identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
74BB3C187CA859B09B5946A46756B7D9.text	74BB3C187CA859B09B5946A46756B7D9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cleonardo compassionate Lörz & Engel & Jereczek & Ćwierz & Jażdżewska 2026	<div><p>Cleonardo compassionate sp. nov.</p><p>Figs 5, 6, 7, 8, 9</p><p>Eusiridae ADF 7841: Jażdżewska and Mamos 2019: fig. 2 A.</p><p>Type material.</p><p>Holotype: Pacific • mature female, 10 mm, SMF 62814, COI: PQ 734425, body remnants and one slide with appendages; Clarion-Clipperton Zone, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-117.32&amp;materialsCitation.latitude=12.251" title="Search Plazi for locations around (long -117.32/lat 12.251)">OMS exploration contract area</a>, R / V Thompson, ABYSSLINE- 2, EBS, AB 2 - EB 06, 01/03/2015, 12°15.06'N, 117°19.2'W, 4137 m . Paratype: Pacific • mature male, 10 mm (urosome broken off), SMF 62815, COI: PQ 734442, Clarion-Clipperton Zone, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.22193&amp;materialsCitation.latitude=11.832967" title="Search Plazi for locations around (long -116.22193/lat 11.832967)">BGR exploration contract area</a>, R / V Sonne, MANGAN 2018, EBS, SO 262-150, 07/05/2018, 11°50.009'N, 116°14.780'W – 11°49.978'N, 116°13.316'W, 4074–4095 m .</p><p>Additional material.</p><p>Pacific • Clarion-Clipperton Zone; BGR exploration contract area, several stations • abyss adjacent to Kuril-Kamchatka Trench, one station (Table 2).</p><p>Type locality.</p><p>Abyssal Pacific Ocean, Clarion-Clipperton Zone, 12°15.06'N, 117°19.2'W, 4137 m.</p><p>Description.</p><p>Body (Figs 5, 6): dorsally smooth.</p><p>Head (Figs 5, 6): rostrum slightly curved and 1 / 3 length of peduncle 1 of antenna 1; head lobes rounded, slightly produced.</p><p>Eyes (Figs 5, 6): present, small, rounded, white in preserved specimen.</p><p>Antenna 1 (Fig. 7): ~ 1 / 2 body length, same length as antenna 2; article 1 of antenna 1 length 2 × as long as broad and extended into a blunt point; article 2 thinner and longer than article 1 and ending in a dentate extension; simple and plumose setae on articles 1 and 2; article 3 short, but longer than first segment of flagellum; accessory flagellum uni-articulated, thin and as long as first article of flagellum; flagellum 69 - articulated and longer than articles 1, 2, and 3 combined, articles of flagellum short except for first article which is as long as next four articles, and last six articles which are longer than previous articles, with the last one the longest; calceoli on posterior margin of flagellum, no calceoli on last six segments; short simple setae along flagellum.</p><p>Antenna 2 (Fig. 7): ~ 1 / 2 body length; articles 1–3 short, with plumose setae; article 3 length 1 / 5 of article 4; article 4 broader and slightly longer than article 5, both articles with simple and plumose setae as well as calceoli on posterior margin; flagellum 46 - articulated, shorter than articles 4 and 5 combined; articles short except for first article which is as long as next four articles, and last three articles longer than previous ones; calceoli on posterior margin of flagellum, but no calceoli on last four articles; short setae on flagellum.</p><p>Mandibles (Fig. 7): palp tri-articulated, article 1 length 1 / 3 of article 2; article 2 of left mandible with many short setae on outer margin; article 3 little shorter than article 2; article 3 with one strong pectinated setae on lateral side and at apex; articles 2 and 3 with long setae and additionally with few plumose setae on tip of article 3; right lacinia mobilis tridentate, incisor is one long tooth plate; mandible corpus with five dentated setae and one normal thin seta, left lacinia six-dentate, mandible corpus with row of four dentated setae and one thin normal one; molar tubercle of left mandible prominent with dozens of denticles, and carrying a small plumose setae at the lower corner; molar of right mandible missing.</p><p>Maxillae 1 (Fig. 7): inner lobe of right maxilla 1 missing; inner lobe of left maxilla 1 not reaching length of outer lobe with two plumose setae; outer lobe with 11 spine-teeth, first row with four spine-teeth of which three are six-dentate and one uni-dentate, second row with seven spine-teeth of which six are more than ten-dentate and one uni-dentate.</p><p>Maxillae 2 (Fig. 7): inner lobe broader and reaching length of outer; inner lobe with short smooth setae and plumose setae, additionally one stronger plumose seta; posterior and lower anterior margins with very thin setae; outer lobe with long simple setae on tip and long plumose ones on inner margin.</p><p>Maxilliped (Fig. 7): inner lobes nearly reaching length of article 1 of palp with three teeth and thick plumose setae; outer lobes reaching 1 / 2 of article 2 of palp, with many strong setae along inner margin, setae on apex and posterior margin longer and plumose; palp four-articulated; article 1 length 1 / 2 of article 3; articles 2 and 3 with longer plumose setae; dactylus with seven strong setae.</p><p>Upper lip (Fig. 7): entire, ventral margin broadly rounded, with fine setae.</p><p>Lower lip (Fig. 7): broad, outer lobes with little incision.</p><p>Pereon: Gnathopod 1 (Fig. 8): slightly smaller than gnathopod 2 but of a similar shape, both strongly subchelate; coxa approx. same size as following coxae, strongly widening distally, axe-shaped, anterior corner rounded, slightly produced, posterior corner rounded, little notch on posterior and anterior corner, surface of coxa with few small spines; basis long and elongated, with many plumose and smooth setae; merus with long plumose setae; carpal lobe broad, slightly produced, but not reaching palmar angle, carpal lobe with eight rows of ~ 8 plumose setae, the first row with only two smooth setae the other rows with eight plumose setae; subovate propodus with few setae on posterior margin and apex; palm with plumose setae, shorter smooth setae and seven spines; palmar angle with nine spines of different sizes (two of these are broken and seem to have been very long like in other Cleonardo species); dactylus with eight small spines.</p><p>Gnathopod 2 (Fig. 8): larger than gnathopod 1; coxa nearly rectangular, anteroproximal corner produced, anterodistal corner rounded, with one notch that carries one thin seta, posterior margin straight with one stronger spine on upper half, posterodistal corner with two notches, each carrying one seta, surface of coxa with small spines; basis longer and a bit wider than in gnathopod 1; carpal lobe is more elongated and nearly reaching palmar angle, with eight rows of setae, with the first row only consisting of one smooth seta, the other seven consisting of eight plumose setae; propodus more elongated and more egg-shaped, palm with seven strong spines and shorter smooth and plumose setae, palmar angle with ten strong spines; dactylus broken off.</p><p>Pereopod 3 (Fig. 9): coxa similar to coxa 2, with straight anterior and posterior margins and one little notch at anterodistal corner and two at posterodistal corner, with strong spine proximally on posterior margin, surface of coxa with many small spines; basis long and slender, with short setae on inner, and long setae on outer and inner margins; merus 2 / 3 length of basis, with long plumose setae and short smooth ones; next articles broken off.</p><p>Pereopod 4 (Fig. 9): coxa 4 anterior margin straight, rounded below, posterior margin excavated and more pointed, surface of coxa with many small spines; similar to pereopod 3; basis is slender and long, with many smooth and plumose setae; merus shorter than basis but with longer plumose setae; next articles broken off.</p><p>Pereopod 5 (Figs 5, 6): coxa 5: bilobed, with anterior lobe slightly shorter than posterior lobe; subsequent articles missing.</p><p>Pereopod 6 (Fig. 9): coxa 2 × as broad as long, corners rounded, little notch on anterior margin; basis large and ovate, anterior margin serrated and with spines, surface of basis with spines and row of long setae, posterodistal corner produced along ischium; merus is damaged; carpus longer than propodus; dactylus longer than propodus, with setae along its margins and tip.</p><p>Pereopod 7 (Fig. 9): coxa anteriorly produced, nearly oval, corners rounded; similar to pereopod 6, but with slightly longer articles; basis large ovate, serrated on posterior margin, with little spines along the margins, surface of basis with row of long setae, posterodistal corner of basis drawn into lobe exceeding ischium; merus shorter than basis, with spines; carpus longer than propodus; dactylus longer than propodus, with setae along its margins and tip.</p><p>Pleon: Epimeral plate 1 (Figs 5, 6): anterior and posterior margins rounded, ventral margin convex; plate with small spines.</p><p>Epimeral plate 2 (Figs 5, 6): anterior margin concave, anterodistal corner rounded and produced, posterior margin convex, posterodistal corner rectangular; plate with small spines.</p><p>Epimeral plate 3 (Figs 5, 6): anterior margin less concave, anterodistal corner rounded, posterior margin slightly rounded, posterodistal corner slightly acute; plate with small spines.</p><p>Urosome: Urosomites 1–3 (Figs 5, 6): urosomite 1 longer than other two.</p><p>Uropod 1 (Fig. 8): lanceolate; peduncle longer than rami, dorsolateral and dorsomedial margins with eight and nine spines, respectively; inner ramus longer than outer; margins of rami and peduncle with spines on dorsolateral and dorsomedial margins with 11 and 15 spines, respectively; slightly shorter than telson.</p><p>Uropod 2 (Fig. 8): lanceolate; peduncle as long as outer ramus, with inner corner acute, dorsolateral and dorsomedial margins with four and nine spines, respectively; inner ramus longer than outer; dorsolateral and dorsomedial margins with 14 and 17 spines, respectively; barely exceeding telson.</p><p>Uropod 3 (Fig. 8): lanceolate; peduncle short, inner corner acute, dorsolateral and dorsomedial margins with four spines; inner ramus longer than outer dorsolateral and dorsomedial margins with eight and nine spines, respectively; barely exceeding telson.</p><p>Telson (Fig. 9): deeply cleft, ~ 80 %, narrowing distally, lobes slightly separating towards the tip.</p><p>Oostegite (Fig. 9): slender and long; serrated and with long setae.</p><p>Remarks.</p><p>Cleonardo compassionate sp. nov. shows the most similarity to Cleonardo daniela sp. nov., Cleonardo biscayensis Chevreux, 1908 and Cleonardo longipes Stebbing, 1888, but multiple differences are still present, as summarized in Table 3.</p><p>Further differences to C. longipes are: eyes present (vs absent); antenna 1, same length as antenna 2 (vs antenna 1 shorter than 2), article 2 with lateral dentate extension (vs no lateral dentation), accessory flagellum thin and as long as first article of flagellum (vs accessory flagellum shorter), flagellum 69 - articulated with first article of flagellum as long as the three following articles, articles of flagellum very thin (vs first article as long as five or six subsequent articles and not thin); antenna 2, articles 4 and 5 equal in length and both with calceoli on posterior margin (vs article 4 longer than article 5 and calceoli on anterior margin); mandible, palp article 1 longer, ~ 1 / 3 of article 2, article 3 shorter than article 2 (vs article 2 shorter); maxilliped, dactylus with 7 short strong setae (vs 10 setae); coxa 1, no spines on hind margin (vs one spine on hind margin); gnathopod 1, carpal lobe rounded and weakly produced (vs carpal lobe more elongated and nearly reaching palmar angle); gnathopod 2, carpal lobe more elongated than in gnathopod 1, but still quite short (vs carpal lobe similar as in gnathopod 1); pereopods 5–7, carpus longer than propodus, dactylus same length as propodus (vs dactylus 2 / 3 of length of propodus and carpus shorter than propodus); uropod 3 without plumose setae (vs with plumose setae). The differences between the new species and C. biscayensis are as follows: eyes present (vs absent); antenna 1, same length as antenna 2 (vs antenna 1 longer than 2); antenna 2 with calceoli on posterior margin (vs calceoli on anterior margin), flagellum shorter than articles 4 and 5 combined (vs flagellum much longer); mandible, palp article 1 longer, ~ 1 / 3 of article 2, article 3 shorter than article 2 (vs article 2 shorter than article 3); gnathopod 1, carpal lobe rounded and weakly produced (vs carpal lobe more elongated and nearly reaching palmar angle); pereopods 5–7, carpus longer than propodus, dactylus same length as propodus (vs carpus not longer than propodus); uropod 3 with both margins of both rami with multiple spines (vs uropod 3 with one additional strong spine on inner ramus).</p><p>Variation.</p><p>This species shows sexual dimorphism. Antennae 1 and 2 in males are longer than in females, article 1 of antenna 1 has long rows of brush setae (vs no brush setae in female). Additionally, males have calceoli on flagellum (Figs 11, 12).</p><p>Etymology.</p><p>The name ‘ compassionate ’ is used as a noun in apposition, defined as the feeling that arises when you are confronted with another’s suffering and feel motivated to relieve that suffering.</p><p>Distribution.</p><p>Abyssal Pacific Ocean, Clarion-Clipperton Zone in 4074–4352 m depth and Northwest Pacific Ocean, Kuril-Kamchatka Trench area in 5222–5223 m.</p><p>Molecular identification.</p><p>Following the definition given by Pleijel et al. (2008), the sequence of the holotype male of Cleonardo compassionate sp. nov. (SMF 62814, GenBank accession number PQ 734425) is designed as a hologenophore of all obtained sequences. The sequences of the paratypes and additional individuals of the species are also available in GenBank (Table 2). Furthermore, the species has received a Barcode Index Number from Barcode of Life Data Systems: BOLD: ADF 7841 (https://doi.org/10.5883/BOLD:ADF7841).</p></div>	https://treatment.plazi.org/id/74BB3C187CA859B09B5946A46756B7D9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lörz, Anne-Nina;Engel, Laura;Jereczek, Halina;Ćwierz, Nikodem;Jażdżewska, Anna M.	Lörz, Anne-Nina, Engel, Laura, Jereczek, Halina, Ćwierz, Nikodem, Jażdżewska, Anna M. (2026): Eusiridae (Crustacea, Amphipoda) of the Clarion-Clipperton Zone in the abyssal east Pacific with descriptions of five new species. ZooKeys 1274: 35-77, DOI: 10.3897/zookeys.1274.140610
4C6552D5A0165A6692A3BC876E09E712.text	4C6552D5A0165A6692A3BC876E09E712.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cleonardo daniela Lörz & Engel & Jereczek & Ćwierz & Jażdżewska 2026	<div><p>Cleonardo daniela sp. nov.</p><p>Figs 1, 2, 3, 4</p><p>Type material.</p><p>Holotype: Pacific • mature male, 6.5 mm, SMF 62810, COI: PQ 734433, body remnants and one slide with appendages; Clarion-Clipperton Zone; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.55&amp;materialsCitation.latitude=12.367" title="Search Plazi for locations around (long -116.55/lat 12.367)">UKSR- 1 exploration contract area</a>, R / V Thompson, ABYSSLINE- 2, EBS, AB 2 - EB 01, 18/02/2015, 12°22.02'N, 116°33'W, 4209 m . Paratypes: Pacific • immature male, 7 mm, SMF 62811, COI: PQ 734235, Clarion-Clipperton Zone; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.7&amp;materialsCitation.latitude=12.36" title="Search Plazi for locations around (long -116.7/lat 12.36)">UKSR- 1 exploration contract area</a>, R / V Thompson, ABYSSLINE- 2, EBS, AB 2 - EB 09, 10/03/2015, 12°21.6'N, 116°42'W, 4170 m • two individuals: mature male, 12 mm, SMF 62812, COI: PQ 734289, juvenile male, 5.5 mm, SMF 62813, COI: PQ 734766, Clarion-Clipperton Zone; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.29458&amp;materialsCitation.latitude=11.309075" title="Search Plazi for locations around (long -116.29458/lat 11.309075)">BGR exploration contract area</a>, R / V Kilo Moana, MANGAN 2023, EBS, KM 23-50, 01/05/2023, 11°17.7919'N, 116°18.8626'W – 11°18.5445'N, 116°17.6747'W, 4185– 4182 m .</p><p>Additional material.</p><p>Pacific • Clarion-Clipperton Zone; BGR exploration contract area, several stations (Table 2).</p><p>Type locality.</p><p>Abyssal Pacific Ocean, Clarion-Clipperton Zone, 12°22.02'N, 116°33'W, 4209 m.</p><p>Description.</p><p>Body (Fig. 1): dorsally smooth.</p><p>Head (Fig. 1): rostrum acute and curved, reaching 1 / 3 of length of first article of peduncle of antenna 1; head lobes rounded and not covered by coxa 1.</p><p>Eyes: present, small, rounded, white in preserved specimen.</p><p>Antenna 1 (Fig. 2): slightly shorter than antenna 2, weakly setose, reaching 1 / 2 the body; first article broad, extending to form two small teeth distally, same length as article 2; article 2 ending in acute extension distolaterally; few long plumose setae on article 1 and 2; article 3 shorter than first segment of flagellum with short setae; accessory flagellum uni-articulated, 1 / 2 the length of first segment of flagellum, with few thin setae on apex; flagellum 39 - articulated, slightly longer than articles 1–3 combined, segments very narrow with first segment as long as the five subsequent ones, last three segments longer than previous ones, few setae on both sides and calceoli posteriorly.</p><p>Antenna 2 (Fig. 2): weakly setose, reaching 1 / 2 the body; peduncular article 3 1 / 2 the length of article 4; article 4 longer and broader than article 5; article 5 slender; articles 4 and 5 with short normal and long plumose setae; calceoli on posterior margin of article 5 and flagellum; article 4 only with two or three calceoli at lower half; flagellum shorter than articles 4 and 5 combined, with few setae and 26 - articulated, first segment as long as nearly three of the subsequent ones and last three articles longer than the previous ones.</p><p>Mandibles (Fig. 2): palp tri-articulated, article 3 shorter than article 2; articles 2 and 3 with long setae on inner margins and each with few plumose setae apical of article 3, second article of palp of right mandible additionally with many short setae on outer margin, article 3 additionally with one strong pectinated setae on lateral side and another one on apex; left lacinia mobilis 7 - dentate, right lacinia 3 - dentate and bifid; accessory spine row 6 - dentate setae on the left and five on the right mandible, each with one additional normal long setae; molar triturative with dozens of denticles, and carrying a small plumose seta.</p><p>Maxillae 1 (Fig. 2): palp bi-articulated, exceeding outer lobe; outer lobe with nine or ten apical spine-teeth; inner lobe of left maxilla 1 missing; inner lobe of right maxilla not very broad, reaching length of outer lobe, apex with two plumose setae.</p><p>Maxillae 2 (Fig. 2): inner lobe 2 × as broad as outer, reaching height of outer lobe; thick setae on inner lobe reaching from apex to nearly the end of front margin; apex of outer lobe with setae that are&gt; 2 × length of setae of inner lobe.</p><p>Maxilliped (Fig. 2): inner lobes missing; outer lobe slightly serrated, with row of plumose setae on inner margins, reaching 1 / 2 the length of article 2 of palp, apex with four long plumose setae; palp four-articulated; article 2 broader and longer than other articles, inner margin with long setae; article 3 with rows of plumose setae; inner margin of dactylus with four setae-like spines.</p><p>Upper lip (Fig. 2): entire, ventral margin broadly rounded, with fine setae.</p><p>Lower lip (Fig. 2): broad, inner lobes weak, outer lobes larger; with many thin setae.</p><p>Pereon: Gnathopod 1 (Fig. 3): slightly smaller than gnathopod 2 but of similar shape, both strongly subchelate; coxa broadly rounded anteriorly, strongly widening distally, axe shaped, not covering head lobes; posterior margin almost straight, with small notch armed with one spine, as deep as broad; basis elongated, with few long setae; ischium and merus short with few setae; carpus not broad, posterior edge with few setae, carpal lope slightly elongated with four rows of four or five pectinated setae; subovate propodus with tuft of setae on upper hind margin and apex, palm curved, with simple and plumose setae and four spines, palmar angle with seven spines with one very long; dactylus with ten small stout setae on inner margin and two longer ones on tip.</p><p>Gnathopod 2 (Fig. 3): coxa rectangular, nearly as long as coxa 1; anterior and posterior lower angles with little notch; posterior margin proximally with one strong spine, basis with more long plumose setae than gnathopod 1 and with one pappose seta; carpal lobe less elongated, with four rows of seven or eight plumose setae; propodus more elongated and upper hind margin nearly straight, palm similar to gnathopod 1 but with four spines, palmar angle with six spines with one 2 × length of other spines; dactylus with 12 small stout setae on inner margin and two longer ones on tip.</p><p>Pereopod 3 (Fig. 4): coxa rectangular, rounded anteriorly, posterior angle with little notch and spine, with strong spine proximally on posterior margin; basis long and elongated; basis, ischium and merus with long plumose setae of different length; carpus same length as propodus but shorter than merus; dactylus same length as propodus and with simple setae.</p><p>Pereopod 4 (Fig. 4): coxa narrowing distally, rounded, posterior margin weakly excavated, anterior edge nearly straight; basis, ischium and merus similar to pereopod 3; carpus and subsequent articles broken off.</p><p>Pereopod 5 (Fig. 4): coxa &lt;1 / 2 length of coxa 4, rounded, broader than long; basis very broad, margins weakly serrated and with spines, anteroventral lobe small, posteroventral lobe very short; merus shorter than basis; carpus and following articles broken off, but as seen in paratype SMF 62812; propodus longer than carpus, dactylus 3 / 4 of length of propodus, inner margin of dactylus and propodus with row of small spines.</p><p>Pereopod 6 (Fig. 4): coxa short, with broadly rounded posterior lobe, posteroventral lobe reaching end of ischium, subsequent articles broken off, but as seen in paratype SMF 62812; basis to propodus similar to pereopod 5.</p><p>Pereopod 7 (Fig. 4): coxa as long as broad, nearly quadrate but upper margin slightly longer than lower and with rounded corners; basis ovate, with spines on margins and long setae in a midline, posterior margin with five serrations each with inset small spine, posteroventral lobe reaching end of ischium; ischium with one spine at anterior edge; merus same length as basis, with elongated posterior edge, that ends with two spines, shorter anterior edge with two spines; carpus to dactylus broken off, but as seen in paratype SMF 62812; propodus is longer than carpus.</p><p>Pleon: Epimeral plate 1 (Fig. 1): anterior and posterior margins rounded, ventral margin convex.</p><p>Epimeral plate 2 (Fig. 1): anterior margin concave with produced rounded edge, posterior margin rounded with acute corner, ventral margin nearly straight.</p><p>Epimeral plate 3 (Fig. 1): anterior margin similar to epimeral plate 2, posteroventral corner with small tooth, ventral margin convex.</p><p>Urosome: Urosomites 1–3 (Fig. 1): urosomite 1 slightly longer than urosomites 2 and 3 combined; urosomite 2 shortest.</p><p>Uropod 1 (Fig. 3): lanceolate; peduncle slightly longer than inner ramus, dorsolateral and dorsomedial margins with 12 and five spines, respectively; outer ramus 1 / 5 shorter than inner ramus, dorsolateral and dorsomedial margins with seven and two spines, respectively; inner ramus nearly reaching outer ramus of uropod 2, dorsolateral and dorsomedial margins with four and five spines, respectively.</p><p>Uropod 2 (Fig. 3): lanceolate; peduncle same length as outer ramus, dorsolateral and dorsomedial margins with five and two spines, respectively; outer ramus ~ 3 / 4 length of inner ramus, dorsolateral and dorsomedial margins with five and three spines, respectively.</p><p>Uropod 3 (Fig. 3): lanceolate; peduncle short,&gt; 1 / 2 length of inner ramus, with few spines; outer edge forming acute tip; outer ramus 3 / 4 length of inner ramus, dorsolateral and dorsomedial margins with three and six spines, respectively.</p><p>Telson (Fig. 4): deeply cleft, ~ 75 %; elongate, narrowing distally, with few spines, lobes nearly acute.</p><p>Remarks.</p><p>Cleonardo daniela sp. nov. is most similar to Cleonardo longipes Stebbing, 1888 but differs in the following characters: eyes present, antenna 1 shorter than antenna 2, corners of article 1 not produced, but forming two small teeth on lateral side, article 2 forming acute extension on outer lateral side nearly reaching 1 / 2 of article 3, uni-articulated accessory flagellum broader and nearly reaching 1 / 2 of first article of flagellum, first article of flagellum shorter than in Cleonardo longipes . Morphological differences to C. longipes and other closely related species are summarized in Table 3.</p><p>Etymology.</p><p>This species is named for Daniela Engel, the mother of the author LE. The name is used as a noun in apposition.</p><p>Distribution.</p><p>Abyssal Pacific Ocean, Clarion-Clipperton Zone in 4170 m – 4209 m depth.</p><p>Molecular identification.</p><p>Following the definition given by Pleijel et al. (2008), the sequence of the holotype male of Cleonardo daniela sp. nov. (SMF 62810, GenBank accession number PQ 734433) is designed as a hologenophore of all obtained sequences. The COI sequences of the single paratype and additional individuals are also available in GenBank (Table 2). Furthermore, the species has received a Barcode Index Number from Barcode of Life Data Systems: BOLD: AEB 2179 (https://doi.org/10.5883/BOLD:AEB2179).</p></div>	https://treatment.plazi.org/id/4C6552D5A0165A6692A3BC876E09E712	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lörz, Anne-Nina;Engel, Laura;Jereczek, Halina;Ćwierz, Nikodem;Jażdżewska, Anna M.	Lörz, Anne-Nina, Engel, Laura, Jereczek, Halina, Ćwierz, Nikodem, Jażdżewska, Anna M. (2026): Eusiridae (Crustacea, Amphipoda) of the Clarion-Clipperton Zone in the abyssal east Pacific with descriptions of five new species. ZooKeys 1274: 35-77, DOI: 10.3897/zookeys.1274.140610
89E71B3D794E50D5BE39F7F8E7FDBD40.text	89E71B3D794E50D5BE39F7F8E7FDBD40.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cleonardo Stebbing 1888	<div><p>Genus Cleonardo Stebbing, 1888</p><p>Type species.</p><p>Cleonardo longipes Stebbing, 1888 .</p></div>	https://treatment.plazi.org/id/89E71B3D794E50D5BE39F7F8E7FDBD40	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lörz, Anne-Nina;Engel, Laura;Jereczek, Halina;Ćwierz, Nikodem;Jażdżewska, Anna M.	Lörz, Anne-Nina, Engel, Laura, Jereczek, Halina, Ćwierz, Nikodem, Jażdżewska, Anna M. (2026): Eusiridae (Crustacea, Amphipoda) of the Clarion-Clipperton Zone in the abyssal east Pacific with descriptions of five new species. ZooKeys 1274: 35-77, DOI: 10.3897/zookeys.1274.140610
85D274B79E805028B451AF128F3FA8AF.text	85D274B79E805028B451AF128F3FA8AF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dorotea Corbari, Frutos & Sorbe 2019	<div><p>Genus Dorotea Corbari, Frutos &amp; Sorbe, 2019</p><p>Type species.</p><p>Dorotea papuana Corbari, Frutos &amp; Sorbe, 2019 (original designation).</p><p>Included species.</p><p>Dorotea aberrantis (Bellan-Santini &amp; Ledoyer, 1987); D. papuana Corbari, Frutos &amp; Sorbe, 2019; Dorotea elizae sp. nov.</p><p>Diagnosis.</p><p>(updated after Corbari et al. 2019). Body not strongly compressed, dorsally smooth. Rostrum short. Head lateral cephalic lobe weakly produced. Eyes lacking. Antennae medium size. Antenna 1 probably slightly longer than antenna 2, peduncular article 1 subequal to article 2, accessory flagellum uni-articulated, linear. Antenna 2, peduncular article 4 broader than article 5, flagellum not shortened. Upper lip evenly rounded distally. Lower lip, inner lobes weak. Mandible, incisor with long incurved cutting edge, poorly dentate or smooth; lacinia mobilis five-dentate; setal row with six or seven setae; palp, article 3 longer than article 2, slightly falciform, tapering distally; article 2 broader than article 3. Maxilla 1, inner plate with 1–3 subapical setae; outer plate with ten or 11 apical stout setae; palp slender, distal article longest. Maxilla 2, inner plate broader and shorter than outer plate. Maxilliped palp, articles ordinary; outer plate large; inner plate with three apical teeth. Coxae 1–4 regular, medium; coxa 1 broadly rounding, not produced; coxa 4 slightly excavate posteriorly. Coxae 5 and 6 bilobate. Gnathopods 1 and 2 subsimilar, both strongly subchelate; carpal lobes broad; propodus large, posterior margin short, palm oblique, margin spinose. Pereopods 3 and 4 slender, merus slightly longer than carpus, dactylus simple, stout and medium length. Pereopods 5–7 homopodous, subequal in form and size, not greatly elongate; bases broad; dactylus simple, stout and medium length. Epimera 1–3 regular, not serrate on posterior margin. Uropods 1 and 2 rami lanceolate, outer ramus shorter than inner ramus. Uropod 3 rami broadly lanceolate, outer ramus shorter than inner ramus. Telson elongate, lobate, weakly cleft. Brood plates on pereopods 2–5. Coxal gills on pereopods 2–7.</p><p>The genus Dorotea was created for Dorotea papuana Corbari, Frutos &amp; Sorbe, 2019 and also includes D. aberrantis (Bellan-Santini &amp; Ledoyer, 1987) previously assigned to Eusiroides (Bellan-Santini and Ledoyer 1987) . Our individual matches the generic diagnosis in most characters. However, seven characters appear to vary between species assigned to this genus (see Table 4).</p><p>Each of the species of Dorotea is known only from a unique individual of a different sex ( D. papuana from the female, for the other two species only males are known), so there is no possibility to infer the intraspecific variation. With the exception of the shape of the upper lip these characters need to be treated as species specific ones. There is a large difference in shape of the upper lip between our specimen and the other two Dorotea species described; however, due to the three-dimensional nature of this appendage the differences in observed shape may be associated with positioning during slide preparation. The lack of more individuals of any of the species prevents from decision how stable is this character; this is why the shape of the upper lip was left unchanged in the generic diagnosis.</p></div>	https://treatment.plazi.org/id/85D274B79E805028B451AF128F3FA8AF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lörz, Anne-Nina;Engel, Laura;Jereczek, Halina;Ćwierz, Nikodem;Jażdżewska, Anna M.	Lörz, Anne-Nina, Engel, Laura, Jereczek, Halina, Ćwierz, Nikodem, Jażdżewska, Anna M. (2026): Eusiridae (Crustacea, Amphipoda) of the Clarion-Clipperton Zone in the abyssal east Pacific with descriptions of five new species. ZooKeys 1274: 35-77, DOI: 10.3897/zookeys.1274.140610
2B8536E984315136BA6B274294C6F72A.text	2B8536E984315136BA6B274294C6F72A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dorotea elizae Lörz & Engel & Jereczek & Ćwierz & Jażdżewska 2026	<div><p>Dorotea elizae sp. nov.</p><p>Figs 13, 14, 15, 16</p><p>Type material.</p><p>Holotype: Pacific • male, 16 mm, SMF 62816, COI: PQ 734718, body remnants and two slides with appendages, Clarion-Clipperton Zone, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-117.32&amp;materialsCitation.latitude=12.251" title="Search Plazi for locations around (long -117.32/lat 12.251)">OMS exploration contract area</a>, R / V Thompson, ABYSSLINE-2, EBS, AB 2 - EB 06, 01/03/2015, 12°15.06'N, 117°19.2'W, 4137 m .</p><p>Type locality.</p><p>Abyssal Pacific Ocean, Clarion-Clipperton Zone, 12°15.06'N, 117°19.2'W, 4137 m.</p><p>Description.</p><p>Based on male, 16 mm, SMF 62816.</p><p>Body (Fig. 13): dorsally smooth.</p><p>Head (Fig. 13): almost as long as deep, as long as pereonites 1 and 2 combined; no eyes or ocular pigment visible; rostrum vestigial; interantennal lobe weak, subquadrate.</p><p>Antenna 1 (Fig. 14): long, as long as pereon and pleon segments combined (checked before dissection), length ratios of peduncle articles 1–3 1: 1: 0.3; article 1 with seven groups of slender setae and a few additional displaced long setae; article 2 with eight rows of slender setae and additional setae at the distal margin; article 3 with one group of short, slender setae and a few setae at the distal margin; flagellum ≥ 100 - articulated, a few distal ones broken off, almost every flagellar article with a few long, slender setae on ventral side, additional group of medial setae every few articles, calceoli present; accessory flagellum uni-articulated, reaching 2 / 3 of the first flagellar article, with three short distal setae.</p><p>Antenna 2 (Figs 13, 14): shorter than antenna 1, as long as head and pereon segments combined; article 4 length 2 × article 5; peduncular article 4 with nine rows of brush setae and at the distal margin; peduncle article 5 with four rows of short setae and group of long setae at the distal margin; flagellum longer than peduncle, ≥ 50 - articulated (distal part broken off), short setae placed distally on flagellar articles, calceoli present.</p><p>Upper lip (Fig. 14): entire, wider than long, apically emarginate, with submarginal row of setules. Epistome bluntly pointed.</p><p>Mandible (Fig. 14): incisor margins curved, with smooth cutting edge; left lacinia mobilis five-dentate; right lacinia mobilis narrower with indistinct teeth; accessory spine rows with seven narrow spines; molar columnar, strongly triturative, denticulate, one associated seta at left molar (at right molar probably broken off); palp tri-articulated, article 1 short, article 2 length 0.6 × article 3, with 11 posterodistal setae, article 3 slightly tapering and curved, seven (left) or eight (right) long setae on medial surface proximally, posterior margin with a row of ~ 70 setae of different lengths.</p><p>Lower lip (Fig. 14): outer lobes broadly rounded, mandibular lobes bluntly pointed; inner lobes small, separate; inner and outer lobes setulose apically.</p><p>Maxilla 1 (Fig. 14): inner plate subtriangular (partly damaged at right Mx), with two tiny, subapical setae; outer plate with 11 spine-teeth; palp bi-articulated, longer than outer plate, slender, rounded apically, article 1 length 0.5 × article 2, article 2 with 12 (right) or 14 (left) apical / subapical setae in two rows and one lateral seta.</p><p>Maxilla 2 (Fig. 14): left – lost, right – plates subequal in length, inner plate 2 × as wide as outer, with 19 setae apically and subapically organized in two rows, fine setules on the surface; outer plate rounded with ~ 15 apical setae.</p><p>Maxilliped (Fig. 15): strongly setose; inner plate subrectangular, reaching ~ 1 / 2 basal article of palp, apical margin with three blunt spines and eight longer setae; outer plate slightly curved, long, reaching ~ 1 / 2 length of palp article 2; palp four-articulated, strong; article 2 widening distally; article 3 expanded mediodistally, with long setae at the edge and on the surface; article 4 strong, slightly curved, with spine on its tip and four seta-like spines; length ratios of articles 1–4 1: 2.4: 1.8: 1.</p><p>Pereon: Pereonites 1–5 (Fig. 13) of similar length, pereonites 6 and 7 slightly longer.</p><p>Gnathopod 1 (Fig. 15): coxa subquadrate, slightly narrowing posteriorly; basis straight, slightly broadening distally, with a group of long setae at the surface at 1 / 3 of its length, two rows of medium length setae posteriorly at the distal 1 / 2 of the article and a group of setae at the distal margin; merus posterodistally with 15 setae; carpus expanded, subtriangular, anterior margin with three groups of setae, posterior lobe rounded with setae organized in six rows and a few displaced setae, two rows of five setae each on the surface; propodus subchelate, anterior margin with three groups of long setae, palm oblique, strongly setose with six large and one smaller spines along the margin and additional two at dactylus insertion; dactylus curved, as long as palm with a row of short setules along posterior margin.</p><p>Gnathopod 2 (Fig. 15): coxa subquadrate, apex rounded, ventral margin naked; basis straight, a group of long setae at the surface at in proximal part, posterior margin with irregularly placed setae of different length, a group of three medium length setae at anterodistal corner and of four setae at posterodistal corner; merus, posterodistally with 18 setae, additional group of three setae on the medial surface; carpus expanded, subtriangular, anterior margin with three groups of setae, posterior lobe rounded with setae organized in seven rows, two rows of four setae each on the surface; propodus subchelate, anterior margin with four groups of long setae, palm oblique, strongly setose with four large and one small spines along the margin and additional two below dactylus insertion; dactylus curved, slightly shorter than palm.</p><p>Pereopod 3 (Fig. 16): coxa subrectangular, ventral margin naked; basis long and narrow, length 6.9 × width, three setae along anterior margin, posterior margin with four medium length setae, two short and one medium length setae at posterodistal corner; merus slightly expanded distally, short setae in four groups along posterior margin, one longer seta at posterodistal corner; carpus, posteriorly armed with short setae irregularly placed; propodus posterior margin with eight groups of short setae and spines, one at the posterodistal corner; dactylus thin, slightly curved, with three setae along anterior margin; length ratio of articles 2–7: 1: 0.1: 0.55: 0.47: 0.7: 0.35.</p><p>Pereopod 4 (Fig. 16): coxa wider than deep, with a small posterodistal lobe; remaining articles similar to pereopod 3 except length ratio of articles 2–7: 1: 0.1: 0.57: 0.5: 0.7: 0.38.</p><p>Pereopod 5 (Fig. 16): coxa bilobed with weak posterior excavation, posterior lobe deeper than anterior lobe; basis weakly expanded, without posterodistal lobe, length 3 × width, traces of six stouter setae along anterior margin, trace of one seta at anterodistal corner, posterior margin slightly crenulated; merus length 0.6 × basis, with traces of two setae along anterior margin; carpus – dactylus broken off.</p><p>Pereopod 6 (Figs 13, 16): coxa bilobed, anterior lobe significantly smaller than long posterior lobe; weakly expanded, without posterodistal lobe, length 3 × width, nine short setae along anterior margin, one short seta at anterodistal corner, posterior margin slightly crenulated with short setules in the crenulations; merus length 0.7 × basis, anterior margin with short spines in four groups, some setae along posterior margin; carpus – dactylus broken off.</p><p>Pereopod 7 (Fig. 16): coxa small, subtriangular; basis weakly expanded anteriorly and tapering distally, without posterodistal lobe, length 2.8 × width, anterior margin with seven short setae, posterior margin slightly crenulated; merus length 0.7 × basis, with some setae and small spines along anterior and posterior margins, groups of two spines at both anterodistal and posterodistal corners; carpus – dactylus broken off.</p><p>Pleon: Pleonites 1–3 (Fig. 13): dorsally smooth.</p><p>Epimeral plates 1–3 (Fig. 13): anteriorly evenly rounded, posteriorly subquadrate.</p><p>Pleopods (Fig. 13): powerful, peduncles and rami long.</p><p>Urosome: Urosomites 1–3 (Fig. 13): dorsally smooth, urosomite 1 as long as urosomites 2 and 3 combined, the latter of the same length.</p><p>Uropod 1 (Fig. 16): lanceolate; peduncle subequal in length to inner ramus, dorsolateral and dorsomedial margins with nine and 11 spines, respectively, blunt distoventral process bearing a distal spine; rami of similar length, outer ramus slightly shorter than peduncle, dorsolateral and dorsomedial margins with six and eight spines, respectively.</p><p>Uropod 2 (Fig. 16): lanceolate; shorter than uropod 1, peduncle length 0.75 × inner ramus, dorsolateral and dorsomedial margins with two and five spines, respectively; inner ramus 1.2 × length of outer ramus, both rami bearing dorsolateral and dorsomedial spines.</p><p>Uropod 3 (Figs 13, 16): lanceolate; peduncle short, length 0.5 × inner ramus, dorsolateral and dorsomedial margins with none and seven spines, respectively; outer ramus probably slightly shorter than inner ramus (left: outer ramus broken at 1 / 5, right: outer ramus broken just before the tip; Fig. 13), dorsolateral and dorsomedial margins both with seven spines.</p><p>Telson (Fig. 16): narrowing distally, longer than peduncle of uropod 3, reaching 1 / 5 of inner ramus (checked before dissection), length 2.75 × width, cleft almost 40 %, lobes apically rounded but tips slightly damaged, a few setules on dorsal surface.</p><p>Variation.</p><p>The individual is unique so no intraspecific variation may be provided.</p><p>Remarks.</p><p>Apart from the morphological differences between already described species and D. elizae sp. nov. listed in Table 4, the new species differs from them in the length of dactyli of pereopods 3 and 4, the shape of bases of pereopods 5–7, the length of rami of uropod 3, and the size of cleft of the telson (Table 5). The recently published article by Weston et al. (2024) provided description of a new genus Dulcibella Weston &amp; González in Weston, González, Escribano &amp; Ulloa, 2024 closely related to the Dorotea . The morphological description of the new species of this genus, Dulcibella camanchaca (Weston &amp; González in Weston, González, Escribano and Ulloa 2024), was supplemented by the COI sequence. These authors mention in their work a “ non-public genetic record on BOLD from the Clarion-Clipperton Zone being ~ 90 % similar ” to their species (Weston et al. 2024). Indeed, the cited record is of Dorotea elizae sp. nov. and the p-distance reaches 0.095; however, generic placement of D. camanchaca is far from clear and warrants further study. Additional work may result in the synonymisation of the new genus with Dorotea as the characters used for its separation seem rather species-specific than being valid at the genus level. Also, the molecular study of Weston et al. (2024) does not fully support Dulcibella genus erection. Such work requires supplementary analyses and is beyond the scope of this paper. For this reason we retain the genus allocation as Dorotea herein. Due to the fact that the barcode of D. papuana was provided (Corbari et al. 2019) it was possible to also make a molecular comparison between that species and D. elizae sp. nov. The two available barcodes present 0.155 p-distance that strongly exceeds the threshold used to molecularly delimit amphipod species (e. g., Tempestini et al. 2018). Unfortunately, the remaining species D. aberrantis has not yet been barcoded. The present finding of Dorotea elizae sp. nov. extends the known geographic distribution of the genus to central East Pacific and also its deepest record to 4137 m.</p><p>Etymology.</p><p>The species is named for Eliza Kordula, the girlfriend of the author NĆ.</p><p>Distribution.</p><p>Known only from type locality, Abyssal Pacific Ocean, Clarion-Clipperton Zone, 4137 m.</p><p>Molecular identification.</p><p>Following the definition given by Pleijel et al. (2008), the sequence of the holotype male of Dorotea elizae sp. nov. (SMF 62816, GenBank accession number PQ 734718) is designed as a hologenophore of all obtained sequences. The species has also received the Barcode Index Number from Barcode of Life Data Systems: BOLD: AEB 4129 (https://doi.org/10.5883/BOLD:AEB4129).</p></div>	https://treatment.plazi.org/id/2B8536E984315136BA6B274294C6F72A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lörz, Anne-Nina;Engel, Laura;Jereczek, Halina;Ćwierz, Nikodem;Jażdżewska, Anna M.	Lörz, Anne-Nina, Engel, Laura, Jereczek, Halina, Ćwierz, Nikodem, Jażdżewska, Anna M. (2026): Eusiridae (Crustacea, Amphipoda) of the Clarion-Clipperton Zone in the abyssal east Pacific with descriptions of five new species. ZooKeys 1274: 35-77, DOI: 10.3897/zookeys.1274.140610
B898A958174F5CF1A02C0BAC388411D2.text	B898A958174F5CF1A02C0BAC388411D2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhachotropis clarionclippertoni Lörz & Engel & Jereczek & Ćwierz & Jażdżewska 2026	<div><p>Rhachotropis clarionclippertoni sp. nov.</p><p>Figs 17, 18, 19, 20</p><p>Type material.</p><p>Holotype: Pacific • female, 8 mm, SMF 62817, COI: PQ 734529, body remnants and two slides with appendages, Clarion-Clipperton Zone, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-117.237&amp;materialsCitation.latitude=12.038" title="Search Plazi for locations around (long -117.237/lat 12.038)">OMS exploration contract area</a>, R / V Thompson, ABYSSLINE- 2, EBS, AB 2 - EB 11, 14/03/2015, 12°2.28'N, 117°14.22'W, 4097 m . Paratype: Pacific • female, 8.5 mm, SMF 62818, COI: PQ 734352, Clarion-Clipperton Zone, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-117.4868&amp;materialsCitation.latitude=11.8307" title="Search Plazi for locations around (long -117.4868/lat 11.8307)">BGR exploration contract area</a>, R / V Kilo Moana, MANGAN 2016, EBS, Ma 16-91, 09/05/2016, 11°49.792'N, 117°30.458'W – 11°49.842'N, 117°29.208'W, 4344– 4344 m .</p><p>Additional material.</p><p>Pacific • Clarion-Clipperton Zone; BGR exploration contract area, several stations (Table 2).</p><p>Type locality.</p><p>Abyssal Pacific Ocean, Clarion-Clipperton Zone, 12°2.28'N, 117°14.22'W, 4097 m.</p><p>Description.</p><p>Based on holotype, female, 8 mm, SMF 62817.</p><p>Body (Figs 17, 18): all pereonites smooth; pleonites 1 and 2 smooth, pleonite 3 with a minute dorsal spine.</p><p>Head (Fig. 18): dorsally smooth, no eyes visible, rostrum short, slightly downcurved.</p><p>Antenna 1 (Fig. 18): peduncle article 2 narrower and longer than article 1,&gt; 2 × as long as article 3; flagellum eight-articulated; accessory flagellum uni-articulated.</p><p>Antenna 2 (Fig. 18): peduncle articles 4 and 5 equal in length; flagellum broken off.</p><p>Upper lip (Fig. 19): circular in shape, ventral margins rounded, lateral margins finely setose.</p><p>Mandible (Fig. 19): incisor process strongly dentate, well-developed; left lacinia mobilis denticulate; molar process conical, edge with row of strong spines; article 3 longer than article 2, articles 2 and 3 inner margin with long slender setae, plumose setae at apical tip of palp.</p><p>Lower lip (Fig. 19): setose anterodistally, outer lobes separated by broad gap.</p><p>Maxilla 1 (Fig. 19): inner plate bearing two subterminal plumose setae; outer plate with nine denticulate spines; article 1 of palp 1 / 3 length of second article, article 2 of palp with several slender setae.</p><p>Maxilla 2 (Fig. 19): inner and outer plates subequal in length, margins bearing stout and slender setae; inner plate slightly wider than outer plate.</p><p>Maxilliped (Fig. 19): inner plate distally with three strong spines; outer plate 2 × as long as inner plate, reaching almost 1 / 2 of article 2 of maxillipedal palp, outer plate margins and palp article setose.</p><p>Pereon: Gnathopod 1 (Fig. 18): coxa 1 anteriorly extended; gnathopods similar in shape, subchelate; gnathopod 1 slightly smaller than gnathopod 2; basis bearing small spines at posteroventral corner; ischium and merus with long setae at posteroventral corner; carpal lobe extending width of propodus, spines at distal end of lobe and along posterior margin; propodus widened, oval; dactylus slender, reaching end of palm.</p><p>Gnathopod 2 (Fig. 18): coxa 2 subquadrate; gnathopod 2 basis 1.2 × as wide as basis of gnathopod 1; ischium to dactylus similar to gnathopod 1, except carpal lobe longer and narrower.</p><p>Pereopods 3 and 4 (Fig. 20): coxae 3 and 4 subquadrate. Pereopod 3 articles long and narrow; carpus, propodus and dactylus in ratio of 1: 0.6: 0.96. Pereopod 4 similar to pereopod 3, carpus, propodus and dactylus broken off.</p><p>Pereopod 5 (Fig. 20): coxa 5 lobate; basis subrectangular; remaining pereopod missing.</p><p>Pereopod 6 (Fig. 20): coxa 6 lobate; basis subrectangular with small posteroventral lobe; merus longer than basis; remaining pereopod missing.</p><p>Pereopod 7 (Fig. 20): pereopod 7 longer than pereopods 5 and 6; basis slightly expanded, subovate; merus longer than basis; rest of pereopod missing.</p><p>Pleon: Epimera 1–3 (Figs 17, 18): epimeral plates ventrally rounded; epimeral plate 3 posterior margin and posteroventral corner minutely serrated.</p><p>Pleopods (Fig. 20): pleopod 2 (right side) rami with 12 and 15 articles; rami similar length, as long as peduncle.</p><p>Urosome: Urosomites 1–3 (Figs 17, 18): urosome segment 1 dorsolaterally with a small spine, urosomites 2 and 3 smooth.</p><p>Uropod 1 (Fig. 20): lanceolate; outer and inner ramus same length; rami shorter than peduncle.</p><p>Uropod 2 (Fig. 20): lanceolate; outer ramus shorter than inner ramus; peduncle shorter than outer ramus and longer than inner ramus.</p><p>Uropod 3 (Fig. 20): lanceolate; outer and inner ramus same length, nearly 2 × as long as peduncle.</p><p>Telson (Fig. 18): Telson&gt; 3 × as long as wide, with V-shaped cleft of ~ 15 %.</p><p>Remarks.</p><p>The combination of the following characters found in Rhachotropis clarionclippertoni sp. nov. (produced first coxa, cleft telson, rostrum shorter than 1 / 2 of first peduncle of antenna 1) is shared with the following species: R. americana Bousfield &amp; Hendrycks, 1995, R. antarctica K. H. Barnard, 1932, R. gubilata J. L. Barnard, 1964, R. palporum Stebbing, 1908, R. portoricana J. L. Barnard, 1964, and R. rossi Lörz, 2010, but only the new species and R. portoricana have no lateral teeth on pleonites 1–3. Rhachotropis clarionclippertoni sp. nov. is very similar to R. portoricana but can be distinguished by the following characters (characters of R. portoricana in parentheses): rounded ventrolateral margin of urosomite 1 (vs ventrolateral margin shows a drawn out process), rostrum reaching only 1 / 3 of the length of the cephalic lobe (vs rostrum reaching 1 / 2 length of the cephalic lobe), length to width ratio of basis of pereopod 6 is 0.4 (vs 0.6), length to width ratio of basis of pereopod 7 is 0.54 (vs 0.65), no posterodistal lobe on basis of pereopod 7 (vs small but distinct lobe), telson cleft 15 % (vs telson cleft 8 %).</p><p>Etymology.</p><p>The name clarionclippertoni refers to the Clarion-Clipperton Zone in the abyssal east Pacific, a marine area known for the high density of manganese nodules, amongst which this new amphipod was collected.</p><p>Distribution.</p><p>Abyssal Pacific Ocean, Clarion-Clipperton Zone in 4097–4430 m depth.</p><p>Molecular identification.</p><p>Following the definition given by Pleijel et al. (2008), the sequence of the holotype male of Rhachotropis clarionclippertoni (SMF 62817, GenBank accession number PQ 734529) is designed as a hologenophore of all obtained sequences. The sequences of the paratype and additional individuals of the species are also available in GenBank (Table 2). Furthermore, the species has received a Barcode Index Number from Barcode of Life Data Systems: BOLD: AEB 0147 (https://doi.org/10.5883/BOLD:AEB0147).</p></div>	https://treatment.plazi.org/id/B898A958174F5CF1A02C0BAC388411D2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lörz, Anne-Nina;Engel, Laura;Jereczek, Halina;Ćwierz, Nikodem;Jażdżewska, Anna M.	Lörz, Anne-Nina, Engel, Laura, Jereczek, Halina, Ćwierz, Nikodem, Jażdżewska, Anna M. (2026): Eusiridae (Crustacea, Amphipoda) of the Clarion-Clipperton Zone in the abyssal east Pacific with descriptions of five new species. ZooKeys 1274: 35-77, DOI: 10.3897/zookeys.1274.140610
61535576FB6B5413BC6239DA11E7E33A.text	61535576FB6B5413BC6239DA11E7E33A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhachotropis laure Lörz & Engel & Jereczek & Ćwierz & Jażdżewska 2026	<div><p>Rhachotropis laure sp. nov.</p><p>Figs 21, 22, 23, 24</p><p>Type material.</p><p>Holotype: Pacific • female, 10.5 mm, SMF 62819, COI: PQ 734767, body remnants and two slides with appendages, Clarion-Clipperton Zone, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.98624&amp;materialsCitation.latitude=11.8317" title="Search Plazi for locations around (long -116.98624/lat 11.8317)">BGR exploration contract area</a>, R / V Kilo Moana, MANGAN 2016, EBS, Ma 16-28, 01/05/2016, 11°49.654'N, 117°00.299'W – 11°49.902'N, 116°59.174'W, 4143– 4133 m . Paratype: Pacific • female, 10 mm, SMF 62820, COI: PQ 734754, Clarion-Clipperton Zone, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-117.79462&amp;materialsCitation.latitude=11.794617" title="Search Plazi for locations around (long -117.79462/lat 11.794617)">BGR exploration contract area</a>, R / V Sonne, MANGAN 2018, EBS, SO 262-155, 09/05/2018, 11°47.436'N, 117°32.213'W – 11°47.677'N, 117°47.677'W, 4352– 4351 m .</p><p>Additional material.</p><p>Pacific • Clarion-Clipperton Zone; BGR and OMS exploration contract areas, several stations (Table 2).</p><p>Type locality.</p><p>Abyssal Pacific Ocean, Clarion-Clipperton Zone, 11°49.654'N, 117°00.299'W – 11°49.902'N, 116°59.174'W, 4143– 4133 m.</p><p>Description.</p><p>Based on holotype, female, 10.5 mm, SMF 62819.</p><p>Body (Figs 21, 22): all pereonites smooth; pleonites 1–3 with a dorsal spine, pleonites 2 and 3 with a carina.</p><p>Head (Figs 21, 22): dorsally smooth, no eyes visible; rostrum short, slightly downcurved; cephalic lobes rounded.</p><p>Antennae 1 and 2 (Fig. 22): antennae very long, almost as long as body. Antenna 1 peduncle article 2 narrower and same length as article 1, longer than article 3; flagellum broken off; accessory flagellum uni-articulated. Antenna 2 peduncle articles 4 and 5 equal in length, several plumose setae on fourth article; article 5 long, 1 / 4 length of article 4.</p><p>Upper lip (Fig. 23): circular in shape, two setose areas anteriorly (ventral margin rounded, lateral margins finely setose).</p><p>Mandible (Fig. 23): Mandible with incisor process well-developed; left lacinia mobilis denticulate; molar process conical, large molar, edge with row of strong spines; palp article 1 short, ~ 1 / 3 length of article 2, article 3 almost 2 × as long as article 2, articles 2 and 3 with long slender setae, plumose setae at apical tip of left mandible palp.</p><p>Lower lip (Fig. 23): setose anterodistally, outer lobes separated by broad gap.</p><p>Maxilla 1 (Fig. 23): inner plate bearing one subterminal plumose setae; outer plate with nine denticulate spines; article 1 of palp 1 / 2 length of second article, article 2 of palp with several slender setae.</p><p>Maxilla 2 (Fig. 23): inner and outer plates subequal in length, margins bearing stout and slender setae; inner plate slightly wider than outer plate.</p><p>Maxilliped (Fig. 23): inner plate distally with short, thick spines; outer plate 2 × as long as inner plate, reaching 1 / 2 of article 2 of maxillipedal palp, outer plate margins strongly setose; palp article strongly setose.</p><p>Pereon: Gnathopod 1 (Fig. 22): coxa 1 anteriorly slightly drawn out; gnathopods similar in shape, subchelate; gnathopod 1 slightly smaller than gnathopod 2; basis bearing small spines at anteroventral corner; ischium and merus with long setae at posteroventral corner; carpus lobe extending width of propodus, spines at distal end of lobe and along the posterior margin; propodus widened, oval, palmar corner with six spines; dactylus slender, reaching end of palm.</p><p>Gnathopod 2 (Fig. 22): coxa 2 subquadrate; gnathopod 2 basis wider than basis of gnathopod 1; ischium to dactylus similar to gnathopod 1.</p><p>Pereopods 3 and 4 (Fig. 24): coxa 3 subquadrate, coxa 4 posteriorly drawn out. Pereopods 3 and 4 articles long and narrow; pereopod 3 broken off at basis; pereopod 4 broken off at merus.</p><p>Pereopod 5 (Fig. 24): coxa 5 lobate. Pereopod 5 slender, basis subrectangular, posteroventral angle notched, merus long; carpus, propodus, and dactylus broken off.</p><p>Pereopod 6 (Fig. 24): coxa 6 slightly lobate, pereopod 6 larger, but similar in shape to pereopod 5, merus long; carpus, propodus, and dactylus broken off.</p><p>Pereopod 7 (Fig. 24): coxa 7 posterior margin drawn out. Pereopod 7 longer than pereopods 5 and 6; basis with rounded at posterodistal corner; merus posteroventral angle produced, remaining pereopod missing.</p><p>Pleon: Epimera (Figs 21, 22): epimeral plates ventrally rounded, epimeral plates 1 and 2 posteriorly smooth, epimeral plate 3 serrated.</p><p>Pleopods (Fig. 24): pleopod 1 (right side) rami with 15 and 18 articles; rami same length, as long as peduncle.</p><p>Urosome: Urosomites (Figs 21, 22): urosome segment 1 with carina and dorsal spine, urosomes 2 and 3 smooth.</p><p>Uropod 1 (Fig. 24): lanceolate; outer rami broken off; inner ramus shorter than peduncle.</p><p>Uropod 2 (Fig. 24): lanceolate; outer ramus 0.3 × shorter than inner ramus; peduncle longer than outer ramus.</p><p>Uropod 3 (Fig. 24): lanceolate; inner and outer ramus approx. the same length, rami nearly 2 × as long as peduncle.</p><p>Telson (Fig. 22): Telson 3 × as long as wide, with a V-shaped cleft of ~ 10 %.</p><p>Remarks.</p><p>Rhachotropis laure sp. nov. is most similar to Rhachotropis gislii Thurston, 1980, R. distincta (Holmes, 1908), and R. gracilis Bonnier, 1896; these four species share the character combination of coxa 1 not being produced, cleft telson, and serrated epimeral plate 3. Rhachotropis laure sp. nov. can be distinguished from R. gislii by the latter bearing lateral teeth on the pleon, a rounded cephalic lobe, and a smooth urosomite 1. Rhachotropis gracilis and R. distincta do not show a strong excavation of coxa 4 as presented in R. laure sp. nov. Characters separating the morphologically close Rhachotropis species are listed in Table 6.</p><p>Another morphologically very similar species to Rhachotropis laure sp. nov. is Rhachotropis arii Thurston, 1980, but the latter has a smooth third epimeral plate, whereas in R. laure sp. nov. it is distinctly serrated (Table 6).</p><p>Etymology.</p><p>This species is named after Laure de Montety who works at the Marine and Freshwater Research Insitute of Iceland, Hafrannsóknastofnun, for illuminating many benthic organisms with her taxonomic expertise and for being a friend. It is used as a noun in apposition.</p><p>Distribution.</p><p>Abyssal Pacific Ocean, Clarion-Clipperton Zone in 4026–4368 m depth.</p><p>Molecular identification.</p><p>Following the definition given by Pleijel et al. (2008), the sequence of the holotype male of Rhachotropis laure (SMF 62819, GenBank accession number PQ 734767) is designed as a hologenophore of all obtained sequences. The sequences of the paratype and additional individuals of the species are also available in GenBank (Table 2). Furthermore, the species has received a Barcode Index Number from Barcode of Life Data Systems: BOLD: AEB 2577 (https://doi.org/10.5883/BOLD:AEB2577).</p></div>	https://treatment.plazi.org/id/61535576FB6B5413BC6239DA11E7E33A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lörz, Anne-Nina;Engel, Laura;Jereczek, Halina;Ćwierz, Nikodem;Jażdżewska, Anna M.	Lörz, Anne-Nina, Engel, Laura, Jereczek, Halina, Ćwierz, Nikodem, Jażdżewska, Anna M. (2026): Eusiridae (Crustacea, Amphipoda) of the Clarion-Clipperton Zone in the abyssal east Pacific with descriptions of five new species. ZooKeys 1274: 35-77, DOI: 10.3897/zookeys.1274.140610
AF3BC43CCE82556E98B89C587E39B69B.text	AF3BC43CCE82556E98B89C587E39B69B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhachotropis S. I. Smith 1883	<div><p>Genus Rhachotropis S. I. Smith, 1883</p><p>Rhachotropis S. I. Smith, 1883: 222. – Stebbing 1906: 847. – Shoemaker 1930: 317. – Gurjanova, 1951: 706. – J. L. Barnard 1969: 229. – Barnard and Karaman 1991: 337, figs 59 F, 61 H, 62 B, 63 B.</p><p>Gracilipes Holmes, 1908: 526 .</p><p>Type species.</p><p>Oniscus aculeata Lepechin, 1780 .</p></div>	https://treatment.plazi.org/id/AF3BC43CCE82556E98B89C587E39B69B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lörz, Anne-Nina;Engel, Laura;Jereczek, Halina;Ćwierz, Nikodem;Jażdżewska, Anna M.	Lörz, Anne-Nina, Engel, Laura, Jereczek, Halina, Ćwierz, Nikodem, Jażdżewska, Anna M. (2026): Eusiridae (Crustacea, Amphipoda) of the Clarion-Clipperton Zone in the abyssal east Pacific with descriptions of five new species. ZooKeys 1274: 35-77, DOI: 10.3897/zookeys.1274.140610
