identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
2AB6F26126435B55A47051547AD055E8.text	2AB6F26126435B55A47051547AD055E8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myllocerina Pierce 1913	<div><p>Subtribe Myllocerina Pierce, 1913</p><p>Diagnosis.</p><p>Prothorax in lateral view with anterior margin straight and lacking vibrissae.</p></div>	https://treatment.plazi.org/id/2AB6F26126435B55A47051547AD055E8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wijesinghe, Dilshara D.;Hong, Ki-Jeong	Wijesinghe, Dilshara D., Hong, Ki-Jeong (2026): Bridging biodiversity gaps: Updated taxonomy and diversity of Sri Lankan Myllocerus (Coleoptera, Curculionidae, Entiminae). ZooKeys 1284: 87-111, DOI: 10.3897/zookeys.1284.191034
30F4CA722B9A58F0920E1039810404B2.text	30F4CA722B9A58F0920E1039810404B2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myllocerus angulatipes Marshall 1916	<div><p>Myllocerus angulatipes Marshall, 1916</p><p>Fig. 2 A, B</p><p>Myllocerus angulatipes Marshall, 1916: 316 . Type locality: Ceylon-Dikoya; Type repository: NHMUK. Type examined.</p><p>Material examined.</p><p>Paratype. Sri Lanka • 1 ♀; Central Province, Dikoya, 3800–4200 ft; 6 Dec. 1881 – 16 Jan. 1882; G. Lewis leg.; NHMUK, NHMUK 013663874 .</p><p>Diagnosis.</p><p>Body dark brown; dorsal surface with brown scaling, variegated with darker patches on the elytra. Head bearing a broad lateral green stripe ventral to eye, continued along prothorax and elytra; on elytra stripe broader and extending to elytral stria 6 near base and to elytral stria 5 posterior to mid-length, irregular and variable. Head finely striolate, the striolation visible through thin scaling; eyes lateral and convex; interocular distance broader than length of eye. Rostrum about as long as broad in ♂ and slightly shorter in ♀, narrowed from base to beyond mid-length; apical emargination moderately deep, forming a right angle. Antennae reddish brown; scape with sparse minute pale scales and curved black setae; funicular antennomeres 1 and 2 equal, funicular antennomere 3 longer than 4; funicular antennomere 7 longer than broad. Prothorax transverse; sides rounded and broadest at mid-length; shallowly constricted and transversely impressed anteriorly and posteriorly; posterior margin truncate and not broader than anterior margin, which is rounded dorsally. Elytra subtruncate at base, parallel-sided in ♂ and slightly dilated posteriorly in ♀; dorsal surface almost flat; disc with a small transverse flattened area posterior to scutellar shield, followed posteriorly by a shallow, broad, oblique impression extending from elytra humeri to suture. Legs reddish brown; femora with a single tooth; inner margin of protibiae with a distinct angulation proximal to mid-length, weaker on metathoracic pair.</p><p>Remarks.</p><p>This species is similar to M. zeylanicus Marshall, 1916, but differs by the following characters: head bearing a broad lateral green stripe ventral to eye, continued along prothorax and elytra; rostrum with apical emargination forming a right angle, median carina reaching to frons; inner margin of protibiae with a distinct angulation proximal to mid-length.</p><p>Plant association.</p><p>Unknown.</p><p>Distribution.</p><p>Sri Lanka.</p></div>	https://treatment.plazi.org/id/30F4CA722B9A58F0920E1039810404B2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wijesinghe, Dilshara D.;Hong, Ki-Jeong	Wijesinghe, Dilshara D., Hong, Ki-Jeong (2026): Bridging biodiversity gaps: Updated taxonomy and diversity of Sri Lankan Myllocerus (Coleoptera, Curculionidae, Entiminae). ZooKeys 1284: 87-111, DOI: 10.3897/zookeys.1284.191034
1EC4932D55455163AC5946A7F9C76352.text	1EC4932D55455163AC5946A7F9C76352.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myllocerus curvicornis (Fabricius 1792)	<div><p>Myllocerus curvicornis (Fabricius, 1792)</p><p>Fig. 2 C, D</p><p>Curculio curvicornis Fabricius, 1792: 488 . Type locality: India orientalis; Type repository: Zoological Museum, University of Copenhagen. Type not examined.</p><p>= Myllocerus transmarinus Boheman, 1834: 429; Marshall 1916: 343; Ramamurthy and Ghai 1988: 455. Type locality: India orientalis; Type repository: Swedish Museum of Natural History, Stockholm.</p><p>Material examined.</p><p>Sri Lanka • 1 ♂ 1 ♀; North Central Province, Dambewatana, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=80.560165&amp;materialsCitation.latitude=7.9366665" title="Search Plazi for locations around (long 80.560165/lat 7.9366665)">Galkiriyagama</a>; 7°56.2'N, 80°33.61'E; 8 Jan. 2020; KJ. Hong leg.; on Mangifera indica ( Anacardiaceae); SCNU • 1 ♂ 1 ♀; North Central Province, Bakamuna, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=80.84233&amp;materialsCitation.latitude=7.783167" title="Search Plazi for locations around (long 80.84233/lat 7.783167)">Sirikanduyaya</a>; 7°46.99'N, 80°50.54'E; 20 Feb. 2024; DD. Wijesinghe leg.; on Gliricidia sepium ( Fabaceae); SCNU • 1 ♂; Western Province, Colombo; 7–27 Apr. 1882; G. Lewis leg.; NHMUK, NHMUK 013663873 .</p><p>Diagnosis.</p><p>Body black to dark brown, with grey and brown scaling. Prothorax dorsally either dark brown with a narrow median pale brown line, or pale brown with a dark brown, longitudinal, lateral stripes. Elytra dark to pale brown dorsally, irregularly mottled with small grey spots, occasionally partly confluent. Lateral margins of head, prothorax, and elytra, as well as ventral surfaces, grey to whitish. Head with sparse obscure punctation concealed by scaling; eyes sublateral. Antennae dark brown, with depressed grey setae; funicular antennomere 2 distinctly longer than 1, funicular antennomere 3 slightly longer than 4. Prothorax transverse, sides slightly rounded, posterior margin moderately bisinuate, not broader than anterior margin. Elytra rather broad, separately rounded at base, deeply puncto-striate, punctures clearly visible through scaling; setae very short and recumbent. Legs dark brown with grey scaling; femora unidentate; tibiae simple.</p><p>Remarks.</p><p>This species is closely related to M. subfasciatus Guérin-Méneville, 1843, but differs by the following characters: funicular antennomere 2 distinctly longer than 1, funicular antennomere 3 slightly longer than 4; funicular antennomeres 2–6 without conspicuous white setae; metafemora unidentate.</p><p>Plant association.</p><p>Psophocarpus tetragonolobus ( Fabaceae), Vachellia nilotica (= Acacia arabica) ( Fabaceae), Acacia decurrens ( Fabaceae), Melastoma malabathricum ( Melastomataceae), Theobroma cacao ( Malvaceae), Camellia sinensis ( Theaceae), Cocos nucifera ( Arecaceae), Ziziphus mauritiana ( Rhamnaceae), Rosa spp. ( Rosaceae), Erythrina lithosperma ( Fabaceae), Gliricidia sepium ( Fabaceae), Mangifera indica ( Anacardiaceae) (Ramamurthy and Ghai 1988; Shanthichandra et al. 1990; Marshall 1916).</p><p>Distribution.</p><p>Sri Lanka, India.</p></div>	https://treatment.plazi.org/id/1EC4932D55455163AC5946A7F9C76352	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wijesinghe, Dilshara D.;Hong, Ki-Jeong	Wijesinghe, Dilshara D., Hong, Ki-Jeong (2026): Bridging biodiversity gaps: Updated taxonomy and diversity of Sri Lankan Myllocerus (Coleoptera, Curculionidae, Entiminae). ZooKeys 1284: 87-111, DOI: 10.3897/zookeys.1284.191034
60E76275F9785730BF76BF46F2D95916.text	60E76275F9785730BF76BF46F2D95916.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myllocerus delicatulus Boheman 1842	<div><p>Myllocerus delicatulus Boheman, 1842</p><p>Myllocerus delicatulus Boheman, 1842: 6 . Type locality: Pondichery, India orientalis; Type repository: Swedish Museum of Natural History, Stockholm. Type not examined.</p><p>= Phyllobius mimicus Walker, 1859: 263; Marshall 1916: 304; Ramamurthy and Ghai 1988: 419. Type locality: Ceylon; Type repository: NHMUK.</p><p>Material examined.</p><p>No specimens examined; data based on published records.</p><p>Diagnosis.</p><p>Body dark brown, uniformly clothed with light yellowish green scaling. Head subconical; eyes dorsal, widely separated, interocular distance nearly equal to width of eye; frons without median impression. Rostrum as long as head; as long as or slightly broader than long, gradually narrowed anteriorly, sides straight; scrobes entirely dorsal, interscrobal distance slightly broader than interocular distance. Antennae black to dark brown; funicular antennomeres 1 and 2 elongate and subequal; funicular antennomere 3 slightly longer than 4; funicular antennomere 4 longer than 5; funicular antennomere 5 yellowish. Prothorax transverse, subcylindrical, sides parallel posteriorly and slightly narrowed only on anterior half; posterior sinuation shallow. Elytra with bases weakly rounded; strial punctures scarcely visible through scaling; elytral interstriae with scales circular, convex, disc-shaped. Legs yellowish; femora with a single small tooth (Marshall 1916).</p><p>Remarks.</p><p>This species can be distinguished from M. viridanus (Fabricius, 1775) by the following characters: head subconical, eyes more widely separated; interocular distance nearly as broad as width of eye; frons not impressed; antennae yellowish; prothorax weakly narrowed anteriorly; elytra with base distinctly rounded. It is also closely related to M. thompsoni Ramamurthy &amp; Ghai, 1988, but differs by the following characters: interocular distance nearly as broad as width of eye; presence of a longitudinal ridge around eyes; funicular antennomeres 1 and 2 subequal; funicular antennomeres 3–6 not equal, progressively elongate; and elytral interstriae with scales distinctly smaller (Ramamurthy and Ghai 1988).</p><p>Plant association.</p><p>Morus spp. ( Moraceae), Litchi chinensis ( Sapindaceae), Lantana camara ( Verbenaceae) (Ramamurthy and Ghai 1988).</p><p>Distribution.</p><p>Sri Lanka, India.</p></div>	https://treatment.plazi.org/id/60E76275F9785730BF76BF46F2D95916	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wijesinghe, Dilshara D.;Hong, Ki-Jeong	Wijesinghe, Dilshara D., Hong, Ki-Jeong (2026): Bridging biodiversity gaps: Updated taxonomy and diversity of Sri Lankan Myllocerus (Coleoptera, Curculionidae, Entiminae). ZooKeys 1284: 87-111, DOI: 10.3897/zookeys.1284.191034
18B846E1FCEC55A0B60DE93D52AAA535.text	18B846E1FCEC55A0B60DE93D52AAA535.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myllocerus dentifer (Fabricius 1792)	<div><p>Myllocerus dentifer (Fabricius, 1792)</p><p>Fig. 3 A, B</p><p>Curculio dentifer Fabricius, 1792: 488 . Type locality: India orientalis; Type repository: Zoological Museum, University of Copenhagen. Type not examined.</p><p>Material examined.</p><p>Sri Lanka • 1 ♀, Southern Province, Galle; 27 Nov. – 4 Dec. 1881; G. Lewis leg.; NHMUK, NHMUK 013663882 .</p><p>Diagnosis.</p><p>Body black, clothed with greyish brown to pale brown scaling, sometimes mottled with small darker spots along elytral striae. Head with shallow, indistinct punctation concealed by scaling; eyes widely separated, interocular distance greater than interscrobal distance. Rostrum slightly longer than head, sides weakly narrowed anteriorly, dorsally nearly flat, with a shallow median impression with a fine carina extending onto frons. Antennae with scape only slightly exceeding anterior margin of prothorax; funicular antennomeres 1 and 2 subequal. Prothorax transverse; sides subparallel or slightly rounded in anterior half, then strongly constricted posteriorly; posterolateral angles acute; posterior margin moderately bisinuate, as wide as anterior margin; dorsal surface with large shallow punctures obscured by scaling and with a shallow transverse impression anterior to posterior margin. Elytra subparallel-sided; elytral striae narrow and finely punctate as visible through scaling; elytral interstriae almost flat, bearing very short, dense, sub depressed setae with circular scales. Legs dark brown; profemora bidentate, metafemora tridentate; tibiae with inner margin deeply emarginate proximally, especially metathoracic legs.</p><p>Remarks.</p><p>This species is extremely close to M. discolor Boheman, 1834 in general structure, but differs in the following characters: rostrum very slightly narrow anteriorly, slightly longer than head; prothorax transverse, posterior margin moderately bisinuate; elytral striae narrow, finely punctate as visible through scaling. Also, it is similar to M. sabulosus Marshall, 1916 in general appearance but easily distinguished, by having only one tooth on metafemora instead of three (Marshall 1916).</p><p>Plant association.</p><p>Zea mays ( Poaceae), Pennisetum glaucum ( Poaceae), Sorghum bicolor ( Poaceae), Setaria italica ( Poaceae), Gossypium spp. ( Malvaceae), Vigna mungo ( Fabaceae), Glycine max ( Fabaceae), Vigna unguiculata ( Fabaceae), Trifolium alexandrinum ( Fabaceae), Cyamopsis tetragonoloba ( Fabaceae), Abelmoschus esculentus ( Malvaceae), Luffa acutangula ( Cucurbitaceae), Prunus dulcis ( Rosaceae), Crotalaria juncea ( Fabaceae), Alcea rosea ( Malvaceae), Rosa spp. ( Rosaceae); attracted to light (Ramamurthy and Ghai 1988).</p><p>Distribution.</p><p>Sri Lanka, Afghanistan, India.</p></div>	https://treatment.plazi.org/id/18B846E1FCEC55A0B60DE93D52AAA535	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wijesinghe, Dilshara D.;Hong, Ki-Jeong	Wijesinghe, Dilshara D., Hong, Ki-Jeong (2026): Bridging biodiversity gaps: Updated taxonomy and diversity of Sri Lankan Myllocerus (Coleoptera, Curculionidae, Entiminae). ZooKeys 1284: 87-111, DOI: 10.3897/zookeys.1284.191034
D375FCE4A5675DA3B87192B85AFF560A.text	D375FCE4A5675DA3B87192B85AFF560A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myllocerus discolor Boheman 1834	<div><p>Myllocerus discolor Boheman, 1834</p><p>Fig. 3 C, D</p><p>Myllocerus discolor Boheman, 1834: 428 . Type locality: Bengal; Type repository: Swedish Museum of Natural History, Stockholm. Type not examined.</p><p>= Myllocerus variegatus Boheman, 1842: 9; Marshall 1916: 348; Ramamurthy and Ghai 1988: 459. Type locality: Bengal; Type repository: Swedish Museum of Natural History, Stockholm.</p><p>= Myllocerus uniformis Marshall, 1916: 349; Ramamurthy and Ghai 1988: 459. Type locality: India-Mundali, Jaunsar; Type repository: NHMUK.</p><p>= Myllocerus canescens Marshall, 1916: 350; Ramamurthy and Ghai 1988: 459. Type locality: Ceylon-Halupahani; Type repository: NHMUK.</p><p>= Myllocerus persicus Zumpt, 1938: 84; Ramamurthy and Ghai 1988: 459. Type locality: Japan; Type repository: Natural History Museum in Basel, Switzerland.</p><p>Material examined.</p><p>Sri Lanka • Uva Province, Halupahani, Haldummulle; NHMUK, NHMUK 013663894 .</p><p>Diagnosis.</p><p>Integument black, densely covered with greyish white scales, rendering elytra almost entirely of this color, except for a faint, narrow pale-brown stripe extending anteriorly from elytral humeri. Head and broad lateral stripes of the prothorax clothed with pale-brown scales. Eyes widely separated and convex. Rostrum slightly widened apically; posterior interocular distance greater than interscrobal distance. Antennae with scape only slightly exceeding anterior margin of prothorax, funicular antennomeres 1 and 2 subequal. Prothorax weakly transverse, nearly as long as broad, with posterior margin deeply bisinuate. Elytral striae broad, with coarse punctures; setae markedly longer and more conspicuous on the disc. Legs robust; metafemora distinctly tridentate.</p><p>Remarks.</p><p>For comparison with M. dentifer (Fabricius, 1792), see above.</p><p>Plant association.</p><p>Triticum aestivum ( Poaceae), Oryza sativa ( Poaceae), Zea mays ( Poaceae), Sorghum bicolor ( Poaceae), Pennisetum glaucum ( Poaceae), Eleusine coracana ( Poaceae), Gossypium spp. ( Malvaceae), Saccharum officinarum ( Poaceae), Corchorus spp. ( Malvaceae), Vigna aconitifolia ( Fabaceae), Ipomoea batatas ( Convolvulaceae), Mangifera indica ( Anacardiaceae), Ficus carica ( Moraceae), Ziziphus mauritiana ( Rhamnaceae), Musa spp. ( Musaceae), Litchi chinensis ( Sapindaceae), Aegle marmelos ( Rutaceae), Citrus limon ( Rutaceae), Psidium guajava ( Myrtaceae), Eriobotrya japonica ( Rosaceae), Morus spp. ( Moraceae), Abelmoschus esculentus ( Malvaceae), Cucurbita spp. ( Cucurbitaceae), Moringa oleifera ( Moringaceae), Medicago sativa ( Fabaceae), Tectona grandis ( Lamiaceae), Sesbania bispinosa ( Fabaceae), Sesbania spp. ( Fabaceae), Vachellia nilotica (= Acacia intsia) ( Fabaceae), Dalbergia sissoo and Dalbergia paniculata ( Fabaceae), Ziziphus jujuba ( Rhamnaceae), Citrus spp. ( Rutaceae), Solanum melongena ( Solanaceae), Glycine max ( Fabaceae) (Marshall 1916; Ramamurthy and Ghai 1988).</p><p>Distribution.</p><p>Sri Lanka, India, Iran, Pakistan.</p></div>	https://treatment.plazi.org/id/D375FCE4A5675DA3B87192B85AFF560A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wijesinghe, Dilshara D.;Hong, Ki-Jeong	Wijesinghe, Dilshara D., Hong, Ki-Jeong (2026): Bridging biodiversity gaps: Updated taxonomy and diversity of Sri Lankan Myllocerus (Coleoptera, Curculionidae, Entiminae). ZooKeys 1284: 87-111, DOI: 10.3897/zookeys.1284.191034
327F0DA6221A51318EDD87EC8AEC8C47.text	327F0DA6221A51318EDD87EC8AEC8C47.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myllocerus equinus Ramamurthy & Ghai 1988	<div><p>Myllocerus equinus Ramamurthy &amp; Ghai, 1988</p><p>Myllocerus equinus Ramamurthy &amp; Ghai, 1988: 464 . Type locality: Sri Lanka; Type repository: State Museum of Zoology, Dresden. Type not examined.</p><p>Material examined.</p><p>No specimens examined; data based on published records.</p><p>Diagnosis.</p><p>Body dark brown to black, densely clothed with dull white scales; elytra dull whitish, each bearing nine irregularly shaped dark brown-black spots. Head with median area densely covered with white scales; eyes lateral, interocular distance shorter than length of eye and nearly equal to interscrobal distance. Rostrum slightly longer than head, apically deeply and acutely emarginate; median carina distinct and well developed. Antennae with scape stout, barely surpassing anterior margin of prothorax; funicular antennomere 2 slightly longer than 1, funicular antennomere 3 longer than 4. Prothorax transverse, sides weakly narrowing anteriorly, strongly constricted posteriorly; posterior margin distinctly bisinuate and broader than anterior margin. Elytra with striae originating from base; strial punctures deep, oblong, and well defined; elytral interstriae bearing distinct scaly vestiture with opaque ridges, imparting a ribbed appearance, lateral elytral interstriae clearly demarcated. Legs dark brown with dull white scaling; profemora bidentate, metafemora tridentate (Ramamurthy and Ghai 1988).</p><p>Remarks.</p><p>This species differs from M. undecimpustulatus undatus Marshal, 1916 in elytra with strial punctures originating from base, punctures deep and oblong; elytral interstriae bearing distinct scaly vestiture with opaque ridges, imparting a ribbed appearance, lateral interstriae clearly demarcated; prothorax weakly narrowed anteriorly; elytral humeri not projecting laterally; posterior interocular distance nearly equal to interscrobal distance (Ramamurthy and Ghai 1988).</p><p>Plant association.</p><p>Unknown.</p><p>Distribution.</p><p>Sri Lanka.</p></div>	https://treatment.plazi.org/id/327F0DA6221A51318EDD87EC8AEC8C47	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wijesinghe, Dilshara D.;Hong, Ki-Jeong	Wijesinghe, Dilshara D., Hong, Ki-Jeong (2026): Bridging biodiversity gaps: Updated taxonomy and diversity of Sri Lankan Myllocerus (Coleoptera, Curculionidae, Entiminae). ZooKeys 1284: 87-111, DOI: 10.3897/zookeys.1284.191034
ED9F92A280DA5B4B985ACF176487B58A.text	ED9F92A280DA5B4B985ACF176487B58A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myllocerus fringilla Faust 1897	<div><p>Myllocerus fringilla Faust, 1897</p><p>Myllocerus fringilla Faust, 1897: 356 . Type locality: Malacca; Type repository: State Museum of Zoology, Dresden. Type not examined.</p><p>Material examined.</p><p>No specimens examined; data based on published records.</p><p>Diagnosis.</p><p>Antennal scape not reaching middle of pronotum. Long rostrum with a distinct median impression, sometimes with a fine glabrous median line extending onto frons. Ventral surface distinctly and darkly punctate, similar to dorsal surface. Antennae and legs vary in color, ranging from yellowish red to dark brown. Pronotum and elytra more distinctly punctate; elytral strial punctures becoming larger and deeper towards base, elytral interstriae weakly convex; elytral apex strongly beak-shaped produced downward; femora bidentate (Faust 1897; Marshall 1916).</p><p>Remarks.</p><p>This species is closely related to M. lineatocollis (Boheman, 1842), but differs by the following characters: rostrum being longer with a distinct impression; elytra produced downwards at apex; having two femoral teeth (Marshall 1916).</p><p>Ramamurthy and Ghai (1988) listed Myllocerus fringilla Faust, 1897 as a synonym of M. lineatocollis var. frontalis Faust, 1897, citing Faust (1897: 363). However, the original description by Faust (1897) established only M. frontalis Faust, 1897, and not a form under M. lineatocollis; the latter thus appears to be a subsequent misinterpretation. Myllocerus lineatocollis (Boheman, 1842) is currently treated as a valid species, with M. frontalis as its synonym. Comparison of the original descriptions indicates that M. fringilla is distinct, particularly in the structure of the rostrum and the form of the elytral apex. Therefore, the treatment by Wijesinghe and Hong (2025), which synonymized M. lineatocollis var. frontalis with M. fringilla, is based on an invalid name and is considered erroneous.</p><p>Plant association.</p><p>Unknown.</p><p>Distribution.</p><p>Sri Lanka, Malacca.</p></div>	https://treatment.plazi.org/id/ED9F92A280DA5B4B985ACF176487B58A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wijesinghe, Dilshara D.;Hong, Ki-Jeong	Wijesinghe, Dilshara D., Hong, Ki-Jeong (2026): Bridging biodiversity gaps: Updated taxonomy and diversity of Sri Lankan Myllocerus (Coleoptera, Curculionidae, Entiminae). ZooKeys 1284: 87-111, DOI: 10.3897/zookeys.1284.191034
3288CA03A19D5DB39C24871B06586A6B.text	3288CA03A19D5DB39C24871B06586A6B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myllocerus Schonherr 1823	<div><p>Genus Myllocerus Schönherr, 1823</p><p>Myllocerus Schönherr, 1823: 1144 . [Type species: Curculio curvicornis Fabricius, 1792, by original designation].</p><p>= Macrocorynus Schönherr, 1823: 1144 . [Type species: Curculio discoideus Olivier, 1807].</p><p>= Epherina Pascoe, 1869: 102 . [Type species: Epherina longicornis Pascoe, 1869].</p><p>= Idaspora Pascoe, 1869: 101 . [Type species: Idaspora terrea Pascoe, 1869].</p><p>= Proxyrus Pascoe, 1870: 437 . [Type species: Proxyrus abstersus Pascoe, 1870].</p><p>= Hyperstylus Roelofs, 1873: 171 . [Type species: Hyperstylus pallipes Roelofs, 1873].</p><p>= Proxyrodes Blackburn, 1892: 48 . [Type species: Proxyrodes maculatus Blackburn, 1892].</p><p>= Exmyllocerus Voss, 1937: 230 . [Type species: Myllocerus ginfushanensis Voss, 1933].</p><p>= Hypomyllocerus Voss, 1937: 231 . [Type species: Myllocerus inflatus Voss, 1937].</p><p>= Pachymyllocerus Voss, 1937: 235 . [Type species: Myllocerus durus Voss, 1937].</p><p>= Isomyllocerus Marshall, 1954: 214 . [Type species: Curculio dorsatus Fabricius, 1798].</p><p>= Pseudocanoixus Voss, 1958: 23 . [Type species: Macrocorynus fallaciosus Voss, 1958].</p><p>= Calomyllocerus Voss, 1959: 408 . [Type species: Myllocerus liesenfeldti Voss, 1959].</p><p>= Mylloceroversus Hoffmann, 1961: 644 . [Type species: Myllocerus acaciae Hoffmann, 1961].</p><p>Diagnosis.</p><p>General color varies to a great extent, ranging from pale white to bright metallic with stripes, blotches, and spots of various patterns. Body elongate, more than twice as long as broad; sometimes cylindrical, more commonly oblong to ovate; convex dorsally, flat ventrally. Head wider than long; eyes dorsal to sublateral, variable in convexity. Rostrum stout, variable in shape; epistomal emargination ranging from deep and triangular to shallow and rounded; epistome bounded posteriorly by distinct carina. Scrobes dorso-lateral, visible from above, very deep anteriorly, abruptly shallow posteriorly, curved downwards and fading anteriorly of eye; genae dilated or not. Rostral carina transverse or longitudinal, distinct up to base or fading before reaching it. Antennae inserted near apex of rostrum; scape elongate, curved, extending beyond anterior margin of prothorax; funicle 7 - jointed, funicular antennomeres 1 and 2 longer than others; club compact, spindle-shaped, 4 - jointed. Prothorax subcylindrical to subtrapezoid; with posterior margin truncate or bisinuate, without ocular lobe or vibrissae on anterolateral margin. Elytra oblong, distinctly broader than prothorax at elytral humeri, distinct elytral humeri with ten punctate elytral striae; sides generally subparallel from elytral humeri to beyond mid-length, margins gently sinuate above metacoxa. Legs with procoxa contiguous, metacoxa not reaching margin of elytra; femora clavate and dentate (with 1–3 teeth), always on distal half; tibiae simple or occasionally basally sinuate, protibiae sometimes internally angulate or bisinuate; metatibial corbels absent; tarsi slender, tarsomere 1 elongate, 3 broadly ovate and bilobed; claws free. Abdominal ventrite 1 with intercoxal process ogival and narrower than metacoxa; abdominal ventrite 2 as long as or longer than 3 and 4 combined, separated from ventrite 1 by a curved incision which is shallower medially.</p><p>Distribution.</p><p>Afrotropical and Oriental Regions, southern and eastern Palaearctic and Australia (Alonso-Zarazaga and Lyal 1999; Alonso-Zarazaga et al. 2023); one species is adventive in the Nearctic Region (Hunsberger 2003).</p></div>	https://treatment.plazi.org/id/3288CA03A19D5DB39C24871B06586A6B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wijesinghe, Dilshara D.;Hong, Ki-Jeong	Wijesinghe, Dilshara D., Hong, Ki-Jeong (2026): Bridging biodiversity gaps: Updated taxonomy and diversity of Sri Lankan Myllocerus (Coleoptera, Curculionidae, Entiminae). ZooKeys 1284: 87-111, DOI: 10.3897/zookeys.1284.191034
9C1592079F9557629F3975ACEF2D1653.text	9C1592079F9557629F3975ACEF2D1653.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myllocerus subfasciatus Guerin-Meneville 1843	<div><p>Myllocerus subfasciatus Guérin-Méneville, 1843</p><p>Fig. 4 A, B</p><p>Myllocerus subfasciatus Guérin-Méneville, 1843: 54 . Type locality: India-Neelgheries; Type repository: Swedish Museum of Natural History, Stockholm. Type not examined.</p><p>= Myllocerus spurcatus Walker, 1859: 263; Marshall 1916: 345; Ramamurthy and Ghai 1988: 456. Type locality: Ceylon; Type repository: NHMUK.</p><p>Material examined.</p><p>Sri Lanka • 1 ♂, Western Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=80.026665&amp;materialsCitation.latitude=7.0326667" title="Search Plazi for locations around (long 80.026665/lat 7.0326667)">Weliweriya</a>, 7°1.96'N, 80°1.6'E; 25 Jul. 2023; KJ. Hong leg.; on Solanum melongena ( Solanaceae); SCNU • 1 ♂; Central Province, Dikoya, 3800–4200 ft; 6 Dec. 1881 – 16 Jan. 1882; G. Lewis leg.; NHMUK, NHMUK 013663889 .</p><p>Diagnosis.</p><p>Body black, clothed with coarse brown scaling; prothorax usually paler laterally and bearing a paler median line; elytra variably irrorated with greyish or whitish scales, typically forming faint transverse bands, occasionally more distinct and conspicuous. Head with a distinct median fovea; eyes lateral; interocular distance broader than eye width and slightly broader than interscrobal distance. Rostrum slightly longer than head, weakly dilated apically, dorsally rather deeply impressed, with a narrow smooth median line extending onto frons. Antennae stout; scape bearing dense suberect setae; funicular antennomere 2 longer than 1, funicular antennomeres 3 and 4 subequal; funicular antennomeres 2–6 with conspicuous white setae. Prothorax transverse, sides strongly rounded, deeply constricted and transversely impressed posteriorly, more shallowly constricted anteriorly; posterior margin distinctly bisinuate; a shallow rounded lateral impression present posterior to mid-length. Elytra relatively broad, elytral humeri rectangularly rounded; elytral striae broad, deeply punctate; vestiture composed of dense, short, sub depressed, squamiform setae. Legs dark brown with dense grey or brown scaling; femora bearing two unequal teeth; inner margin of tibia with a slight angular projection.</p><p>Remarks.</p><p>For comparison with M. curvicornis (Fabricius, 1792), see above.</p><p>Plant association.</p><p>Gossypium spp. ( Malvaceae), Corchorus spp. ( Malvaceae), Arachis hypogaea ( Fabaceae), Solanum melongena ( Solanaceae), Solanum tuberosum ( Solanaceae), Malus domestica ( Rosaceae), Prunus spp. ( Rosaceae), Boehmeria nivea ( Urticaceae), Alcea rosea ( Malvaceae); attracted to light (Ramamurthy and Ghai 1988; Shanmugam et al. 2018).</p><p>Distribution.</p><p>Sri Lanka, India, Myanmar.</p></div>	https://treatment.plazi.org/id/9C1592079F9557629F3975ACEF2D1653	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wijesinghe, Dilshara D.;Hong, Ki-Jeong	Wijesinghe, Dilshara D., Hong, Ki-Jeong (2026): Bridging biodiversity gaps: Updated taxonomy and diversity of Sri Lankan Myllocerus (Coleoptera, Curculionidae, Entiminae). ZooKeys 1284: 87-111, DOI: 10.3897/zookeys.1284.191034
DA3F2FAC0EEC5F1DB7B4BD665A2334D5.text	DA3F2FAC0EEC5F1DB7B4BD665A2334D5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myllocerus undecimpustulatus subsp. undatus Marshall 1916	<div><p>Myllocerus undecimpustulatus undatus Marshall, 1916</p><p>Fig. 5 A – D</p><p>Myllocerus undatus Marshall, 1916: 350 .</p><p>Myllocerus undecimpustulatus undatus: O’Brien et al. 2006: 1. Type locality: Ceylon-Weligama; Type repository: NHMUK. Type not examined.</p><p>Material examined.</p><p>Sri Lanka • 1 ♀, Central Province, Kandy; Jun. 1905; GE. Bryant leg.; NHMUK, NHMUK 013663888 ; • 1 ♂, Western Province, Colombo; Sep. 1890; “ HP. Green ” [EE. Green] leg.; NHMUK, NHMUK 013663891 .</p><p>Diagnosis.</p><p>Body maculate, clothed with dull white and black scales, often with yellowish scales on rostrum and head; prothorax bearing three darker, partially denuded longitudinal stripes, lateral stripes interrupted; elytra with more or less distinct large dark brown blotches, usually arranged into three irregular oblique bands. Eyes sublateral; posterior interocular distance greater than interscrobal distance. Antennae stout; scape with dense suberect setae; funicular antennomere 2 longer than 1, funicular antennomeres 3 and 4 subequal. Rostrum slightly longer than head, weakly dilated apically, with a narrow smooth median line confined to posteriorly. Prothorax weakly rounded laterally, constricted posteriorly; posterior margin bisinuate. Elytral humeri strongly angulate; projecting laterally and distinctly broader than prothorax. Legs with profemora and mesofemora bidentate, metafemora distinctly tridentate.</p><p>Remarks.</p><p>For comparison with M. equinus Ramamurthy &amp; Ghai, 1988, see above. This subspecies is closely related to Myllocerus undecimpustulatus undecimpustulatus Faust, 1891, but can be distinguished primarily by its narrower interocular distance, which is not broader than the length of an eye.</p><p>Plant association.</p><p>Oryza sativa ( Poaceae), Zea mays ( Poaceae), Cajanus cajan ( Fabaceae), Gossypium spp. ( Malvaceae), Corchorus spp. ( Malvaceae), Helianthus annuus ( Asteraceae), Mangifera indica ( Anacardiaceae), Punica granatum ( Lythraceae), Ziziphus mauritiana ( Rhamnaceae), Fragaria × ananassa ( Rosaceae), Malus domestica ( Rosaceae), Medicago sativa ( Fabaceae), Sesbania bispinosa ( Fabaceae), Dalbergia sissoo ( Fabaceae), Imperata cylindrica ( Poaceae), Psophocarpus tetragonolobus ( Fabaceae), Citrus spp. ( Rutaceae), Solanum melongena ( Solanaceae), Daucus carota ( Apiaceae), Ipomoea batatas ( Convolvulaceae), Conocarpus erectus ( Combretaceae), Bauhinia × blakeana ( Fabaceae), Chrysobalanus icaco ( Chrysobalanaceae), Phoenix roebelenii ( Arecaceae), Prunus persica ( Rosaceae), Lagerstroemia indica ( Lythraceae), Capsicum spp. ( Solanaceae), Litchi chinensis ( Sapindaceae), Muntingia calabura ( Muntingiaceae), Fragaria × ananassa ( Rosaceae) and Solanum melongena ( Solanaceae) (Butani 1979; Ramamurthy and Ghai 1988; Neal 2013; Rajan and Ghosh 2019).</p><p>Distribution.</p><p>Sri Lanka, adventive in the Southeastern United States (Hunsberger 2003; O’Brien et al. 2006).</p></div>	https://treatment.plazi.org/id/DA3F2FAC0EEC5F1DB7B4BD665A2334D5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wijesinghe, Dilshara D.;Hong, Ki-Jeong	Wijesinghe, Dilshara D., Hong, Ki-Jeong (2026): Bridging biodiversity gaps: Updated taxonomy and diversity of Sri Lankan Myllocerus (Coleoptera, Curculionidae, Entiminae). ZooKeys 1284: 87-111, DOI: 10.3897/zookeys.1284.191034
DA91AA605926598E8E3EB1BAA6C981DB.text	DA91AA605926598E8E3EB1BAA6C981DB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myllocerus viridanus (Fabricius 1775)	<div><p>Myllocerus viridanus (Fabricius, 1775)</p><p>Curculio viridanus Fabricius, 1775: 155 . Type locality: India-Tranquebar; Type repository: Zoological Museum, University of Copenhagen. Type not examined.</p><p>= Myllocerus angustifrons Faust, 1897: 356; Marshall 1916: 301; Ramamurthy and Ghai 1988: 414. Type locality: India; Type repository: State Museum of Zoology, Dresden.</p><p>Material examined.</p><p>No specimens examined; data based on published records.</p><p>Diagnosis.</p><p>Body covered with dense, uniform light-green scales, sometimes varying to pale greenish white; in paler specimens a chalky-white efflorescence may be present. Head tinged with yellow, bearing metallic green scales at apex of rostrum; narrowed anteriorly, with straight sides. Eyes dorsal causing external margins of both eyes plainly visible from above at same time and converging slightly anteriorly, frons with a median, longitudinal impression. Rostrum evidently longer, broader than head; sides straight; scrobes entirely dorsal, interscrobal distance slightly broader than interocular distance. A low, rounded longitudinal ridge originates from anteroventral margin of eye and curves indistinctly inward near mid-length. Antennae with funicular antennomeres 1 and 2 elongate and subequal; funicular antennomere 3 slightly longer than 4, and funicular antennomere 4 longer than 5. Prothorax subconical, widest posteriorly and strongly narrowed anteriorly; sides straight; posterior margin deeply bisinuate; dorsal surface bearing deep, well-separated punctures concealed by scaling. Elytra almost parallel-sided from elytral humeri to posterior to mid-length in ♂, gradually rounded toward the apex and with laterally prominent elytral humeri in ♀; vestiture with scales on elytral interstriae more oval than circular. Legs black with green scaling; femora bidentate, outer tooth consistently small (Marshall 1916).</p><p>Remarks.</p><p>For comparison with M. delicatulus Boheman, 1842, see above.</p><p>Plant association.</p><p>Oryza sativa ( Poaceae), Corchorus spp. ( Malvaceae), Arachis hypogaea ( Fabaceae), Ricinus communis ( Euphorbiaceae), Linum usitatissimum ( Linaceae), Helianthus annuus ( Asteraceae), Ipomoea batatas ( Convolvulaceae), Trichosanthes cucumerina ( Cucurbitaceae), Lablab purpureus ( Fabaceae), Abelmoschus esculentus ( Malvaceae), Citrus limon ( Rutaceae), Mangifera indica ( Anacardiaceae), Psidium guajava ( Myrtaceae), Carica papaya ( Caricaceae), Manilkara zapota ( Sapotaceae), Ficus carica ( Moraceae), Prunus persica ( Rosaceae), Theobroma cacao ( Malvaceae), Camellia sinensis ( Theaceae), Rosa spp. ( Rosaceae), Indigofera spp. ( Fabaceae), Plumbago zeylanica ( Plumbaginaceae), Sesbania spp. ( Fabaceae), Tectona grandis ( Lamiaceae), Eucalyptus spp. ( Myrtaceae), Acalypha spp. ( Euphorbiaceae), Lantana camara ( Verbenaceae); attracted to light (Ramamurthy and Ghai 1988).</p><p>Distribution.</p><p>Sri Lanka, India.</p></div>	https://treatment.plazi.org/id/DA91AA605926598E8E3EB1BAA6C981DB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wijesinghe, Dilshara D.;Hong, Ki-Jeong	Wijesinghe, Dilshara D., Hong, Ki-Jeong (2026): Bridging biodiversity gaps: Updated taxonomy and diversity of Sri Lankan Myllocerus (Coleoptera, Curculionidae, Entiminae). ZooKeys 1284: 87-111, DOI: 10.3897/zookeys.1284.191034
FB66E0581A6158448095B11A8FFB8920.text	FB66E0581A6158448095B11A8FFB8920.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myllocerus zeylanicus Marshall 1916	<div><p>Myllocerus zeylanicus Marshall, 1916</p><p>Fig. 4 C, D</p><p>Myllocerus zeylanicus Marshall, 1916: 315 . Type locality: Ceylon-Kandy; Type repository: NHMUK. Type not examined.</p><p>Material examined.</p><p>Sri Lanka • 1 ♂, Central Province, Kandy; 1550–1700 ft; Jun. 1908; GE. Bryant leg.; NHMUK, NHMUK 013663875 .</p><p>Diagnosis.</p><p>Body dark reddish brown, clothed with grey to pale grey-green scales, very sparse dorsally but much denser on lateral surfaces and ventrally. Head minutely striolate beneath scaling; eyes large and lateral; interocular distance nearly as broad as interscrobal distance. Rostrum slightly longer than head and nearly as long as broad; sides narrowed from base to mid-length and slightly dilated anteriorly; apical emargination moderately deep, forming an acute angle. Dorsal surface of rostrum shallowly impressed, with faint lateral carinae and a very fine median carina extending onto frons. Antennae dark reddish brown; funicular antennomeres 1 and 2 equal in length, funicular antennomere 3 distinctly longer than 4; funicular antennomere 7 longer than broad; club elongate, club antennomere 1 conical, as long as broad and not shorter than 2. Prothorax about as long as broad, with sides slightly rounded; shallowly constricted and transversely impressed near both posterior and anterior margins; posterior margin subtruncate; anterior margin of uniform width, oblique laterally; dorsal surface densely punctate. Elytra subtruncate at posterior margin, parallel-sided in ♂ and broader posteriorly in ♀; dorsally almost flat; elytral humeri roundly rectangular; elytral striae deeply punctate; elytral interstriae bearing short, suberect, pale, curved setae. Legs dark reddish brown; femora with a sharp tooth.</p><p>Remarks.</p><p>This species is closely related to M. angulatipes Marshall, 1916 see above. Also, it is closely related to M. hispidus Marshall, 1916, but differs by the following characters: rostral apex with apical emargination moderately deep, forming an acute angle; funicular antennomere 7 longer than broad; antennal club elongate, with club antennomere 1 conical, as long as broad and not shorter than 2; and elytral interstriae bearing short, suberect, pale, curved setae, whereas in M. hispidus setae are longer, erect, and black (Marshall 1916).</p><p>Plant association.</p><p>Unknown.</p><p>Distribution.</p><p>Sri Lanka.</p></div>	https://treatment.plazi.org/id/FB66E0581A6158448095B11A8FFB8920	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wijesinghe, Dilshara D.;Hong, Ki-Jeong	Wijesinghe, Dilshara D., Hong, Ki-Jeong (2026): Bridging biodiversity gaps: Updated taxonomy and diversity of Sri Lankan Myllocerus (Coleoptera, Curculionidae, Entiminae). ZooKeys 1284: 87-111, DOI: 10.3897/zookeys.1284.191034
