid, inner margin straight, without concavity, outer margin smooth, ventral margin smooth and with distinct rounded projection, basal ventral process absent; dorsal aedeagal valve narrow and long with two pairs of apical lobes, outer lobe with outer margin smooth, inner margin smooth in dorsal view, apical portion with two pairs of teeth, ventrad and posterad, and two smooth filaments, ventro-posterad and ventro-anterad, basal portion ventral margin smooth, dorsal margin with a straight tooth anterad, inner lobe membranous, gibbous, rounded, larger than outer lobe, its dorsal area and ventral area not covered by outer lobe when seen in lateral view, dorsal margin smooth, ventral margin smooth; basal cover plate, trapezoidal, and not divided, posterior margin strongly incurved medially, anterior margin straight or nearly so; apodema without apical membrane, not extending anterior to basal plate. Female. Unknown. Variations. Pronotum smoother; tergal process with tubercles fully mesad, depression deep, its surrounding area with tufts of setae.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFAAFFB88DF366F2B27B9B8B.taxon	diagnosis	Differential diagnosis. This species differs from all other species of vallensis species-group by having the outer lobe of the dorsal aedeagal valves with two pairs of filament projections (ventro-posterad and ventro-anterad) and the ventral aedeagal valves with rounded projection on its ventral margin.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFAAFFB88DF366F2B27B9B8B.taxon	materials_examined	Material examined. Holotype, ♂, BRAZIL, Rondônia, Itapuã do Oeste, FLONA Jamari, 9 º 15 ’ 36 ’’ S 62 º 54 ’ 46 ’’ W, 11 – 25. v. 2017, arm. Malaise, Z. F. M. Silva & J. A. Rafael col. (INPA). Paratypes: BRAZIL, Rondônia, Itapuã do Oeste, FLONA Jamari, 09 º 15 ’ 36 ’’ S 62 º 54 ’ 46 ’’ W, 15 ♂ ix. 2016, 9 ♂, xii. 2016, 1 ♂, i. 2017, 5 ♂ iii. 2017, 4 ♂, 15 – 28. iv. 2017, 1 ♂, 11 – 25. v. 2017, arm. Malaise, Z. F. M. Silva & J. A. Rafael col. (INPA *); 7 ♂, Porto Velho, ESEC, Morro Três Irmãos, i. 2017, arm. Malaise, V. S. Silva & J. A. Rafael col. (INPA *).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFAAFFB88DF366F2B27B9B8B.taxon	etymology	Etymology. The specific epithet refers to the Primarch Perturabo from the Warhammer 40 k media.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFAAFFB88DF366F2B27B9B8B.taxon	distribution	Distribution. Brazil: Rondônia.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFAAFFBE8DF3635EB22E9EB3.taxon	description	(Fig. 8 A – D)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFAAFFBE8DF3635EB22E9EB3.taxon	diagnosis	Diagnosis. Male. Head. Mandible with three distal teeth; clypeus with one median carina. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad, depression with surrounding area setose; posterior hypopygeal margin slightly incurved medially. Genitalia. Harpe with apical margin narrow and slightly posterad, outer margin rounded, inner margin rounded subapically; apex of digitus short, about 2 × as long as wide; basivolsela ventral margin rounded; ventral aedeagal valve with apex anterior to apex of dorsal aedeagal valve, wide, abruptly narrowing apicad, outer margin sinuous, not diverging apicad, apex acute and bifid, inner margin smooth, outer margin smooth, ventral margin smooth and without teeth, basal ventral process absent; aedeagal dorsal valve narrow and long with three pairs of apical lobes, outer lobe, with outer margin smooth, inner margin smooth in dorsal view, apical portion with two pairs of teeth, ventrad, basal portion ventral margin smooth, dorsal margin smooth, median lobe with teeth ventrad, inner lobe membranous, gibbous, about as large as outer lobe and almost covered by outer lobe when seen in lateral view, dorsal margin smooth, ventral margin smooth; basal cover plate, trapezoidal, anterior margin slightly outcurved medially, posterior margin slightly incurved medially, and not divided; apodema not extending anterior to basal plate. Female. Unknown.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFAAFFBE8DF3635EB22E9EB3.taxon	materials_examined	Material revised. Holotype, ♂, BRAZIL, Pernambuco, Caruaru, iv. 1972, M. Alvarenga col. (PMAE). Paratypes: BRAZIL, Pernambuco, 3 ♂, Caruaru, IV – 1972, M. Alvarenga (PMAE) (UFES nº 13303, 13304, 13305). New material. BRAZIL, Bahia, 1 ♂, Buerarema, Faz. Puaia, seringueira, 14 º 57 ’ S 39 º 7 ’ W, 10. III. 2001, arm. Malaise, J. A. Cardoso & J. R. Maia (CPDC); 2 ♂, Itajuípe, Cachoeira, 15 º 17 ’ S 39 º 03 ’ W, 27. XI. 2002, arm. Malaise (CPDC *); 1 ♂, Santa Luzia, Faz. Conceição, 15 º 20 ’ S 39 º 26 ’ W, 14. III. 2001, arm. Malaise, J. A. Cardoso & J. R. Maia (CPDC); 1 ♂, Uruçuca, Faz. Brasil, 14 º 33 ’ S 39 º 21 ’ W, 19. VIII. 2002, arm. Malaise (CPDC); 1 ♂, Uruçuca, Faz. Guarany, 14 º 34 ’ S 39 º 19 ’ W, 20. VIII. 2002, arm. Malaise (CPDC).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFAAFFBE8DF3635EB22E9EB3.taxon	distribution	Distribution. This species was known from Brazil: Pernambuco (Azevedo 1999 a). Now, it is recorded for the first time from the Brazilian state of Bahia.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFACFFBF8DF36447B44C9ECF.taxon	description	(Fig. 8 E – H)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFACFFBF8DF36447B44C9ECF.taxon	diagnosis	Diagnosis. Male. Head. Mandible with three distal teeth; clypeus with one median carina. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad, depression with surrounding area setose; posterior hypopygeal margin straight or nearly so. Genitalia. Harpe with apical margin narrow and slightly posterad, outer margin rounded, inner margin angled subapically; apex of digitus elongated, about 4 × as long as wide; basivolsela ventral margin rounded; ventral aedeagal valve with apex anterior to apex of dorsal aedeagal valve, wide, progressively narrowing apicad, outer margin outcurved, not diverging apicad, apex rounded and not bifid, inner margin smooth, and incurved, with weak concavity, outer margin smooth, ventral margin smooth and without teeth, basal ventral process absent; aedeagal dorsal valve narrow and long, with two pairs of apical lobes, outer lobe, with outer margin smooth, inner margin with apical filaments in dorsal view, apical portion with four pairs of teeth, three posterad and one anterad, latter in a distinct ventrad projection, apical filament ramified, posterad, and fully smooth, basal portion ventral margin smooth, dorsal margin with a straight tooth anterad, inner lobe membranous, gibbous, larger than outer lobe and its ventral area not covered by outer lobe when seen in lateral view, ventral margin smooth; basal cover plate, trapezoidal, anterior margin slightly outcurved medially, posterior margin strongly incurved medially, and medially divided; apodema not extending anterior to basal plate. Female. Unknown.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFACFFBF8DF36447B44C9ECF.taxon	discussion	Remarks. Previous revisions did not mention or illustrate the presence of a ramified filamentous projection on the outer lobe of the aedeagal dorsal valve. In addition, the number of apical teeth in the outer lobe apical half was not accurately illustrate.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFACFFBF8DF36447B44C9ECF.taxon	materials_examined	Material revised. Holotype, ♂, BRAZIL, Amazonas, Manaus, Reserve 1208, 17. xi. 1986, Malaise trap, B. Klein col. (INPA). Paratypes: BRAZIL, Amazonas, Manaus, Malaise, B. Klein col., 1 ♂, 26. II. 1985 (UFES nº 13521); 1 ♂, 26. VI. 1985 (UFES nº 13536); 1 ♂, 21. I. 1986 (UFES nº 13524); 1 ♂, 11. IV. 1986 (UFES nº 13525); 2 ♂, 22. IV. 1986 (UFES nº 13527, 13528); 1 ♂, 13. V. 1986 (UFES nº 13526); 1 ♂, 12. VI. 1986 (UFES nº 13533); 1 ♂, 17. VI. 1986 (UFES nº 13529); 1 ♂, 22. VII. 1986 (UFES nº 13517); 1 ♂, 24. IX. 1986 (UFES nº 13523); 1 ♂, 15. X. 1986 (UFES nº 13520); 1 ♂, 21. I. 1987 (UFES nº 13522); 1 ♂, Reserva 1208, 3. VI. 1986 (UFES nº 13530); 1 ♂, Reserva 1210, 10. VII. 1986 (UFES nº 13532); 1 ♂, Res. 1112, 3. II. 1986 (UFES nº 13519); 1 ♂, Reserva 1301, 13. V. 1985 (UFES nº 13535); 1 ♂, Reserva 1310, 9. II. 1986 (UFES nº 13537); 1 ♂, Res. Duche 1 m, 17. VII. 1992, adesiva, Videl & Videl col. (UFES nº 13538); 2 ♂, Res. Duche 50 m, 29. VII. 1992, adesiva, Vidal & Vidal col. (UFES nº 13539, 13540). New material. BRAZIL, Amazonas, Manaus, 1 ♂, Reserva 1208, Central norte, 09. X. 1984, rede, B. Klein col., Molecular B nº 00411 (INPA); 1 ♂, Reserva 1112, Central oeste, 09. X. 1984, rede, B. Klein col. (INPA); 1 ♂, Reserva 1112, Lateral oeste, 24. X. 1984, rede, Bert Klein col. (INPA); 1 ♂, Reserva 1208, Lateral leste, 9. X. 1984, rede, B. Klein col. (INPA); 1 ♂, Reserva 1208, 13. V. 1987, Malaise, B. Klein col. (INPA); 1 ♂, Reserva 1208, 14. I. 1986, Malaise, B. Klein col. (INPA); 1 ♂, Reserva 1208, Central norte, 09. X. 1984, rede, B. Klein col. (INPA); 1 ♂, Reserva 1401, Gavião, 10 – 25. XI. 1994, arm. Malaise (INPA); 1 ♂, Reserva 1401, Gavião, 18 – 30. IX. 1995, arm. Malaise (INPA); 1 ♂, Reserva Ducke, Rod. AM 010, Km 26, mata, [02 ° 53 ’ 29 ” S 59 ° 57 ’ 54 ” W], IX. 2001, arm. Malaise, J. F. Vidal col. (INPA); Amazonas / Pará {border}, 1 ♂, Rio Nhamundá, Jg. Areivas, 17 – 20. v. 2008, 01 º 35 ’ 11.5 ’’ S 57 º 27 ’ 32 ’’ W, Malaise, J. A. Rafael et al. col. (INPA); Pará, Oriximiná, Porto das Trombetas, 01 º 27 ’ S 56 º 30 ’ W, arm. Malaise, J. Cardoso & G. Costa col., 1 ♂, 14 – 24. XII. 2006 (UFES nº 106531); 3 ♂, 24. XII. 2006 – 3. I. 2007 (UFES nº 108874, 108950, 108984); 2 ♂, 25. XII. 2006 – 4. I. 2007 (UFES nº 106590, 106651); 2 ♂, 16 – 26. II. 2007 (UFES nº 107145, 107150); 2 ♂, 24. II – 6. III. 2007 (UFES nº 107211, 107216); 1 ♂, 25. II – 7. III. 2007 (UFES nº 106706); 3 ♂, 26. II – 8. III. 2007 (UFES nº 106664, 106674, 106679); FRENCH GUIANA, Saül, 1 ♂, 7 Km N 62 Km SE of Eaux Claires, Mt. Fuméc, 490 m, [03 ° 36 ’ N 53 ° 35 ’ W], 1 – 8. VI. 1997, FIT, J. Ashe, R. Brooks (CNCI); SURINAME, Marowijne, 2 ♂, Nassan Mountain, 500 m, 4 º 48 ’ 36 ’’ N 54 º 31 ’ 16 ’’ W, 2 – 4. VI. 1999, FIT, Z. H. Falin & B. DeDijn (CNCI *).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFACFFBF8DF36447B44C9ECF.taxon	distribution	Distribution. This species was known from Brazil: Amapá and Amazonas (Azevedo 1999 a). Now, it is recorded for the first time from Suriname, French Guiana, and the Brazilian state of Pará.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFADFFBF8DF3679BB27B996B.taxon	description	(Fig. 8 I – L) Description. Male. Head. Mandible with three distal teeth; median clypeal lobe subtrapezoidal, with one angulate tooth, median clypeal carina high and straight in profile or nearly so, complete or nearly so; frons weakly coriaceous, punctures small, and sparse; vertex crest strongly outcurved in dorsal view. Mesosoma. Dorsal pronotal area with anterior margin coriaceous. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad, depression deep with surrounding area setose. Genitalia. Harpe with apical margin wide and mesad, outer margin rounded, inner margin angled subapically; apex of digitus short, about 2 × as long as wide; ventral aedeagal valve with apex aligned with apex of dorsal aedeagal valve, entirely narrow, outer margin sinuous, diverging apicad, apex acute and not bifid, inner margin straight, without concavity, outer margin spinose, ventral margin strongly serrated and without teeth, basal ventral process filamentous, and sinuous, originating laterally; dorsal aedeagal valve narrow and long with two pairs of apical lobes, outer lobe with outer margin smooth, inner margin smooth in dorsal view, apical portion with four pairs of teeth, three ventrad and one dorsad, basal portion ventral margin smooth, dorsal margin smooth, inner lobe membranous, gibbous, rounded, larger than outer lobe and its ventral area not covered by outer lobe when seen in lateral view, dorsal margin with arched tooth anterad; basal cover plate, trapezoidal, and not divided, posterior margin strongly incurved medially; apodema without apical membrane, not extending anterior to basal plate. Female. Unknown.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFADFFBF8DF3679BB27B996B.taxon	diagnosis	Differential diagnosis. This species is similar to D. vulkani sp. nov. by having at least two pairs of apical teeth ventrad on the outer lobe of the dorsal aedeagal valves, basal ventral process filamentous and a ventral aedeagal valve ventral margin strongly serrated. However, D. roboutei sp. nov. differs from it by having four pairs of apical teeth, with none of them posterad, whereas D. vulkani sp. nov. has three pairs of apical teeth with one of them posterad.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFADFFBF8DF3679BB27B996B.taxon	materials_examined	Material examined. Holotype, ♂, BRAZIL, Rondônia, 1 ♂, Itapuã do Oeste, FLONA Jamari, 09 º 15 ’ 36 ’’ S 62 º 54 ’ 46 ’’ W, ix. 2016, arm. Malaise, Z. F. M Silva & J. A. Rafael col. (INPA). Paratypes: BRAZIL, Rondônia, Itapuã do Oeste, FLONA Jamari, 09 º 15 ’ 36 ’’ S 62 º 54 ’ 46 ’’ W, 1 ♂ 11 – 25. v. 2017, 2 ♂ 15 – 28. iv. 2017, 2 ♂ i. 2017, 1 ♂ iii. 2017, 6 ♂ ix. 2016, arm. Malaise, Z. F. M Silva & J. A. Rafael col. (INPA *).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFADFFBF8DF3679BB27B996B.taxon	etymology	Etymology. The specific epithet refers to the Primarch Roboute, from the Warhammer 40 k media.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFADFFBF8DF3679BB27B996B.taxon	distribution	Distribution. Brazil: Rondônia.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFADFFBC8DF3603FB35E9CC7.taxon	description	(Fig. 8 M – P)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFADFFBC8DF3603FB35E9CC7.taxon	description	Description. Male. Head. Mandible with three distal teeth; median clypeal lobe subtrapezoidal, with one angulate tooth, median clypeal carina high and angled in profile, complete or nearly so; frons coriaceous, punctures small, and dense; vertex crest weakly outcurved in dorsal view. Mesosoma. Dorsal pronotal area with anterior margin coriaceous. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad, depression deep with surrounding area setose; posterior hypopygeal margin straight or nearly so. Genitalia. Harpe with apical margin wide and mesad, outer margin slightly angulate, inner margin angled subapically; apex of digitus short, about 2 × as long as wide; ventral aedeagal valve with apex anterior to apex of dorsal aedeagal valve, entirely narrow, outer margin sinuous, not diverging apicad, apex acute and bifid, inner margin straight, without concavity, outer margin spinose, ventral margin basal half strongly spinose and without teeth, basal ventral process filamentous, and sinuous, originating laterally; dorsal aedeagal valve narrow and long with two pairs of apical lobes, outer lobe with outer margin smooth, inner margin smooth in dorsal view, apical portion with three pairs of teeth, dorsad, ventrad and postero-ventrad, with latter 2 × longer, basal portion ventral margin with projection ventrad, dorsal margin smooth, inner lobe membranous, gibbous, rounded, about as large as outer lobe and almost covered by outer lobe when seen in lateral view, dorsal margin with arched tooth anterad, ventral margin smooth; basal cover plate, trapezoidal, and not divided, posterior margin strongly incurved medially, anterior margin straight or nearly so; apodema without apical membrane, not extending anterior to basal plate. Female. Unknown.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFADFFBC8DF3603FB35E9CC7.taxon	diagnosis	Differential diagnosis. This species is similar to D. scavatus by having three pairs of apical teeth on the outer lobe of the dorsal aedeagal valves, ventrad, posterad and dorsad. However, D. rogali sp. nov. has the posterad pair of apical teeth 2 × times the size of the others and a filamentous ventral process, whereas D. scavatus has all three pairs of apical teeth about the same size and a lamellar basal ventral process.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFADFFBC8DF3603FB35E9CC7.taxon	materials_examined	Material examined. Holotype, ♂, BRAZIL, Amazonas, Barcelos, Rio Padauari, Com. Ararinha, 00 º 30 ’ 18 ’’ N 64 º 03 ’ 30 ’’ W, 05 – 08. vi. 2010, R. Machado, P. Dias, Malaise, J. A. Rafael, (INPA). Paratypes: BOLIVIA, Beni. 1 ♂, Rio Itenez at mouth of Rio Baures, X – 10 – 1964, J. K. Bouseman, J. Lussenhop col. (UFES nº 14196), previously determined as D. stellatus; PERU, San Martin, 1 ♂, 23 km S. Picota, Conservacíon Mun. Zona Barreal, 335 m, 7 – 14. III. 2005, 7 º 04.88 ’ S 76 º 18.89 ’’ W, M. E. Irwin, J. D. Vasquez, Malaise in tropical deciduous forest [PE 11 – 02] (MUSM *); 1 ♂, SM, Picota, 23 km S Picota: Quinillal, 335 m, 0705 / 7619, ME Irwin, JD Vasquez (MUSM).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFADFFBC8DF3603FB35E9CC7.taxon	etymology	Etymology. The specific epithet refers to the Primarch Rogal, from the Warhammer 40 k media.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFADFFBC8DF3603FB35E9CC7.taxon	distribution	Distribution. Brazil: Amazonas; Bolivia; and Peru.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFAEFF828DF3658EB4129FB7.taxon	description	(Fig. 9 A – D)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFAEFF828DF3658EB4129FB7.taxon	diagnosis	Diagnosis. Male. Head. Mandible with three distal teeth; clypeus with one median carina. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad, depression with surrounding area glabrous; posterior hypopygeal margin straight or nearly so. Genitalia. Harpe with apical margin narrow and slightly posterad, outer margin rounded, inner margin angled subapically; apex of digitus short, about 2 × as long as wide; basivolsela ventral margin rounded; ventral aedeagal valve with apex anterior to apex of dorsal aedeagal valve, entirely narrow, outer margin sinuous, diverging apicad, apex acute and not bifid, inner margin smooth, outer margin smooth, ventral margin smooth and without teeth, basal ventral process unidentate, originating medially; aedeagal dorsal valve narrow and long, with two pairs of apical lobes, outer lobe, with outer margin serrated, inner margin with apical filaments in dorsal view, apical portion with two pairs of teeth, ventrad, with subapical one serrated, apical filament, ventro-posterad, and fully smooth, basal portion ventral margin smooth, dorsal margin smooth, inner lobe membranous, gibbous, larger than outer lobe and its ventral area not covered by outer lobe when seen in lateral view, dorsal margin with arched tooth anterad, ventral margin smooth; basal cover plate, trapezoidal, anterior margin slightly outcurved medially, posterior margin slightly incurved medially, and not divided; apodema not extending anterior to basal plate. Female. Unknown.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFAEFF828DF3658EB4129FB7.taxon	discussion	Remarks. Previous revisions mention that this species had three pairs of apical lobes (inner, median and outer lobes), however, after reanalyzing the holotype, we found that the projections in the outer lobe were confused with additional lobes. This species is now bilobate, with two pairs of lobes, as is usual for vallensis species. Variations. Basal ventral process apex rounded.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFAEFF828DF3658EB4129FB7.taxon	materials_examined	Material revised. Holotype, ♂, BRAZIL, Espírito Santo, Santa Maria de Jetibá, Fazenda Clarindo Krüger, 20 º 03 ’ S 40 º 44 ’ O, 29. XI – 06. XII. 2002, Armadilha Malaise, [M. T.] Tavares, [C. O.] Azevedo e eq [uipe] col. (UFES nº 14371). New material. BRAZIL, Rio de Janeiro, Teresópolis, 22 º 26 ’ S 42 º 56 ’ W, arm. Möricke, A. L. Peronti & M. L. Silva col., 1 ♂ 19. VI. 2004 (UFES nº 80543), 1 ♂ VIII. 2005 (UFES nº 80571); Teresópolis 1 ♂, Faz. Marcos Palmeiras, 22 º 26 ’ S 42 º 56 ’ W, v. 2005, arm. Möricke, Borda B, Peronti col. (UFES nº 172358); São Paulo, 1 ♂, Serra Bocaina, [22 ° 40 ’ S 44 ° 36 ’ W], 13 – 17. I. 1969 (MCZH).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFFAEFF828DF3658EB4129FB7.taxon	distribution	Distribution. This species was known from Brazil: Espírito Santo (Colombo & Azevedo 2016). Now, it is recorded for the first time from the Brazilian states of Rio de Janeiro and São Paulo.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF90FF828DF3677EB22E9B05.taxon	description	(Fig. 9 E – G)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF90FF828DF3677EB22E9B05.taxon	diagnosis	Diagnosis. Male. Head. Mandible with three distal teeth; clypeus with one median carina. Metasoma. Metasomal segment II with median tergal process tubercles strongly mesad, depression with surrounding area setose; posterior hypopygeal margin straight or nearly so. Genitalia. Harpe with apical margin narrow and slightly posterad, outer margin rounded, inner margin angled subapically; apex of digitus short, about 2 × as long as wide; basivolsela ventral margin rounded; ventral aedeagal valve with apex anterior to apex of dorsal aedeagal valve, entirely narrow, outer margin sinuous, diverging apicad, apex dilated, acute and not bifid, inner margin smooth, outer margin smooth, ventral margin smooth and without teeth, basal ventral process unidentate, originating medially; aedeagal dorsal valve narrow and long with two pairs of apical lobes, outer lobe, with outer margin smooth, inner margin smooth in dorsal view, apical portion with three pairs of teeth, two ventro-anterad and one ventro-posterad, subapical projection strongly serrated, and subcircular, basal portion ventral margin smooth, dorsal margin smooth, inner lobe membranous and gibbous, dorsal margin smooth, ventral margin smooth; basal cover plate absent; apodema not extending anterior to basal plate. Female. Unknown. Variations. Outer lobe of the dorsal aedeagal valve with subcircular subapical lamellar serrated projection narrower.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF90FF828DF3677EB22E9B05.taxon	materials_examined	Material revised. Holotype, ♂, BRAZIL, Rio de Janeiro, Silva Jardim, viii. 1974, F. M. Oliveira col. (CNCІ). New material. BRAZIL, Bahia, 2 ♂, Coaraci, Faz. São José, 14 º 38 ’ S 39 º 32 ’ W, 26. XI. 2002, arm. Malaise (CPDC *).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF90FF828DF3677EB22E9B05.taxon	distribution	Distribution. This species was known from Brazil: Rio de Janeiro (Azevedo 1999 a). Now, it is recorded for the first time from the Brazilian state of Bahia.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF90FF838DF362CBB3589D03.taxon	description	(Fig. 9 H – K)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF90FF838DF362CBB3589D03.taxon	description	Description. Male. Head. Mandible with three distal teeth; median clypeal lobe subtrapezoidal, with one angulate tooth, median clypeal carina high and straight in profile or nearly so, complete or nearly so; frons coriaceous, punctures small, and dense; vertex crest straight or nearly so in dorsal view. Mesosoma. Dorsal pronotal area with anterior margin coriaceous. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad, depression deep with surrounding area setose; posterior hypopygeal margin straight or nearly so. Genitalia. Harpe with apical margin wide and mesad, outer margin rounded, inner margin angled subapically; apex of digitus short, about 2 × as long as wide; ventral aedeagal valve with apex aligned with apex of dorsal aedeagal valve, entirely narrow, outer margin sinuous, diverging apicad, apex acute and bifid, inner margin straight, without concavity, outer margin spinose, ventral margin smooth and without teeth, basal ventral process lamellar, occupying only base of ventral aedeagal valve, originating medially; dorsal aedeagal valve narrow and long with two pairs of apical lobes, outer lobe with outer margin smooth, inner margin with teeth on basal portion in dorsal view, apical portion with two pairs of teeth, ventrad and postero-dorsad, latter with short protrusion, basal portion ventral margin with projection ventro-posterad fully smooth, dorsal margin smooth, inner lobe membranous, gibbous, rounded, larger than outer lobe and its dorsal area not covered by outer lobe when seen in lateral view, dorsal margin with arched tooth anterad, ventral margin smooth; basal cover plate, trapezoidal, and not divided, posterior margin strongly incurved medially, anterior margin slightly outcurved medially; apodema without apical membrane, not extending anterior to basal plate. Female. Unknown. Variations. Basal ventral process filamentous; outer lobe of the dorsal aedeagus valve without small protrusion on apical teeth.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF90FF838DF362CBB3589D03.taxon	diagnosis	Differential diagnosis. This species differs from all other species of vallensis species-group by having two pairs of apical teeth on the outer lobe of the dorsal aedeagal valves, ventrad and dorso-posterad, the latter having a small protrusion.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF90FF838DF362CBB3589D03.taxon	materials_examined	Material examined. Holotype, ♂, BRAZIL, Maranhão, 1 ♂, São Pedro da Água Branca, Fazenda Santa Rosa, 05 ° 07 ’ 07 ” S 48 ° 15 ’ 19 ” W, 06. XII. 2001, arm. Malaise, J. A. Rafael & equipe col. (INPA). Paratypes: BOLIVIA, Cochabamba, 2 ♂, 67.5 km E Villa Tunari, Est. Biol. Valle del Saita, 300 m, 17 º 6 ’ 19 ’’ S 64 º 46 ’ 57 ’’ W, 7 – 9. II. 1999, FIT, rain forest, F. Génier col. (CNCI *); 2 ♂, Villa Tunari, 16 º 54 ’ 55 ’’ S 65 º 22 ’ 06 ’’ W, 6. X. 2001, trop. for., MT,. H. Heider (CNCI); 1 ♂, trop. for., 16 ° 54 ’ 55 ” S 65 ° 22 ’ 06 ” W, 5 – 11. XI. 2001, MT, H. Heider (CNCI); Santa Cruz, 1 ♂, Rio Ichigo, (Locality B), July 24, 1965, See Bouseman field notes in Mammology Department, J. K. Bouseman Collector, (UFES nº 14198), previously determined as D. stellatus; BRAZIL, Goiás, 2 ♂, Araguaia River expedition, 162 Km, N. of Aruana, [13 ° 53 ’ 26 ” S 50 ° 50 ’ 07 ” W], 29 agosto 1992, Malaise, Scott R. Shaw coll. (ESUW).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF90FF838DF362CBB3589D03.taxon	etymology	Etymology. The specific epithet refers to the Primarch Sanguinius from the Warhammer 40 k media.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF90FF838DF362CBB3589D03.taxon	distribution	Distribution. Bolivia and Brazil: Goiás, Maranhão.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF91FF838DF364D7B19E99D5.taxon	description	(Fig. 9 L – N)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF91FF838DF364D7B19E99D5.taxon	diagnosis	Diagnosis. Male. Head. Mandible with three distal teeth; clypeus with one median carina. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad, depression with surrounding area setose; posterior hypopygeal margin straight or nearly so. Genitalia. Harpe with apical margin wide and mesad, outer margin rounded, inner margin rounded subapically; apex of digitus short, about 2 × as long as wide; basivolsela ventral margin rounded; ventral aedeagal valve with apex posterior to apex of dorsal aedeagal valve, entirely narrow, outer margin sinuous, diverging apicad, apex acute and bifid, inner margin smooth, outer margin spinose, in ventral view, ventral margin without teeth, basal ventral process lamellar, occupying only base of ventral aedeagal valve, originating laterally; dorsal aedeagal valve narrow and long with two pairs of apical lobes, outer lobe, with outer margin spinose, inner margin with teeth on median portion in dorsal view, apical portion with three pairs of teeth, ventro-posterad, postero-ventrad and dorso-posterad, basal portion ventral margin smooth, dorsal margin smooth, inner lobe membranous, gibbous, about as large as outer lobe and almost covered by outer lobe when seen in lateral view, dorsal margin with arched tooth anterad, ventral margin smooth; basal cover plate, trapezoidal, anterior margin straight or nearly so, posterior margin strongly incurved medially, and not divided; apodema not extending anterior to basal plate. Female. Unknown.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF91FF838DF364D7B19E99D5.taxon	materials_examined	Material revised. Holotype, ♂, PERU, Madre de Dios, Puerto Maldonado, 9. i. 1984, L. Huggert col. (PMAE). New material. BRAZIL, Acre, 1 ♂, Bujari, FES, Antimary, 09 º 20 ’ 01 ’’ S 68 º 19 ’ 17 ’’ W, 18 – 31. viii. 2017, arm. Malaise grande, Z. F. M. Silva & J. A. Rafael col. (INPA); Manoel Urbano, RESEX Cazumbá-Iracema, Castanheira, 9 º 08 ’ 31.1 ’’ S 68 º 56 ’ 44.0 ’’ W, 1 ♂ xi. 2017, 1 ♂ v. 2018, arm. Malaise, E. F. Morato col. (INPA); Manoel Urbano, RESEX Cazumbá-Iracema, Santo António, 1 ♂ Janeiro 2018, 1 ♂ iv. 2018, 2 ♂, viii. 2017 (INPA *); Rondonia, 2 ♂, Itapuã do Oeste, FLONA Jamari, 09 º 15 ’ 36 ’’ S 62 º 54 ’ 46 ’’ W, 15 – 28. iv. 2017, arm. Malaise, Z. F. M. Silva & J. A. Rafael col. (INPA); PERU, Madre de Dios, 4 ♂, Cocha Salvador, 310 m, [11 ° 59 ’ 53.2 ” S 71 ° 13 ’ 50.1 ” W], 20 – 21. X. 2000, FIT, R. Brooks (CNCI *).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF91FF838DF364D7B19E99D5.taxon	distribution	Distribution. This species was known from Peru (Azevedo 1999 a). Now, it is recorded for the first time from Brazil: Acre and Rondônia.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF91FF808DF3616CB54E9BA7.taxon	description	(Fig. 10 A – D)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF91FF808DF3616CB54E9BA7.taxon	diagnosis	Diagnosis. Male. Head. Mandible with three distal teeth; clypeus with one median carina. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad, depression with surrounding area setose; posterior hypopygeal margin straight or nearly so. Genitalia. Harpe with apical margin wide and mesad, outer margin rounded, inner margin angled subapically; apex of digitus short, about 2 × as long as wide; smooth; basivolsela ventral margin rounded; ventral aedeagal valve with apex posterior to apex of dorsal aedeagal valve, entirely narrow, outer margin sinuous, diverging apicad, apex acute and not bifid, inner margin smooth, outer margin serrated, in ventral view, ventral margin serrated and without teeth, basal ventral process filamentous, and rounded, originating laterally; aedeagal dorsal valve narrow and long, with two pairs of apical lobes, outer lobe, with outer margin smooth, inner margin smooth in dorsal view, apical portion with three pairs of teeth, ventro-posterad, ventro-anterad and posterad, former two significantly distant from latter one., basal portion ventral margin with projection ventro-anterad fully smooth, dorsal margin smooth, inner lobe membranous, gibbous, larger than outer lobe and its ventral area not covered by outer lobe when seen in lateral view, dorsal margin with arched tooth anterad, ventral margin smooth; basal cover plate, trapezoidal, anterior margin straight or nearly so, posterior margin strongly incurved medially, and not divided; apodema not extending anterior to basal plate. Female. Unknown.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF91FF808DF3616CB54E9BA7.taxon	materials_examined	Material revised. Holotype, ♂, BRAZIL, São Paulo, São Paulo, lbitinga, rubber-tree crop, 6. ix. 1988, Möricke trap (UFES nº 37751). Paratypes: BRAZIL, São Paulo, Ibitinga, 1 ♂, 3. V. 1989, BB {= white Möricke} (UFES nº 14190); 1 ♂, 22. III. 1989, BB {= white Möricke} (UFES nº 14193); 1 ♂, 22. III. 1985, BA {= yellow Möricke} (UFES nº 14194); 1 ♂, Cultura de Seringueira, 10. viii. 1988, Möricke (UFES nº 14188); 1 ♂, 18. v. 1988, Bac. Am. {= yellow Möricke} (UFES nº 14187). New material. ARGENTINA, Salta, 1 ♂, 30 km E Salta, Campo Quijano, El Alisal for., [24 º 54 ’ 36 ” S 65 º 38 ’ 27 ” W], 18 – 28. II. 1992, FIT, S. A. Marshall (CNCI); BRAZIL, Goiás, 2 ♂, Itumbiara, Faz. Fac. Agron- 429, Bacia Amarela, P. M., 13. XII. 1998, C. H. Marchiori col. (UFES nº 16254, 16255); São Paulo, Jaboticabal, 1 ♂, 21 º 19 ’ 30 ’’ S 48 º 13 ’ 05 ’’ W, 6 m, 20 – 26. i. 2024, S. R. Viel. col. (UFES nº 172360); 1 ♂, 21 º 19 ’ S 48 º 13 ’ W, 150 m, 16 – 22. xi. 2003, S. R. Viel col. (UFES nº 172361); PARAGUAY, Cazaapá, Hermosa, San Rafael Res., 1 ♂, López family, bank Rio Tebicuary, 80 m, 26 ° 17 ’ 23 ” S 55 ° 43 ’ 7 ” W, 1 – 4. XII. 2000, Z. H. Falin, FIT (CNCI); Cazaapá, Hermosa, San Rafael Res., 2 ♂, López family, 80 m, 26 ° 18 ’ 29 ” S, 55 ° 45 ’ 3 ” W, 1 – 3. XII. 2000, FIT, Z. H. Falin (CNCI); Cazaapá, Hermosa, San Rafael Res., 5 ♂, Sosa family, 90 m, 26 ° 19 ’ 15 ” S, 55 ° 44 ’ 55 ” W, 3 – 6. XII. 2000, FIT, Z. H. Falin (CNCI *); Puerto Stroessner, 5 ♂, [25 ° 31 ’ S 54 ° 37 ’ W], 18. XII. 1971, L. Peña (CNCI *); SURINAME, Marowijne, 15 ♂, Perica 70 Km E Paramaribo on East, West rd, 5 m, 5 º 40 ’ 28 ’’ N 54 º 36 ’ 31 ’’ W, 31. V – 5. IV. 1999, FIT, E. H. Falin, B. DeDijn (CNCI *); Para, 1 ♂, Carolina creek, 11 km SE Zaderij Airport 30 m, 5 º 23 ’ 36 ’’ N 55 º 09 ’ 29 ’’ W, 18 – 20. vi. 1999, FIT, Z. H. Falin & A. Gangadin (CNCI).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF91FF808DF3616CB54E9BA7.taxon	distribution	Distribution. This species was known from Bolivia, Brazil (Rondônia, São Paulo), Paraguay, and Suriname (Azevedo 1999 a). Now, it is recorded for the first time from Argentina and the Brazilian state of Goiás.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF92FF818DF36373B37F9ECF.taxon	description	(Fig. 10 E – G)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF92FF818DF36373B37F9ECF.taxon	diagnosis	Diagnosis. Male. Head. Mandible with three distal teeth; clypeus with one median carina. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad, depression with surrounding area setose; posterior hypopygeal margin straight or nearly so. Genitalia. Harpe with apical margin narrow and slightly posterad, outer margin rounded, inner margin rounded subapically; apex of digitus short, about 2 × as long as wide; basivolsela ventral margin rounded; ventral aedeagal valve with apex anterior to apex of dorsal aedeagal valve, entirely narrow, outer margin straight or nearly so, not diverging apicad, apex acute and not bifid, inner margin smooth, outer margin smooth, ventral margin smooth and without teeth, basal ventral process absent; aedeagal dorsal valve narrow and long, with two pairs of apical lobes, outer lobe medially divided horizontally, with outer margin smooth, inner margin smooth in dorsal view, apical portion divided horizontally, ventral margin serrated, basal portion ventral margin smooth, dorsal margin smooth, inner lobe membranous and gibbous, dorsal margin smooth, ventral margin smooth; basal cover plate absent; apodema extending anterior to basal plate. Female. Unknown.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF92FF818DF36373B37F9ECF.taxon	materials_examined	Material revised. Holotype, ♂, BRAZIL, Paraná, Colombo, Embrapa, Br 476, Km 20, 15. vi. 1987, Malaise trap; Survey Profaupar (DZUP). Paratype: BRAZIL, Paraná, 1 ♂, Telêmaco Borba, Res. Samuel Klabin, 15. XII. 1986, Lev. Ent. PROFAUPAR, Malaise (UFES nº 14208). Vouchers: BRAZIL, Paraná, 1 ♂, Res. Samuel, Klabin, 11. VII. 1988, Lev. Ent. PROFAUPAR Malaise, Telêmaco Borba (UFES nº 14207); São Paulo, 1 ♂, Salesópolis, Est. Biol. Boracéia, 23 º 39 ’ 06,5 ’’ S 45 º 53 ’ 48 ’’ W, 1 – 3. IV. 2001, Moericke, STP Amarante & Eq. col. (UFES nº 14206). New material. BRAZIL, Paraná, 1 ♂, Piraquara, Mananciais da Serra Mar, [25 º 29 ’ 46 ” S 48 º 58 ’ 54 ” W], 08. XII. 2006, arm. Malaise, J. A. Rafael col. (INPA); São Paulo, 1 ♂, São Rosué, 12. III. 1991, M. T. Tavares, col. (UFES nº 62216); 1 ♂, Serra da Bocaina, 1680 m, [22 ° 40 ’ S 44 ° 36 ’ W], III. 1973 (CNCI); Rio de Janeiro, 1 ♂, Serra dos Órgãos, [22 ° 27 ’ 18 ” S 42 ° 59 ’ 06 ” W], Teresopolis, R. J., I – 21 – 26 – 69, C. Porter, A. García (MCZH); 1 ♂, Faz. Marcos Palmeiras, 22 º 26 ’ S 42 º 56 ’ W, v 2005, arm. Möricke, Borda B, Peronti col. (UFES nº 172362).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF92FF818DF36373B37F9ECF.taxon	distribution	Distribution. This species was known from Brazil: Paraná and São Paulo (Azevedo 1999 a). Now, it is recorded for the first time from the Brazilian state of Rio de Janeiro.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF93FF818DF3679BB3679B15.taxon	description	(Fig. 10 H – J)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF93FF818DF3679BB3679B15.taxon	diagnosis	Diagnosis. Male. (based on literature). Head. Mandible with three distal teeth; clypeus with one median carina. Posterior hypopygeal margin straight or nearly so. Genitalia. Harpe with apical margin narrow and slightly posterad, outer margin slightly angulate, inner margin rounded subapically; apex of digitus short, about 2 × as long as wide; basivolsela ventral margin sinuous; ventral aedeagal valve with apex anterior to apex of dorsal aedeagal valve, entirely narrow, outer margin sinuous, not diverging apicad, apex rounded and not bifid, inner margin smooth, outer margin smooth, ventral margin smooth and without teeth, basal ventral process lamellar, occupying wholly basal half of ventral aedeagal valve, originating medially; aedeagal dorsal valve narrow and long, with two pairs of apical lobes, outer lobe, with outer margin smooth, inner margin smooth in dorsal view, apical portion with two pairs of teeth, ventrad, basal portion ventral margin smooth, dorsal margin smooth, inner lobe membranous and gibbous, dorsal margin smooth, ventral margin smooth; basal cover plate, trapezoidal, anterior margin slightly outcurved medially, posterior margin strongly incurved medially, and not divided; apodema not extending anterior to basal plate. Female. Unknown.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF93FF818DF3679BB3679B15.taxon	distribution	Distribution. Brazil: Maranhão (Colombo et al. 2018).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF93FF818DF362ACB2C198E8.taxon	description	(Fig. 10 K – N)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF93FF818DF362ACB2C198E8.taxon	diagnosis	Diagnosis. Male. Head. Mandible with three distal teeth; clypeus with one median carina. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad, depression with surrounding area setose; posterior hypopygeal margin straight or nearly so. Genitalia. Harpe with apical margin wide and mesad, outer margin rounded, inner margin rounded subapically; apex of digitus short, about 2 × as long as wide; basivolsela ventral margin rounded; ventral aedeagal valve with apex anterior to apex of dorsal aedeagal valve, entirely narrow, outer margin straight or nearly so, diverging apicad, apex acute and bifid, inner margin smooth, outer margin spinose, in ventral view, ventral margin smooth and without teeth, basal ventral process filamentous, and rounded, originating laterally; aedeagal dorsal valve narrow and long, with two pairs of apical lobes, outer lobe, with outer margin smooth, inner margin smooth in dorsal view, apical portion with three pairs of teeth, anterad, ventrad and posterad, basal portion ventral margin smooth, dorsal margin smooth, inner lobe membranous, gibbous, about as large as outer lobe and its dorsal and ventral area not covered by outer lobe when seen in lateral view, dorsal margin smooth, ventral margin smooth; apodema not extending anterior to basal plate. Female. Unknown.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF93FF818DF362ACB2C198E8.taxon	materials_examined	Material revised. Holotype, ♂, ECUADOR, Napo, Laguna Jattungocha, 20 km S Nuevo Roca Juerte, Rio Yasuni, 7 – 9. ii. 1986, Malaise trap, A. T. Finnamore col. (PMAE).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF93FF818DF362ACB2C198E8.taxon	distribution	Distribution. Ecuador (Azevedo 1999 a).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF95FF878DF366F2B34A992E.taxon	description	(Fig. 11 A – D)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF95FF878DF366F2B34A992E.taxon	diagnosis	Diagnosis. Male. Head. Mandible with three distal teeth; clypeus with one median carina. Metasoma. Metasomal segment II with median tergal process tubercles strongly mesad, depression with surrounding area setose; posterior hypopygeal margin slightly incurved medially. Genitalia. Harpe with apical margin wide and mesad, outer margin rounded, inner margin rounded subapically; apex of digitus short, about 2 × as long as wide; basivolsela ventral margin rounded; ventral aedeagal valve with apex anterior to apex of dorsal aedeagal valve, wide, progressively narrowing apicad, outer margin sinuous, diverging apicad, apex acute and not bifid, inner margin smooth, outer margin smooth, ventral margin smooth and without teeth, basal ventral process absent; aedeagal dorsal valve narrow and long with two pairs of apical lobes, outer lobe, with outer margin smooth, inner margin smooth in dorsal view, apical portion with three pairs of teeth, ventrad, basal portion ventral margin smooth, dorsal margin smooth, inner lobe membranous, gibbous, about as large as outer lobe and almost covered by outer lobe when seen in lateral view, dorsal margin smooth, ventral margin smooth; basal cover plate, trapezoidal, anterior margin strongly incurved medially, posterior margin strongly incurved medially, and not divided; apodema extending anterior to basal plate.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF95FF878DF366F2B34A992E.taxon	description	Female. Unknown.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF95FF878DF366F2B34A992E.taxon	discussion	Remarks. Previous revisions did not illustrate the presence of three pairs of apical teeth ventrad on the outer lobe of the dorsal aedeagal valves. Although, the presence of a “ serrated ” ventrad margin in the outer lobe was mentioned. Variations. Ventral aedeagal valve basal half wider and with apex less divergent.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF95FF878DF366F2B34A992E.taxon	materials_examined	Material revised. Holotype, ♂, DOMINICAN REPUBLIC, 4.7 km South Santo Domingo; A. Forsyth col. (MCZH). Vouchers: ECUADOR, Pichincha, Rio Palanque Res. Sta., 1 ♂, 5 – V – 25 – VII – 1985, 250 m, S. & J. Peck (UFES nº 14252); Tinalandia, 4 ♂, 16 km se. Sto. Domingo 500 m, VI – VIII – 1985, S. & J. Peck (UFES nº 14248 – 14251); Tinalandia, 5 ♂, 14 – VI – 1976, S. + J. Peck (UFES nº 14241 – 14245). New material. ECUADOR. Esmeraldas, Bilsa Biol. Station, 0 ° 34 ’ N 79 ° 71 ’ W, 500 m, 1 ♂ 28. IV – 10. V. 1996, 1 ♂ 10. V – 5. VI. 1996, FIT, P. Hibbs (CNCI); Santo Doming de los Tsáchilas, Otongachi, La Unión del Toachi, 9 ♂ V. 2003, 5 ♂ VI. 2003 leg Giovanni Onore, Museo Genova, coll. G. Pagliano, Acquisto 2012 (MCSN *); Pichincha, 1 ♂, Tinalandia, 16 km SE Sto Domingo, 15 - VI- 1975, S. + J. Peck (PMAE); 1 ♂, 16 km SE Sto Domingo, Tinalandia, [0 ° 16 ’ 3 ” S 79 ° 3 ’ 39 ” W], 680 m, 15 – 30. VI. 1976, J. Peck (CNCI); 2 ♂, 16 km SE Sto Domingo, Tinalandia, 750 m, 0 ° 16 ’ 3 ’’ S 79 ° 3 ’ 39 ’’ W, 26 – 27. III. 1999, R. Brooks & D. Brzoska (CNCI); 6 ♂, 47 km S Sto Domingo, Rio Palenque, [0 ° 39 ’ S 79 ° 18 ’ W], 1 – 14. VII. 1975, S. Forsyth (CNCI *); 3 ♂, 47 km S Sto Domingo, Rio Palenque, 200 m, 18 – 30. V. 1975, S. Peck (CNCI); 1 ♂, 47 km S Sto Domingo, Rio Palenque Sta, 22 – 31. VII. 1976, S + J Peck (CNCI); 6 ♂, 49 km S Sto. Domingo, M. T., J. Belwood (CNCI *); 1 ♂, Maquipucuna Biol. Res. 0 ° 12 ’ N 78 ° 63 ’ W, 1300 m, 3 – 18. IV. 1996, FIT, P. J. Hibbs (CNCI); 4 ♂, Maquipucuna Biol. Sta. 0 ° 7 ’ 22 ’’ N 78 ° 39 ’ 0 ’’ W, 27 – 29. X. 1999, FIT, Z. H. Falin (CNCI *); 2 ♂, Rio Palenque, [0 ° 30 ’ S 79 ° 20 ’ W], 9. II. 1976, H. Howden (CNCI); 3 ♂, Rio Palenque, [0 ° 30 ’ S 79 ° 20 ’ W], 21 – 25. II. 1976 (CNCI); 6 ♂, Rio Palenque Station, 250 m, [0 ° 30 ’ S 79 ° 20 ’ W], 22 – 31. VII. 1973, S. + J. Peck (CNCI *); PANAMA, Darién, 1 ♂, P. N. Darién, Pirre (Malaise, Est. Rancho Frio 80 m, 16 nov 00 – 17 ene 2001, R. Cambra, A. Santos (MIUP).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF95FF878DF366F2B34A992E.taxon	distribution	Distribution. This species was known from The Dominican Republic (Azevedo 1999 a) and Ecuador (Azevedo 2003). Now, it is recorded for the first time from Panama.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF95FF858DF360F8B2719E97.taxon	description	(Fig. 11 E – H)	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF95FF858DF360F8B2719E97.taxon	diagnosis	Diagnosis. Male. Head. Mandible with two distal teeth; clypeus with one median carina. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad, depression with surrounding area setose; posterior hypopygeal margin slightly incurved medially. Genitalia. Harpe with apical margin narrow and slightly posterad, outer margin rounded, inner margin rounded subapically; apex of digitus short, about 2 × as long as wide; basivolsela ventral margin rounded; ventral aedeagal valve with apex aligned with apex of dorsal aedeagal valve, wide, abruptly narrowing apicad, outer margin sinuous, diverging apicad, apex strongly setose, rounded and not bifid, inner margin smooth, outer margin smooth, ventral margin without teeth, basal ventral process absent; aedeagal dorsal valve narrow and long, with two pairs of apical lobes, outer lobe, with outer margin smooth, inner margin smooth in dorsal view, apical portion strongly reduced, without any modifications, basal portion ventral margin smooth, dorsal margin smooth, inner lobe membranous, gibbous, larger than outer lobe and its entire area not covered by outer lobe when seen in lateral view, dorsal margin smooth, ventral margin smooth; basal cover plate, trapezoidal, anterior margin slightly outcurved medially, posterior margin strongly incurved medially, and not divided; apodema not extending anterior to basal plate. Female. Unknown.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF95FF858DF360F8B2719E97.taxon	discussion	Remarks. Previous revisions did not mention or illustrate the apex of the ventral aedeagal valves as strongly setose, an unusual and unique character in the vallensis species-group.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF95FF858DF360F8B2719E97.taxon	materials_examined	Material revised. Holotype, ♂, COLOMBIA, Valle [del Cauca], Peñas Blancas, 10 km W Cali, 26 – 28.?. 1975, Malaise trap, R. Wilkerson (FSCA). Paratype 1 ♂, COLOMBIA, Valle [del Cauca], Peñas Blancas, 12 – 14. II. 1975 – 1, R. Wilkerson, Malaise Trap (UFES nº 14336). Voucher: BRAZIL, Espírito Santo, 1 ♂, S. Teresa, Est. Biol. Santa Lúcia, 10. XII. 2001, varredura, Azevedo & Kawada col. (UFES nº 14338). New material. BRAZIL, Espírito Santo, 1 ♂, Santa Maria de Jetibá, Faz. Clarindo Kruger, 20 º 04 ’ 27.9 ’’ S 40 º 44 ’ 51.3 ’’ O, 06 – 13. xii. 2002, armadilha Malaise, M. T. Tavares, C. O. Azevedo e eq. col. (UFES nº 14636).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF95FF858DF360F8B2719E97.taxon	distribution	Distribution. Colombia (Azevedo 1999 a), Brazil: Espirito Santo (Redighieri & Azevedo 2004), Panama and Colombia (Brito & Azevedo 2017).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF97FF858DF36423B27B9917.taxon	description	(Fig. 11 I – L) Description. Male. Head. Mandible with three distal teeth; median clypeal lobe subtrapezoidal, with one angulate tooth, median clypeal carina high and straight in profile or nearly so, incomplete; frons weakly coriaceous, punctures small, and sparse; vertex crest weakly outcurved in dorsal view. Mesosoma. Dorsal pronotal area with anterior margin coriaceous. Metasoma. Metasomal segment II with median tergal process tubercle weakly mesad, depression deep with surrounding area setose. Genitalia. Harpe with apical margin wide and mesad, outer margin rounded, inner margin rounded subapically; apex of digitus short, about 2 × as long as wide; ventral aedeagal valve with apex anterior to apex of dorsal aedeagal valve, entirely narrow, outer margin sinuous, diverging apicad, apex acute and not bifid, inner margin straight, without concavity, outer margin spinose, ventral margin strongly serrated and without teeth, basal ventral process filamentous, and sinuous, originating laterally; dorsal aedeagal valve narrow and long with two pairs of apical lobes, outer lobe with outer margin smooth, inner margin smooth in dorsal view, apical portion with three pairs of teeth, two ventrad and one posterad, basal portion ventral margin smooth, dorsal margin smooth, inner lobe membranous, gibbous, rounded, larger than outer lobe and its dorsal area not covered by outer lobe when seen in lateral view, dorsal margin with arched tooth anterad, ventral margin smooth; basal cover plate, trapezoidal, and not divided, posterior margin strongly incurved medially, anterior margin straight or nearly so; apodema without apical membrane, not extending anterior to basal plate. Female. Unknown.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF97FF858DF36423B27B9917.taxon	diagnosis	Differential diagnosis. This species is similar to D. roboutei sp. nov. by having at least two pairs of apical teeth ventrad on the outer lobe of the dorsal aedeagal valves, a filamentous basal ventral process and the ventral margin of the ventral aedeagal valves strongly serrated. However, D. vulkani sp. nov. has three pairs of apical teeth on the outer lobe of the aedeagus dorsal valves, with a distinct posterad pair, whereas, D. roboutei sp. nov. has four pairs of apical teeth on the outer lobe, without a posterad pair.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF97FF858DF36423B27B9917.taxon	materials_examined	Material examined. Holotype, ♂, BRAZIL, Rondônia, Porto Velho, ESEC, Morro dos Três Irmãos, i. 2017, arm. Malaise, V. S. Silva & J. A. Rafael col (INPA). Paratypes: BRAZIL, Rondônia, Itapuã do Oeste, FLONA Jamari, 09 º 15 ’ 36 ’’ S 62 º 54 ’ 46 ’’ W, 2 ♂ ix. 2016, 1 ♂ xii. 2016, 6 ♂ i. 2017, arm. Malaise, Z. F. M. Silva & J. A. Rafael col. (INPA *); Porto Velho, 3 ♂, ESEC, Morro dos Três Irmãos, i. 2017, arm. Malaise, V. S. Silva & J. A. Rafael col (INPA).	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF97FF858DF36423B27B9917.taxon	etymology	Etymology. The specific epithet refers to the Primarch Vulkan, from the Warhammer 40 k media.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
8625131CFF97FF858DF36423B27B9917.taxon	distribution	Distribution. Brazil: Rondônia.	en	Pereira, Leonardo R., De Brito, Chirlei D., Azevedo, Celso O. (2026): Revision of the Neotropical vallensis species-group (Hymenoptera, Bethylidae, Dissomphalus), with the discovery of fifteen new species. Zootaxa 5785 (3): 401-444, DOI: 10.11646/zootaxa.5785.3.1, URL: https://doi.org/10.11646/zootaxa.5785.3.1
