taxonID	type	description	language	source
873C87B0C931FFC4FC5EFB3514420206.taxon	materials_examined	AUSTRALIA: WESTERN AUSTRALIA, NORTHERN TERRITORY. There are records from various caves (Moulds, 2004; Scholtz, 1986 b). Four of the seven Omorgus alatus specimens available to Scholtz (1986 b) originated from caves or rock shelters, leading him to speculate that this species may have adapted to life in sheltered situations, thus relieving the beetles from the necessity to burrow into soil for protection. Hamilton-Smith et al. (1989: 20) concluded that this “ extremely rare ” species seems to be “ more or less confined to cave environments, ” while Moulds (2004) classified it as a troglophile and perhaps a guanophile.	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C931FFC4FC56F98815790C06.taxon	materials_examined	AUSTRALIA: WESTERN AUSTRALIA, SOUTH AUSTRALIA. Omorgus amictus has only been recorded from the Nullarbor Plain in southern Australia (Scholtz, 1986 b), from where are numerous records from various caves (Moulds, 2004; Richards, 1971; Scholtz, 1986 b). It has been collected from both the entrance zone and the dark zone of mostly shallow caves, but the species has also been taken from the soil surface and from rabbit warrens (Richards, 1971). Richards (1971) found that Omorgus amictus is in caves associated with bird guano, but not with bat guano. Outside caves, it has also been found on carcasses and skins. Since they can survive equally on the surface and in caves, it should be considered a troglophile (Moulds, 2004; Richards, 1971). Since its association with bird (not bat) guano is facultative, it is also categorised as a (bird-) guanophile (Moulds, 2004).	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C931FFC4FFEDF95513C50C1D.taxon	materials_examined	SOUTH AFRICA: WESTERN CAPE. Specimens were collected in the dark zone of the Table Mountain caves on the Cape Peninsula. Microchiropteran bats were present throughout the dark zone of the cave where the beetles were collected (Sharratt et al., 2000). We interpret this as an incidental occurrence of the Trogidae in a cave.	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C931FFC4FC70FF1A1490067D.taxon	description	(Fig. 1 B)	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C931FFC4FC70FF1A1490067D.taxon	materials_examined	DEMOCRATIC REPUBLIC OF THE CONGO: HAUT-KATANGA. Scholtz (1980) reported an undisclosed number of Phoberus pusillus specimens from the Kakontwe and Mwanga Caves in the vicinity of Likasi. These beetles were collected by Narcisse Leleup (1912 – 2001), unrivalled collector of soil insects, but also a collector of bats in search of their ectoparasites (Van Cakenberghe et al., 2017). No fewer than 18 bat species have been recorded from these two caves, represented by both insect-feeders and fruit-eaters (Van Cakenberghe et al., 2017). We interpret this as that Phoberus pusillus is a troglophile, and probably a bat-guanophile.	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C931FFC4FE9BFE4510B90693.taxon	materials_examined	UNITED STATES OF AMERICA: GEORGIA. The species was reported to “ apparently live ” in bat guano in a cave (Reeves et al., 2000: 175). Slay et al. (2012) considered this to be an accidental collection since there is only this single record from a cave. Trox aequalis, widespread through the Nearctic Region, is usually associated with nests of various mammals and birds (Vaurie, 1955) and not with caves or guano, supporting the contention of Slay et al. (2012).	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C931FFC4FEB2FD15138707EE.taxon	materials_examined	UNITED STATES OF AMERICA: IOWA, ARKANSAS. This scarce American species is almost exclusively found in fox burrows (Robinson, 1941; Vaurie, 1955). Peck & Christiansen (1990) presented a single record from a cave, themselves denoting it as an accidental collection, something with which Slay et al. (2012) agreed. Robinson (1941) reported four individuals of Trox laticollis which were feeding on a dead fox in a cave; we suspect that the common denominator is the fox, and not the cave.	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C931FFC4FEB5FC2010D901B6.taxon	description	(Fig. 1 A)	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C931FFC4FEB5FC2010D901B6.taxon	materials_examined	SPAIN; UNITED STATES OF AMERICA: MISSOURI. This widepread Palaearctic species has spread to the New World and to Australasia (Pittino & Bezděk, 2016); it has been denoted as subcosmopolitan, among others by Zidek (2013). Besides feeding on animal carcasses, this species is often found in various bird and mammal nests and burrows (e. g. Vaurie, 1955; Hicks, 1959). Trox scaber has been recorded from caves in both Western Europe and North America (Gardner, 1986; Tinaut et al., 2008; Pérez et al., 2011). We call into question the categorisation by Slay et al. (2012) of this species being only accidentally in caves; these authors overlooked the reports from Europe.	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C930FFC5FF48FF1813FD050B.taxon	materials_examined	AUSTRALIA: QUEENSLAND, NEW SOUTH WALES. Scholtz (1986 b: 35) reported that this species “ is frequently taken in caves. ” It may hence indeed be a troglophile; no other relevant information exists.	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C930FFC5FF5CFEBD13B606D3.taxon	materials_examined	UNITED STATES OF AMERICA: NEW MEXICO. Vaurie (1955) reported a specimen of Omorgus carinatus from ‘ Carlsbad Cavern, ’ but from only the reported label data it is impossible to tell whether that specimen was collected inside the cave system itself, or from the wider Carlsbad Caverns National Park. Cokendolpher & Polyak (1996: 190) reported a single individual from the “ twilight / dark junction ” of a cave. Omorgus carinatus appears to be associated with nests or middens of woodrats (Neotoma Say & Ord, Cricetidae) (Loomis, 1922; Slay et al., 2012), while woodrats are commonly found in caves (e. g. Tweet et al., 2012). The foregoing facts lead Slay et al. (2012) to categorise the presence of this beetle species in a cave to be accidental.	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C930FFC5FF62FCD510DF00D3.taxon	description	(Fig. 1 C)	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C930FFC5FF62FCD510DF00D3.taxon	materials_examined	AUSTRALIA; INDONESIA; MALAYSIA. This species is widespread through the Oriental and Australasian Regions (Scholtz, 1986 b; Strümpher & Kalawate, 2023). It is often encountered in caves, where it has repeatedly been recorded feeding on, burrowing into, and breeding in bat guano (Blair, 1929; Chapman, 1983; Leefmans, 1932; McClure et al., 1967; Moulds, 2004; Scholtz, 1986 b). McClure et al. (1967) reportedthat adults, larvae and pupae of the species lived inside the guano of a Malaysian cave and was rarely seen on the surface. Arrow (1927) additionally reported that Omorgus costatus was collected from bat guano in hollow trees in Malaysia. Moulds (2004) categorised this trogid species as a troglophile and possibly a guanophile; we confirm here that the species is indeed bat-guanophilic, since narrow associations with bat guano have been observed repeatedly.	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C930FFC5FF68FAD510230195.taxon	materials_examined	MEXICO: MORELOS. Muñiz Vélez (2001) reported fragments of one specimen of Omorgus fuliginosus from an archaeological site inside a cave. We consider this to be an incidental occurrence.	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C930FFC5FEBCFA1A10C402E5.taxon	materials_examined	MALAYSIA; INDONESIA. All specimens of Omorgus mollis available or known to Scholtz (1986 b) were collected in bat guano in hollow trees. If this observational pattern persists, Omorgus mollis may uniquely be a non-cavernicolous bat-guanobiont. Presently this seems to be an unlikely life history style — more field observations and collecting of this rare species are required, as well as an open mind that Nature contains secrets which are unlikely but — in fact — true.	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C930FFC5FF56F92A135503BB.taxon	materials_examined	UNITED STATES OF AMERICA: CALIFORNIA. A single specimen was trapped near an entrance to a cave (Aalbu, 1990). Aalbu (1990: 9) considered this an “ accidental cavernicolous ” beetle. Slay et al. (2012) overlooked this report.	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C930FFC5FF61F86D13CB0C1B.taxon	materials_examined	MEXICO: MORELOS. Muñiz Vélez (2001) reported fragments of one specimen of Omorgus rubricans from an archaeological site inside a cave. We consider this to be an incidental occurrence.	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C930FFC5FC78FF18142E06B6.taxon	materials_examined	UNITED STATES OF AMERICA: TEXAS; BRAZIL: TOCANTINS. Omorgus suberosus is an abundant, generalist species, ubiquitous throughout the New World (Vaurie, 1962), and adventive and widespread in Australasia, Oceania and the Oriental Region (Cartwright & Gordon, 1971; Miquel, 2019; Scholtz, 1986 b). Specimens were collected in two Texan caves, in one of the caves on bat guano (Kohls and Jellison, 1948). Slay et al. (2012: 196) considered these to be “ accidental collections. ” Muñiz Vélez (2001) reported fragments of several Omorgus suberosus individuals from an archaeological site inside a Mexican cave, while Costa-Silva et al. (2021) reported a specimen, bearing a label with the locality ‘ Casa de Pedra Cave, ’ in central Brazil, but without ecobiological remarks. We consider these also to be incidental occurrences.	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C93FFFCAFC57F9D2151302D3.taxon	description	(Fig. 1 E)	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C93FFFCAFC57F9D2151302D3.taxon	materials_examined	BRAZIL: PARÁ. A number of specimens were collected from bat guano in a cave (Correa et al., 2022). In the opinion of Correa et al. (2022), this species may truly be associated with caves. We agree and consider Omorgus batesi as a troglophile and bat-guanophile.	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C93FFFCAFF71FECA104B0C1B.taxon	description	(Fig. 1 D)	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C93FFFCAFF71FECA104B0C1B.taxon	description	Omorgus lindemannae clearly is a troglophile. Its relation to guano has not yet been reported, but we cautiously suggest that it may be a bat-guanophile, based on the enormous deposits of bat guano in both these caves. Pittino (2005: 75) denoted Omorgus lindemannae to be a “ very rare species. ” These are new records of a trogid species in a cave and possibly associated with bat guano. The collecting and other label data of the Omorgus lindemannae museum specimens discovered by us are presented below. The first four entries represent the five cave specimens. The entries as a whole define a further range extension of this trogid (e. g. Frey, 2009). Ever since its two descriptions (Petrovitz, 1975; Scholtz, 1980, as synonym), this species had been known only from two localities in south-central Tanzania (Lindi and Rukwa Regions), one specimen from each locality. Pittino (2005) reported the next known specimen, from the Limpopo province of South Africa, which represented a huge range extension. Our newly detected specimens further extend the known distribution of Omorgus lindemannae to Botswana and to the KwaZulu-Natal province of South Africa.	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C93FFFCAFF71FECA104B0C1B.taxon	materials_examined	BOTSWANA North West District – 1 ♀ Gcwihaba Cave [as “ Drotsky's Cave ”], - 20.024 °, 21.354 °, 1969 - 10 - 03, W Haacke & O Prozesky || Trox zumpti Haaf, det. CH Scholtz, 1980 (TMSA). — Misidentified as Omorgus zumpti by Scholtz (1980: 36). New country record. SOUTH AFRICA Limpopo – 1 ♂ Kruger National Park, Luvuvhu River, Lanner Gorge Cave [as “ Cave N 2, Pafuri ”], ca. - 22.450 °, 31.150 °, 1983 - 08 - 29, L Braack (TMSA). – 1 ♂ Kruger National Park, Luvuvhu River, Lanner Gorge Cave [as “ Lanner Gorge, Cave N 2 ”], ca. - 22.450 °, 31.150 °, 1983 - 09 - 26, L Braack (SANC). – 2 ♀ Kruger National Park, Luvuvhu River, Lanner Gorge Cave [as “ Pafuri cave ”], ca. - 22.450 °, 31.150 °, 1993 - 12, CH Scholtz (TMSA). Mpumalanga – 1 ♀ Kruger National Park, Skukuza, - 25.00 °, 31.59 °, 1995 - 02 - 25, UV light trap, S Endrödy-Younga (TMSA). New provincial record. KwaZulu-Natal – 1 ♀ Ndumo Game Reserve, ca. - 26.87 °, 32.25 ° [centroid], 1960 - 12, G van Son || Trox zumpti Haaf, det. CH Scholtz, 1980 (TMSA). — Misidentified as Omorgus zumpti by Scholtz (1980: 36). New provincial record.	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C93FFFCAFC50FF18151801AE.taxon	materials_examined	SOUTH AFRICA: LIMPOPO. Braack (1989) surveyed Lanner Gorge Cave, a remote cave in the far north of the Kruger National Park, and described its macro-arthropod fauna. He estimated that more than 2,400 adult individuals of Omorgus zumpti were simultaneously present on the cave floor, mostly in the twilight zone. This beetle species is widespread in southern Africa, but rarely collected (Scholtz, 1980). Lanner Gorge Cave is inhabited by a large, permanent colony of fruit bats, the entire cave floor being covered with bat guano, in its proximal half more than a metre deep (Braack, 1989). Despite the abundance of bat guano, Braack (1989) did not report any interaction between the Trogidae and the guano, but rather reported that adult beetles were on several occasions observed feeding on dead bats. He reported this circumstance both in his text and in a schematic representation of trophic interactions in the cave (his Figure 2). We consider it unlikely that the Trogidae would not feed on the abundance of bat excrement, hence we suggest that Omorgus zumpti perhaps indeed is bat-guanophilic, while clearly being a troglophile.	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C93FFFC8FC07F8ED11D304C9.taxon	description	(Fig. 1 F)	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C93FFFC8FC07F8ED11D304C9.taxon	materials_examined	BRAZIL: MINAS GERAIS. Several specimens were sampled from bat guano in caves (Correa et al., 2022). Correa et al. (2022) found this species to have been the most abundant scarab species in their survey of Scarabaeoidea in Brazilian caves. They argued that this species may also be truly associated with caves, and suggested that the species may be classified as a troglophile. Necessarily it also seems to be bat-guanophilic. the species was captured inside the cave or in the vicinity of the cave. False record of a trogid species being associated with bat guano	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
873C87B0C93DFFC8FEBBFEFD11D30035.taxon	materials_examined	AUSTRIA; FRANCE. Numerous instances of specimens from woodpecker nests secondarily occupied by starlings or “ bats ” [sic!] were reported by Petrovitz (1969: 104). From Petrovitz’s short report it cannot confidently be concluded that the associated vertebrates were bats indeed: it may have been only birds, or both birds and bats. Byk et al. (2019) comprehensively reviewed the ecology of Trox perrisii without adding any information about this species being associated with either bats or caves, besides misinterpreting Petrovitz’s (1969) report therein that they categorically stated that Petrovitz’s woodpecker hollows were secondarily occupied by bats, not birds. We disagree that such a bat association can be confirmed from existing literature. Byk et al. (2019) confirmed that this beetle species inhabits the nests of several bird species, as well as hollows in trees, with or without bird inhabitants. Trox nidicola Bonnaire, 1881 is a junior synonym of Trox perrisii: the description of this nominal species was based on a small number of specimens found among debris in a bird nest in a tree hollow (Bonnaire, 1881). There is — in our view — no unambiguous evidence that Trox perrisii is associated either with caves or with bat guano. We suggest that Trox perrisii be deleted from the corpus of bat-guano-associated Trogidae. The species does seem to be bird-guano-associated. It is not a troglophile.	en	Strümpher, Werner P., Stals, Riaan (2025): A world list of known cave- and bat guano-associated Trogidae (Coleoptera: Scarabaeoidea), and further range extension of Omorgus lindemannae (Petrovitz). Faunitaxys 13 (9): 1-11, DOI: 10.57800/faunitaxys-13(09), URL: http://dx.doi.org/10.1186/s40360-025-00930-2
