identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
50F8274568D152AEA5F505FCC6668C0D.text	50F8274568D152AEA5F505FCC6668C0D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Marasmius flavosulcatus Chun Y. Deng 2026	<div><p>Marasmius flavosulcatus Chun Y. Deng sp. nov.</p><p>Fig. 2</p><p>Chinese name.</p><p>黄沟纹小皮伞 (huang gou wen xiao pi san).</p><p>Etymology.</p><p>Flavosulcatus refers to the yellow, plicate pileus.</p><p>Holotype.</p><p>China • Guizhou province: Tongren City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.21908&amp;materialsCitation.latitude=28.65361" title="Search Plazi for locations around (long 108.21908/lat 28.65361)">Guizhou Mayanghe National Nature Reserve</a>, at 28°39'13.00"N, 108°13'8.67"E, alt. 667.82 m, 23 Sep. 2025, HGASMF 01-32728 (DCY 10537) .</p><p>Diagnosis.</p><p>Marasmius flavosulcatus is characterized by a plicate, pale-yellow pileus, a long reddish-yellow stipe, large basidiospores, globulare, hymeniform pilepellis, and the absence of pleurocystidia and caulocystidia. It differs from Marasmius grandiviridis by small yellow basidiomes lacking olive coloration, more regularly clavate, bulbous cheilocystidia (Wannathes et al. 2009 a).</p><p>Description.</p><p>Basidiomata medium to large. Pileus 1–4 cm broad, hemispherical when young, plano-convex with age, umbilicate, yellowish white (3 A 2, 4 A 2), deeply sulcate and striate. Lamellae adnexed, subdistant, L = 9–13 (l = 0–1), 3–5 mm broad, thick, non-marginate, non-intervenose, yellowish white (3 A 2); edge smooth and entire, Context thin, concolorous with pileus. Stipe 50–95 × 1–2 mm, central, cylindrical, hollow; surface dry, non-insititious, apex whitish, becoming yellowish white (3 A 2), light orange (6 A 5), greyish orange (6 B 5). Taste and odor not distinctive.</p><p>Basidiospores 27–34 × 3.8–5.4 µm, Q = 5.4–8.5, on average 29.45 [± 1.82 SD] × 4.73 [± 0.45 SD] µm, Qm = 6.29 [± 0.91 SD], N = 40 from 4 collections, elongate clavate to fusoid, often curved in profile, smooth, hyaline, inamyloid, thin-walled. Basidia 30–51 × 7–11 µm, clavate, 2 - or 4 - spored. Cheilocystidia abundant, 17–30 × 7–15 µm, pyriform, clavate. Pileipellis a hymeniform layer of Globulares-type cells, 14–28 × 7–15 µm, broadly clavate to pyriform, hyaline, inamyloid, thin- to thick-walled, non-gelatinous. Lamellar trama regular; hyphae 2.5–10 μm diam., cylindrical, hyaline, dextrinoid, thin-walled. Stipitipellis hyphae 4–8 (– 10) µm diam., subparallel, cylindrical, yellow to greenish yellow, smooth, dextrinoid. Caulocystidia absent. Clamp connections present in all tissues.</p><p>Habitat.</p><p>Gregarious on humus in forest.</p><p>Additional collection examined.</p><p>China • Guizhou Province: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.706894&amp;materialsCitation.latitude=26.621143" title="Search Plazi for locations around (long 106.706894/lat 26.621143)">Guiyang City</a>, 26°37'16.118"N, 106°42'24.825"E, alt. 1290.15 m, 21 Jun. 2024, HKAS 154778 (DCY 8225, paratype) ; ibid, HGASMF 01-28414 (DCY 8213); China • Guizhou province: Tongren City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.21796&amp;materialsCitation.latitude=28.656872" title="Search Plazi for locations around (long 108.21796/lat 28.656872)">Guizhou Mayanghe National Nature Reserve</a>, 28°39'24.74"N, 108°13'4.63"E, alt. 643.30 m, 7 Jun. 2025, HGASMF 01-31659 (DCY 10011) .</p><p>Notes.</p><p>A basepair comparison between the holotype sequence of the new species and the closely related species Marasmius grandiviridis Wannathes, Desjardin &amp; Lumyongs, revealed 92.82 % identity with 45 substitutions / indels in ITS, and 98.53 % identity with 13 substitutions / indels in LSU, supporting its recognition as a distinct species. Based on the phylogenetic tree, the new species is nested in subg. Globulares ser. Purpureostriatus (Oliveira et al. 2020; Oliveira et al. 2024). Compared to the medium- to large-sized basidiomata and sulcate pileus, the new species is morphologically similar to M. bekolacongoli Beeli and M. campestris N. K. Zeng, Zhi Q. Liang &amp; M. S. Su, Marasmius galbinus T. H. Li &amp; Chun Y. Deng and M. grandiviridis . Marasmius bekolacongoli has a red or grayish red pileus, with violet to dark violaceous tints in suici, and is described as having smaller basidiospores, 20 × 4.5 µm (Antonín 2007). Marasmius campestris differs by its purple-tinged pileus, shorter stipe, and presence of caulocystidia (Liang et al. 2017). Marasmius galbinus differs by its smaller, paler, conical pileus and shorter basidiospores (14–16 vs. 27–34 μm) (Deng and Li 2011). Marasmius grandiviridis has a larger, yellowish-green pileus with darker olive-green plicate and irregularly clavate to lageniform-mucronate cheilocystidia (Wannathes et al. 2009 a).</p></div>	https://treatment.plazi.org/id/50F8274568D152AEA5F505FCC6668C0D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Deng, Chun-Ying;Li, Tai-Shun;Xiang, Zhun;Zhao, Qi	Deng, Chun-Ying, Li, Tai-Shun, Xiang, Zhun, Zhao, Qi (2026): Three new species of Marasmius (Agaricales, Basidiomycota) from southwestern China. MycoKeys 136: 233-250, DOI: 10.3897/mycokeys.136.196875
0B6C7BC5B7975DE485D800D412214AFF.text	0B6C7BC5B7975DE485D800D412214AFF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Marasmius griseobrunneus Chun Y. Deng 2026	<div><p>Marasmius griseobrunneus Chun Y. Deng sp. nov.</p><p>Fig. 3</p><p>Chinese name.</p><p>灰褐小皮伞 (hui he xiao pi san).</p><p>Etymology.</p><p>“ griseobrunneus ” refers to its pileus becoming grayish brown when mature.</p><p>Holotype.</p><p>China • Yunnan Province: Kunming City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.74155&amp;materialsCitation.latitude=25.13521" title="Search Plazi for locations around (long 102.74155/lat 25.13521)">Heilongtan Park</a>, 25°08'06.76"N, 102°44'29.57"E, alt. 1949.09 m. 22 Oct. 2025, collected by Chunying Deng, HKAS 154776 (DCY 10741).</p><p>Diagnosis.</p><p>Marasmius griseobrunneus is characterized by gymnopoid basidiomata, small to medium-sized, applanate, occasionally shallowly infundibuliform, grayish brown pileus, medium-sized basidiospores, and pileipellis composed of hymeniform Globulares-type cells. It differs from Marasmius wynneae by a smaller pileus (3 cm vs. 5 cm), subdistant lamellae, an olivaceous brown pileus, and dense caulocystidia.</p><p>Description.</p><p>Basidiomata small to medium-sized. Pileus 1–3 cm, broadly convex when young, applanate at maturity, sometimes centrally depressed to form a shallow funnel, white to cream when young, becoming orange-gray (6 B 2) to grayish brown (6 D 3), glabrous to weakly striate, slightly hygrophanous at the margin. Lamellae adnate, subdistant, L = 15–18 (l = 2–4), 1–2 mm broad, white, reticulate or intervenose. Context white, thin. Stipe 35–40 × 2–5 mm, central, cylindrical, tapering toward the base, apex white, becoming reddish brown (8 D 5, 8 F 5), hollow, fibrous, covered with white furfuraceous scales. Base mycelium white. Taste and odor not distinctive.</p><p>Basidiospores 6.3–7.8 × 3.2–4.4 µm, Q = 1.5–2.3, mean 7.06 [± 0.42 SD] × 3.91 [± 0.34 SD] µm, Qm = 1.82 [± 0.22 SD], N = 60 from three collections, ellipsoid, thin-walled, smooth, inamyloid. Basidia 24–35 × 4–6 µm, clavate, 2 - or 4 - spored. Basidioles fusoid, cylindrical to clavate. Cheilocystidia common, 14–21 × 3.5–12 µm, broadly clavate to pyriform or subcylindrical, thin-walled. Pleurocystidia absent. Pileipellis a hymeniform layer of Globulares-type cells; terminal elements 13–32 × 9.3–15 μm, broadly clavate to pyriform or subcylindrical, thin-walled. Lamellae trama interwoven, with hyphae 3.5–7 μm in diam, hyaline, dextrinoid, thin-walled. Stipitipellis a cutis composed of cylindrical hyphae 3–7 μm wide, parallel, smooth. Caulocystidia cluster or single with clavate, subclavate, globulares to pyriform cells, 10–31 × 6.8–11 µm. Clamp connections present.</p><p>Habitat.</p><p>Scattered to gregarious on humus in mixed coniferous and broad-leaved forests.</p><p>Additional collection examined.</p><p>China • Yunnan Province: Kunming City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.74155&amp;materialsCitation.latitude=25.13521" title="Search Plazi for locations around (long 102.74155/lat 25.13521)">Heilongtan Park</a>, 25°08'06.76"N, 102°44'29.57"E, alt. 1949.09 m. 22 Oct. 2025, collected by Chunying Deng, HKAS 154775 (DCY 10692) ; • ibid, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.77113&amp;materialsCitation.latitude=25.086134" title="Search Plazi for locations around (long 102.77113/lat 25.086134)">Jindian Park</a>, 25°05'10.08"N, 102°46'16.08"E, alt. 1999.84 m. 23 Oct. 2025, HKAS 154777 (DCY 10758, paratype) . China • Guizhou Province: Tongren City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.1811&amp;materialsCitation.latitude=28.815166" title="Search Plazi for locations around (long 108.1811/lat 28.815166)">Guizhou Mayanghe National Nature Reserve</a>, 28°48'54.60"N, 108°10'51.96"E, alt. 1027.13 m, 31 May 2025, HGASMF 01-31535 (DCY 9854) . China • Guizhou Province: Zunyi City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.90887&amp;materialsCitation.latitude=28.1652" title="Search Plazi for locations around (long 105.90887/lat 28.1652)">Guizhou Xishui National Nature Reserve</a>, 28°09'54.72"N, 105°54'31.92"E, alt. 735.29 m, 21 Oct. 2024, HGASMF 01-29990 (DCY 8989) .</p><p>Notes.</p><p>A basepair comparison between the holotype sequence of the new species and the closely related species Marasmius wynneae Berk. &amp; Broome (specimen voucher: DMS- 9344190), revealed 92.62 % identity with 52 substitutions / indels in ITS, and 98.94 % identity with 9 substitutions / indels in LSU, supporting its recognition as a distinct species. Based on molecular phylogenetic and morphological analyses, Marasmius griseobrunneus belongs to subg. Globulares ser. Wynnearum (Oliveira et al. 2020; Oliveira et al. 2024). Within this series, Marasmius wynneae, M. nivicola and M. albogriseus (Peck) Singer are morphologically similar to the new species. However, M. wynneae lacks caulocystidia (Antonín et al. 2010 a). Marasmius nivicola differs by a whitish pileus, crowded and distinctly intervenose lamellae, a solid mycelium mat around the stipe base, and slender cheilocystidia (Takahashi 2000). Marasmius albogriseus is distinguished by the lighter colored pileus, a well-developed tomentum at the stipe base, and gregarious and caespitose habitat (Halling et al. 1985).</p></div>	https://treatment.plazi.org/id/0B6C7BC5B7975DE485D800D412214AFF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Deng, Chun-Ying;Li, Tai-Shun;Xiang, Zhun;Zhao, Qi	Deng, Chun-Ying, Li, Tai-Shun, Xiang, Zhun, Zhao, Qi (2026): Three new species of Marasmius (Agaricales, Basidiomycota) from southwestern China. MycoKeys 136: 233-250, DOI: 10.3897/mycokeys.136.196875
A0D648F9E9225739BB0F4AE4E85F5210.text	A0D648F9E9225739BB0F4AE4E85F5210.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Marasmius variegatus Chun Y. Deng & Qi Zhao 2026	<div><p>Marasmius variegatus Chun Y. Deng &amp; Qi Zhao sp. nov.</p><p>Fig. 4</p><p>Chinese name.</p><p>杂色小皮伞 (za se xiao pi san).</p><p>Etymology.</p><p>“ variegatus ” refers to its white pileus mottled with irregular yellowish-brown areas.</p><p>Holotype.</p><p>China • Yunnan Province: Puer City, Jingdong Yi Autonomous County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.020004&amp;materialsCitation.latitude=24.534721" title="Search Plazi for locations around (long 101.020004/lat 24.534721)">Ailao Mountains</a>, 24°32'05"N, 101°01'12"E, alt. 2512 m, 25 Aug. 2025, collected by Tai-Shun Li, HKAS 154773 (LTS 131)</p><p>Diagnosis.</p><p>Marasmius variegatus is characterized by a pileus with uneven coloration, subdistant, narrow and branched lamellae, a stiff stipe, medium-sized basidiospores, pileipellis and cheilocystidia composed of two cell types, and relatively long caulocystidia. It differs from M. ochroleucus by its crowed and close lamellae, and the presence of two-types of cells in both cheilocystidia and pileipellis.</p><p>Description.</p><p>Basidiomata medium-sized. Pileus 0.8–3 cm, broadly convex to applanate, white with light orange (5 A 5), and grayish orange patches (5 B 4, 5 B 6), hygrophanous, faintly striate, margin recurved with age. Lamellae adnate, subdistant, L = 11–12 (l = 2–5), white, narrow, frequently forked. Stipe 40–60 × 1–1.5 mm, central, cylindrical, white when young, gradually becoming grayish orange (5 B 6), golden yellow (5 B 7, 6 B 7), reddish golden (6 C 7), base with thick, robust mycelium, brownish orange (6 C 8).</p><p>Basidiospores 9–12 × 4–5.3 µm, Q = 1.8–2.8, mean 10.28 [± 0.98 SD] × 4.66 [± 0.34 SD] µm, Qm = 2.22 [± 0.28 SD], N = 60 from three collections, ellipsoid, thin-walled, smooth, inamyloid. Basidia 27–35 × 5–7.8 µm, clavate, 2 - or 4 - spored. Basidioles fusoid, cylindrical to clavate, hyaline, thin-walled, inamyloid. Pleurocystidia absent. Cheilocystidia similar to the cells of the pileipellis: 1) Siccus-type cells, hyaline when grouped; main body 12–28 × 6–15 µm, clavate to pyriform, thin-walled, hyaline, inamyloid; diverticula or setulae apical, short and somewhat divergent, 1–2.5 × 0.6–1 µm, verruciform to cylindrical, digitiform, regular in outline or a little monilioid, many times vesiculose-rounded or balloon-shaped, abundant to scarce, solid, apex obtuse and rounded; 2) Globulares-type cells, 10–24 × 6–13 µm, more thin-walled. Lamellae trama inamyloid, irregular, hyphae cylindrical, 1.5–5 µm diam., regular in outline, hyaline, branched, smooth, thin-walled. Pileipellis hymeniform, composed of two cellular types: 1) Siccus-type broom cells, main body 15–24 × 6–12 µm, pyriform to cylindrical-clavate, hyaline, thin-walled; setulae apical, erect, elongate to very short, 1–3 × 1–2 µm, cylindrical, digitiform or verruciform, regular or irregular in outline, abundant or scarce, solid or with lumen, apex acute to obtuse; 2) Globulares-type smooth cells, more numerous than the first type at the mature basidiomata, 16.3–26.3 × 12.5–17.5 µm, pyriform or balloon-shaped, vesiculose to clavate, or irregular in outline, hyaline, smooth or apical portion with thicker walls, dextrinoid. Caulocystidia 15–70 × 5–10 µm, clavate, fusoid, irregular in outline, obtuse, hyaline, smooth, inamyloid. Clamp connections present in all tissues.</p><p>Habitat.</p><p>Solitary or gregarious on well-rotted bark or wood.</p><p>Additional collection examined.</p><p>China • Yunnan Province: Puer City, Jingdong Yi Autonomous County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.02778&amp;materialsCitation.latitude=24.540556" title="Search Plazi for locations around (long 101.02778/lat 24.540556)">Ailao Mountains</a>, 24°32'26"N, 101°01'40"E, alt. 2603 m, 25 Aug. 2022, collected by Jin-Rong Lu, KHAS 135842 (LJR-210) ; • ibid, 24°32'11"N, 01°01'56"E, alt. 2606 m, 18 Jul. 2024, Tai-Shun Li, HKAS 154774 (LTS 313, paratype) .</p><p>Notes.</p><p>A basepair comparison between the holotype sequence of the new species and the closely related species Marasmius ochroleucus Desjardin &amp; E. Horak (specimen voucher: LE 295978), revealed 96.17 % identity with 26 substitutions / indels in ITS, and 98.95 % identity with 9 substitutions / indels in LSU, supporting its recognition as a distinct species. Based on molecular phylogenetic evidence, Marasmius variegatus belongs to subg. Globulares ser. Luteoli (Oliveira et al. 2020; Oliveira et al. 2024). The new species forms an independent clade and is the sister taxon of M. ochroleucus . The latter species has only one type of broom cell in the pileipellis and cheilocystidia, as well as densely crowded lamellae. Comparing the cream, hygrophanous pileus, the new species is similar to Marasmius pseudopellucidus Wannathes and M. strobiluriformis Antonin, R. Ryoo &amp; H. D. M. pseudopellucidus has larger basidiospores (Xm = 11 × 4.1 µm), grows gregariously on bamboo debris and possesses only siccus-type cheilocystidia (Wannathes et al. 2009 b). Marasmius strobiluriformis differs by the pure white, smaller pileus, more crowded lamellae, and rostrate cheilocystidia (Antonín et al. 2010 b).</p></div>	https://treatment.plazi.org/id/A0D648F9E9225739BB0F4AE4E85F5210	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Deng, Chun-Ying;Li, Tai-Shun;Xiang, Zhun;Zhao, Qi	Deng, Chun-Ying, Li, Tai-Shun, Xiang, Zhun, Zhao, Qi (2026): Three new species of Marasmius (Agaricales, Basidiomycota) from southwestern China. MycoKeys 136: 233-250, DOI: 10.3897/mycokeys.136.196875
