taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
25B9D093FE735D338059EF8A819707B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690137	https://doi.org/10.3897/asp.84.e184638.figure1	Figure 1. Male antenna of Tritonaclia kefersteinii, ms – main seta, as – additional seta (see text).	Figure 1. Male antenna of Tritonaclia kefersteinii, ms – main seta, as – additional seta (see text).	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
25B9D093FE735D338059EF8A819707B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690138	https://doi.org/10.3897/asp.84.e184638.figure2	Figure 2. Phylogenetic relationship of Tritonaclia (blue clade) and selected other “ black-and-white ” Madagascan Syntomini species, inferred based on Maximum Likelihood analysis of eight markers, in IQ-TREE 3. Nodes support values represent SH-like / Ultrafast Bootstrap results, respectively.	Figure 2. Phylogenetic relationship of Tritonaclia (blue clade) and selected other “ black-and-white ” Madagascan Syntomini species, inferred based on Maximum Likelihood analysis of eight markers, in IQ-TREE 3. Nodes support values represent SH-like / Ultrafast Bootstrap results, respectively.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
25B9D093FE735D338059EF8A819707B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690139	https://doi.org/10.3897/asp.84.e184638.figure3	Figure 3. Tritonaclia adults. A T. kefersteinii male (MNHN, EL 83627 – CLV _ 1); B T. kefersteinii female (MNHN, EL 83626 – CLV _ 2); C T. kefersteinii female with reduced wing pattern (NHMUK, NHMUK 015109820), see text; D T. quinquepunctata, male (MNHN, EL 74630). Scale bars: 1 cm.	Figure 3. Tritonaclia adults. A T. kefersteinii male (MNHN, EL 83627 – CLV _ 1); B T. kefersteinii female (MNHN, EL 83626 – CLV _ 2); C T. kefersteinii female with reduced wing pattern (NHMUK, NHMUK 015109820), see text; D T. quinquepunctata, male (MNHN, EL 74630). Scale bars: 1 cm.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
25B9D093FE735D338059EF8A819707B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690140	https://doi.org/10.3897/asp.84.e184638.figure4	Figure 4. Male genitalia of Tritonaclia. A, B Ventral view and phallus of T. kefersteinii (MNHN, preparation ♂ nr 124); C, D ventral view and phallus of T. quinquepunctata (MNHN, slide P. Griveaud, No. 501).	Figure 4. Male genitalia of Tritonaclia. A, B Ventral view and phallus of T. kefersteinii (MNHN, preparation ♂ nr 124); C, D ventral view and phallus of T. quinquepunctata (MNHN, slide P. Griveaud, No. 501).	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
25B9D093FE735D338059EF8A819707B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690141	https://doi.org/10.3897/asp.84.e184638.figure5	Figure 5. Female genitalia of Tritonaclia kefersteinii. A MNHN, EL 83626 - CLV _ 2, preparation ♀ No. 123; B – D NHMUK, NHMUK 015109820, slide NHMUK, NHMUK 014331971.	Figure 5. Female genitalia of Tritonaclia kefersteinii. A MNHN, EL 83626 - CLV _ 2, preparation ♀ No. 123; B – D NHMUK, NHMUK 015109820, slide NHMUK, NHMUK 014331971.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
25B9D093FE735D338059EF8A819707B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690142	https://doi.org/10.3897/asp.84.e184638.figure6	Figure 6. Tritonaclia tollini adults. A – C Original specimens of G. A. Keferstein, males; A, B, F, G paralectotypes (MLUH - ZNS), upper- and underside; C, H lectotype (MLUH - ZNS), upper- and underside; D lectotype of Tritonaclia melania syn. nov. (NHMUK, NHMUK 010620989); E original illustration of T. tollini in Keferstein (1870); I, J male, upper- and underside (ISEA PAS, DL _ 2959); K male, lateral view of abdomen (ISEA PAS, DL _ 01-216). Scale bars: 1 cm.	Figure 6. Tritonaclia tollini adults. A – C Original specimens of G. A. Keferstein, males; A, B, F, G paralectotypes (MLUH - ZNS), upper- and underside; C, H lectotype (MLUH - ZNS), upper- and underside; D lectotype of Tritonaclia melania syn. nov. (NHMUK, NHMUK 010620989); E original illustration of T. tollini in Keferstein (1870); I, J male, upper- and underside (ISEA PAS, DL _ 2959); K male, lateral view of abdomen (ISEA PAS, DL _ 01-216). Scale bars: 1 cm.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
25B9D093FE735D338059EF8A819707B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690143	https://doi.org/10.3897/asp.84.e184638.figure7	Figure 7. Tritonaclia adults. A – E T. vonifotsy Wiorek, sp. nov., males; A, B holotype, upper- and underside (ISEA PAS, DL _ 1864); C Lectotype, upperside (MNHN, EL 74622); D, E lectotype, upper- and underside (MNHN, EL 74624); F T. vonifotsy Wiorek, sp. nov., female, lectotype (NHMUK, NHMUK 010918662 – DL 1863); G T. tollini, female (ISEA PAS, DL _ 2981); H, I unidentified female of T. tollini / T. vonifotsy Wiorek, sp. nov., upper- and underside (NHMUK, NHMUK 015109812). Scale bars: 1 cm.	Figure 7. Tritonaclia adults. A – E T. vonifotsy Wiorek, sp. nov., males; A, B holotype, upper- and underside (ISEA PAS, DL _ 1864); C Lectotype, upperside (MNHN, EL 74622); D, E lectotype, upper- and underside (MNHN, EL 74624); F T. vonifotsy Wiorek, sp. nov., female, lectotype (NHMUK, NHMUK 010918662 – DL 1863); G T. tollini, female (ISEA PAS, DL _ 2981); H, I unidentified female of T. tollini / T. vonifotsy Wiorek, sp. nov., upper- and underside (NHMUK, NHMUK 015109812). Scale bars: 1 cm.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
25B9D093FE735D338059EF8A819707B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690144	https://doi.org/10.3897/asp.84.e184638.figure8	Figure 8. Tritonaclia adults. A – C T. stephania; A female holotype (NHMUK, NHMUK 010620988); B, C male, upper- and underside (ISEA PAS, MAD _ 166); D – F T. ombilahy Wiorek & Przybyłowicz, sp. nov., male holotype, upper-, underside and lateroventral view of the abdomen (MNHN, EL 74649); G T. ombilahy Wiorek & Przybyłowicz, sp. nov. female, originally “ neallotype ” of T. stephania (MNHN, EL 65139, see text). Scale bars: 1 cm.	Figure 8. Tritonaclia adults. A – C T. stephania; A female holotype (NHMUK, NHMUK 010620988); B, C male, upper- and underside (ISEA PAS, MAD _ 166); D – F T. ombilahy Wiorek & Przybyłowicz, sp. nov., male holotype, upper-, underside and lateroventral view of the abdomen (MNHN, EL 74649); G T. ombilahy Wiorek & Przybyłowicz, sp. nov. female, originally “ neallotype ” of T. stephania (MNHN, EL 65139, see text). Scale bars: 1 cm.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
25B9D093FE735D338059EF8A819707B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690145	https://doi.org/10.3897/asp.84.e184638.figure9	Figure 9. Male genitalia of Tritonaclia tollini. A, B Specimen ISEA PAS, DL _ 01-216, slide S 554; C – E specimen ISEA PAS, DL _ 2959, slide S 555; A – C ventral, dorsal and lateral view, note the shape of tegumen (arrowhead in B), and uncus; D, E phallus; F, G protruding genitalia of the lectotype (MLUH - ZNS), note shape of uncus bifurcation (arrowhead).	Figure 9. Male genitalia of Tritonaclia tollini. A, B Specimen ISEA PAS, DL _ 01-216, slide S 554; C – E specimen ISEA PAS, DL _ 2959, slide S 555; A – C ventral, dorsal and lateral view, note the shape of tegumen (arrowhead in B), and uncus; D, E phallus; F, G protruding genitalia of the lectotype (MLUH - ZNS), note shape of uncus bifurcation (arrowhead).	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
25B9D093FE735D338059EF8A819707B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690146	https://doi.org/10.3897/asp.84.e184638.figure10	Figure 10. Male genitalia of Tritonaclia vonifotsy Wiorek, sp. nov. A Ventral view, arrowhead indicates protrusions of juxta; B dorsal view, arrowhead indicates lateral protrusion of uncus; C phallus; D – F lateral view, note variation in uncus length, arrowhead in D indicates protrusion of sacculus; A, B, D paratype, MNHN, EL 74622, préparation ♂ no. 155; C, E paratype, MNHN, EL 74624, préparation ♂ no. 156; F holotype, ISEA PAS, DL _ 1864, slide S 553.	Figure 10. Male genitalia of Tritonaclia vonifotsy Wiorek, sp. nov. A Ventral view, arrowhead indicates protrusions of juxta; B dorsal view, arrowhead indicates lateral protrusion of uncus; C phallus; D – F lateral view, note variation in uncus length, arrowhead in D indicates protrusion of sacculus; A, B, D paratype, MNHN, EL 74622, préparation ♂ no. 155; C, E paratype, MNHN, EL 74624, préparation ♂ no. 156; F holotype, ISEA PAS, DL _ 1864, slide S 553.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
25B9D093FE735D338059EF8A819707B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690147	https://doi.org/10.3897/asp.84.e184638.figure11	Figure 11. Male genitalia of Tritonaclia stephania, specimen ISEA PAS, MAD _ 166, slide S 556. A Ventral view; B dorsal view; C phallus.	Figure 11. Male genitalia of Tritonaclia stephania, specimen ISEA PAS, MAD _ 166, slide S 556. A Ventral view; B dorsal view; C phallus.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
25B9D093FE735D338059EF8A819707B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690148	https://doi.org/10.3897/asp.84.e184638.figure12	Figure 12. Male genitalia of Tritonaclia ombilahy Wiorek & Przybyłowicz, sp. nov., holotype, MNHN, EL 74649, préparation ♂ no. 157. A Ventral view, arrowhead indicates elongate arms of juxta; B dorsal view; C lateral view, arrowhead indicates lateral protrusion of tegumen; D, E phallus.	Figure 12. Male genitalia of Tritonaclia ombilahy Wiorek & Przybyłowicz, sp. nov., holotype, MNHN, EL 74649, préparation ♂ no. 157. A Ventral view, arrowhead indicates elongate arms of juxta; B dorsal view; C lateral view, arrowhead indicates lateral protrusion of tegumen; D, E phallus.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
25B9D093FE735D338059EF8A819707B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690149	https://doi.org/10.3897/asp.84.e184638.figure13	Figure 13. Female genitalia of Tritonaclia tollini, specimen ISEA PAS, DL _ 2981, slide S 557. A Ventral view; B dorsal view; C magnification of distal part of genitalia, note coiled ductus bursae; D magnification of signum.	Figure 13. Female genitalia of Tritonaclia tollini, specimen ISEA PAS, DL _ 2981, slide S 557. A Ventral view; B dorsal view; C magnification of distal part of genitalia, note coiled ductus bursae; D magnification of signum.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
25B9D093FE735D338059EF8A819707B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690150	https://doi.org/10.3897/asp.84.e184638.figure14	Figure 14. Female genitalia of Tritonaclia vonifotsy Wiorek, sp. nov., lectotype NHMUK, NHMUK 010918662 - DL 1863, genitalia slide NHMUK 010316589. A Ventral view; B magnification of distal part of genitalia, arrowhead indicates lateral protrusion of lamella postvaginalis (see text); C magnification of coiled ductus bursae; D magnification of signum.	Figure 14. Female genitalia of Tritonaclia vonifotsy Wiorek, sp. nov., lectotype NHMUK, NHMUK 010918662 - DL 1863, genitalia slide NHMUK 010316589. A Ventral view; B magnification of distal part of genitalia, arrowhead indicates lateral protrusion of lamella postvaginalis (see text); C magnification of coiled ductus bursae; D magnification of signum.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
25B9D093FE735D338059EF8A819707B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690151	https://doi.org/10.3897/asp.84.e184638.figure15	Figure 15. Female genitalia of Tritonaclia ombilahy Wiorek & Przybyłowicz, sp. nov., specimen MNHN, EL 65139, slide P. Griveaud No. 291, originally “ neallotype ” of T. stephania (see text).	Figure 15. Female genitalia of Tritonaclia ombilahy Wiorek & Przybyłowicz, sp. nov., specimen MNHN, EL 65139, slide P. Griveaud No. 291, originally “ neallotype ” of T. stephania (see text).	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
5EF7B3DD661E5297B9352701CCED6A32.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690137	https://doi.org/10.3897/asp.84.e184638.figure1	Figure 1. Male antenna of Tritonaclia kefersteinii, ms – main seta, as – additional seta (see text).	Figure 1. Male antenna of Tritonaclia kefersteinii, ms – main seta, as – additional seta (see text).	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
5EF7B3DD661E5297B9352701CCED6A32.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690139	https://doi.org/10.3897/asp.84.e184638.figure3	Figure 3. Tritonaclia adults. A T. kefersteinii male (MNHN, EL 83627 – CLV _ 1); B T. kefersteinii female (MNHN, EL 83626 – CLV _ 2); C T. kefersteinii female with reduced wing pattern (NHMUK, NHMUK 015109820), see text; D T. quinquepunctata, male (MNHN, EL 74630). Scale bars: 1 cm.	Figure 3. Tritonaclia adults. A T. kefersteinii male (MNHN, EL 83627 – CLV _ 1); B T. kefersteinii female (MNHN, EL 83626 – CLV _ 2); C T. kefersteinii female with reduced wing pattern (NHMUK, NHMUK 015109820), see text; D T. quinquepunctata, male (MNHN, EL 74630). Scale bars: 1 cm.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
5EF7B3DD661E5297B9352701CCED6A32.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690140	https://doi.org/10.3897/asp.84.e184638.figure4	Figure 4. Male genitalia of Tritonaclia. A, B Ventral view and phallus of T. kefersteinii (MNHN, preparation ♂ nr 124); C, D ventral view and phallus of T. quinquepunctata (MNHN, slide P. Griveaud, No. 501).	Figure 4. Male genitalia of Tritonaclia. A, B Ventral view and phallus of T. kefersteinii (MNHN, preparation ♂ nr 124); C, D ventral view and phallus of T. quinquepunctata (MNHN, slide P. Griveaud, No. 501).	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
5EF7B3DD661E5297B9352701CCED6A32.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690141	https://doi.org/10.3897/asp.84.e184638.figure5	Figure 5. Female genitalia of Tritonaclia kefersteinii. A MNHN, EL 83626 - CLV _ 2, preparation ♀ No. 123; B – D NHMUK, NHMUK 015109820, slide NHMUK, NHMUK 014331971.	Figure 5. Female genitalia of Tritonaclia kefersteinii. A MNHN, EL 83626 - CLV _ 2, preparation ♀ No. 123; B – D NHMUK, NHMUK 015109820, slide NHMUK, NHMUK 014331971.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
5EF7B3DD661E5297B9352701CCED6A32.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690152	https://doi.org/10.3897/asp.84.e184638.figure16	Figure 16. Distributional maps of Tritonaclia. A T. kefersteinii; B T. quinquepunctata; C T. tollini, violet dot – doubtful locality, red dot – collecting locality of the rediscovered type series, see text; D. T. vonifotsy Wiorek, sp. nov., violet dot – unidentified females of T. tollini / T. vonifotsy Wiorek, sp. nov., see text; E. T. stephania; F. T. ombilahy Wiorek & Przybyłowicz, sp. nov., violet dot – doubtful locality, see text.	Figure 16. Distributional maps of Tritonaclia. A T. kefersteinii; B T. quinquepunctata; C T. tollini, violet dot – doubtful locality, red dot – collecting locality of the rediscovered type series, see text; D. T. vonifotsy Wiorek, sp. nov., violet dot – unidentified females of T. tollini / T. vonifotsy Wiorek, sp. nov., see text; E. T. stephania; F. T. ombilahy Wiorek & Przybyłowicz, sp. nov., violet dot – doubtful locality, see text.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
A382B77B392351D2B9A721A274A119A5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690144	https://doi.org/10.3897/asp.84.e184638.figure8	Figure 8. Tritonaclia adults. A – C T. stephania; A female holotype (NHMUK, NHMUK 010620988); B, C male, upper- and underside (ISEA PAS, MAD _ 166); D – F T. ombilahy Wiorek & Przybyłowicz, sp. nov., male holotype, upper-, underside and lateroventral view of the abdomen (MNHN, EL 74649); G T. ombilahy Wiorek & Przybyłowicz, sp. nov. female, originally “ neallotype ” of T. stephania (MNHN, EL 65139, see text). Scale bars: 1 cm.	Figure 8. Tritonaclia adults. A – C T. stephania; A female holotype (NHMUK, NHMUK 010620988); B, C male, upper- and underside (ISEA PAS, MAD _ 166); D – F T. ombilahy Wiorek & Przybyłowicz, sp. nov., male holotype, upper-, underside and lateroventral view of the abdomen (MNHN, EL 74649); G T. ombilahy Wiorek & Przybyłowicz, sp. nov. female, originally “ neallotype ” of T. stephania (MNHN, EL 65139, see text). Scale bars: 1 cm.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
A382B77B392351D2B9A721A274A119A5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690148	https://doi.org/10.3897/asp.84.e184638.figure12	Figure 12. Male genitalia of Tritonaclia ombilahy Wiorek & Przybyłowicz, sp. nov., holotype, MNHN, EL 74649, préparation ♂ no. 157. A Ventral view, arrowhead indicates elongate arms of juxta; B dorsal view; C lateral view, arrowhead indicates lateral protrusion of tegumen; D, E phallus.	Figure 12. Male genitalia of Tritonaclia ombilahy Wiorek & Przybyłowicz, sp. nov., holotype, MNHN, EL 74649, préparation ♂ no. 157. A Ventral view, arrowhead indicates elongate arms of juxta; B dorsal view; C lateral view, arrowhead indicates lateral protrusion of tegumen; D, E phallus.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
A382B77B392351D2B9A721A274A119A5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690151	https://doi.org/10.3897/asp.84.e184638.figure15	Figure 15. Female genitalia of Tritonaclia ombilahy Wiorek & Przybyłowicz, sp. nov., specimen MNHN, EL 65139, slide P. Griveaud No. 291, originally “ neallotype ” of T. stephania (see text).	Figure 15. Female genitalia of Tritonaclia ombilahy Wiorek & Przybyłowicz, sp. nov., specimen MNHN, EL 65139, slide P. Griveaud No. 291, originally “ neallotype ” of T. stephania (see text).	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
A382B77B392351D2B9A721A274A119A5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690152	https://doi.org/10.3897/asp.84.e184638.figure16	Figure 16. Distributional maps of Tritonaclia. A T. kefersteinii; B T. quinquepunctata; C T. tollini, violet dot – doubtful locality, red dot – collecting locality of the rediscovered type series, see text; D. T. vonifotsy Wiorek, sp. nov., violet dot – unidentified females of T. tollini / T. vonifotsy Wiorek, sp. nov., see text; E. T. stephania; F. T. ombilahy Wiorek & Przybyłowicz, sp. nov., violet dot – doubtful locality, see text.	Figure 16. Distributional maps of Tritonaclia. A T. kefersteinii; B T. quinquepunctata; C T. tollini, violet dot – doubtful locality, red dot – collecting locality of the rediscovered type series, see text; D. T. vonifotsy Wiorek, sp. nov., violet dot – unidentified females of T. tollini / T. vonifotsy Wiorek, sp. nov., see text; E. T. stephania; F. T. ombilahy Wiorek & Przybyłowicz, sp. nov., violet dot – doubtful locality, see text.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
EA035FA42FB858EEBC220F34725AFADE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690139	https://doi.org/10.3897/asp.84.e184638.figure3	Figure 3. Tritonaclia adults. A T. kefersteinii male (MNHN, EL 83627 – CLV _ 1); B T. kefersteinii female (MNHN, EL 83626 – CLV _ 2); C T. kefersteinii female with reduced wing pattern (NHMUK, NHMUK 015109820), see text; D T. quinquepunctata, male (MNHN, EL 74630). Scale bars: 1 cm.	Figure 3. Tritonaclia adults. A T. kefersteinii male (MNHN, EL 83627 – CLV _ 1); B T. kefersteinii female (MNHN, EL 83626 – CLV _ 2); C T. kefersteinii female with reduced wing pattern (NHMUK, NHMUK 015109820), see text; D T. quinquepunctata, male (MNHN, EL 74630). Scale bars: 1 cm.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
EA035FA42FB858EEBC220F34725AFADE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690140	https://doi.org/10.3897/asp.84.e184638.figure4	Figure 4. Male genitalia of Tritonaclia. A, B Ventral view and phallus of T. kefersteinii (MNHN, preparation ♂ nr 124); C, D ventral view and phallus of T. quinquepunctata (MNHN, slide P. Griveaud, No. 501).	Figure 4. Male genitalia of Tritonaclia. A, B Ventral view and phallus of T. kefersteinii (MNHN, preparation ♂ nr 124); C, D ventral view and phallus of T. quinquepunctata (MNHN, slide P. Griveaud, No. 501).	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
EA035FA42FB858EEBC220F34725AFADE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690152	https://doi.org/10.3897/asp.84.e184638.figure16	Figure 16. Distributional maps of Tritonaclia. A T. kefersteinii; B T. quinquepunctata; C T. tollini, violet dot – doubtful locality, red dot – collecting locality of the rediscovered type series, see text; D. T. vonifotsy Wiorek, sp. nov., violet dot – unidentified females of T. tollini / T. vonifotsy Wiorek, sp. nov., see text; E. T. stephania; F. T. ombilahy Wiorek & Przybyłowicz, sp. nov., violet dot – doubtful locality, see text.	Figure 16. Distributional maps of Tritonaclia. A T. kefersteinii; B T. quinquepunctata; C T. tollini, violet dot – doubtful locality, red dot – collecting locality of the rediscovered type series, see text; D. T. vonifotsy Wiorek, sp. nov., violet dot – unidentified females of T. tollini / T. vonifotsy Wiorek, sp. nov., see text; E. T. stephania; F. T. ombilahy Wiorek & Przybyłowicz, sp. nov., violet dot – doubtful locality, see text.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
84FCADDFDF2053DCA5F99BA37F0C1C00.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690144	https://doi.org/10.3897/asp.84.e184638.figure8	Figure 8. Tritonaclia adults. A – C T. stephania; A female holotype (NHMUK, NHMUK 010620988); B, C male, upper- and underside (ISEA PAS, MAD _ 166); D – F T. ombilahy Wiorek & Przybyłowicz, sp. nov., male holotype, upper-, underside and lateroventral view of the abdomen (MNHN, EL 74649); G T. ombilahy Wiorek & Przybyłowicz, sp. nov. female, originally “ neallotype ” of T. stephania (MNHN, EL 65139, see text). Scale bars: 1 cm.	Figure 8. Tritonaclia adults. A – C T. stephania; A female holotype (NHMUK, NHMUK 010620988); B, C male, upper- and underside (ISEA PAS, MAD _ 166); D – F T. ombilahy Wiorek & Przybyłowicz, sp. nov., male holotype, upper-, underside and lateroventral view of the abdomen (MNHN, EL 74649); G T. ombilahy Wiorek & Przybyłowicz, sp. nov. female, originally “ neallotype ” of T. stephania (MNHN, EL 65139, see text). Scale bars: 1 cm.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
84FCADDFDF2053DCA5F99BA37F0C1C00.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690147	https://doi.org/10.3897/asp.84.e184638.figure11	Figure 11. Male genitalia of Tritonaclia stephania, specimen ISEA PAS, MAD _ 166, slide S 556. A Ventral view; B dorsal view; C phallus.	Figure 11. Male genitalia of Tritonaclia stephania, specimen ISEA PAS, MAD _ 166, slide S 556. A Ventral view; B dorsal view; C phallus.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
84FCADDFDF2053DCA5F99BA37F0C1C00.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690152	https://doi.org/10.3897/asp.84.e184638.figure16	Figure 16. Distributional maps of Tritonaclia. A T. kefersteinii; B T. quinquepunctata; C T. tollini, violet dot – doubtful locality, red dot – collecting locality of the rediscovered type series, see text; D. T. vonifotsy Wiorek, sp. nov., violet dot – unidentified females of T. tollini / T. vonifotsy Wiorek, sp. nov., see text; E. T. stephania; F. T. ombilahy Wiorek & Przybyłowicz, sp. nov., violet dot – doubtful locality, see text.	Figure 16. Distributional maps of Tritonaclia. A T. kefersteinii; B T. quinquepunctata; C T. tollini, violet dot – doubtful locality, red dot – collecting locality of the rediscovered type series, see text; D. T. vonifotsy Wiorek, sp. nov., violet dot – unidentified females of T. tollini / T. vonifotsy Wiorek, sp. nov., see text; E. T. stephania; F. T. ombilahy Wiorek & Przybyłowicz, sp. nov., violet dot – doubtful locality, see text.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
9B1BBF61D9FF5ACDA573131C324DBCC3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690142	https://doi.org/10.3897/asp.84.e184638.figure6	Figure 6. Tritonaclia tollini adults. A – C Original specimens of G. A. Keferstein, males; A, B, F, G paralectotypes (MLUH - ZNS), upper- and underside; C, H lectotype (MLUH - ZNS), upper- and underside; D lectotype of Tritonaclia melania syn. nov. (NHMUK, NHMUK 010620989); E original illustration of T. tollini in Keferstein (1870); I, J male, upper- and underside (ISEA PAS, DL _ 2959); K male, lateral view of abdomen (ISEA PAS, DL _ 01-216). Scale bars: 1 cm.	Figure 6. Tritonaclia tollini adults. A – C Original specimens of G. A. Keferstein, males; A, B, F, G paralectotypes (MLUH - ZNS), upper- and underside; C, H lectotype (MLUH - ZNS), upper- and underside; D lectotype of Tritonaclia melania syn. nov. (NHMUK, NHMUK 010620989); E original illustration of T. tollini in Keferstein (1870); I, J male, upper- and underside (ISEA PAS, DL _ 2959); K male, lateral view of abdomen (ISEA PAS, DL _ 01-216). Scale bars: 1 cm.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
9B1BBF61D9FF5ACDA573131C324DBCC3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690143	https://doi.org/10.3897/asp.84.e184638.figure7	Figure 7. Tritonaclia adults. A – E T. vonifotsy Wiorek, sp. nov., males; A, B holotype, upper- and underside (ISEA PAS, DL _ 1864); C Lectotype, upperside (MNHN, EL 74622); D, E lectotype, upper- and underside (MNHN, EL 74624); F T. vonifotsy Wiorek, sp. nov., female, lectotype (NHMUK, NHMUK 010918662 – DL 1863); G T. tollini, female (ISEA PAS, DL _ 2981); H, I unidentified female of T. tollini / T. vonifotsy Wiorek, sp. nov., upper- and underside (NHMUK, NHMUK 015109812). Scale bars: 1 cm.	Figure 7. Tritonaclia adults. A – E T. vonifotsy Wiorek, sp. nov., males; A, B holotype, upper- and underside (ISEA PAS, DL _ 1864); C Lectotype, upperside (MNHN, EL 74622); D, E lectotype, upper- and underside (MNHN, EL 74624); F T. vonifotsy Wiorek, sp. nov., female, lectotype (NHMUK, NHMUK 010918662 – DL 1863); G T. tollini, female (ISEA PAS, DL _ 2981); H, I unidentified female of T. tollini / T. vonifotsy Wiorek, sp. nov., upper- and underside (NHMUK, NHMUK 015109812). Scale bars: 1 cm.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
9B1BBF61D9FF5ACDA573131C324DBCC3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690145	https://doi.org/10.3897/asp.84.e184638.figure9	Figure 9. Male genitalia of Tritonaclia tollini. A, B Specimen ISEA PAS, DL _ 01-216, slide S 554; C – E specimen ISEA PAS, DL _ 2959, slide S 555; A – C ventral, dorsal and lateral view, note the shape of tegumen (arrowhead in B), and uncus; D, E phallus; F, G protruding genitalia of the lectotype (MLUH - ZNS), note shape of uncus bifurcation (arrowhead).	Figure 9. Male genitalia of Tritonaclia tollini. A, B Specimen ISEA PAS, DL _ 01-216, slide S 554; C – E specimen ISEA PAS, DL _ 2959, slide S 555; A – C ventral, dorsal and lateral view, note the shape of tegumen (arrowhead in B), and uncus; D, E phallus; F, G protruding genitalia of the lectotype (MLUH - ZNS), note shape of uncus bifurcation (arrowhead).	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
9B1BBF61D9FF5ACDA573131C324DBCC3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690149	https://doi.org/10.3897/asp.84.e184638.figure13	Figure 13. Female genitalia of Tritonaclia tollini, specimen ISEA PAS, DL _ 2981, slide S 557. A Ventral view; B dorsal view; C magnification of distal part of genitalia, note coiled ductus bursae; D magnification of signum.	Figure 13. Female genitalia of Tritonaclia tollini, specimen ISEA PAS, DL _ 2981, slide S 557. A Ventral view; B dorsal view; C magnification of distal part of genitalia, note coiled ductus bursae; D magnification of signum.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
9B1BBF61D9FF5ACDA573131C324DBCC3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690152	https://doi.org/10.3897/asp.84.e184638.figure16	Figure 16. Distributional maps of Tritonaclia. A T. kefersteinii; B T. quinquepunctata; C T. tollini, violet dot – doubtful locality, red dot – collecting locality of the rediscovered type series, see text; D. T. vonifotsy Wiorek, sp. nov., violet dot – unidentified females of T. tollini / T. vonifotsy Wiorek, sp. nov., see text; E. T. stephania; F. T. ombilahy Wiorek & Przybyłowicz, sp. nov., violet dot – doubtful locality, see text.	Figure 16. Distributional maps of Tritonaclia. A T. kefersteinii; B T. quinquepunctata; C T. tollini, violet dot – doubtful locality, red dot – collecting locality of the rediscovered type series, see text; D. T. vonifotsy Wiorek, sp. nov., violet dot – unidentified females of T. tollini / T. vonifotsy Wiorek, sp. nov., see text; E. T. stephania; F. T. ombilahy Wiorek & Przybyłowicz, sp. nov., violet dot – doubtful locality, see text.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
9B1BBF61D9FF5ACDA573131C324DBCC3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690153	https://doi.org/10.3897/asp.84.e184638.figure17	Figure 17. Wing patches of modified, scent scales in Tritonaclia. A T. tollini hindwing upperside; B T. tollini forewing underside; C T. vonifotsy Wiorek, sp. nov. hindwing upperside; D T. vonifotsy Wiorek, sp. nov. forewing underside.	Figure 17. Wing patches of modified, scent scales in Tritonaclia. A T. tollini hindwing upperside; B T. tollini forewing underside; C T. vonifotsy Wiorek, sp. nov. hindwing upperside; D T. vonifotsy Wiorek, sp. nov. forewing underside.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
9B1BBF61D9FF5ACDA573131C324DBCC3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690154	https://doi.org/10.3897/asp.84.e184638.figure18	Figure 18. Wing venation, A – C Tritonaclia tollini, D – F T. vonifotsy Wiorek, sp. nov. A Male forewing, frame indicates the area magnified in B; B terminal bifurcation of vein 1 A + 2 A; C male hindwing, note the complete fusion of veins Sc + Rs + M 1 and presence of vein 3 A.; D male hindwing, slide ISEA PAS, S 554, frames indicate areas magnified in E and F; E terminal bifurcation of veins Sc + Rs + M 1; F basal, vestigial bifurcation of 3 A.	Figure 18. Wing venation, A – C Tritonaclia tollini, D – F T. vonifotsy Wiorek, sp. nov. A Male forewing, frame indicates the area magnified in B; B terminal bifurcation of vein 1 A + 2 A; C male hindwing, note the complete fusion of veins Sc + Rs + M 1 and presence of vein 3 A.; D male hindwing, slide ISEA PAS, S 554, frames indicate areas magnified in E and F; E terminal bifurcation of veins Sc + Rs + M 1; F basal, vestigial bifurcation of 3 A.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
9B1BBF61D9FF5ACDA573131C324DBCC3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690155	https://doi.org/10.3897/asp.84.e184638.figure19	Figure 19. SEM visualisation of male wing scent (androconial) scales in Tritonaclia tollini. A, B Upper lamina of forewing underside scale (in A basal part of scale in bottom right); C, D upper lamina of hindwing upperside scale (in C basal part of scale in bottom right).	Figure 19. SEM visualisation of male wing scent (androconial) scales in Tritonaclia tollini. A, B Upper lamina of forewing underside scale (in A basal part of scale in bottom right); C, D upper lamina of hindwing upperside scale (in C basal part of scale in bottom right).	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
9B1BBF61D9FF5ACDA573131C324DBCC3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690138	https://doi.org/10.3897/asp.84.e184638.figure2	Figure 2. Phylogenetic relationship of Tritonaclia (blue clade) and selected other “ black-and-white ” Madagascan Syntomini species, inferred based on Maximum Likelihood analysis of eight markers, in IQ-TREE 3. Nodes support values represent SH-like / Ultrafast Bootstrap results, respectively.	Figure 2. Phylogenetic relationship of Tritonaclia (blue clade) and selected other “ black-and-white ” Madagascan Syntomini species, inferred based on Maximum Likelihood analysis of eight markers, in IQ-TREE 3. Nodes support values represent SH-like / Ultrafast Bootstrap results, respectively.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
94740EE0EC1656DD9786AAC0B8B61853.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690143	https://doi.org/10.3897/asp.84.e184638.figure7	Figure 7. Tritonaclia adults. A – E T. vonifotsy Wiorek, sp. nov., males; A, B holotype, upper- and underside (ISEA PAS, DL _ 1864); C Lectotype, upperside (MNHN, EL 74622); D, E lectotype, upper- and underside (MNHN, EL 74624); F T. vonifotsy Wiorek, sp. nov., female, lectotype (NHMUK, NHMUK 010918662 – DL 1863); G T. tollini, female (ISEA PAS, DL _ 2981); H, I unidentified female of T. tollini / T. vonifotsy Wiorek, sp. nov., upper- and underside (NHMUK, NHMUK 015109812). Scale bars: 1 cm.	Figure 7. Tritonaclia adults. A – E T. vonifotsy Wiorek, sp. nov., males; A, B holotype, upper- and underside (ISEA PAS, DL _ 1864); C Lectotype, upperside (MNHN, EL 74622); D, E lectotype, upper- and underside (MNHN, EL 74624); F T. vonifotsy Wiorek, sp. nov., female, lectotype (NHMUK, NHMUK 010918662 – DL 1863); G T. tollini, female (ISEA PAS, DL _ 2981); H, I unidentified female of T. tollini / T. vonifotsy Wiorek, sp. nov., upper- and underside (NHMUK, NHMUK 015109812). Scale bars: 1 cm.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
94740EE0EC1656DD9786AAC0B8B61853.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690146	https://doi.org/10.3897/asp.84.e184638.figure10	Figure 10. Male genitalia of Tritonaclia vonifotsy Wiorek, sp. nov. A Ventral view, arrowhead indicates protrusions of juxta; B dorsal view, arrowhead indicates lateral protrusion of uncus; C phallus; D – F lateral view, note variation in uncus length, arrowhead in D indicates protrusion of sacculus; A, B, D paratype, MNHN, EL 74622, préparation ♂ no. 155; C, E paratype, MNHN, EL 74624, préparation ♂ no. 156; F holotype, ISEA PAS, DL _ 1864, slide S 553.	Figure 10. Male genitalia of Tritonaclia vonifotsy Wiorek, sp. nov. A Ventral view, arrowhead indicates protrusions of juxta; B dorsal view, arrowhead indicates lateral protrusion of uncus; C phallus; D – F lateral view, note variation in uncus length, arrowhead in D indicates protrusion of sacculus; A, B, D paratype, MNHN, EL 74622, préparation ♂ no. 155; C, E paratype, MNHN, EL 74624, préparation ♂ no. 156; F holotype, ISEA PAS, DL _ 1864, slide S 553.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
94740EE0EC1656DD9786AAC0B8B61853.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690150	https://doi.org/10.3897/asp.84.e184638.figure14	Figure 14. Female genitalia of Tritonaclia vonifotsy Wiorek, sp. nov., lectotype NHMUK, NHMUK 010918662 - DL 1863, genitalia slide NHMUK 010316589. A Ventral view; B magnification of distal part of genitalia, arrowhead indicates lateral protrusion of lamella postvaginalis (see text); C magnification of coiled ductus bursae; D magnification of signum.	Figure 14. Female genitalia of Tritonaclia vonifotsy Wiorek, sp. nov., lectotype NHMUK, NHMUK 010918662 - DL 1863, genitalia slide NHMUK 010316589. A Ventral view; B magnification of distal part of genitalia, arrowhead indicates lateral protrusion of lamella postvaginalis (see text); C magnification of coiled ductus bursae; D magnification of signum.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
94740EE0EC1656DD9786AAC0B8B61853.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690152	https://doi.org/10.3897/asp.84.e184638.figure16	Figure 16. Distributional maps of Tritonaclia. A T. kefersteinii; B T. quinquepunctata; C T. tollini, violet dot – doubtful locality, red dot – collecting locality of the rediscovered type series, see text; D. T. vonifotsy Wiorek, sp. nov., violet dot – unidentified females of T. tollini / T. vonifotsy Wiorek, sp. nov., see text; E. T. stephania; F. T. ombilahy Wiorek & Przybyłowicz, sp. nov., violet dot – doubtful locality, see text.	Figure 16. Distributional maps of Tritonaclia. A T. kefersteinii; B T. quinquepunctata; C T. tollini, violet dot – doubtful locality, red dot – collecting locality of the rediscovered type series, see text; D. T. vonifotsy Wiorek, sp. nov., violet dot – unidentified females of T. tollini / T. vonifotsy Wiorek, sp. nov., see text; E. T. stephania; F. T. ombilahy Wiorek & Przybyłowicz, sp. nov., violet dot – doubtful locality, see text.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
94740EE0EC1656DD9786AAC0B8B61853.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690153	https://doi.org/10.3897/asp.84.e184638.figure17	Figure 17. Wing patches of modified, scent scales in Tritonaclia. A T. tollini hindwing upperside; B T. tollini forewing underside; C T. vonifotsy Wiorek, sp. nov. hindwing upperside; D T. vonifotsy Wiorek, sp. nov. forewing underside.	Figure 17. Wing patches of modified, scent scales in Tritonaclia. A T. tollini hindwing upperside; B T. tollini forewing underside; C T. vonifotsy Wiorek, sp. nov. hindwing upperside; D T. vonifotsy Wiorek, sp. nov. forewing underside.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
94740EE0EC1656DD9786AAC0B8B61853.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690154	https://doi.org/10.3897/asp.84.e184638.figure18	Figure 18. Wing venation, A – C Tritonaclia tollini, D – F T. vonifotsy Wiorek, sp. nov. A Male forewing, frame indicates the area magnified in B; B terminal bifurcation of vein 1 A + 2 A; C male hindwing, note the complete fusion of veins Sc + Rs + M 1 and presence of vein 3 A.; D male hindwing, slide ISEA PAS, S 554, frames indicate areas magnified in E and F; E terminal bifurcation of veins Sc + Rs + M 1; F basal, vestigial bifurcation of 3 A.	Figure 18. Wing venation, A – C Tritonaclia tollini, D – F T. vonifotsy Wiorek, sp. nov. A Male forewing, frame indicates the area magnified in B; B terminal bifurcation of vein 1 A + 2 A; C male hindwing, note the complete fusion of veins Sc + Rs + M 1 and presence of vein 3 A.; D male hindwing, slide ISEA PAS, S 554, frames indicate areas magnified in E and F; E terminal bifurcation of veins Sc + Rs + M 1; F basal, vestigial bifurcation of 3 A.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
94740EE0EC1656DD9786AAC0B8B61853.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690156	https://doi.org/10.3897/asp.84.e184638.figure20	Figure 20. SEM visualisation of male wing scent (androconial) scales in Tritonaclia vonifotsy Wiorek, sp. nov. A Upper lamina of forewing underside scale (basal part in top right); B inner structure of forewing underside scale; C hindwing upperside scale, upper lamina in top left (basal part of scale bottom left), lower lamina in bottom right (basal part of scale top right); D magnification of upper lamina of hindwing upperside scale; E cross section of hindwing upperside scale; F magnification of inner structure of hindwing upperside scale.	Figure 20. SEM visualisation of male wing scent (androconial) scales in Tritonaclia vonifotsy Wiorek, sp. nov. A Upper lamina of forewing underside scale (basal part in top right); B inner structure of forewing underside scale; C hindwing upperside scale, upper lamina in top left (basal part of scale bottom left), lower lamina in bottom right (basal part of scale top right); D magnification of upper lamina of hindwing upperside scale; E cross section of hindwing upperside scale; F magnification of inner structure of hindwing upperside scale.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
94740EE0EC1656DD9786AAC0B8B61853.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690142	https://doi.org/10.3897/asp.84.e184638.figure6	Figure 6. Tritonaclia tollini adults. A – C Original specimens of G. A. Keferstein, males; A, B, F, G paralectotypes (MLUH - ZNS), upper- and underside; C, H lectotype (MLUH - ZNS), upper- and underside; D lectotype of Tritonaclia melania syn. nov. (NHMUK, NHMUK 010620989); E original illustration of T. tollini in Keferstein (1870); I, J male, upper- and underside (ISEA PAS, DL _ 2959); K male, lateral view of abdomen (ISEA PAS, DL _ 01-216). Scale bars: 1 cm.	Figure 6. Tritonaclia tollini adults. A – C Original specimens of G. A. Keferstein, males; A, B, F, G paralectotypes (MLUH - ZNS), upper- and underside; C, H lectotype (MLUH - ZNS), upper- and underside; D lectotype of Tritonaclia melania syn. nov. (NHMUK, NHMUK 010620989); E original illustration of T. tollini in Keferstein (1870); I, J male, upper- and underside (ISEA PAS, DL _ 2959); K male, lateral view of abdomen (ISEA PAS, DL _ 01-216). Scale bars: 1 cm.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
94740EE0EC1656DD9786AAC0B8B61853.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1690149	https://doi.org/10.3897/asp.84.e184638.figure13	Figure 13. Female genitalia of Tritonaclia tollini, specimen ISEA PAS, DL _ 2981, slide S 557. A Ventral view; B dorsal view; C magnification of distal part of genitalia, note coiled ductus bursae; D magnification of signum.	Figure 13. Female genitalia of Tritonaclia tollini, specimen ISEA PAS, DL _ 2981, slide S 557. A Ventral view; B dorsal view; C magnification of distal part of genitalia, note coiled ductus bursae; D magnification of signum.	2026-06-23	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz		Zenodo	biologists	Wiorek, Marcin;Lees, David C.;Wahlberg, Niklas;Przybyłowicz, Łukasz			
