identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
8A3C1C105A46559589252C6A8D0C996E.text	8A3C1C105A46559589252C6A8D0C996E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cabalodontia albofulva Westphalen & Gugliotta 2026	<div><p>Cabalodontia albofulva Westphalen &amp; Gugliotta sp. nov.</p><p>Figs 7 A, 7 B, 11 A, 11 J</p><p>Etymology.</p><p>Refers to the white basidiomes that become yellowish when dried.</p><p>Diagnosis.</p><p>Differs from C. queletii by thinner and more fragile aculei and smaller basidiospores.</p><p>Typification.</p><p>Brazil. Rio Grande do Sul: São Francisco de Paula, Parador Hampel, 19 Jun. 2017, M. C. Westphalen 563 / 17 (SP 467080) .</p><p>Description.</p><p>Basidiomes adnante, annual, resupinate, not detaching, forming small patches on the substrate, very soft and brittle; sterile margins white, membranaceous, up to 1 mm wide. Hymenophore odontioid, white to cream when fresh, becoming yellowish to beige upon drying. Aculei thin, up to 0.8 mm long, somewhat clustered and very brittle upon drying, 6–8 per mm. Subiculum, white to cream, homogeneous, cottony, thin and fragile, up to 0.5 mm thick.</p><p>Hyphal system monomitic, hyphae loosely arranged; generative hyphae clamped, thin- to slightly thick-walled, hyaline, often branching near the septa, 3–5 μm wide, CB –. Skeletocystidia abundant, arising from sclerified generative hyphae in the subiculum and projecting into the trama and above the hymenium, clavate, usually tapering towards the apex, encrusted portion 25–60 (– 70) × 6–11 μm; thin-walled leptocystidia present on tips of the aculei, clavate to capitate, 4–7 μm wide. Basidia clavate, tetrasterigmate, 12–15 × 3.5–5 μm. Basidiospores ellipsoid, hyaline, thin-walled, IKI –, CB –, (4 –) 4.3–5.1 (– 5.3) × 2.4–3.0 (– 3.1) μm.</p><p>Habitat and distribution.</p><p>Known only from the type locality in Araucaria forests in Rio Grande do Sul State. Growing on dead logs of unidentified angiosperms.</p><p>Notes.</p><p>Cabalodontia albofulva is characterized by fragile basidiomes with thin aculei measuring 0.5–0.8 mm long and ellipsoid basidiospores measuring (4 –) 4.3–5.1 (– 5.3) × 2.4–3.0 (– 3.1) µm. Cabalodontia queletii (Bourdot &amp; Galzin) Piątek is morphologically similar but differs by having thicker aculei and larger basidiospores, measuring 5–6 × 3–3.5 µm (Bernicchia and Gorjón 2010). Phylogenetically, C. albofulva is closely related to C. lincangense, from which it differs by approximately eight bp in the ITS sequences, four in the tef 1 - α coding region, and five in the rpb 1 coding region. Morphologically, however, C. lincangense is distinct, displaying more widely spaced and longer aculei (2–4 per mm, 1–1.5 mm long) and shorter basidiospores measuring (3.5 –) 3.8–4.2 (– 4.5) × (2.3 –) 2.5–2.8 µm (Dong et al. 2024). Cabalodontia albofulva is currently known only from its type locality in southern Brazil. Despite extensive sampling in the surrounding region, including adjacent Araucaria forests, no additional specimens have been collected, suggesting that it could represent a rare taxon.</p></div>	https://treatment.plazi.org/id/8A3C1C105A46559589252C6A8D0C996E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Westphalen, Mauro Carpes;Minosso, Nathalia Michele Martins;Regio, Nicolas do Carmo;Gugliotta, Adriana de Mello;Rajchenberg, Mario;Silveira, Rosa Mara Borges da	Westphalen, Mauro Carpes, Minosso, Nathalia Michele Martins, Regio, Nicolas do Carmo, Gugliotta, Adriana de Mello, Rajchenberg, Mario, Silveira, Rosa Mara Borges da (2026): Unveiling the hidden diversity of neotropical Steccherinum and allied genera (Steccherinaceae, Basidiomycota). IMA Fungus 17: e 182915, DOI: 10.3897/imafungus.17.182915
D92BD90AE1425A88AFB46AB82AD3C90D.text	D92BD90AE1425A88AFB46AB82AD3C90D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cabalodontia brunnea Westphalen & Regio 2026	<div><p>Cabalodontia brunnea Westphalen &amp; Regio sp. nov.</p><p>Figs 7 C, 7 D, 11 B</p><p>Etymology.</p><p>Refers to the brownish basidiomes.</p><p>Diagnosis.</p><p>Characterized by the pale brown basidiomes with short, irregular spines, hyaline to yellow hyphae, and basidiospores (3.7 –) 4–4.7 (– 4.9) × 2.4–3.0 (– 3.1) μm.</p><p>Typification.</p><p>Brazil. Rio Grande do Sul: Caxias do Sul, Cânion Palanquinhos, 18 Sep. 2017, M. C. Westphalen 600 / 17 (SP 512588) .</p><p>Description.</p><p>Basidiomes adnate, resupinate, annual, not detaching, forming confluent large patches on the substrate, soft and somewhat brittle; sterile margins absent or very thin, fimbriate to byssoid, up to 1 mm wide. Hymenophore pale brown to ochraceous when fresh, unchanging or slightly darker upon drying, formed by small irregular spines, up to 0.5 mm long, often somewhat cluttered, very brittle upon drying, 6–9 per mm. Subiculum beige to pale yellowish brown, homogeneous, thin and fragile, cottony, up to 0.5 mm thick.</p><p>Hyphal system monomitic, hyphae more densely arranged towards the trama and looser near the substratum; generative hyphae clamped, thin- to thick-walled, often branching near the septa or at clamp connections, 2.5–5 μm wide, hyaline to pale yellow, golden yellowish-brown in mass, CB +. Skeletocystidia abundant, heavily encrusted, arising from sclerified generative hyphae in the subiculum and projecting toward the trama and above the hymenium, clavate or more commonly tapering towards the apex, encrusted portion 25–70 (– 100) × 5–9 (– 12) μm. Basidia clavate, tetrasterigmate, 13–16 × 4–5 μm. Basidiospores ellipsoid, hyaline, thin-walled, (3.7 –) 4–4.7 (– 4.9) × 2.4–3.0 (– 3.1) μm, IKI –, CB –.</p><p>Habitat and distribution.</p><p>Known only from the type locality in Araucaria forests in Rio Grande do Sul State. Growing on dead logs of unidentified angiosperms.</p><p>Notes.</p><p>Cabalodontia brunnea is distinguished within the genus by its pale brown basidiomes with irregular short spines and a somewhat farinaceous appearance due to the encrusted cystidia. Microscopically, it presents yellowish hyphae and ellipsoid basidiospores measuring (3.8 –) 4–4.7 (– 4.9) × 2.4–3.0 (– 3.1) μm. As in the case of C. albofulva, this species is known only from its type locality in southern Brazil. Despite extensive sampling in the region, including nearby Araucaria forests, no additional specimens have been found, suggesting that it could represent a rare taxon.</p><p>Phylogenetically, C. brunnea is closely related to C. delicata, a species also found in Araucaria forests and high-altitude Atlantic rainforest areas (Westphalen et al. 2019). However, C. delicata is relatively common in these habitats and differs from C. brunnea by its pale white to cream-colored basidiomes, shorter aculei, and smaller, subglobose to broadly ellipsoid basidiospores.</p></div>	https://treatment.plazi.org/id/D92BD90AE1425A88AFB46AB82AD3C90D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Westphalen, Mauro Carpes;Minosso, Nathalia Michele Martins;Regio, Nicolas do Carmo;Gugliotta, Adriana de Mello;Rajchenberg, Mario;Silveira, Rosa Mara Borges da	Westphalen, Mauro Carpes, Minosso, Nathalia Michele Martins, Regio, Nicolas do Carmo, Gugliotta, Adriana de Mello, Rajchenberg, Mario, Silveira, Rosa Mara Borges da (2026): Unveiling the hidden diversity of neotropical Steccherinum and allied genera (Steccherinaceae, Basidiomycota). IMA Fungus 17: e 182915, DOI: 10.3897/imafungus.17.182915
468D4A4267705C5595F407B47008A6D5.text	468D4A4267705C5595F407B47008A6D5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cabalodontia lincangense (J. H. Dong & C. L. Zhao) Westphalen & Regio 2026	<div><p>Cabalodontia lincangense (J. H. Dong &amp; C. L. Zhao) Westphalen &amp; Regio comb. nov.</p><p>Basionym.</p><p>Steccherinum lincangense J. H. Dong &amp; C. L. Zhao, Mycosphere 15 (1): 1252. 2024.</p><p>Notes.</p><p>Cabalodontia lincangense was originally described based on a phylogeny that did not adequately represent Cabalodontia and other Steccherinaceae lineages, resulting in its placement in Steccherinum . In our analysis, the species nests within Cabalodontia, sharing morphological features such as an odontioid hymenophore, a monomitic hyphal system with clamped hyphae, and skeletocystidia tapering toward the apex. It is closely related to C. albofulva but can be differentiated by more widely spaced and longer aculei (2–4 per mm, 1–1.5 mm long) and shorter basidiospores measuring (3.5 –) 3.8–4.2 (– 4.5) × (2.3 –) 2.5–2.8 µm (Dong et al. 2024).</p></div>	https://treatment.plazi.org/id/468D4A4267705C5595F407B47008A6D5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Westphalen, Mauro Carpes;Minosso, Nathalia Michele Martins;Regio, Nicolas do Carmo;Gugliotta, Adriana de Mello;Rajchenberg, Mario;Silveira, Rosa Mara Borges da	Westphalen, Mauro Carpes, Minosso, Nathalia Michele Martins, Regio, Nicolas do Carmo, Gugliotta, Adriana de Mello, Rajchenberg, Mario, Silveira, Rosa Mara Borges da (2026): Unveiling the hidden diversity of neotropical Steccherinum and allied genera (Steccherinaceae, Basidiomycota). IMA Fungus 17: e 182915, DOI: 10.3897/imafungus.17.182915
ABB50256A31B533BB19C5DB64668F5C7.text	ABB50256A31B533BB19C5DB64668F5C7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cabalodontia tenuissima (C. L. Zhao & Y. X. Wu 2021) Westphalen & Regio 2026	<div><p>Cabalodontia tenuissima (C. L. Zhao &amp; Y. X. Wu) Westphalen &amp; Regio comb. nov.</p><p>Basionym.</p><p>Steccherinum tenuissimum C. L. Zhao &amp; Y. X. Wu, PLoS ONE 16 (1): e 0244520, 7. 2021.</p><p>Notes.</p><p>Cabalodontia tenuissima was originally described based on a phylogeny that included only Steccherinum spp., where it nested in a separate clade with S. xanthum, a species shown by our analyses to be a synonym of C. albofibrillosa . Morphologically, C. tenuissima differs from all other confirmed Cabalodontia spp. by presenting a dimitic hyphal system, although it shares other features of the genus, such as fragile and thin basidiomes with an odontioid hymenophore (Wu et al. 2021 a). It is possible that the skeletal hyphae reported by the authors represent undifferentiated lower portions of skeletocystidia, since the incrustations are restricted to the apical region. This interpretation is also supported by the drawings presented in the original description, where clamped hyphae are abundant and few thick-walled skeletal hyphae are shown, which is not common in truly dimitic species. Such thick-walled hyphal segments are also observed in other species of the genus, such as C. brunnea and C. albofulva, but they are scarce and, when carefully observed, give rise to cystidia. Further examination of the specimens is required to confirm this hypothesis. Nevertheless, because our phylogenetic analyses are well supported and include sequences from the type material (CLZhao 3153), we chose to transfer this species to Cabalodontia . Together with Irpex oreophilus (Lindsay &amp; Gilb.) Niemelä, these are the only two dimitic taxa recovered within the genus.</p></div>	https://treatment.plazi.org/id/ABB50256A31B533BB19C5DB64668F5C7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Westphalen, Mauro Carpes;Minosso, Nathalia Michele Martins;Regio, Nicolas do Carmo;Gugliotta, Adriana de Mello;Rajchenberg, Mario;Silveira, Rosa Mara Borges da	Westphalen, Mauro Carpes, Minosso, Nathalia Michele Martins, Regio, Nicolas do Carmo, Gugliotta, Adriana de Mello, Rajchenberg, Mario, Silveira, Rosa Mara Borges da (2026): Unveiling the hidden diversity of neotropical Steccherinum and allied genera (Steccherinaceae, Basidiomycota). IMA Fungus 17: e 182915, DOI: 10.3897/imafungus.17.182915
17B1C77BD1D852CEA66BFD9B1EF5B93C.text	17B1C77BD1D852CEA66BFD9B1EF5B93C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Steccherinum bononiae Westphalen & Gugliotta 2026	<div><p>Steccherinum bononiae Westphalen &amp; Gugliotta sp. nov.</p><p>Figs 8 A, 9 A, 11 C</p><p>Etymology.</p><p>In honor of Dr. Vera Bononi, for her contribution to the knowledge of Steccherinum in Brazil.</p><p>Diagnosis.</p><p>Distinguished by the combination of resupinate to effused-reflexed basidiomes with waxy spines upon drying, elongated skeletocystidia protruding above the hymenium, aculei measuring up to 0.75 mm, and basidiospores 2.6–3.6 × 1.8–2.5 µm.</p><p>Typification.</p><p>Brazil. São Paulo: São Luis do Paraitinga, Parque Estadual da Serra do Mar, Núcleo Santa Virgínia, Trilha Poço do Pito, 06 Jun. 2017, M. C. Westphalen 557 / 17 (SP 512686) .</p><p>Description.</p><p>Basidiomes adnate, resupinate to effused-reflexed, easily separable from the substratum and usually detaching upon drying, membranaceous to papery and pliable when fresh, turning corky and somewhat waxy upon drying; pilei when present arising from effused parts of the basidiomes and at the margins, usually imbricate and narrow, up to 8 mm wide, pileus surface cream to beige, fimbriate, faintly zonate; sterile margins entire, smooth, pelliculose, up to 2 mm wide, white to cream. Hymenophore hydnoid, pale salmon to pale yellowish when fresh, turning beige to tan upon drying, aculei 0.25–0.75 × 0.1–0.25 mm, with acute apices, with a pilose appearance from the protruding cystidia, solitary or more rarely laterally fused, somewhat crowded, 6–9 per mm. Subiculum cream to beige, homogeneous, slightly dense, up to 0.8 mm thick.</p><p>Hyphal system dimitic; subicular hyphae compact, not agglutinated; aculei tramal hyphae intertwined, parallel; generative hyphae clamped, thin to slightly thick-walled, hyaline, occasionally branched, 2–4 µm wide, more abundant in the base of the subiculum; skeletal hyphae thick-walled to almost solid, hyaline to slightly yellowish, 2–4.5 µm wide. Skeletocystidia abundant, arising from the trama and protruding above the hymenium, somewhat elongated, covered with medium-sized crystals, (20–35 – 65 × 5–9 (– 10) µm; leptocystidia present, often scattered and more abundant at the apical portion of the aculei, clavate to fusoid, smooth to apically encrusted. Basidia clavate, tetrasterigmate, 10–14 × 4–5 µm. Basidiospores broadly ellipsoid, hyaline, thin-walled IKI –, CB –, 2.6–3.6 × 1.8–2.5 µm.</p><p>Mating system.</p><p>Tetrapolar. Monosporic cultures obtained from specimen NR 71.</p><p>Habitat and distribution.</p><p>Known from southern, southeastern, and northeastern Brazil. Likely widespread in the Brazilian Atlantic Rainforest.</p><p>Specimens examined.</p><p>Brazil • Pernambuco: Olinda, 7 GAC - Batalhão do Exército, 16 Jun. 2018, R. S. Chikowski RC 1625 (URM 93107) . • Rio Grande do Sul: São Francisco de Paula, FLONA, 12 Mar. 2022, M. C. Westphalen 726 / 22 (ICN 213868) ; • Dom Pedro de Alcântara, RPPN Mata do Prof. Baptista, 17 Nov. 2022, N. C. Regio NR 71 (ICN 213869) . • São Paulo: São Paulo, Parque Estadual das Fontes do Ipiranga, 09 May 2014, A. M. Gugliotta 1615 (SP 512683) ; • Parque Estadual Cantareira, Núcleo Engordador, Trilha da Cachoeira, 24 Apr. 2017, M. C. Westphalen 547 / 17 (SP 512685) ; • Ribeirão Grande. Parque Estadual Intervales, Trilha roda d’água, 07 Jul. 2015, V. Motato-Vásquez MV 446 (SP 512675) .</p><p>Notes.</p><p>Basidiomes of S. bononiae exhibit considerable variation, ranging from completely effused to effused-reflexed, and aculei vary in size from 0.25 to 0.75 mm long. The basidiospore size and shape in this species resemble those of S. molle and S. resinaceum . However, S. molle can be distinguished by its softer, membranaceous basidiomes, slightly shorter spines, and wider skeletocystidia, which are more deeply embedded in the trama and covered with larger, chunky crystals. Steccherinum resinaceum, in turn, can be distinguished by its denser basidiomes, laterally fused aculei that are often bifurcated at the apices, and embedded cystidia covered with large crystals. Phylogenetically, S. bononiae forms a sister clade to S. elegantissimum, but the latter can be distinguished by its thinner, shorter aculei and larger basidiospores. Additionally, S. elegantissimum typically grows on thin branches, forming small concrescent patches, whereas S. bononiae usually forms basidiomes as a single patch. Both species share the presence of elongated and projecting cystidia covered with a somewhat organized cap of crystals, but these crystals are slightly larger in S. bononiae (Fig. 9).</p></div>	https://treatment.plazi.org/id/17B1C77BD1D852CEA66BFD9B1EF5B93C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Westphalen, Mauro Carpes;Minosso, Nathalia Michele Martins;Regio, Nicolas do Carmo;Gugliotta, Adriana de Mello;Rajchenberg, Mario;Silveira, Rosa Mara Borges da	Westphalen, Mauro Carpes, Minosso, Nathalia Michele Martins, Regio, Nicolas do Carmo, Gugliotta, Adriana de Mello, Rajchenberg, Mario, Silveira, Rosa Mara Borges da (2026): Unveiling the hidden diversity of neotropical Steccherinum and allied genera (Steccherinaceae, Basidiomycota). IMA Fungus 17: e 182915, DOI: 10.3897/imafungus.17.182915
EA3F572F522856E3B8A27CAA67DB4127.text	EA3F572F522856E3B8A27CAA67DB4127.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Steccherinum elegantissimum Westphalen & R. M. Silveira 2026	<div><p>Steccherinum elegantissimum Westphalen &amp; R. M. Silveira sp. nov.</p><p>Figs 8 B, 9 B, 11 D, 11 L</p><p>Etymology.</p><p>Refers to the delicate basidiomes with thin aculei and the elongated projecting cystidia.</p><p>Diagnosis.</p><p>Characterized by effused basidiome with small aculei (up to 0.5 mm long), a cottony subiculum, elongated and thin skeletocystidia protruding above the hymenium, and basidiospores with (3.4 –) 3.5–5.0 (– 5.1) × 2.2–2.9 (– 3.0) µm.</p><p>Typification.</p><p>Brazil. São Paulo: Ribeirão Grande. Parque Estadual Intervales, 28 Feb. 2018, M. C. Westphalen 633 / 18 (SP 512681) .</p><p>Description.</p><p>Basidiomes adnate, effused, usually not detaching when dried, but easily separable if pulled from the substratum, formed by the coalescence of several small patches, membranaceous and pliable when fresh, turning papery and fragile after drying; sterile margins entire, byssoid, smooth to finely fimbriate, thin, up to 0.5 mm wide. Hymenophore hydnoid, cream to pale salmon when fresh, more or less unchanging or slightly duller upon drying, aculei 0.3–0.5 (0.55) × 0.1–0.2 mm, with acute apices, pilose from the protruding cystidia, solitary or rarely fused at the base, sub-distant, 5–8 per mm. Subiculum white, homogeneous, with a loose cottony and soft consistency, up to 0.3 mm thick.</p><p>Hyphal system dimitic, subicular hyphae loosely arranged, tramal hyphae intertwined and somewhat more densely arranged; generative hyphae clamped, thin to slightly thick-walled, hyaline, occasionally branched, 2–3 µm wide; skeletal hyphae thick-walled to almost solid, abundant throughout the basidiome, hyaline, 2–4 µm wide. Skeletocystidia abundant, arising from the trama and protruding above the hymenium, somewhat thin and elongated, covered with small crystals, 20–70 (– 75) × 4–8, but rarely above 6 µm wide; leptocystidia present, often scattered and more abundant at the apical portion of the aculei, clavate to fusoid, smooth. Basidia clavate, tetrasterigmate, 13–15 × 4.5–5 µm. Basidiospores ellipsoid to narrowly ellipsoid, hyaline, thin-walled, IKI –, CB –, (3.4 –) 3.5–5.0 (– 5.1) × 2.2–2.9 (– 3.0) µm.</p><p>Mating system.</p><p>Tetrapolar. Monosporic cultures obtained from specimens 720 / 21 and 721 / 21.</p><p>Habitat and distribution.</p><p>Known from Araucaria forest and Atlantic Rainforest areas in southern and southeastern Brazil.</p><p>Specimens examined.</p><p>Brazil • Rio Grande do Sul: Canela, FLONA, 22 Oct. 2021, M. C. Westphalen 720 / 21 and 721 / 21 (ICN 213870 and 213871) .</p><p>Notes.</p><p>This species can be recognized by its small, sub-distant aculei and basidiomes formed by several concrescent small patches, typically growing on thin branches. Microscopically, the basidiospores are larger compared to other Neotropical Steccherinum species, measuring (3.4 –) 3.5–5.0 (– 5.1) × 2.2–2.9 (– 3.0) µm. Additionally, S. elegantissimum has the longest and thinnest cystidia in the group, covered by a layer of thin crystals (Figs 9 B, 11 L). The basidiomes with a cottony subiculum resemble those of S. perparvulum and S. molle, but the latter two species have smaller basidiospores and typically form larger patches that cover wider branches or logs.</p></div>	https://treatment.plazi.org/id/EA3F572F522856E3B8A27CAA67DB4127	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Westphalen, Mauro Carpes;Minosso, Nathalia Michele Martins;Regio, Nicolas do Carmo;Gugliotta, Adriana de Mello;Rajchenberg, Mario;Silveira, Rosa Mara Borges da	Westphalen, Mauro Carpes, Minosso, Nathalia Michele Martins, Regio, Nicolas do Carmo, Gugliotta, Adriana de Mello, Rajchenberg, Mario, Silveira, Rosa Mara Borges da (2026): Unveiling the hidden diversity of neotropical Steccherinum and allied genera (Steccherinaceae, Basidiomycota). IMA Fungus 17: e 182915, DOI: 10.3897/imafungus.17.182915
374EBD760E2951DEAD9ACBEE1650DB00.text	374EBD760E2951DEAD9ACBEE1650DB00.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Steccherinum molle Westphalen & Minosso 2026	<div><p>Steccherinum molle Westphalen &amp; Minosso sp. nov.</p><p>Figs 8 C, 9 C, 11 E, 11 K</p><p>Etymology.</p><p>Refers to the soft and pliable consistency of the basidiomes.</p><p>Diagnosis.</p><p>Recognized by soft basidomes with a cottony subiculum, small aculei up to 0.7 mm long, skeletocystidia usually embedded in the trama and encrusted with large crystals, and basidiospores 2.5–3.5 (– 3.6) × 1.8–2.4 (– 2.5) µm.</p><p>Typification.</p><p>Brazil. Rio Grande do Sul: Dom Pedro de Alcântara, RPPN Mata do Prof. Baptista, 10. Sep. 2022, M. C. Westphalen 734 / 22 (ICN 213874) .</p><p>Description.</p><p>Basidiomes adnate, resupinate, easily separable from the substratum, usually detaching upon drying, coalescing to form large patches, soft and membranaceous when fresh, unchanging to slightly papery upon drying, but remaining soft and pliable, easily tearing; sterile margins thin, cottony, smooth to finely floccose, up to 1.5 mm wide. Hymenophore hydnoid, cream to pale orange when fresh, unchanged upon drying, aculei (0.3 –) 0.4–0.5 (– 0.7) × 0.1–0.2 (– 0.25) mm, with acute apices, sometimes slightly asperulate from the protuding cystidia, solitary or more rarely fused at the base, sub-distant, 5–7 per mm. Subiculum white, homogeneous, loose, and cottony, very thin, up to 0.4 mm thick.</p><p>Hyphal system dimitic, subicular hyphae very loosely arranged, tramal hyphae intertwined and somewhat densely arranged, subparallel; generative hyphae clamped, thin to slightly thick-walled, hyaline, occasionally branched, 2–3 µm wide; skeletal hyphae thick-walled to almost solid, hyaline, 2–4 µm wide. Skeletocystidia abundant, immersed in the trama or protruding into the hymenium, coarsely encrusted with large crystals, (20 –) 30–60 × 7–13 (– 15); leptocystidia present, more commonly seen on the apices of the aculei, smooth to finely encrusted, mostly clavate. Basidia clavate, tetrasterigmate, 10–14 × 4–5 µm. Basidiospores ellipsoid, hyaline, thin-walled, IKI –, CB –, 2.5–3.5 (– 3.6) × 1.8–2.4 (– 2.5) µm.</p><p>Mating system.</p><p>Tetrapolar. Monosporic cultures obtained from specimen 734 / 22.</p><p>Habitat and distribution.</p><p>Known from Atlantic Rainforest areas in southeastern and northeastern Brazil and Araucaria Forests in southern Brazil. Mostly found in high-altitude regions above 700 m. Likely widespread throughout the Atlantic rainforest biome.</p><p>Specimens examined.</p><p>Brazil • Rio Grande do Sul: São Francisco de Paula, Hotel Parador Hampel, Trilha, 19 Jun. 2017, M. C. Westphalen 568 / 17 (SP 512692) ; • ibid., FLONA, 14 May 2018, M. C. Westphalen 641 / 18 (SP 512689); • Farroupilha, Parque dos Pinheiros, 20 Apr. 2018, M. C. Westphalen 661 / 18 (SP 512690) ; • ibid., 27 Mar. 2019 M. C. Westphalen 687 / 19 (SP 512691); • ibid., 21 Jan. 2023 M. C. Westphalen 739 / 23 (ICN 213875) . • São Paulo: Ribeirão Grande, Parque Estadual Intervales, Trilha da gruta, 07 Jul. 2015, V. Motato-Vásquez MV 450 (SP 512672) . • Sergipe: Areia Branca, Parque Nacional Serra de Itabaiana, 25 Jul. 2025, R. S. Souza RSS 236 (URM) .</p><p>Notes.</p><p>This species can be recognized mainly by the soft and pliable basidiomes with a cottony subiculum, usually forming large patches on the substratum, small aculei, and embedded cystidia encrusted with large crystals. Steccherinum resinaceum presents similar basidiospores and cystidia but can be distinguished by the harder, waxy basidiomes and laterally fused or bifurcate aculei. Steccherinum perparvulum is similar macroscopically but presents papery basidiomes upon drying, not soft or pliable, shorter basidiospores, and slightly thinner cystidia encrusted with smaller crystals.</p></div>	https://treatment.plazi.org/id/374EBD760E2951DEAD9ACBEE1650DB00	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Westphalen, Mauro Carpes;Minosso, Nathalia Michele Martins;Regio, Nicolas do Carmo;Gugliotta, Adriana de Mello;Rajchenberg, Mario;Silveira, Rosa Mara Borges da	Westphalen, Mauro Carpes, Minosso, Nathalia Michele Martins, Regio, Nicolas do Carmo, Gugliotta, Adriana de Mello, Rajchenberg, Mario, Silveira, Rosa Mara Borges da (2026): Unveiling the hidden diversity of neotropical Steccherinum and allied genera (Steccherinaceae, Basidiomycota). IMA Fungus 17: e 182915, DOI: 10.3897/imafungus.17.182915
0A01D71BAC6755D28EB9CF34490005C2.text	0A01D71BAC6755D28EB9CF34490005C2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Steccherinum perparvulum Hjortstam & Ryvarden	<div><p>Steccherinum perparvulum Hjortstam &amp; Ryvarden</p><p>Figs 8 D, 9 D, 11 F</p><p>Description.</p><p>Full description in: Hjortstam and Ryvarden (2008).</p><p>Mating system.</p><p>Tetrapolar. Monosporic cultures obtained from specimens 710 / 20 and 742 / 23.</p><p>Habitat and distribution.</p><p>Widespread in southern to southeastern Brazil, mainly in high-altitude areas.</p><p>Specimens examined.</p><p>Brazil • Rio Grande do Sul: Nova Roma do Sul, ponte velha, 05 Apr. 2017, M. C. Westphalen 524 / 17 (SP 512658) ; • São Francisco de Paula, CPCN Pró-Mata, 17 May 2018, M. C. Westphalen 659 / 18 (SP 512661) ; • ibid., 22. Apr. 2023 M. C. Westphalen 744 / 23 (ICN 213878); • ibid., 19 Apr; 2024, N. C. Regio NR 186 (ICN 213879); • ibid., FLONA, 19 Apr. 2023, M. C. Westphalen 742 / 23 (ICN 213877); • Caxias do Sul, Parque da Gruta Nossa Senhora de Lourdes, 28 Mar. 2019, M. C. Westphalen 692 / 19; ibid., 09. Nov. 2020, M. C. Westphalen 710 / 20 (ICN 213876) . • São Paulo: Moji-Guaçu, Fazenda Campininha, 29–30 Jan 1987, D. Pegler, K. Hjortstam &amp; L. Ryvarden 24589 (O - holotype) ; • Parelheiros, Parque Estadual da Serra do Mar, Núcleo Curucutu, 16 Nov. 2016, V. Motato-Vásquez MV 815 (SP 512656) ; • São Paulo, Parque CIENTEC, 07 Jul. 2016, V. Motato-Vásquez MV 728 (SP 512657) ; • ibid., Parque Estadual Cantareira, Núcleo Engordador, Trilha da Cachoeira, 24 Apr. 2017, M. C. Westphalen 543 / 17 (SP 512659) ; • Santo André, Reserva Biológica do Alto da Serra de Paranapiacaba, 25 Aug. 2017, M. C. Westphalen 592 / 17 (SP 512660) .</p><p>Notes.</p><p>This species is primarily distinguished by having the smallest basidiospores (2.3–3.1 × 1.7–2.2 µm) of all Neotropical hydnoid / odontioid Steccherinum, usually measuring under 3 µm long (Table 5). Macroscopically, it typically forms large basidiomes with short aculei and a papery consistency when dried. While most of the specimens examined were strictly resupinate, some exhibited small pilei along the margins. We examined the holotype of S. perparvulum and found several specimens with basidiospore sizes and general morphological characteristics consistent with the species. However, our phylogenetic analyses revealed that these specimens comprise at least three distinct lineages (Figs 1, 3). These lineages show minor differences in the ITS region, although subtle variations also occur among specimens within the same lineage (Fig. 12).</p><p>To determine whether the genetic differences observed represent distinct biological species or reflect broader molecular variability within the ITS region of S. perparvulum, we conducted mating tests with monosporic cultures from three different specimens (Table 6). Two of these specimens belonged to the same lineage (710 and 744, lineage 1), whereas the third represented a different lineage (742, lineage 3). The mating tests showed positive results only between specimens of the same lineage, with clamp connections present in 11 of 12 monosporic pairings. In contrast, all confrontations with specimen 742 were negative, with only simple-septate hyphae present. This further supports the hypothesis that the observed molecular differences represent separate species within the group. Unfortunately, cultures from lineage 2 were not available for compatibility testing. Further studies incorporating additional monosporic cultures would be valuable to explore potential intercompatibility within the group, especially considering that lineage 2 is phylogenetically very close to lineage 1.</p><p>While our studies corroborate at least two different species within S. perparvulum, no morphological or biogeographical evidence was found to support their segregation. Therefore, we chose to retain these taxa under the same name for the time being, treating it as a species complex. Considering their morphological and phylogenetic affinities, this approach aims to facilitate the taxonomy of the group rather than complicating it by increasing the number of species and adding morphologically indistinguishable taxa. In addition, at present, it is not possible to define which of the lineages represents S. perparvulum s. s. Sequences from the type specimen or the type locality could further elucidate this issue and help clarify the taxonomy of this species complex.</p></div>	https://treatment.plazi.org/id/0A01D71BAC6755D28EB9CF34490005C2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Westphalen, Mauro Carpes;Minosso, Nathalia Michele Martins;Regio, Nicolas do Carmo;Gugliotta, Adriana de Mello;Rajchenberg, Mario;Silveira, Rosa Mara Borges da	Westphalen, Mauro Carpes, Minosso, Nathalia Michele Martins, Regio, Nicolas do Carmo, Gugliotta, Adriana de Mello, Rajchenberg, Mario, Silveira, Rosa Mara Borges da (2026): Unveiling the hidden diversity of neotropical Steccherinum and allied genera (Steccherinaceae, Basidiomycota). IMA Fungus 17: e 182915, DOI: 10.3897/imafungus.17.182915
589A4DB46D955DF6AA259B7FAD240669.text	589A4DB46D955DF6AA259B7FAD240669.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Steccherinum resinaceum Westphalen & Minosso 2026	<div><p>Steccherinum resinaceum Westphalen &amp; Minosso sp. nov.</p><p>Figs 8 E, 9 E, 11 G</p><p>Etymology.</p><p>Refers to the hard and waxy consistency of the basidiomes when dried.</p><p>Diagnosis.</p><p>Distinguished from other species in the genus mainly by the waxy and dense basidiomes upon drying, the laterally fused aculei that bifurcate at apices, and the skeletocystidia encrusted with large crystals, usually embedded in the trama.</p><p>Typification.</p><p>Brazil. São Paulo: São Luís do Paraitinga, Parque Estadual da Serra do Mar, Núcleo Santa Virgínia, Trilha Pirapitinga, 05 Jun. 2017, M. C. Westphalen 551 / 17 (SP 512670) .</p><p>Description.</p><p>Basidiomes adnate, resupinate, easily separable from the substratum and usually detaching upon drying, membranaceous to papery and pliable when fresh, turning waxy and somewhat rigid upon drying; sterile margins entire, smooth, pelliculose, up to 1 mm wide. Hymenophore hydnoid, cream to pale yellowish when fresh, turning beige to ochraceous upon drying, aculei (0.3 –) 0.4–0.75 × 0.15–0.3 (– 0.5) mm, usually with straight to bifurcate apices with a pilose appearance from the protruding cystidia, solitary or more commonly laterally fused, somewhat crowded, 5–6 per mm. Subiculum cream to beige, homogeneous, dense, up to 0.3 mm thick.</p><p>Hyphal system dimitic, subicular hyphae very compact, tramal hyphae intertwined, subparallel; generative hyphae clamped, thin to slightly thick-walled, hyaline, occasionally branched, 2–4 µm wide, more abundant in the base of the subiculum; skeletal hyphae thick-walled to almost solid, hyaline to slightly yellowish, 2–4.5 µm wide. Skeletocystidia abundant, immersed in the trama or protruding into the hymenium, coarsely encrusted with large crystals, 25–45 (– 50) × (7 –) 9–11 (– 12) µm, some thinner and longer cystidia also observed immersed in the subiculum, up to 60 µm long and 6–8 µm wide; leptocystidia present, abundant in the apices of the aculei, clavate, ventricose, or capitate, smooth or with a crown of crystals. Basidia clavate, tetrasterigmate, 10–12 × 3.5–5 µm. Basidiospores broadly ellipsoid, hyaline, thin-walled IKI –, CB –, 2.7–3.4 (– 3.5) × 1.7–2.4 µm.</p><p>Habitat and distribution.</p><p>Known only from Atlantic Rainforest areas in southeastern Brazil.</p><p>Specimens examined.</p><p>Brazil • São Paulo: São Paulo, Parque Estadual Cantareira, Núcleo Engordador, Trilha da Cachoeira, 24 Apr. 2017, M. C. Westphalen 540 / 17 (SP 512671) ; • São Luís do Paraitinga, Parque Estadual da Serra do Mar, Núcleo Santa Virgínia, Trilha Olho d’ água, 13 Feb. 2019 M. C. Westphalen 665 / 19 (SP 512687, ICN 213880) ; • São Paulo, Parque Estadual das Fontes do Ipiranga, 18 Feb. 2019, M. C. Westphalen 679 / 19 (SP 512669) .</p><p>Notes.</p><p>Steccherinum resinaceum is characterized by its waxy and somewhat hard basidiomes when dried, a hymenophore composed of laterally fused, often bifurcating aculei, and coarsely encrusted cystidia with large crystals. Macroscopically, the basidiomes of S. bononiae are somewhat similar but differ in having solitary aculei with a pilose appearance due to prominently protruding skeletocystidia, whereas in S. resinaceum the skeletocystidia are visible only at the apices of the aculei (Fig. 8 E). Additionally, the cystidia in S. bononiae are longer and typically covered with small- to medium-sized crystals.</p><p>Steccherinum molle shares similar cystidial morphology and basidiospore size with S. resinaceum . Nonetheless, it can be distinguished by its softer basidiomes with a cottony subiculum and slightly shorter and thinner aculei. Unfortunately, we could not obtain monosporic cultures of S. resinaceum to confirm its mating system.</p></div>	https://treatment.plazi.org/id/589A4DB46D955DF6AA259B7FAD240669	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Westphalen, Mauro Carpes;Minosso, Nathalia Michele Martins;Regio, Nicolas do Carmo;Gugliotta, Adriana de Mello;Rajchenberg, Mario;Silveira, Rosa Mara Borges da	Westphalen, Mauro Carpes, Minosso, Nathalia Michele Martins, Regio, Nicolas do Carmo, Gugliotta, Adriana de Mello, Rajchenberg, Mario, Silveira, Rosa Mara Borges da (2026): Unveiling the hidden diversity of neotropical Steccherinum and allied genera (Steccherinaceae, Basidiomycota). IMA Fungus 17: e 182915, DOI: 10.3897/imafungus.17.182915
E4650FA5EAB75498B9966BF1F05720C9.text	E4650FA5EAB75498B9966BF1F05720C9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Steccherinum subochraceum Bononi & Hjortstam	<div><p>Steccherinum subochraceum Bononi &amp; Hjortstam</p><p>Figs 8 F, 9 F, 10 A, 11 H</p><p>Description.</p><p>Full description in Hjortstam and Bononi (1986).</p><p>Mating system.</p><p>Tetrapolar. Monosporic cultures obtained from specimen 730 / 22.</p><p>Habitat and distribution.</p><p>Known from the Atlantic Rainforest in southern and southeastern Brazil.</p><p>Specimens examined.</p><p>Brazil • Rio Grande do Sul: São Salvador, Montenegro, 04 Apr. 1945, Rick s. n. (PACA 22824 - holotype of Iprex hydenus Rick) ; • São Francisco de Paula, FLONA, 12 Mar. 2022, M. C. Westphalen 730 / 22 (ICN 213881) ; • ibid., 25 Nov. 2023, M. C. Westphalen 748 / 23 (ICN 213883); • Farroupilha, Parque dos Pinheiros, 22 Jul. 2023, M. C. Westphalen 746 / 23 (ICN 213882) ; • Canela, FLONA, 13 Jul. 2024, Westphalen 761 / 24 (ICN 313884) . • São Paulo: São Paulo, Parque Estadual das Fontes do Ipiranga, 06 Oct. 1966, H. Requejo s. n. (SP 97591 - holotype) .</p><p>Notes.</p><p>S. subochraceum was described from southeastern Brazil to provide a valid name for Iprex hydenus, originally described by Rick (1959) but considered invalid due to the lack of a designated type specimen. Among Neotropical hydnoid Steccherinum species, S. subochraceum is readily distinguished by the effused-reflexed basidiomes with notably long aculei (Figs 8 F, 10 A), usually measuring over 1.5 mm long (0.85–2.25 × 0.15–0.45 mm) (Table 4). Microscopically, it presents thin, elongated cystidia [25–60 (– 70) × 5–8 (– 10) µm] immersed in the trama, sometimes protruding at the apices of the aculei, and subglobose basidiospores measuring (3.2 –) 3.3–4.2 × 2.5–3.8 µm. Steccherinum larssonii and Steccherinum basibadium share a similar basidiospore size range (3.5–4.5 × 3.0–3.5 µm and 3.6–4.5 × 3.1–3.4 µm, respectively) but can be distinguished by their significantly shorter aculei, which reach a maximum length of 0.75 mm. In addition, Steccherinum basibadium forms more developed pilei with a brownish surface, whereas S. larssonii often produces completely resupinate basidiomes or only small pilei at the margins (Maas Geesteranus 1974; Westphalen et al. 2021).</p><p>The subicular generative hyphae of S. subochraceum are regularly clamped, although simple-septate hyphae were observed in the trama of some specimens. Simple-septate hyphae were readily observed in specimen 746 / 23 but were less common in 730 / 22. Phylogenetically, S. subochraceum forms a sister clade to S. undigerum, a Neotropical species with somewhat similar basidiome morphology (effused-reflexed with imbricate pilei) and that also exhibits a strongly fimbriate mycelium in culture. However, S. undigerum differs in having a poroid hymenophore with dentate dissepiments and slightly larger basidiospores (4–5 × 3.5–4.5 μm).</p></div>	https://treatment.plazi.org/id/E4650FA5EAB75498B9966BF1F05720C9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Westphalen, Mauro Carpes;Minosso, Nathalia Michele Martins;Regio, Nicolas do Carmo;Gugliotta, Adriana de Mello;Rajchenberg, Mario;Silveira, Rosa Mara Borges da	Westphalen, Mauro Carpes, Minosso, Nathalia Michele Martins, Regio, Nicolas do Carmo, Gugliotta, Adriana de Mello, Rajchenberg, Mario, Silveira, Rosa Mara Borges da (2026): Unveiling the hidden diversity of neotropical Steccherinum and allied genera (Steccherinaceae, Basidiomycota). IMA Fungus 17: e 182915, DOI: 10.3897/imafungus.17.182915
8202208943AC52D2806711C6C4DAE67C.text	8202208943AC52D2806711C6C4DAE67C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Steccherinum undulatum Westphalen & R. M. Silveira 2026	<div><p>Steccherinum undulatum Westphalen &amp; R. M. Silveira sp. nov.</p><p>Figs 8 G, 9 G, 10 B, 11 I</p><p>Etymology.</p><p>Refers to the basidiomes with small, wavy pilei.</p><p>Diagnosis.</p><p>Recognized mainly by the combination of basidiomes formed by several small, wavy, imbricate pilei, aculei with 0.75–1 mm long, and ellipsoid basidiospores 3.3–4.1 × 2.0–2.7 (– 2.8) µm.</p><p>Typification.</p><p>Brazil. Rio Grande do Sul: São Francisco de Paula, CPCN Pró-Mata, 20 Apr. 2023, M. C. Westphalen 743 / 23 (ICN 213872) .</p><p>Description.</p><p>Basidiomes adnate, concrescent, effused-reflexed, with several small, wavy, imbricate pilei, formed by the coalescence of several small pilei with conjoined reflexed bases, usually not detaching when dried, but easily separable if pulled from the substratum, somewhat membranaceous and pliable when fresh, turning papery to corky upon drying and somewhat waxy; pilei small, up to 4 mm wide and 9 mm in length, pilear surface cream to beige, tomentose, sulcate, and sometimes faintly zonate; sterile margins entire, somewhat cottony, smooth to finely fimbriate, up to 1.5 mm wide. Hymenophore hydnoid, at first orange in young and fresh specimens, then turning pale salmon to beige with age and upon drying, aculei 0.75–1.0 × 0.2–0.4 mm, with acute to round apices, sometimes slightly asperulate from the protruding cystidia, solitary or more rarely laterally fused, crowded, 4–6 per mm. Subiculum white, homogeneous, up to 0.6 mm thick.</p><p>Hyphal system dimitic, subicular hyphae compact, not agglutinated, tramal hyphae parallel, intertwined, and densely arranged; generative hyphae clamped, thin to slightly thick-walled, hyaline, occasionally branched, 2–4 µm wide; skeletal hyphae thick-walled to almost solid, abundant throughout the basidiome, hyaline, 2–5 µm wide. Skeletocystidia abundant, immersed in the trama or with the apices protruding into the hymenium, covered with a thick cap of small crystals, 20–60 × 7–10 µm; leptocystidia present, often scattered and somewhat inconspicuous, mostly clavate or with a rounded apex, smooth. Basidia clavate, tetrasterigmate, 12–16 × 4–5 µm. Basidiospores ellipsoid, hyaline, thin-walled, IKI-, CB-, 3.3–4.1 × 2.0–2.7 (– 2.8) µm.</p><p>Habitat and distribution.</p><p>Known only from Araucaria forests in southern Brazil.</p><p>Specimens examined.</p><p>Brazil • Rio Grande do Sul: Canela, FLONA, 13 Jul. 2024, M. C. Westphalen 760 / 24 (ICN 213873) .</p><p>Notes.</p><p>This species can be distinguished by its basidiomes consisting of small, wavy, imbricate pilei (Fig. 10 B). One of the collections studied (760 / 24) exhibited a bright orange coloration when fresh, whereas the other (743 / 23) displayed a more faded salmon hue. This suggests that only young, fresh specimens display brighter colors, which gradually turn paler as they mature or dry. The aculei of S. undulatum are the second largest in the group, being smaller only than those of S. subochraceum . Microscopically, the species also features slightly larger basidiospores when compared to S. molle, S. resinaceum, and S. bononiae, but smaller than those of S. elegantissimum .</p><p>Although surveys for Steccherinum species have been ongoing since 2017, only two specimens of S. undulatum have been found, both in Araucaria forests in southern Brazil, approximately 45 km apart. This suggests that it is likely rarer than other species in the group, which have been more frequently collected and are more widely distributed.</p></div>	https://treatment.plazi.org/id/8202208943AC52D2806711C6C4DAE67C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Westphalen, Mauro Carpes;Minosso, Nathalia Michele Martins;Regio, Nicolas do Carmo;Gugliotta, Adriana de Mello;Rajchenberg, Mario;Silveira, Rosa Mara Borges da	Westphalen, Mauro Carpes, Minosso, Nathalia Michele Martins, Regio, Nicolas do Carmo, Gugliotta, Adriana de Mello, Rajchenberg, Mario, Silveira, Rosa Mara Borges da (2026): Unveiling the hidden diversity of neotropical Steccherinum and allied genera (Steccherinaceae, Basidiomycota). IMA Fungus 17: e 182915, DOI: 10.3897/imafungus.17.182915
