taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
5B30B6DB947A5684AC114F192403A675.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1661248	https://doi.org/10.3897/fr.29.189975.figure1	Figure 1. Schematic drawing of MUPA-ATZ 0103, postcranial skeleton of the paratype of Paludidraco multidentatus, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain), in field top (A 1) and field bottom (A 2) views, corresponding to the anatomically dorsolateral and dorsoventral views, respectively. The figure shows the skeletal elements disarticulated from the specimen (in colour) and their position in the field. Asterisk (*) indicates an estimated position of the bones in the field. Dashed lines and areas represent broken bones. Scale bar: 100 mm.	Figure 1. Schematic drawing of MUPA-ATZ 0103, postcranial skeleton of the paratype of Paludidraco multidentatus, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain), in field top (A 1) and field bottom (A 2) views, corresponding to the anatomically dorsolateral and dorsoventral views, respectively. The figure shows the skeletal elements disarticulated from the specimen (in colour) and their position in the field. Asterisk (*) indicates an estimated position of the bones in the field. Dashed lines and areas represent broken bones. Scale bar: 100 mm.	2026-05-29	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán		Zenodo	biologists	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán			
5B30B6DB947A5684AC114F192403A675.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1661249	https://doi.org/10.3897/fr.29.189975.figure2	Figure 2. Block containing most of the preserved postcranial skeleton (MUPA-ATZ 0103) of the paratype of Paludidraco multidentatus, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain), in field top view (corresponding to the anatomically dorsolateral view). Dashed lines and areas represent broken bones. Scale bar: 100 mm.	Figure 2. Block containing most of the preserved postcranial skeleton (MUPA-ATZ 0103) of the paratype of Paludidraco multidentatus, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain), in field top view (corresponding to the anatomically dorsolateral view). Dashed lines and areas represent broken bones. Scale bar: 100 mm.	2026-05-29	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán		Zenodo	biologists	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán			
5B30B6DB947A5684AC114F192403A675.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1661250	https://doi.org/10.3897/fr.29.189975.figure3	Figure 3. Block containing most of the preserved postcranial skeleton (MUPA-ATZ 0103) of the paratype of Paludidraco multidentatus, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain), in field bottom view (corresponding to the anatomically ventrolateral view). Dashed lines and areas represent broken bones. Scale bar: 100 mm.	Figure 3. Block containing most of the preserved postcranial skeleton (MUPA-ATZ 0103) of the paratype of Paludidraco multidentatus, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain), in field bottom view (corresponding to the anatomically ventrolateral view). Dashed lines and areas represent broken bones. Scale bar: 100 mm.	2026-05-29	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán		Zenodo	biologists	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán			
5B30B6DB947A5684AC114F192403A675.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1661251	https://doi.org/10.3897/fr.29.189975.figure4	Figure 4. Cervical vertebral elements of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain) (see Suppl. material 1: figs S 1, S 2 for precise position of the disarticulated elements in the field). A. Schematic drawing of MUPA-ATZ 0103 indicating the position of the articulated cervical elements. B 1 – C 12. Isolated cervical elements: B 1 – B 10. cc # 1; C 1 – C 12. cc # 2. D – F 2. Articulated cervical elements C # 3 to C # 5?. Cervical elements shown in: B 1 – B 4, C 1 – C 4. Cranial or caudal; B 5, B 6, C 5 – C 8, D, F 1, F 2. Lateral; B 7, B 8, C 9, C 10, E. Dorsal; B 9, B 10, C 11, C 12. Ventral view. Red asterisks indicate pre-infraprezygapophyses, blue asterisks indicate postzygapophyses, and purple asterisks indicate infrapostzygapophyses. Dashed lines and areas represent broken bones and grey areas represent sediment. Scale bars: 100 mm (A); 20 mm (B 1 – D); 10 mm (E – F 2).	Figure 4. Cervical vertebral elements of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain) (see Suppl. material 1: figs S 1, S 2 for precise position of the disarticulated elements in the field). A. Schematic drawing of MUPA-ATZ 0103 indicating the position of the articulated cervical elements. B 1 – C 12. Isolated cervical elements: B 1 – B 10. cc # 1; C 1 – C 12. cc # 2. D – F 2. Articulated cervical elements C # 3 to C # 5?. Cervical elements shown in: B 1 – B 4, C 1 – C 4. Cranial or caudal; B 5, B 6, C 5 – C 8, D, F 1, F 2. Lateral; B 7, B 8, C 9, C 10, E. Dorsal; B 9, B 10, C 11, C 12. Ventral view. Red asterisks indicate pre-infraprezygapophyses, blue asterisks indicate postzygapophyses, and purple asterisks indicate infrapostzygapophyses. Dashed lines and areas represent broken bones and grey areas represent sediment. Scale bars: 100 mm (A); 20 mm (B 1 – D); 10 mm (E – F 2).	2026-05-29	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán		Zenodo	biologists	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán			
5B30B6DB947A5684AC114F192403A675.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1661252	https://doi.org/10.3897/fr.29.189975.figure5	Figure 5. Dorsal vertebrae of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain). A. Schematic drawing of MUPA-ATZ 0103 indicating the position of the articulated dorsal vertebrae shown in the close-ups; B – H. Close-up of several regions of interest in the dorsal vertebral series where discussed characters are observable. Arrow heads in (D – E) indicate the concave posterior margin of the neural spines described in the text. Dashed lines and areas in (A) represent broken bones. Scale bars: 100 mm (A); 20 mm (B – H).	Figure 5. Dorsal vertebrae of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain). A. Schematic drawing of MUPA-ATZ 0103 indicating the position of the articulated dorsal vertebrae shown in the close-ups; B – H. Close-up of several regions of interest in the dorsal vertebral series where discussed characters are observable. Arrow heads in (D – E) indicate the concave posterior margin of the neural spines described in the text. Dashed lines and areas in (A) represent broken bones. Scale bars: 100 mm (A); 20 mm (B – H).	2026-05-29	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán		Zenodo	biologists	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán			
5B30B6DB947A5684AC114F192403A675.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1661253	https://doi.org/10.3897/fr.29.189975.figure6	Figure 6. Posterior to posteriormost dorsal vertebrae of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain) (see Suppl. material 1: figs S 1, S 2 for precise position of the disarticulated elements in the field): A 1 – A 10, D # 33; B 1 – B 10, D # 34; C 1 – C 6, D # 35; D 1 – D 12, D # 36. Neural arches shown in: A 1, A 2, B 1, B 2, C 1, C 2, D 1, D 2. Cranial; A 3, B 3, D 5, D 6. Dorsal; A 4, A 5, B 4, B 5, C 3, C 4, D 9 – D 12. Lateral; A 6, A 7, B 6, B 7, C 5, C 6, D 3, D 4. Caudal; A 8 – A 10, B 8 – B 10, D 7, D 8. Ventral view. Dotted outline indicates the currently missing distal portion in the rib of vertebra D # 36 (see Suppl. material 1: fig. S 2 G). Purple lines indicate the neurocentral sutures, drawn as dotted lines when not clearly visible. Dashed lines and areas represent broken bones and grey areas represent sediment. Scale bars: 20 mm (A 1 – A 7, A 9 – B 7, B 9 – D 12); 5 mm (A 8, B 8).	Figure 6. Posterior to posteriormost dorsal vertebrae of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain) (see Suppl. material 1: figs S 1, S 2 for precise position of the disarticulated elements in the field): A 1 – A 10, D # 33; B 1 – B 10, D # 34; C 1 – C 6, D # 35; D 1 – D 12, D # 36. Neural arches shown in: A 1, A 2, B 1, B 2, C 1, C 2, D 1, D 2. Cranial; A 3, B 3, D 5, D 6. Dorsal; A 4, A 5, B 4, B 5, C 3, C 4, D 9 – D 12. Lateral; A 6, A 7, B 6, B 7, C 5, C 6, D 3, D 4. Caudal; A 8 – A 10, B 8 – B 10, D 7, D 8. Ventral view. Dotted outline indicates the currently missing distal portion in the rib of vertebra D # 36 (see Suppl. material 1: fig. S 2 G). Purple lines indicate the neurocentral sutures, drawn as dotted lines when not clearly visible. Dashed lines and areas represent broken bones and grey areas represent sediment. Scale bars: 20 mm (A 1 – A 7, A 9 – B 7, B 9 – D 12); 5 mm (A 8, B 8).	2026-05-29	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán		Zenodo	biologists	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán			
5B30B6DB947A5684AC114F192403A675.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1661254	https://doi.org/10.3897/fr.29.189975.figure7	Figure 7. Posteriormost dorsal or anterior sacral vertebrae and sacral neural arch of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain) (see Suppl. material 1: figs S 1, S 2 for precise position of the disarticulated elements in the field). A. Schematic drawing of MUPA-ATZ 0103 indicating the position of vertebra D? # 37. B 1 – B 4, D? # 37; C 1 – C 10, D? # 38; D 1 – D 10. sa # 1. The vertebral elements are shown in: B 1, B 2, C 1, C 2, D 1, D 2. Cranial; B 3, B 4. Craniolateral; C 3, C 4, D 3, D 4. Lateral; C 5, C 6, D 5, D 6. Caudal; C 7, C 8, D 7, D 8. Dorsal; C 9, C 10, D 9, D 10. Ventral view. Purple lines indicate the neurocentral sutures, drawn as dotted lines when not clearly visible. Dashed lines and areas represent broken bones and grey areas represent sediment. Scale bars: 100 mm (A); 20 mm (B 1 – D 10).	Figure 7. Posteriormost dorsal or anterior sacral vertebrae and sacral neural arch of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain) (see Suppl. material 1: figs S 1, S 2 for precise position of the disarticulated elements in the field). A. Schematic drawing of MUPA-ATZ 0103 indicating the position of vertebra D? # 37. B 1 – B 4, D? # 37; C 1 – C 10, D? # 38; D 1 – D 10. sa # 1. The vertebral elements are shown in: B 1, B 2, C 1, C 2, D 1, D 2. Cranial; B 3, B 4. Craniolateral; C 3, C 4, D 3, D 4. Lateral; C 5, C 6, D 5, D 6. Caudal; C 7, C 8, D 7, D 8. Dorsal; C 9, C 10, D 9, D 10. Ventral view. Purple lines indicate the neurocentral sutures, drawn as dotted lines when not clearly visible. Dashed lines and areas represent broken bones and grey areas represent sediment. Scale bars: 100 mm (A); 20 mm (B 1 – D 10).	2026-05-29	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán		Zenodo	biologists	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán			
5B30B6DB947A5684AC114F192403A675.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1661255	https://doi.org/10.3897/fr.29.189975.figure8	Figure 8. Caudal vertebral elements of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain) (see Suppl. material 1: figs S 1, S 2 for precise position of the disarticulated elements in the field): A 1 – A 8. cac # 3 - cac # 4; B 1 – B 12, CA # 1 - CA # 2; C 1 – C 4. caa # 5. The vertebral elements are shown in: A 1, A 2, B 3, B 4. Caudal; A 3, A 4, B 5, B 6, C 1, C 2. Right lateral; A 5, A 6, B 9, B 10, C 3, C 4. Dorsal; A 7, A 8, B 11, B 12. Ventral; B 1, B 2. Cranial; B 7, B 8. Left lateral view. Purple lines indicate the neurocentral sutures, drawn as dotted lines when not clearly visible. Dashed lines and areas represent broken bones and grey areas represent sediment. Scale bar: 20 mm.	Figure 8. Caudal vertebral elements of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain) (see Suppl. material 1: figs S 1, S 2 for precise position of the disarticulated elements in the field): A 1 – A 8. cac # 3 - cac # 4; B 1 – B 12, CA # 1 - CA # 2; C 1 – C 4. caa # 5. The vertebral elements are shown in: A 1, A 2, B 3, B 4. Caudal; A 3, A 4, B 5, B 6, C 1, C 2. Right lateral; A 5, A 6, B 9, B 10, C 3, C 4. Dorsal; A 7, A 8, B 11, B 12. Ventral; B 1, B 2. Cranial; B 7, B 8. Left lateral view. Purple lines indicate the neurocentral sutures, drawn as dotted lines when not clearly visible. Dashed lines and areas represent broken bones and grey areas represent sediment. Scale bar: 20 mm.	2026-05-29	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán		Zenodo	biologists	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán			
5B30B6DB947A5684AC114F192403A675.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1661256	https://doi.org/10.3897/fr.29.189975.figure9	Figure 9. Dorsal ribs of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain). A. Schematic drawing of MUPA-ATZ 0103 indicating the position of the dorsal ribs shown in the close-ups; B – H. Close-up of several regions of interest in the dorsal skeleton where discussed rib characters are observable. Dashed lines and areas in (A) represent broken bones. Scale bars: 100 mm (A), 50 mm (B, D, F); 20 mm (C, E).	Figure 9. Dorsal ribs of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain). A. Schematic drawing of MUPA-ATZ 0103 indicating the position of the dorsal ribs shown in the close-ups; B – H. Close-up of several regions of interest in the dorsal skeleton where discussed rib characters are observable. Dashed lines and areas in (A) represent broken bones. Scale bars: 100 mm (A), 50 mm (B, D, F); 20 mm (C, E).	2026-05-29	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán		Zenodo	biologists	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán			
5B30B6DB947A5684AC114F192403A675.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1661257	https://doi.org/10.3897/fr.29.189975.figure10	Figure 10. Sacral (A 1 – E 4) and caudal (F 1 – I 12) ribs of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain) (see Suppl. material 1: figs S 1, S 2 for precise position of the disarticulated elements in the field): A 1 – A 10. sr # 1; B 1 – B 10. sr # 2; C 1 – C 10. sr # 3; D 1 – D 4. sr # 4; E 1 – E 4. sr # 5; F 1 – F 12. car # 1; G 1 – G 12. car # 2; H 1 – H 12. car # 3; I 1 – I 12. car # 4. The vertebral elements are shown in: A 1, A 2, A 5, A 6, B 1, B 2, B 5, B 6, C 1, C 2, C 5, C 6, D 1, D 2, E 3, E 4, F 1 – F 4, G 1, G 2, G 5, G 6, H 1, H 2, H 5, H 6, I 1–12, I 5, I 6. Cranial or caudal; A 3, A 4, B 3, B 4, C 3, C 4, D 3, D 4, F 7, F 8, G 3, G 4, H 3, H 4, I 3, I 4. Proximal; A 7, A 8, B 9, B 10, C 7, C 8, F 5, F 6, G 11, G 12, H 11, H 12, I 11, I 12. Dorsal; A 9, A 10, B 7, B 8, C 9, C 10, F 9, F 10, G 7, G 8, H 7, H 8, I 7, I 8. Ventral; E 1, E 2, F 11, F 12, G 9, G 10, H 9, H 10, I 9, I 10. Distal view. Dashed lines and areas represent broken bones and grey areas represent sediment. Scale bar: 20 mm.	Figure 10. Sacral (A 1 – E 4) and caudal (F 1 – I 12) ribs of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain) (see Suppl. material 1: figs S 1, S 2 for precise position of the disarticulated elements in the field): A 1 – A 10. sr # 1; B 1 – B 10. sr # 2; C 1 – C 10. sr # 3; D 1 – D 4. sr # 4; E 1 – E 4. sr # 5; F 1 – F 12. car # 1; G 1 – G 12. car # 2; H 1 – H 12. car # 3; I 1 – I 12. car # 4. The vertebral elements are shown in: A 1, A 2, A 5, A 6, B 1, B 2, B 5, B 6, C 1, C 2, C 5, C 6, D 1, D 2, E 3, E 4, F 1 – F 4, G 1, G 2, G 5, G 6, H 1, H 2, H 5, H 6, I 1–12, I 5, I 6. Cranial or caudal; A 3, A 4, B 3, B 4, C 3, C 4, D 3, D 4, F 7, F 8, G 3, G 4, H 3, H 4, I 3, I 4. Proximal; A 7, A 8, B 9, B 10, C 7, C 8, F 5, F 6, G 11, G 12, H 11, H 12, I 11, I 12. Dorsal; A 9, A 10, B 7, B 8, C 9, C 10, F 9, F 10, G 7, G 8, H 7, H 8, I 7, I 8. Ventral; E 1, E 2, F 11, F 12, G 9, G 10, H 9, H 10, I 9, I 10. Distal view. Dashed lines and areas represent broken bones and grey areas represent sediment. Scale bar: 20 mm.	2026-05-29	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán		Zenodo	biologists	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán			
5B30B6DB947A5684AC114F192403A675.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1661258	https://doi.org/10.3897/fr.29.189975.figure11	Figure 11. Gastral ribs of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain). A. Schematic drawing of MUPA-ATZ 0103 indicating the position of the gastral ribs shown in the close-ups; B – E. Close-up of several regions of interest in the dorsal skeleton where discussed characters of the gastralia are observable. Arrow heads in (D – E) indicate the anterior process of some gastral elements described in the text. Dashed lines and areas in (A) represent broken bones. Scale bars: 100 mm (A, B); 50 mm (C, D); 20 mm (E).	Figure 11. Gastral ribs of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain). A. Schematic drawing of MUPA-ATZ 0103 indicating the position of the gastral ribs shown in the close-ups; B – E. Close-up of several regions of interest in the dorsal skeleton where discussed characters of the gastralia are observable. Arrow heads in (D – E) indicate the anterior process of some gastral elements described in the text. Dashed lines and areas in (A) represent broken bones. Scale bars: 100 mm (A, B); 50 mm (C, D); 20 mm (E).	2026-05-29	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán		Zenodo	biologists	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán			
5B30B6DB947A5684AC114F192403A675.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1661259	https://doi.org/10.3897/fr.29.189975.figure12	Figure 12. Pectoral girdle elements of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain) (see Suppl. material 1: figs S 1, S 2 for precise position of the disarticulated elements in the field). A. Schematic drawing of MUPA-ATZ 0103 indicating the position of the right coracoid. B 1 – B 7. Partial left and right clavicles, plus interclavicle; C 1, C 2. Proximal region of the left coracoid; D 1, D 2. Right coracoid; E 1 – E 12. Left scapula; F 1 – F 12. Right scapula; G 1, G 2. Reconstruction of MUPA-ATZ 0103 pectoral girdle in dorsal (G 1) and ventral (G 2) views, where brown areas represent reconstructed bone, grey areas correspond to preserved, but inaccessible bone in that specific view due to its being overlain by skeletal elements and structures highlighted in purple represent presumed characters present in MUPA-ATZ 0103, based on the P. multidentatus holotype (MUPA-ATZ 0101). The girdle elements are shown in: B 1, B 2, B 7, E 7, E 8, F 7, F 8, G 1. Dorsal; B 3, B 4, C 1, D 2, E 11, E 12, F 11, F 12, G 2. Ventral; B 5, B 6, E 9, E 10, F 9, F 10. Distal; E 1, E 2, F 5, F 6. Medial; E 3, E 4, F 3, F 4. Proximal; E 5, E 6, F 1, F 2. Lateral view. Arrowheads in (B 6) indicate the dorsal and ventral striated ridges (purple) and the striae (blue) described in the text. Arrowhead in (B 7) indicates the probable presence of a dorsal clavicular foramen near the clavicular corner. Arrowhead in (D 2) points to the autapomorphic character of an ‘ angled posteromedial margin of the coracoid’ of P. multidentatus. Dashed lines and areas represent broken bones. Scale bars: 100 mm (A); 50 mm (B 1 – B 4, E 1 – F 12, G 1, G 2); 20 mm (C 1, D 2); 10 mm (B 5, B 6); 2 mm (B 7).	Figure 12. Pectoral girdle elements of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain) (see Suppl. material 1: figs S 1, S 2 for precise position of the disarticulated elements in the field). A. Schematic drawing of MUPA-ATZ 0103 indicating the position of the right coracoid. B 1 – B 7. Partial left and right clavicles, plus interclavicle; C 1, C 2. Proximal region of the left coracoid; D 1, D 2. Right coracoid; E 1 – E 12. Left scapula; F 1 – F 12. Right scapula; G 1, G 2. Reconstruction of MUPA-ATZ 0103 pectoral girdle in dorsal (G 1) and ventral (G 2) views, where brown areas represent reconstructed bone, grey areas correspond to preserved, but inaccessible bone in that specific view due to its being overlain by skeletal elements and structures highlighted in purple represent presumed characters present in MUPA-ATZ 0103, based on the P. multidentatus holotype (MUPA-ATZ 0101). The girdle elements are shown in: B 1, B 2, B 7, E 7, E 8, F 7, F 8, G 1. Dorsal; B 3, B 4, C 1, D 2, E 11, E 12, F 11, F 12, G 2. Ventral; B 5, B 6, E 9, E 10, F 9, F 10. Distal; E 1, E 2, F 5, F 6. Medial; E 3, E 4, F 3, F 4. Proximal; E 5, E 6, F 1, F 2. Lateral view. Arrowheads in (B 6) indicate the dorsal and ventral striated ridges (purple) and the striae (blue) described in the text. Arrowhead in (B 7) indicates the probable presence of a dorsal clavicular foramen near the clavicular corner. Arrowhead in (D 2) points to the autapomorphic character of an ‘ angled posteromedial margin of the coracoid’ of P. multidentatus. Dashed lines and areas represent broken bones. Scale bars: 100 mm (A); 50 mm (B 1 – B 4, E 1 – F 12, G 1, G 2); 20 mm (C 1, D 2); 10 mm (B 5, B 6); 2 mm (B 7).	2026-05-29	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán		Zenodo	biologists	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán			
5B30B6DB947A5684AC114F192403A675.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1661260	https://doi.org/10.3897/fr.29.189975.figure13	Figure 13. Pelvic girdle elements of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain) (see Suppl. material 1: figs S 1, S 2 for precise position of the disarticulated elements in the field): A 1 – A 12. Left ilium; B 1 – B 4. Right ilium; C 1 – C 6. Left pubis; D 1 – D 6. Right pubis; E 1 – E 6. Distal region of the probably right pubis; F 1 – F 6. Distal region of the probably left pubis; G 1 – G 6. Left ischium; H 1 – H 6. Right ischium; I. Reconstruction of MUPA-ATZ 0103 pelvic girdle in dorsal (medial) view (ilia not represented). The pelvic elements are shown in: A 1, A 2, B 1, B 2, C 1, C 2, D 5 – E 2, F 5, F 6, G 5 – H 2. Lateral; A 3, A 4, B 3, B 4, C 5 – D 2, E 5 – F 2, G 1, G 2, H 5 – I. Medial; A 5, A 6. Anterior; A 7, A 8. Posterior; A 9, A 10. Dorsal; A 11, A 12. Ventral; C 3, C 4, D 3, D 4, G 3, G 4, H 3, H 4. Proximal; E 3, E 4, F 3, F 4. Distal view. Dashed lines and areas represent broken bones. Scale bar: 50 mm.	Figure 13. Pelvic girdle elements of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain) (see Suppl. material 1: figs S 1, S 2 for precise position of the disarticulated elements in the field): A 1 – A 12. Left ilium; B 1 – B 4. Right ilium; C 1 – C 6. Left pubis; D 1 – D 6. Right pubis; E 1 – E 6. Distal region of the probably right pubis; F 1 – F 6. Distal region of the probably left pubis; G 1 – G 6. Left ischium; H 1 – H 6. Right ischium; I. Reconstruction of MUPA-ATZ 0103 pelvic girdle in dorsal (medial) view (ilia not represented). The pelvic elements are shown in: A 1, A 2, B 1, B 2, C 1, C 2, D 5 – E 2, F 5, F 6, G 5 – H 2. Lateral; A 3, A 4, B 3, B 4, C 5 – D 2, E 5 – F 2, G 1, G 2, H 5 – I. Medial; A 5, A 6. Anterior; A 7, A 8. Posterior; A 9, A 10. Dorsal; A 11, A 12. Ventral; C 3, C 4, D 3, D 4, G 3, G 4, H 3, H 4. Proximal; E 3, E 4, F 3, F 4. Distal view. Dashed lines and areas represent broken bones. Scale bar: 50 mm.	2026-05-29	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán		Zenodo	biologists	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán			
5B30B6DB947A5684AC114F192403A675.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1661261	https://doi.org/10.3897/fr.29.189975.figure14	Figure 14. Fore-limb elements of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain) (see Suppl. material 1: figs S 1, S 2 for precise position of the disarticulated elements in the field): A 1 – A 14. Left humerus; B 1 – B 13. Right humerus. The fore-limb elements are shown in: A 1, A 2, B 1, B 2. Proximal; A 3, A 4, B 3, B 4. Dorsal; A 7, A 8, B 5, B 6. Ventral; A 9, A 10, A 13, A 14, B 7, B 8, B 11, B 12. Pre-axial; A 11, A 12, B 9, B 10, B 13. Postaxial view. Dashed lines and areas represent broken bones. Scale bars: 50 mm (A 1 – A 12, B 1 – B 10); 10 mm (A 14, B 11); 5 mm (A 13, B 12 – B 13).	Figure 14. Fore-limb elements of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain) (see Suppl. material 1: figs S 1, S 2 for precise position of the disarticulated elements in the field): A 1 – A 14. Left humerus; B 1 – B 13. Right humerus. The fore-limb elements are shown in: A 1, A 2, B 1, B 2. Proximal; A 3, A 4, B 3, B 4. Dorsal; A 7, A 8, B 5, B 6. Ventral; A 9, A 10, A 13, A 14, B 7, B 8, B 11, B 12. Pre-axial; A 11, A 12, B 9, B 10, B 13. Postaxial view. Dashed lines and areas represent broken bones. Scale bars: 50 mm (A 1 – A 12, B 1 – B 10); 10 mm (A 14, B 11); 5 mm (A 13, B 12 – B 13).	2026-05-29	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán		Zenodo	biologists	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán			
5B30B6DB947A5684AC114F192403A675.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1661262	https://doi.org/10.3897/fr.29.189975.figure15	Figure 15. Stylopodial hind-limb elements of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain) (see Suppl. material 1: figs S 1, S 2 for precise position of the disarticulated elements in the field): A 1 – A 10. Left femur; B 1 – B 10. Right femur. The hind-limb elements are shown in: A 1, A 2, B 1, B 2. Proximal; A 3, A 4, B 3, B 4. Dorsal; A 5, A 6, B 9, B 10. Pre-axial; A 7, A 8, B 7, B 8. Ventral; A 9, A 10, B 5, B 6. Postaxial. Dashed lines and areas represent broken bones. Scale bar: 50 mm.	Figure 15. Stylopodial hind-limb elements of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain) (see Suppl. material 1: figs S 1, S 2 for precise position of the disarticulated elements in the field): A 1 – A 10. Left femur; B 1 – B 10. Right femur. The hind-limb elements are shown in: A 1, A 2, B 1, B 2. Proximal; A 3, A 4, B 3, B 4. Dorsal; A 5, A 6, B 9, B 10. Pre-axial; A 7, A 8, B 7, B 8. Ventral; A 9, A 10, B 5, B 6. Postaxial. Dashed lines and areas represent broken bones. Scale bar: 50 mm.	2026-05-29	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán		Zenodo	biologists	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán			
5B30B6DB947A5684AC114F192403A675.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1661263	https://doi.org/10.3897/fr.29.189975.figure16	Figure 16. Zeugopodial hind-limb elements of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain) (see Suppl. material 1: figs S 1, S 2 for precise position of the disarticulated elements in the field): A 1 – A 12. Left? tibia; B 1 – B 12. Right? fibula; C 1 – C 12. Left? fibula; D. Reconstruction of the left hind-limb of MUPA-ATZ 0103 in dorsal view (fibula mirrored and reconstructed from the probably right fibula). The hind-limb elements are shown in: A 1, A 2, B 1, B 2, C 1, C 2. Proximal; A 3, A 4, B 3, B 4, C 3, C 4. Lateral; A 5, A 6, B 5, B 6, C 5, C 6. Distal; A 7, A 8, B 7, B 8, C 9, C 10. Pre-axial; A 9, A 10, B 9, B 10, C 7, C 8. Postaxial; A 11, A 12, B 11, B 12, C 11, C 12. Medial view. Dashed lines and areas represent broken bones. Scale bar: 20 mm.	Figure 16. Zeugopodial hind-limb elements of the postcranial skeleton of the paratype of Paludidraco multidentatus, MUPA-ATZ 0103, from the Carnian (Late Triassic) of El Atance (Guadalajara, Central Spain) (see Suppl. material 1: figs S 1, S 2 for precise position of the disarticulated elements in the field): A 1 – A 12. Left? tibia; B 1 – B 12. Right? fibula; C 1 – C 12. Left? fibula; D. Reconstruction of the left hind-limb of MUPA-ATZ 0103 in dorsal view (fibula mirrored and reconstructed from the probably right fibula). The hind-limb elements are shown in: A 1, A 2, B 1, B 2, C 1, C 2. Proximal; A 3, A 4, B 3, B 4, C 3, C 4. Lateral; A 5, A 6, B 5, B 6, C 5, C 6. Distal; A 7, A 8, B 7, B 8, C 9, C 10. Pre-axial; A 9, A 10, B 9, B 10, C 7, C 8. Postaxial; A 11, A 12, B 11, B 12, C 11, C 12. Medial view. Dashed lines and areas represent broken bones. Scale bar: 20 mm.	2026-05-29	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán		Zenodo	biologists	Cabezuelo-Hernández, Alberto;de Miguel Chaves, Carlos;Ortega, Francisco;Pérez-García, Adán			
