taxonID	type	description	language	source
8263FE48FFC34213FF23FE10FB6AFC29.taxon	materials_examined	Type depository: BMNH: holotype (female) (Santos Dias 1956 b, Hoogstraal & El Kammah 1972, Keirans 1985) Type locality: Wandara, Ghana Type host: Redunca arundinum Current status: a synonym of Haemaphysalis rugosa and its original name, as discussed in Hoogstraal & El Kammah (1972). The current status of Haemaphysalis aciculifer rugosa, as Haemaphysalis rugosa, was also established by Hoogstraal & El Kammah (1972). Comments: contrary to the above-cited authors, Keirans & Hillyard (2001) did not recognize the presence of the holotype of Haemaphysalis aciculifer rugosa in their list of types deposited in the BMNH.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFC34213FF23FC41FCF5FA98.taxon	materials_examined	Type depositories: BMNH: a lectotype (male) was designated by Trapido (1965); ENV and MZU: paralectotypes (males) (Trapido 1965, although not named as paralectotypes) Type locality: India Type host: Tragulus meminna Comments: Keirans & Hillyard (2001) listed the type locality as “ Bodd, Indonesia, ” but Haemaphysalis aculeata is not found in Indonesia (Guglielmone et al. 2023). The abbreviation “ Bodd. ” was probably used by Lavarra (1904) for Boddaert and his redescription of Tragulus meminna in Boddaert (1784). Keirans & Hillyard (2001) considered the type host to be Tragulus javanicus or Tragulus napu.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFC34213FF23FAB0FE27F9A1.taxon	materials_examined	Type depository: ENV: syntypes (females) (Nuttall & Warburton 1911) Type locality: Genoa, Italy	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFC24212FF23FF78FB13FE05.taxon	materials_examined	Type depository: ZMB: holotype (female) (Moritz & Fischer 1981) Type locality: Japan Type host: unknown Current status: a synonym of Ixodes acutitarsus and its original name, as stated in Neumann (1911 a). The current status of Haemalastor acutitarsus, as Ixodes acutitarsus, was also established by Neumann (1911 a).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFC24212FF23FDA8FD25FCF5.taxon	description	Feldman-Muhsam, B. (1951) A note on east Mediterranean species of the genus Haemaphysalis. Bulletin of the Research Council of Israel, 1, 96 – 107. Type depository: NTCHU: a paralectotype (female) and paralectotypes (males) were designated by Anderson et al. (2024). Type locality: Ness-Ziona, Central District, Israel	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFC2421DFF23F8DCFB25FF7D.taxon	materials_examined	Type depository: IM: syntypes (females) Type locality: Sasaram, Bihar, India Type host: cattle Current status: a synonym of Hyalomma scupense, as discussed in Apanaskevich et al. (2010).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFCD421DFF23F9B4FE1AF83D.taxon	materials_examined	Type depository: unknown: holotype? (female) Type locality: Egypt Type host: unknown Current status: a preoccupied name changed to Ixodes savignyi Gervais, 1844, a name incertae sedis, as discussed in Guglielmone & Nava (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFCD421DFF23FB44FEEDF985.taxon	materials_examined	Type depository: LC: but lost; UUZM: no type specification (Wallin 1991) Type locality: Oriente Type host: unknown Current status: a synonym of Hyalomma aegyptium and its original name, as listed in Camicas et al. (1998). The current status of Acarus aegyptius, as Hyalomma aegyptium, was established by Koch (1844). Comments: Wallin (1991) discussed the status of many type specimens in the UUZM, but not in the case of Acarus aegyptius.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFCD421DFF23FD84FB02FCF5.taxon	materials_examined	Type depository: undetermined: syntypes (males and females) Type locality: Ouled-Djellal, Algeria Type host: camel Current status: a synonym of Hyalomma dromedarii, as discussed in Apanaskevich et al. (2008 b).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFCD421DFF23FC64FB94FBD5.taxon	materials_examined	Type depository: undetermined: syntypes (males and females) Type locality: Mesopotamia Type host: camel Current status: a synonym of Hyalomma anatolicum, as listed in Camicas et al. (1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFCD421DFF23FEECFEA0FD95.taxon	materials_examined	Type depositories: BMNH and IM: syntypes (males and females) (Keirans & Hillyard 2001) Type locality: Kandeli, Madhya Pradesh, India Type host: buffalo Current status: a synonym of Hyalomma isaaci and its original name. The current status of Hyalomma aegyptium isaaci, as Hyalomma isaaci, was first determined by Kratz (1940) and definitively established by Apanaskevich & Horak (2008 b).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFCC421CFF23FCF0FEB2FBD5.taxon	materials_examined	Type depositories: LMNH, MNHN and ZSH: syntypes (males and females) Type locality: undetermined Type host: most probably Testudo graeca Current status: a synonym of Hyalomma aegyptium, as listed in Camicas et al. (1998). Comments: Neumann (1899) analyzed specimens from several countries, concluding that this tick is adapted to Africa, Asia and southern Europe, but he also included England and Mexico, where Hyalomma affine was found on captive tortoises.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFCC421CFF23FF78FD8BFE5D.taxon	materials_examined	Type depositories: MCZ and USNTC: syntypes (females) (Keirans & Clifford (1984) listed these specimens as “ cotypes ”). Type locality: Berkeley Hills, California, USA. Type host: Otospermophilus beecheyi Current status: a synonym of Ixodes sculptus, as discussed in Guglielmone & Nava (2014). Comments: Keirans & Clifford (1984) did not list the developmental stages deposited in the USNTC, but Ixodes aequalis was described from female ticks.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFCC421CFF23FB45FC1BFA4D.taxon	materials_examined	Type depository: USNPC: syntypes (females) (Keirans & Clifford 1978, named as “ type ”), USNTC: syntype (female) (Cooley & Kohls 1945, named as “ cotype ”; Nava et al. 2023) Type locality: Costa Rica Type host: Felis pardalis Comments: no types of Ixodes affinis are included in the list of types deposited in the USNTC by Keirans & Clifford (1984), but Nava et al. (2023) found three females of the syntype series there.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFCF421FFF23FD5CFB67FC61.taxon	materials_examined	Type depository: not stated: holotype (female) and paratypes (females) Type locality: Akşar, Erzurum, Turkey Type host: Bos taurus Current status: a synonym of Haemaphysalis pospelovashtromae in Guglielmone et al. (2014). Comments: Bursali et al. (2012) treated Haemaphysalis aksarensis as provisionally valid.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFCF421FFF23FC19FD31FB09.taxon	materials_examined	Type depositories: USNTC: holotype (female) and paratypes (females); IEMVPT: paratype: (female). Type locality: Massif du Tasaratanana, Madagascar	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFCF421EFF23F878FE1EFEB5.taxon	materials_examined	Type locality: zoo of Rotterdam, the Netherlands Type host: Trachysaurus rugosus Comments: the syntypes of Amblyomma albolimbatum were collected in the Netherlands, but this is an Australasian species found only in Australia.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFCE421EFF23FEA5FDFBFDB1.taxon	discussion	Comments: Guglielmone et al. (2020, 2021) stated that the morphology of Amblyomma albopictum has been poorly defined and should be reexamined.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFCE421EFF23FBF5FD42FB41.taxon	materials_examined	Type depository: IBU: a lectotype (female) and a paralectotype (female) were designated by Barros-Battesti & Knysak (1999) Type locality: Barro Alto, Minas Gerais, Brazil	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFCE421EFF23F94DFB19F811.taxon	materials_examined	Type depository: FMNH?: holotype and a paratype (females); USNTC: paratypes (females). Type locality: trail of Sentier Botanique, Réserve Spéciale d’Ambohitantely, Analamanga Region, Madagascar Type host: Microgale major Comments: IR visited the FMNH in 2024 and both the holotype and paratypes were not found there.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFC94219FF23FD14FD2DFC0D.taxon	materials_examined	Type depository: USNTC: holotype (female) Type locality: Phoenix Island in the Pacific Ocean	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFC94219FF23F990FDB5F8E5.taxon	materials_examined	Type depository: CMC: lectotype (female) and paralectotypes (females) were designated by Heath & Kwak (2019). Type locality: Ashburton, New Zealand	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFC84218FF23FC41FECCFB41.taxon	description	See also venustus Neumann, 1897 Dermacentor Type depository: USNPC?: holotype? (male) in Keirans (1976), who named the specimen as “ type ” Type locality: Montana, USA Type host: not stated Comments: the “ type ” of Dermacentor andersoni does not appear in the list of types held in the USNPC by Salley et al. (1978).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFC84218FF23F9B4FDB2F83D.taxon	materials_examined	Type depository: SMNH: syntypes (females) but lost (Hoogstraal 1959) Type locality: Ankara, Turkey Type host: donkey Current status: a synonym of Haemaphysalis sulcata, as listed in Camicas et al. (1998). Comments: Guglielmone et al. (2017) stated that there is little justification for the synonymy of Haemaphysalis angorense with Haemaphysalis sulcata.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFC84218FF23FE80FB8EFDC9.taxon	materials_examined	Type depository: IAUP: holotype (male) Type locality: Murzuk, Libya Type host: not stated Current status: a synonym of Hyalomma excavatum, as listed in Camicas et al. (1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFCB421BFF23FF78FBA8FE79.taxon	materials_examined	Type depository: USNTC: a neotype (female) was designated by Robbins & Keirans (1989) Type locality: Pattee Canyon, Missoula County, Montana, USA Type host: nest of Sorex sp. Comments: a neotype was designated because the female used by Neumann (1899) to describe Ixodes angustus was, in fact, a specimen of Ixodes woodi (Robbins & Keirans 1989, 1992). The female neotype was obtained by rearing a nymph collected in a nest of Sorex sp.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFCB421BFF23FE10FBACFCBD.taxon	materials_examined	Type depository: USNTC: holotype (female) (Keirans & Clifford 1978) Type locality: Sabinal, Texas, USA Type host: Neotoma micropus Current status: a synonym of Ixodes woodi (Keirans & Clifford 1984) and its original name. The current status of Ixodes angustus woodi as Ixodes woodi, was established by Cooley & Kohls (1945).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFCB421BFF23FA48FD85F949.taxon	materials_examined	Type depository: unknown: holotype (male?), but lost according to Hooker (1909) Type locality: eastern Florida, USA Type host: Cervus virginianus Current status: a synonym of Rhipicephalus annulatus and its original name (Guglielmone & Nava 2014). The current status of Ixodes annulatus, as Rhipicephalus annulatus, was first established by Neumann (1897) and definitively by Murrell & Barker (2003).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFCA421AFF23FDA8FF04FC29.taxon	materials_examined	Type depository: ZMB: holotype (male) (Moritz & Fischer 1981) Type locality: South Africa Type host: unknown Current status: a synonym of Amblyomma hebraeum, as listed in Camicas et al. (1998). Comments: a female included in the original description of Amblyomma annulipes is lost (Moritz & Fischer 1981).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFCA421AFF23FA6DFBBEF9F9.taxon	materials_examined	Type depository: USNTC: holotype (female) and paratype (female) Type locality: Amboasary-Est, Anjozorobe District, Tananarive Province, Madagascar	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFCA421AFF23FBF4FA74FAFD.taxon	description	In: Teng, K. F. & Cui, Y. Q. (1984) Descriptions of a new species of Haemaphysalis and the male of H. primitiva Teng, 1982 from Yunnan. Acta Zootaxonomica Sinica, 9, 37 – 40. [in Chinese] Type depository: IZAS: holotype (male) Type locality: Lushui County, Yunnan Province, China Type host: the holotype specimen was collected from vegetation Comments: Camicas et al. (1998) and Kolonin (2009) treated Haemaphysalis anomaloceraea as an invalid name.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFD54205FF23F8C4FCF0F87E.taxon	materials_examined	Type depository: not stated: syntypes (a male and nymphs) Type locality: Kremmener Luch, Brandenburg, Germany	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFD44204FF23FB8DFCBDFA99.taxon	materials_examined	Type depositories: USNTC: holotype (male) and paratypes (males and females); BMNH: paratypes (males and females) and DB: a paratype (male) Type locality: Al Hamilee (Al Muroa), Ta’izz Province, Yemen	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFD44204FF23F904FBA7F83D.taxon	description	See ricinus aragaoi Fonseca, 1935 Current status: a synonym of Ixodes fuscipes, as discussed in Labruna et al. (2020).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFD44204FF23FD39FE06FB9D.taxon	materials_examined	Type depositories: BMNH: holotype (male) and paratypes (males and females); OVI and USNTC: paratypes (males and females) Type locality: Kaisho, Karagwe District, Tanzania Type host: Tragelaphus spekei Comments: Walker et al. (2000) stated that Rhipicephalus aquatilis was originally treated as Rhipicephalus sp. III by Yeoman & Walker (1967).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFD74207FF23F94CFE69F875.taxon	materials_examined	Type depository: ZMB: syntypes (males) (Moritz & Fischer 1981) Type locality: Chinchoxo, West Africa Type host: unknown Comments: Kaufman (1972) and Kolonin (2009) treated Amblyomma arcanum (under the former genus Aponomma) as invalid, but it was considered valid by Santos Dias (1993) and provisionally valid by Guglielmone et al. (2014, 2020, 2023) and here.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFD74207FF23FD85FE45FC91.taxon	materials_examined	Type depository: not stated: syntypes (males, females and nymphs) Type locality: Germany	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFD74207FF23FB68FC60FA35.taxon	materials_examined	Type depository: not stated: syntypes (females) Type locality: Hamburg, Germany Type host: the syntype specimens were collected from a nest of Phoenicurus ochruros Current status: a synonym of Ixodes arboricola in Haarløv (1962).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFD74207FF23FA24FC60F9DD.taxon	materials_examined	Type depository: NRS: syntypes (females) (Haarløv 1962) Type locality: Floda, Sweden Type host: Muscicapa grisola Current status: a synonym of Ixodes arboricola in Haarløv (1962).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFD74207FF23FF78FEAEFD95.taxon	materials_examined	Type depositories: ZSI?: holotype (male) and paratypes (a male, a female); USNTC?: paratype (?) (Walker et al. 2000) Type locality: Anand, Gujarat, India Type host: Mus rattus Current status: a synonym of Rhipicephalus ramachandrai, as discussed in Walker et al. (2000). Comments: Hiredougar (1975) listed larvae and nymphs as paratypes of Rhipicephalus arakeri, but only the male and female were described. Walker et al. (2000) stated that the types were not found in the ZSI, and also listed a paratype held in the USNTC, but this specimen does not appear in the catalog of USNTC types by Keirans & Clifford (1984).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFD64206FF23FBD0FDFDFA35.taxon	materials_examined	Type depository: not stated (Neumann 1905): syntypes (males, females and a nymph); BMNH: a syntype (male) (Keirans & Hillyard 2001). Type locality: Argentina Type host: Testudo argentina (Neumann 1905) Comments: this species was first described by Conil (1877) under the name Ixodes testudinis (preoccupied), as discussed in Guglielmone et al. (2001). However, Neumann (1905) described Amblyomma argentinae without considering this tick a synonym or a new name for Ixodes testudinis Conil. 1877. Keirans & Hillyard referred to the type host as Geochelone chilensis.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFD64206FF23FF78FF0DFE5D.taxon	materials_examined	Type depository: USNTC?: holotype (female) Type locality: Pribilof Islands, Alaska, USA Type host: Callorhinus ursinus Current status: a synonym of Ixodes signatus, as stated in Keirans & Clifford (1978). Comments: the holotype of Ixodes arcticus is not listed in the catalog of USNTC types by Keirans & Clifford (1984).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFD64206FF23FA25FD0AF915.taxon	materials_examined	Type depositories: DPZ: holotype (female) and paratypes (females and nymphs); HNMH: paratypes (females, nymphs and a larva) Type locality: Leány Cave, Pilis Mountains, Hungary	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFD14201FF23FC88FB7FFBD5.taxon	materials_examined	Type depository: BMNH: a lectotype (female) and paralectotypes (females) were designated by Keirans & Brewster (1981) Type locality: Quy, England Type host: Arvicola terrestris Current status: a synonym of Ixodes apronophorus, as stated in Keirans & Hillyard (2001).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFD14201FF23FA48FC72F9DD.taxon	materials_examined	Type depositories: NSM: holotype (male) and paratypes (males and females); NIAH: paratypes (males and females) Type locality: Sukumobanare Islands, Amami-oshima Islands, Japan	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFD04200FF23FD38FC65FC45.taxon	materials_examined	Type depository: ZIAC: paratypes (a male and a female) (Filippova 2008) Type locality: Khabarovsk, Russia (Emel’yanova & Kozlovskaya 1967) Type host: the type specimens were collected from vegetation Current status: a synonym of Dermacentor silvarum, as stated in Filippova (2008). Comments: the type locality was not included in Filippova (2008).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFD04200FF23FBF4FB6CFA99.taxon	materials_examined	Type depository: GM: holotype (male) (Hoogstraal & Trapido 1966 a) Type locality: Meteleó, Myanmar Type host: unknown Current status: a synonym of Haemaphyalis asiatica and its original name (Camicas et al. 1998). The current status of Opisthodon asiaticus, as Haemaphysalis asiatica, was established by Neumann (1897).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFD04200FF23FE58FB2EFD01.taxon	materials_examined	Type depository: ZMA: holotype (male) and paratypes (males and females) Type locality: Maralbashi, Xinjiang, China Type host: undetermined Current status: a synonym of Hyalomma asiaticum, as stated in Apanaskevich & Horak (2010).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFD34203FF23FD60FF1BFC0C.taxon	materials_examined	Type depository: BMNH: holotype (male) Type locality: unknown Type host: unknown Current status: a synonym of Dermacentor auratus, as explained in Apanaskevich et al. (2021). Comments: Apanaskevich et al. (2021) redetermined several records of Dermacentor atrosignatus as Dermacentor tricuspis.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFD24202FF23FD61FB61FBB9.taxon	description	Pallas, P. S. (1772) Spicilegia Zoologica. Vol. 40. Fascicule 9. Prostant apud Gottl. August. Lange, 87 pp. Type depository: unknown: holotype? (male) Type locality: unknown Type host: unknown Current status: a synonym of Amblyomma aureolatum and its original name (Camicas et al. 1998). The current status of Acarus aureolatus, as Amblyomma aureolatum, was established by Koch (1844).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFD24202FF23FEECFEA5FDC9.taxon	materials_examined	Type depositories: BMNH and GM: syntypes (females) (Keirans & Hillyard 2001) Type localities: Carin Cheba and Mooleyit, Myanmar Type host: Ursus torquatus and Sus cristatus (Supino 1897) Comments: Wassef & Hoogstraal (1984) stated that Supino described the male and female of Dermacentor auratus, and Keirans & Hillyard (2001) listed a male syntype deposited in the BMNH, but Supino (1897) described the female of Dermacentor auratus, not the male. Keirans & Hillyard (2001) named the type hosts as Ursus tibetanus and Sus scrofa.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFD2420DFF23F894FB4CFEE8.taxon	materials_examined	Type depository: unknown: syntypes (males and females) (Guglielmone et al. 2003, 2021) Type locality: Argentina Type host: Dasypus villosus Current status: a synonym of Amblyomma auricularium and its original name (Lahille 1905, as Amblyomma auriculare). The current status of Ixodes auricularius was also established by Lahille (1905).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFDD420DFF23FD61FD13FBD5.taxon	materials_examined	Type depository: ENV?: syntype (a female) (Keirans & Clifford 1978, named as “ type ”), MNHN?: syntypes (females) (Neumann 1904) Type locality: Punta Arenas, Chile Type host: undetermined species of birds Comments: Ixodes auritulus was first named Ixodes thoracicus by Neumann (1899), based on ticks collected in Chile. Ixodes thoracicus was described by Koch (1844) from specimens collected in Brazil, and Neumann (1904) realized that his Chilean specimens corresponded to a different species that he named Ixodes auritulus. Guglielmone et al. (2014) stated that the types of Ixodes auritulus were not found in the MNHN, and Apanaskevich et al. (2022) did not confirm the presence of a syntype in the ENV. Ixodes auritulus is a name that represents a species complex with a global distribution, as demonstrated by Apanaskevich et al. (2022), but according to these authors Ixodes auritulus sensu stricto is a Neotropical species, bona fide specimens of which were collected in Chile.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFDD420DFF23FB44FF0DFA4D.taxon	materials_examined	Type depositories: MNZT: holotype (male) and paratypes (a female and a nymph); DEDSIR: paratypes (females and nymphs) Type locality: Snare Island, New Zealand Type host: Pelecanoides urinatrix Current status: a synonym of Ixodes zealandicus and its original name, as discussed in Apanaskevich et al. (2022).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFDD420CFF23F8DCFA8AFF51.taxon	materials_examined	Type depository: ZSH: holotype (male) and a paratype (female) Type locality: Sydney, Australia Type host: wombat Current status: a synonym of Bothriocroton auruginans (Klompen et al. 2002) and its original name. The current status of Aponomma auruginans, as Bothriocroton auruginans, was also established by Klompen et al. (2002).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFDC420CFF23FC19FD6EFB09.taxon	materials_examined	Type depository: USNTC: neotype (female) (Estrada-Peña et al. 2012) Type locality: Gatton, Queensland, Australia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFDF420FFF23FF78FE9CFE79.taxon	materials_examined	Type depository: USNTC?: holotype (nymph) Type locality: Trinidad, Trinidad and Tobago Type host: “ Goube-mouche ” Current status: a synonym of Amblyomma longirostre, as listed in Camicas et al. (1998). Comments: Keirans & Clifford (1984) did not include specimens of Amblyomma avicola in their catalog of types held in the USNTC.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFDF420FFF23FB69FC6CFA35.taxon	materials_examined	Type depository: USNTC: a lectotype (female) and paralectotypes (nymphs) were designated by Keirans & Robbins (1987). Type locality: Buru or Sulawesi Islands, Indonesia Type host: Babyrousa babyrussae Comments: Schulze (1933 b) named the type host as Sus babirussa.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFDF420FFF23FCF0FF43FBF1.taxon	materials_examined	Type depositories: BMNH, VRLKK and NMK: syntypes (males and females) (Walker et al. 2000) Type localities: Kiagu, Mbeyo, Mount Kenya and Masai Reserve, Kenya Type hosts: buffalos, rhinoceros and lion Current status: a synonym of Rhipicephalus compositus, as listed in Walker et al. (2000). b	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFDE420EFF23FB1DFD09FA69.taxon	materials_examined	Type depository: BMNH: a lectotype (male) and paralectotypes (females and nymphs) were designated by Keirans & Brewster (1981) Type locality: Burnett District, Queensland, Australia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFDE420EFF23FF79FBB5FE05.taxon	materials_examined	Type depositories: USNTC: holotype (female) and paratypes (males and females); DEEZ, DEP?, DEUA, ECU?, MCZ and USNPC: paratypes (males and females) (Keirans & Clifford 1978) Type locality: Washington County, Arkansas, USA Type host: type specimens were collected from cliffs used by cliff swallows Comments: no paratypes were found in DEP and ECU by Keirans & Clifford (1978).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFDE420EFF23FE35FE74FD01.taxon	materials_examined	Type depository: MCZ: holotype (female) Type locality: Nyika plateau, Malawi Type host: Otomys sp.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFDE4209FF23F8DDFF0DFEE9.taxon	materials_examined	Type depositories: BMNH and USNTC: syntypes (females) (Keirans & Clifford 1978); MCZ: paratypes (females) Type locality: Mammoth Springs, Arkansas, USA Type host: Ondatra zibethicus Comments: Keirans & Clifford (1978) listed paratypes of Ixodes banksi deposited in the BMNH, MCZ and USNTC, a decision supported by Keirans & Clifford (1984) for the USNTC. No holotype was specifically designated for Ixodes banksi, and types of this species are treated here as syntypes, an approach also taken by Keirans & Hillyard (2001).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFD94209FF23FAD5FC32F9A1.taxon	materials_examined	Type depositories: QM: holotype (male); ANIC: paratype (male) Type locality: Peeramon scrub, Atherton Tableland, Queensland, Australia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFD84208FF23FD60FC61FC29.taxon	materials_examined	Type depository: not stated: holotype? (male) Type locality: Keren, Eritrea Type host: unknown Current status: a name incertae sedis (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFDB420BFF23FE59FD26FD01.taxon	materials_examined	Type depositories: USNTC: holotype (male) and paratypes (males, females, nymphs); OVI: paratypes (males, females, nymphs and a larva), BMNH, EAVRO, FMNH and MCZ: paratypes (males, females and nymphs) Type locality: Imurok, Torit County, South Sudan	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFDB420BFF23FBF5FE01FAFD.taxon	materials_examined	Type depository: SAIMR: syntypes (males) (Walker et al. 2000) Type locality: “ Lissenji ” (?) [sic], Central Africa (Zumpt 1950) Type host: Syncerus caffer Comments: Hoogstraal (1956 a) noted that the name of the type locality may be a misspelling of Kisenyi, which is actually a locality in Rwanda.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFDB420BFF23FA6DFD24F985.taxon	materials_examined	Type depository: not stated: holotype (male) and a paratype (female) Type locality: Hubeta, Chercher, Harrar, Ethiopia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFDB420BFF23FF78FB1EFE21.taxon	materials_examined	Type depository: GM (presumed): holotype (female) Type locality: Bhamó, Myanmar Type host: Varanus bengalensis Current status: a nomen dubium, as listed in Camicas et al. (1998) and Guglielmone & Nava (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFDA420AFF23FEA4FE25FD65.taxon	materials_examined	Type depository: unknown Type locality: South America Type host: boa Current status: a nomen dubium (Guglielmone & Nava 2014). Comments: the name Ixodes bibroni Gervais, 1844, has been applied as Amblyomma bibroni (Gervais, 1844) by some workers, instead of Amblyomma dissimile, but this is unjustified. In other cases the name Amblyomma bibroni (Gervais, 1842) has been employed, but this is a nomen nudum, as discussed in Guglielmone & Nava (2014) and Guglielmone et al. (2021).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFDA420AFF23FD14FE6BFC45.taxon	materials_examined	Type depository: ENV: syntypes (females) (Nuttall & Warburton 1911, named as “ types ”) Type locality: Atoyac, State of Guerrero, Mexico Type host: Felis onca Current status: a synonym of Ixodes boliviensis in Kohls (1956 b), treated as a provisional synonym in Guglielmone et al. (2021) and here.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFDA420AFF23FBF5FE6BFB41.taxon	materials_examined	Type locality: Malawi	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFDA420AFF23F9B4FA91F8D9.taxon	materials_examined	Type depository: BMNH?: syntypes (females) (Santos Dias 1958 a) Type locality: southern Africa Type host: Hippopotamus amphibius Current status: a synonym of Cosmiomma hippopotamensis (Guglielmone & Nava 2014). Comments: Keirans & Hillyard (2001) did not list the types of Ixodes bimaculatus as being held in the BMNH.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE54235FF23FD61FF31FC0D.taxon	materials_examined	Type depository: JWC: holotype (male?) but it will probably be transferred to SMF. Type locality: a fossil species found in Burmese amber Type host: unknown Comments: based on its morphology, Guglielmone et al. (2020) believe that the holotype specimen is probably a male.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE54235FF23FC3CFB37FB41.taxon	materials_examined	Type depository: ZIAC: lectotype (male) and paralectotypes (males and a female) (Filippova 2008) Type locality: upper Re-chu River, Basin of Golubaya River, eastern Tibet, China Type host: not stated Current status: a synonym of Dermacentor everestianus, as discussed in Apanaskevich et al. (2014). Comments: Filippova (2008) and Kolonin (2009) treated Dermacentor birulai as a valid species.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE54235FF23FAF8FC45F9A1.taxon	materials_examined	Type depository: ZIAC: lectotype (female) and paralectotypes (females) (Filippova 2008) Type locality: Lake Kuku-Nor, eastern Tibet, China Type host: Equus hemionus Current status: a synonym of Dermacentor nuttalli (Filippova 2008).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE54235FF23FEA5FB59FDC9.taxon	materials_examined	Type depositories: BMNH: lectototype (male) and a paralectotype (female); GM: paralectotypes (males and a female) (Hoogstraal et al. 1965 a) Type locality: Carin-Gheen or Yado Cari Ascinii Cheba, Myanmar Type host: Muntiacus muntjak or Atherurus macrourus (Hoogstraal et al. 1965). Comments: Keirans & Hillyard (2001) treated the specimens deposited in the GM as syntypes.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE54235FF23F9D8FC54F881.taxon	materials_examined	Type depository: USNTC: holotype (female) and paratypes (females, nymphs and larvae) Type locality: Savannah, Georgia, USA Type host: Sigmodon hispidus Current status: a synonym of Ixodes minor (Keirans & Clifford 1984).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE54234FF23F8B9FEB3FF7D.taxon	materials_examined	Type depository: BMNH: a neotype (female) was designated by Trapido (1965) Type locality: State of Madras, India Type host: cattle.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE44234FF23FD84FB96FCF5.taxon	materials_examined	Type depository: BMNH: syntypes (females) Type locality: Venezuela Type host: unknown Current status: a synonym of Amblyomma coelebs, as discussed in Guglielmone et al. (2021). Comments: Keirans & Hillyard (2001) treated Amblyomma bispinosum as a valid name.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE44234FF23FEECFD8DFD95.taxon	discussion	Comments: a lectotype was designated by Hoogstraal & Trapido (1963) under the name Haemaphysalis intermedia, not Haemaphysalis bispinosa intermedia.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE44234FF23FC65FAB5FB09.taxon	materials_examined	Type depositories: CZIITC: holotype (female) and a paratype (male); BMNH?: a paratype (female) Type locality: Porto de Mõs, Leiria, Portugal Type host: Oryctolagus cuniculus Comments: in their catalog of BMNH types, Keirans & Hillyard (2001) found no evidence that a paratype of Ixodes bivari had been deposited in that museum. Ixodes bivari is not treated as valid in Estrada-Peña et al. (2017).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE44234FF23FB20FEADF9A1.taxon	materials_examined	Type depository: not stated: syntypes (nymphs) Type locality: Retalhuleu, Guatemala Type host: Boa imperator Current status: a synonym of: Amblyomma dissimile, as discussed in Guglielmone et al. (2021). Comments: Stoll (1894) described the female of Ixodes boarum, but left open the possibility that the specimens were nymphs because their genital apertures were rudimentary. Neumann (1899) confirmed that the ticks were indeed nymphs.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE44234FF23F9D9FC88F8AD.taxon	materials_examined	Type depository: USNTC: holotype (female) and paratypes (a male and females) Type locality: Timi del Boa, Río Teribe, Bocas del Toro, Panama	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE74237FF23FC88FB5CFBD5.taxon	materials_examined	Type depositories: ISET: holotype (male) and paratypes (males, females, nymphs, a larva); CAN and ENCB: paratypes (males, females, nymphs and larvae) Type locality: Tehuitzla, Morelos, Mexico Type host: unidentified iguana Current status: a provisional synonym of Amblyomma scutatum in Guglielmone et al. (2021).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE74237FF23FA25FCC8F9DD.taxon	materials_examined	Type depository: BMNH: holotype (male) Type locality: Bode River, Sandakan Bay, Sabah, Malaysia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE64236FF23FAB0FC79F915.taxon	materials_examined	Type depositories: MCZ: syntypes (males and females) Type locality: Nicaragua and USA Type hosts: cattle and Dasyprocta Current status: a nomen dubium Comments: Ixodes bovis was treated as a synonym of Rhipicephalus annulatus in Guglielmone & Nava (2014). However, the specimens were collected not just in the USA, but also in Nicaragua where Rhipicephalus annulatus is not present. Ixodes bovis is considered to be a nomen dubium here.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE64236FF23FF79FE19FECD.taxon	materials_examined	Type depositories: ENV: holotype (male) and paratypes (a male and females); LFE: paratypes (a male and a female) Type locality: Agouagon, Benin	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE04230FF23FEA5FB73FD95.taxon	materials_examined	Type depository: USNTC: holotype (female) and paratypes (females, a nymph and a larva) Type locality: Lake Louise, 27 km northwest of Telefomin, West Sepik, Papua New Guinea	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE04233FF23F905FC4CFF7D.taxon	materials_examined	Type depository: IP: holotype (female) (Morel 1965 b) Type locality: Harar, Ethiopia (Nuttall & Warburton 1911, Morel 1965 b) Type host: Procavia brucei (Nuttall & Warburton 1911) Comments: this species is based on a specimen classified by Nuttall & Warburton (1911) as Ixodes ugandanus djaronensis. Keirans (1985) did not recognize Ixodes brumpti as valid and classified this specimen as Ixodes procaviae. Morel (1965 b) named the type host as Heterohyrax brucei.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE34233FF23FEEDFDFFFDB1.taxon	materials_examined	Type depository: ZMB: holotype (female) (Moritz & Fischer, 1981) Type locality: North America Type host: Fringilla albicollis Comments: Ixodes brunneus is a Nearctic species, despite incorrect claims that it is established in other zoogeographic regions (Guglielmone et al. 2021).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE24232FF23FB1CFE4AF984.taxon	description	See also cajennensis Fabricius, 1787, Acarus Type depository: USNTC: a neotype (male) was designated by Nava et al. (2014 a) Type locality: Montalbó hiking trail, Cayenne, French Guiana Type host: the holotype specimen was collected from vegetation Comments: this species was originally named Acarus cajennensis, but the neotype was described under the name Amblyomma cajennense.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE24232FF23F9B4FBA6F8E5.taxon	materials_examined	Type depository: INTAEC: holotype (female) Type locality: Legua 40, La Verde, Chaco, Argentina Type host: Hydrochoerus hydrochaeris Current status: a synonym of Amblyomma dubitatum (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE24232FF23FF78FF47FD95.taxon	materials_examined	Type depository: not stated: syntypes (females), but lost (Estrada-Peña et al. 2018) Type locality: Barcelona, Spain Type host: fox Current status: a synonym of Rhipicephalus pusillus and its original name (Gil Collado 1938). The current status of Rhipicephalus bursa pusillus, as Rhipicephalus pusillus, was also provided by this author. Comments: a neotype was designated under the name Rhipicephalus pusillus, not Rhipicephalus bursa pusillus. c	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE2423DFF23F894FB13FECD.taxon	materials_examined	Type depository: ZMUC: holotype (male), but lost (Nava et al. 2014 a) Type locality: Cayenne, French Guiana Type host: unknown Current status: a synonym of Amblyomma cajennense and its original name (Nava et al. 2014 a). Comments: see Amblyomma cajennense for the designation of a neotype under that name. The specimen deposited in the ZMUC as Acarus cajennensis is, in fact, the specimen of Acarus lineatus (Nava et al. 2014 a).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFED423DFF23FE7DFBC7FDC9.taxon	materials_examined	Type depositories: WAM: holotype (male) and paratypes (a female and a nymph); ANIC: paratypes (males, a female and nymphs) Type locality: 11 km south-southwest of Learmonth, Western Australia, Australia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFED423DFF23FA25FE5EF8AD.taxon	materials_examined	Type depositories: syntypes, BMNH (a female); MNHN (a female), USNTC? (a male), ZSH (females) (Neumann 1899) Type locality: Brazil and Paraguay Type hosts: Myrmecophaga tetradactyla and dog Comments: Keirans & Clifford (1984) did not include specimens of Amblyomma calcaratum in their catalog of types held in the USNTC.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFED423DFF23FBD0FB6BFA35.taxon	materials_examined	Type depositories: BMNH: lectotype (male), paralectotypes (males, a female and a nymph), ZMB: paralectotypes (males and a nymph). The lectotype and paralectotypes were designated by Keirans & Brewster (1981) Type locality: Bade, Central African Republic Type host: unknown Current status: a synonym of Haemaphysalis houyi and its original name (Camicas et al. 1998). The current status of Haemaphysalis calcarata houyi, as Haemaphysalis houyi, was established by Hoogstraal (1954). Comments: Keirans & Hillyard (2001) named as syntypes the specimens held in the ZMB.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFED423CFF23F8DCFB67FF7D.taxon	materials_examined	Type depositories: LP: holotype (male) and a paratype (female); SDC: paratypes (a male and females) Type locality: Piritu, State of Guárico, Venezuela Type host: Tamandua tetradactyla Current status: a synonym of Amblyomma calcaratum, as listed in Camicas et al. (1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFEC423CFF23FDA8FE36FC29.taxon	materials_examined	Type depository: ZMB: syntypes (females) (Moritz & Fischer 1981) Type locality: Sardinia, Italy and Damascus, Syria Type host: cattle Current status: a synonym of Rhipicephalus annulatus (Guglielmone & Nava 2014). Comments: male ticks are included in the original description, but none are listed in the syntype series documented by Mortiz & Fischer, 1981).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFEC423CFF23FC40FBB1FB09.taxon	materials_examined	Type locality: Hostalets de Bas, Gerona, Spain Type host: not stated Current status: a synonym of Rhipicephalus annulatus (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFEC423CFF23FB20FBB1FA69.taxon	materials_examined	Type depository: not stated: holotype (male) and paratypes (males and females) Type locality: Tiberias, Israel Type host: cattle Current status: a synonym of Rhipicephalus annulatus (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFEC423CFF23FA00FBB1F915.taxon	materials_examined	Type locality: Ramian, Iran Type host: not stated Current status: a synonym of Rhipicephalus annulatus (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFEC423CFF23FEECFBB1FDB1.taxon	materials_examined	Type depository: ZIAC: lectotype (a female) and paralectotypes (females) (Filippova 2008), MCZ: syntypes (females) Type locality: Lagodekhi, Province of Tiflis, Georgia Type host: cattle in Birula (1894) Current status: a synonym of Rhipicephalus annulatus (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFEC423CFF23F905FE6BF8D9.taxon	materials_examined	Type locality: Zambia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFEF423FFF23FE59FC03FD65.taxon	materials_examined	Type depository: BMNH: a lectotype (female) and paralectotypes (nymphs and larvae) were designated by Keirans & Brewster (1981) Type locality: Fastcastle, Scotland, Great Britain Type host: the lectotype was collected from rocks under a nest of doves Comments: Guglielmone et al. (2020) treated the morphological definitions of Ixodes caledonicus as uncertain; redescription of the type material may help clarify the status of this species.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFEF423FFF23FD14FBEBFC45.taxon	materials_examined	Type depository: not stated: holotype (female) and paratypes (females and nymphs) Type locality: Munich, Germany Type host: domestic pigeons Current status: a synonym of Ixodes caledonicus (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFEF423FFF23FBF4FB50FA99.taxon	materials_examined	Type depository: MCZ: unknown (nymphs) Type locality: Claremont, California, USA Type host: Toxostoma crissalis Current status: a synonym of Ixodes brunneus (Keirans & Clifford 1978). Comments: the number of female specimens used to describe Ixodes californicus is uncertain.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFEE423EFF23FB68FD63FA4D.taxon	materials_examined	Type depository: FF: holoype (male) and paratypes (females) Type locality: El Prado, Madrid, Spain Type host: Oryctolagus cuniculus Current status: a synonym of Haemaphysalis hispanica and its original name (Hoogstraal & Morel 1970). The current status of Haemaphysalis campanulata hispanica, as Haemphysalis hispanica, was also provided by Gil Collado (1948). Comments: Hoogstraal & Morel (1970) listed these specimens as deposited in the CGC, but Guglielmone et al. (2014) stated that the CGC is held in the FF.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFEE423EFF23F9FCFB52F8AD.taxon	materials_examined	Type depository: ZMB: paratypes (females) (Moritz & Fischer, 1981) Type locality: Beijing, China Type host: dog Current status: a synonym of Haemaphysalis campanulata, as listed in Camicas et al. (1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFEE423EFF23FF78FD60FECD.taxon	materials_examined	Type depository: USNTC: holotype (male) and paratypes (males and females) Type locality: 5 km north of Kolda, Senegal	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFEE4239FF23F8DCFBFBFF7D.taxon	materials_examined	Type depository: IPCAS: holotype (female) Type locality: Qukës, Albania Type host: goat Current status: a synonym of Ixodes gibbosus, as listed in Camicas et al. (1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE94239FF23F9B4FD82F8AD.taxon	materials_examined	Type depository: ZMB: holotype (male) (Walker et al. 2000) Type locality: Cape Colony, South Africa	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE94239FF23FC64FF1AFB6D.taxon	materials_examined	Type depository: BMNH: neotype (female) (Arthur, 1963) Type locality: Ballater, Glen Gavin, Aberdeen, Scotland, Great Britain Type host: dog Comments: Filippova (1977), Teng & Jiang (1991) and Kolonin (2009) treated Ixodes canisuga as a synonym of Ixodes crenulatus, but both species are valid in Guglielmone et al. (2014, 2020, 2023), Estrada-Peña et al. (2017) and here.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE94239FF23FB1CFC66F985.taxon	materials_examined	Type depository: MNHN: holotype (male) (Wilson 1970) Type locality: Port-Hopeful, Kerguelen Islands, Indian Ocean Type host: not stated Current status: a synonym of Ixodes kerguelenensis and its original name (Wilson 1970). The current status of Ixodes canisuga kerguelenensis, as Ixodes kerguelenensis, was established by Zumpt (1952). Comments: Guglielmone et al. (2014) list the holotype as syntype.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE84238FF23FD38FF1AFBB9.taxon	materials_examined	Type depositories: BMNH: holotype (male) and paratypes (males); USNTC: a paratype (male) Type locality: Sungei Opis, Satun, Thailand Type host: Capricornis sumatraensis swettanhami Comments: the type specimens of Haemaphysalis capricornis were originally identified as Haemaphysalis hystricis.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE84238FF23F991FB2CF8D9.taxon	materials_examined	Type depositories: BMNH: holotype (male) and paratypes (males, females, nymphs and larvae); EAVRO, OVI and USNTC: paratypes (males, females, nymphs and larvae) (Keirans & Hillyard 2001) Type locality: Muguga, Kiambu District, Kenya Type host: type specimens were reared in a laboratory from a female tick collected on a lioness.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFE8423BFF23F877FC44FECD.taxon	materials_examined	Type depositories: MCZ: holotype (male); ISET: paratypes (males) and ENCB: paratypes (males) Type locality: Las Majadas, Tehuantepec, Oaxaca, Mexico Type host: unidentified iguana Current status: a synonym of Amblyomma scutatum, as listed in Camicas et al. (1998). Comments. Vargas & Hoffmann (1952) listed the holotype as having been deposited in the ISET, but KLC, IR and RGR visited the MCZ in 2024 and the holotype is found in the MCZ.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFEB423AFF23F904FF1EFEB5.taxon	materials_examined	Type depositories: USNTC: holotype (male) and paratypes (males and females); BMNH?, FMNH: paratypes (a male and a female), MCZ: paratypes (a male and a female), and PNM: paratypes (males and females) Type locality: Myitkyina area, Myanmar Type host: Centropus bengalensis bengalensis Current status: a synonym of Haemaphysalis doenitzi, as listed in Hoogstraal & Wassef (1973) and Camicas et al. (1998). Comments: in their catalog of BMNH tick types, Keirans & Hillyard (2001) did not list specimens of Haemaphysalis centropi.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF54225FF23FB1CFE60FA35.taxon	materials_examined	Type depository: MNHN: syntypes (males and females) Type locality: Congo	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF54225FF23FC41FCACFB6D.taxon	materials_examined	Type depository: ZMB: holotype (female) (Moritz & Fischer, 1981) Type locality: Pará, Brazil Type host: unknown Comments: Kolonin (2009) did not list Haemaphysalis cinnabarina as a valid species, while Guglielmone et al. (2020, 2021, 2023) treated this species as provisionally valid.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF54224FF23F895FCF4FEE9.taxon	materials_examined	Type depository: not stated: holotype (male) and a paratype (female) Type locality: Assagni, Ivory Coast Type host: Syncerus caffer nanus Comments: Rhipicephalus cliffordi is not treated as valid in Walker et al. (2000) but is considered provisionally valid in Guglielmone et al. (2009, 2020, 2023) and here.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF44224FF23F90CFB12F852.taxon	materials_examined	Type depositories: USNTC: holotype (female) and paratypes (male, females, nymphs? and larvae?); AR and BMNH: paratypes (a male? females, nymphs? and larvae?) Type locality: Ambatofinandrahana, Madagascar Type host: Rattus rattus alexandrines Comments: the life stages of some of the paratypes deposited in the above collections are uncertain.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF44224FF23FE81FE45FDB1.taxon	materials_examined	Type depository: ZSH: syntypes (males) Type locality: unknown	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF74227FF23FB20FBFCFA35.taxon	materials_examined	Type depository: LZI: syntypes (males and females) Type locality: Colombia Type host: not stated Current status: a synonym of Dermacentor nitens (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF74227FF23FE59FEDEFCBD.taxon	materials_examined	Type depositories: HNHM: holotype (female); DPZ: paratypes (a female and a nymph), IEBR: a paratype (nymph) Type locality: Ngoc Linh Nature Reserve, Province of Kon Tum, Vietnam Type host: Hipposideros pomona Comments: Guglielmone et al. (2014) listed the DPZ as the depository for the holotype, but this specimen is held in the HNHM.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF74227FF23FCADFE80FB09.taxon	materials_examined	Type depository: ZMB?: holotype (female); USNTC: holotype? (female) (Keirans & Clifford 1984, as “ type ”) Type locality: Watom Island, near New Britain Island, Papua New Guinea Type host: Collocalia spodiopygia reichenowi Comments: in the original description it is stated that the holotype would be transferred to the ZMB, but no specimen of Ixodes collocaliae is listed in the catalog of ZMB types by Moritz & Fischer (1981). Keirans & Clifford (1984) did not specify the developmental stage of the specimen held in the USNTC, but Ixodes collocaliae was described from female ticks.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF74227FF23FA25FB90F931.taxon	materials_examined	Type depository: FMS: holotype (female) and paratypes (nymphs and larvae) Type locality: suburbs of Fukushima, Japan Type host: the holotype molted from a laboratory-reared nymph that had fed on a rabbit.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF64226FF23F96DFC21F877.taxon	materials_examined	Type depository: not stated: syntypes (males) Type locality: West Africa Type host: Manis multiscutata Current status: a synonym of Amblyomma compressum and its original name, as listed in Camicas et al. (1998). The current status of Adenopleura compressum, as Amblyomma compressum, was provided by Schulze (1941) and not by Neumann (1901), as stated in Guglielmone & Nava (2014) and Guglielmone et al. (2017). Comments: Theiler (1962) stated that the type specimens have been lost.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF64226FF23FF78FA69FE5D.taxon	materials_examined	Type depository: ZMB: syntypes (males and females) Type locality: Borneo, Indonesia? Malaysia? Type host: Sus larvatus Comments: Wassef & Hoogstraal (1983) designated a lectotype and paralectotypes from specimens in the Neumann tick collection at the ENV, but it is unclear whether the specimens deposited there belong to the syntype series of Dermacentor compactus. Syntypes are still held at the MZB, as detailed in Moritz & Fischer (1981), and the lectotype and paralectotypes in Wassef & Hoogstraal (1983) are probably invalid, as discussed in Guglielmone et al. (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF64226FF23FDF2FC0BFC36.taxon	materials_examined	Type depository: ZMB?: holotype (male) Type locality: Binaluan, Manila, Philippines Type host: not stated Current status: a synonym of Dermacentor tricuspis and its original name, as discussed in Apanaskevich et al. (2021). The current status of Dermacentor compactus tricuspis, as Dermacentor tricuspis, was also established by these authors. Comments: Schulze (1933 a) stated that the holotype of Indocentor compactus tricuspis was deposited in the ZMB, but it was not included in the list of ZMB types by Moritz & Fischer (1981).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF14221FF23FC18FB85FA99.taxon	materials_examined	Type depository: ZMB: holotype (male) (Moritz & Fischer 1981) Type locality: Queensland, Australia Type host: Hystrix sp. [sic] Current status: a synonym of Bothriocroton concolor and its original name, as discussed in Klompen et al. (2002), who also established its current status. Comments: Neumann (1899) stated that the type specimen was labeled Ixodes coxalis.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF14221FF23FD84FC19FC61.taxon	materials_examined	Type depository: undetermined: syntypes (a male and females); BMNH: a syntype (female) (Keirans & Hillyard 2001) Type locality: Belem, Pará, Brazil Type host: armadillo (Neumann 1899) Current status: generally considered to be a synonym of Amblyomma auricularium, but Guglielmone et al. (2021) believe that this synonymy should be reexamined. Comments: the type host is named Dasypus sp. in Keirans & Hillyard (2001).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF04220FF23FD15FBD0FBB9.taxon	materials_examined	Type depositories: QM: holotype (male) and a paratype (female); ANIC: a paratype (female) and ARI?: paratypes (a male, females) Type locality: Sogeri, Papua New Guinea Type host: “ wallaby ” Comments: Barker et al. (2022) found no paratypes of Ixodes confusus in the ARI.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF04220FF23FBD0FA6AFAD1.taxon	materials_examined	Type depositories: VLM: holotype (male) and paratypes (a male and a female); BMNH, IMT?, MHNM?, MNHN?, OVI? and SAIMR?: paratypes (males and females) Type locality: Ile, Zambézia, Mozambique Type host: buffalo Current status: a synonym of Rhipicephalus longus, as stated in Keirans & Hillyard (2001). Comments: Walker et al. (2000) listed only the VLM and BMNH as type depositories of Rhipicephalus confusus.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF04220FF23F950FE6AF852.taxon	materials_examined	Type depository: USNTC: holotype (male) and paratypes (males and females) Type locality: Boende, province of Équater, Democratic Republic of the Congo Type host: Sus scrofa Comments. Apanaskevich et al. (2013) stated that some paratypes of Rhipicephalus congolensis would be deposited in the OVI and ZIAC.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF04220FF23FA48FBB1F931.taxon	materials_examined	Type depository: MNHN: syntypes (males and females) Type locality: Brazzaville, Congo Type host: cattle Current status: a synonym of Rhipicephalus annulatus (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF34223FF23FF78FC1FFE05.taxon	materials_examined	Type depository: not stated: syntypes (males), but probably in the IOC (Guglielmone et al. 2003) Type localities: Corrego Flor, State of Mato Grosso, and Xingu River, State of Amazonas, Brazil Type host: dog Current status: a synonym of Amblyomma pictum (Guglielmone et al. 2021).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF34223FF23FA6DFC08F949.taxon	materials_examined	Type depository: OVI: a lectotype (male) and paralectotypes (males, a female and nymphs) (Hoogstraal & Wassef 1981) Type locality: Onderstepoort, Pretoria, Province of Gauteng, South Africa Type host: Pedetes capensis Comments: Hoogstraal & Wassef (1981) designated a lectotype of Haemaphysalis cooleyi, but other specimens were named as paratypes, although they should be treated as paralectotypes.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF34223FF23F8E1FE4DF875.taxon	materials_examined	Type locality: Oruro, Bolivia Type host: Ctenomys sp.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF34223FF23FE34FE63FCBD.taxon	materials_examined	Type depository: USNTC: holotype (female) and a paratype (female) Type locality: Pan American Highway, 4.5 km north of Café La Georgina, Cerro de la Muerte, San José, Costa Rica Type host: Turdus sp.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF34223FF23FCADFD90FC45.taxon	materials_examined	Type depository: MCZ: syntype (female) (Cooley & Kohls 1945) Type locality: Salem, Massachusetts, USA	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF24222FF23FF78FE84FE5D.taxon	materials_examined	Type depository: BMNH: a lectotype (male) and paralectotypes (a male and females) were designated by Keirans & Brewster (1981) Type locality: Puerto Cooper, Paraguay Type host: Hydrochoerus hydrochaeris Current status: a synonym of Amblyomma dubitatum (Camicas et al. 1998, Estrada-Peña et al. 2002). Comments: Keirans & Brewster (1981), Keirans & Hillyard (2001) and other authors treated Amblyomma cooperi as a valid species.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF24222FF23FDCCFC44FD65.taxon	materials_examined	Type depositories: USNTC: holotype (female); NWC: paratypes (larvae) Type locality: Whitfield Hall, Penlyne, Saint Thomas Parish, Jamaica	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF24222FF23FBF5FBF6FAA5.taxon	materials_examined	Type depository: LMNH?: holotype (male) Type locality: Sekroe, New Guinea, Indonesia Type host: unknown Comments: Roberts (1970) stated that the holotype was sent from the LMNH to the BM and from there to Australia, with no indication that it was returned to the LMNH (Guglielmone et al. 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF24222FF23FAD4FD4AF9DD.taxon	materials_examined	Type depositories: MG and USNTC: syntypes? (females) (Schulze 1935 a, Keirans & Clifford 1984 as “ types ”) Type locality: Ambon, Indonesia Type host: Phalanger sp. Current status: a synonym of Ixodes cordifer, as listed in Camicas et al. (1998). Comments: Keirans & Clifford (1984) did not specify the developmental stage held in the USNTC, but Ixodes cordifer bibax was described from female ticks.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFFD422DFF23FD5CFE12FBB9.taxon	materials_examined	Type depositories: ENV: a lectotype (male) and a paralectotype (male) (Trapido 1965, who designated only the lectotype); BMNH: a paralectotype (male) (Keirans & Hillyard 2001). Type locality: Borneo Type host: deer Comments: Guglielmone et al. (2014) stated that the lectotype of Haemaphysalis cornigera is deposited in the MNHN but it is not. The alleged specimens of Haemaphysalis cornigera in the MNHN belong to another species, as explained in Trapido (1965).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFFD422DFF23FBD0FF0DFAD1.taxon	materials_examined	Type depository: IM: holotype (male) Type locality: Koderma Station, Chota Nagpur, India Type host: wolf Current status: a synonym of Haemaphysalis anomala and its original name (Camicas et al. 1998). The current status of Haemaphysalis cornigera anomala, as Haemaohysalis anomala, was established by Hoogstraal et al. (1967).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFFD422DFF23FA4EFF33F8EE.taxon	materials_examined	Type depositories: USNTC: holotype (male) and paratypes (a female and others unspecified developmental stages); BMNH?, VRC and ZSI: paratypes (males, females, nymphs and larvae) Type locality: Kyasanur Forest near Barage, Shimoga District, State of Karnataka, India Type host: the holotype specimen was collected from vegetation Current status: a synonym of Haemaphysalis shimoga and its original name (Camicas et al. 1998). The current status of Haemaphysalis cornigera shimoga, as Haemaphysalis shimoga, was established by Hoogstraal & El Kammah (1972). Comments: Keirans & Hillyard (2001) stated that paratypes allegedly delivered to the BMNH were not received there.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFFD422CFF23F89AFF0DFECD.taxon	materials_examined	Type depository: TIUC: syntypes (males and females named as “ types ”) Type localities: Horin, Naimshô and Taito, Taiwan Type hosts: buffalo and zebu Current status: a synonym of Haemaphysalis taiwana and its original name (Camicas et al. 1998). The current status of Haemaphysalis cornigera taiwana, as Haemaphysalis taiwana, was established by Hoogstraal & El Kammah (1972).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFFC422CFF23FE7CFD59FD01.taxon	materials_examined	Type depository: AI (presumed): holotype (male) and a paratype (female) Type locality: Cao Bang, Vietnam Type host: Bos taurus Current status: a synonym of Haemaphysalis shimoga (Camicas et al. 1998). Comments: the above information is provisional.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFFC422CFF23F94CFCB0F875.taxon	materials_examined	Type depositories: USNTC: holotype (male) and paratypes (males and females); BMNH, VCR and ZSI: paratypes (males and females) Type locality: Batangu, Kashmir, India Type host: sheep Comments: the holotype molted from a nymph fed on a sheep.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFFC422CFF23FD38FC63FC61.taxon	materials_examined	Type depository: MNHN: syntypes (females) Type localities: Chile and Ecuador Type host: “ caille ” Comments: Ixodes cornuae was originally named Ixodes auritulus.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFFC422CFF23FC18FD51FAFD.taxon	materials_examined	Type depositories: NM: holotype (male) and a paratype (female); ANIC: paratypes (a male and females), TM: paratypes (females) Type locality: Cox’s Bight, Tasmania, Australia Type host: unknown Comments: Kolonin (2009) doubted the validity of Ixodes cornuatus, while Camicas et al. (1998) treated Ixodes cornuatus Roberts, 1960 as a preoccupied name, although that decision was not accepted by Guglielmone et al. (2009), Guglielmone & Nava (2014) and others.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFFF422FFF23FE34FE3AFC28.taxon	materials_examined	Type depositories: OVI: holotype (female) and paratypes (males, females and nymphs); BMNH and USNTC: paratypes (males, females and nymphs) Type locality. Tsitsikama National Park, South Africa Type host: Aonyx capensis Comments: Keirans & Hillyard (2001) stated that the holotype is held in the USNTC, but Keirans et al. (1982) listed that specimen as deposited in the OVI, and Keirans & Clifford (1984) listed only the presence of paratypes in the USNTC. Larvae are included among paratypes of Ixodes corwini in Keirans et al. (1982) and Keirans & Hillyard (2001), but Keirans et al. (1982) stated that the larva of this tick is unknown; therefore, larvae are excluded as type specimens of Ixodes corwini.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFFF422FFF23FF78FAB9FE05.taxon	materials_examined	Type depository: ZIAC: lectotype (female) (Filippova 2008) Type locality: Peter the First Mountain Range, Tajikistan Type host: Mustela erminea Comments: Hornok et al. (2017) and others treated Ixodes cornutus as a synonym of Ixodes rugicollis, but this name is accepted as provisionally valid in Filippova (1997, 2008), Guglielmone et al. (2014, 2020, 2023) and here.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFFE422EFF23FEECFB46FE5D.taxon	materials_examined	Type depository. MK: syntypes (males and females) Type locality: Egypt Type host: Rhinolophus clivosus Current status: a name incertae sedis in Neumann (1911 a) and Guglielmone & Nava (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFFE422EFF23F974FDD2F8EE.taxon	materials_examined	Type depository: unknown: (syntypes) (male and female) (Guglielmone et al. 2014) Type locality: Germany (Koch 1847)	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF94229FF23FE81FB4AFD01.taxon	materials_examined	Type depository: IZAS: holotype (male) and paratypes (a male and females) Type locality: Pallas, Hashi District, Xinjiang, China Type host: Cricetulus migratorius Comments: the holotype molted from a nymph that had fed on Cricetulus migratorius. Deng et al. (1999) treated Anomalohimalaya cricetuli as a synonym of Anomalohimalaya lotozky, and Chen et al. (2010) did not include this species as a Chinese tick, but it is regarded as valid in Guglielmone et al. (2014, 2020, 2023).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF84228FF23FAD5FE3FF931.taxon	materials_examined	Type depositories: BMNH: lectotype (male) and paralectotypes (males, a female and nymphs) (Keirans & Hillyard 2001); ENV and ZMB: paralectotype (males) (Nuttall & Warburton 1915, Moritz & Fischer 1981) Type locality: Colombo, Sri Lanka Type host: Tragulus meminna Comments: larvae were included among the type specimens in Warburton (1910) and Nuttall & Warburton (1915), and among paralectotypes listed in Keirans & Hillyard (2001), but the larva was not described in Warburton (1910). Nuttall & Warburton (1915) and Moritz & Fischer (1981) named the specimens of Haemaphysalis cuspidata “ type ” and “ paratype, ” respectively.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFF84228FF23FE58FCCFFD65.taxon	materials_examined	Type depository: CGV: syntypes (males) Type locality: Ngômo, Gabon Type host: cattle Current status: Walker et al. (2000) listed Rhipicephalus cuneatus as a probable synonym of Rhipicephalus ziemanni, but Camicas et al. (1998) and Guglielmone & Nava (2014) treated Rhipicephalus cuneatus as a synonym of Rhipicephalus aurantiacus, a position maintained here.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFFB422BFF23FD84FB94FCF5.taxon	materials_examined	Type depository: MNHN: syntypes (males and females) Type locality: Mariana Islands, Pacific Ocean Type host: unknown Current status: a synonym of Amblyomma breviscutatum (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFFB422BFF23FC64FF43FB41.taxon	materials_examined	Type depository: ZMB: holotype (male) and a paratype (female) Type locality: Momanga, north Mindanao, Philippines Type host: unknown Current status: a synonym of Amblyomma breviscutatum (Guglielmone & Nava 2014). d	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFFB422BFF23FAF8FC34F9F9.taxon	materials_examined	Type depository: BMNH: holotype (female) (Keirans & Hillyard 2001) Type locality: Groot Krei, Kei Islands, Indonesia Type host: unknown Current status: Keirans & Hillyard (2001) listed this name as a synonym of Amblyomma cyprium, but it is a synonym of Amblyomma breviscutatum (Santos Dias 1958 a, Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFFB422BFF23F990FA75F83D.taxon	materials_examined	Type depository: USNTC: holotype (female) and paratypes (males, females, nymphs and larvae) Type locality: Nantucket Island, Massachusetts, USA Type host: the holotype was collected from vegetation Current status: a synonym of Ixodes scapularis, as listed in Camicas et al. (1998) and Keirans & Robbins (1999).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFFA422AFF23FF79FB5CFE21.taxon	materials_examined	Type depositories: USNTC: holotype (female) and paratypes (females); MCZ?: a paratype (female) Type locality: Chalco, Ciudad de México, Mexico Type host: Geomys sp. Comments: KLC, IR and RGR visited the MCZ in 2024 and the paratype was not found there.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF854255FF23FF79FD2DFE05.taxon	materials_examined	Type depository: BMNH: syntypes (males and females) (Keirans & Hillyard 2001) Type locality: Fagus Cove, Isabela Island, Galápagos Islands, Ecuador Type host: Amblyrhynchus cristatus Comments: Guglielmone et al. (2014) listed paratypes deposited in the MCZ, but KLC, IR and RGR visited the MCZ in 2024 and paratypes were not found there.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF854255FF23FE34FECCFD59.taxon	materials_examined	Type depository: ZSH: syntypes (males and females) Type locality: Galápagos Islands, Ecuador Type host: Amblyrhynchus sp. Current status: a provisional synonym of Amblyomma darwini in Guglielmone & Nava (2014) and Guglielmone et al. (2021).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF844254FF23FD60FB1FFC61.taxon	materials_examined	Type depository: USNTC?: syntypes? (males, females and a nymph) (Apanaskevich et al. 2008 b) Type locality: Varamin and Tehran, Iran Type host: cattle, camel, sheep Current status: a synonym of Hyalomma dromedarii, as discussed in Apanaskevich et al. (2008 b). Comments: Apanaskevich et al. (2008 b) found specimens of Hyalomma delpyi in the USNTC whose information is mostly consistent with data from the original description and are probably part of the syntype series.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF874257FF23FA49FD2DF8D9.taxon	materials_examined	Type depositories: BMNH: lectotype (male) and a paralectotype (male); LSTMH, USNTC: paralectotypes (males) (Hoogstraal & Trapido 1966 a, who designated the lectotype, Keirans & Hillyard 2001, Apanaskevich 2023) Type locality: Malaysia Type host: Paradoxorus sp. Current status: Haemaphysalis dentipalpis was treated as a synonym of Haemaphysalis asiatica until this species was reinstated by Apanaskevich (2023). Comments: no type specimens of Haemaphysalis dentipalpis are recognized by Keirans & Clifford (1984) as deposited in the USNTC. However, Keirans & Hillyard (2001) stated that a male paralectotype was sent to the USNTC, and Apanaskevich (2023) found that male deposited there. Nuttall & Warburton (1909) listed the type host of Haemaphysalis dentipalpis as Paradoxorus sp?	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF874257FF23FF79FC53FE5D.taxon	materials_examined	Type depositories: AMNH?: holotype (female) and paratypes (male and females); BM: paratypes (a male and females) Type locality: Mount Rawlinson, Huon Peninsula, Morobe District, Papua New Guinea Type host: Dendrolagus matschiei Comments: Guglielmone et al. (2014) stated that no paratypes of Ixodes dendrolagi were found in the ANIC, as subsequently confirmed by B. Halliday (personal communication to R. G. Robbins). KLC visited the AMNH in 2024 and only one paratype (labeled as allotype male) was found there.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF874257FF23FCADFE0FFC45.taxon	materials_examined	Type depository: USNTC: holotype (female) (Keirans & Clifford 1984) Type locality: Maryland, USA	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF864256FF23FF78FC1DFE05.taxon	materials_examined	Type depository: ZMB: holotype (male) (Moritz & Fischer 1981) Type locality: Banat (Koch 1844). Moritz & Fischer (1981) named Hungary as the type locality, while Estrada-Peña (1990) treated Banat as Romanian Type host: not stated Current status: a synonym of Dermacentor marginatus (Estrada-Peña 1990).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF864256FF23FE34FE1FFCBD.taxon	materials_examined	Type depository: ZMB?: syntypes? (a male and a female) Type locality: Canary Islands, Spain Type host: not stated Current status: a synonym of Hyalomma anatolicum (Camicas et al. 1998). Comments: Moritz & Fischer (1981) stated that the type specimens of Hyalomma depressum, supposedly held in the ZMB, were not found there.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF864256FF23FA24FBE3F915.taxon	materials_examined	Type depository: not stated: syntypes (males) Type locality: Damascus, Syria Type host: not stated Current status: a synonym of Hyalomma scupense (Apanaskevich et al. 2010)	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF864256FF23FCACFC78FBF1.taxon	materials_examined	Type depository: ZMB: holotype (male) and paratypes (a male and a female) in Moritz & Fischer (1981), who name the paratypes as syntypes Type locality: “ Golodnaja-Steppe, Transkaspien ” (Kazakhstan?) Type host: not stated Current status: a synonym of Hyalomma scupense (Filippova 2003).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF864256FF23FB68FBC0FA35.taxon	materials_examined	Type depository: ZMB: syntypes (males and females) (Moritz & Fischer 1981) Type locality: Tsingtau, China Type host: cattle Current status: a synonym of Hyalomma scupense (Apanaskevich et al. 2010). Comments: Apanaskevich et al. (2010) found only a female syntype in the ZMB.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF864251FF23F904FA69FF51.taxon	materials_examined	Type depository: not stated in the original description: syntypes (males and females); USNTC?: syntypes? (males) (Apanaskevich et al. 2010) Type locality: Prilep, North Macedonia Type host: cattle Current status: a synonym of Hyalomma scupense (Apanaskevich et al. 2010). Comments: Apanaskevich et al. (2010) found two males in the Schulze Collection, which is at the USNTC, whose locality data match that of Hyalomma detritum dardanicum and that may be part of a syntype series of this taxon.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF814251FF23FC88FE61FB41.taxon	materials_examined	Type depositories: ZIAC?: holotype (male), ZMB?: paratypes (a male and a female) (Schulze 1930 a) Type locality: “ Arycj, ” Kazakhstan Type host: camel Current status: a synonym of Hyalomma scupense (Apanaskevich et al. 2010). Comments: Moritz & Fischer (1981) and Filippova (2008) did not list type specimens of Hyalomma detritum rubrum as deposited in the ZMB and ZIAC, respectively, although Schulze (1930 a) stated that type specimens had been deposited there.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF814251FF23FEC8FB8EFDB1.taxon	materials_examined	Type depository: not stated: syntypes (males) Type locality: Caucasus Type host: not stated Current status: a synonym of Hyalomma excavatum, as listed in Camicas et al. (1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF814251FF23FDA8FBEAFC91.taxon	materials_examined	Type locality: Beijing, China Type host: not stated Current status: a synonym of Hyalomma scupense (Apanaskevich et al. 2010).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF814251FF23F9D9FB02F8AD.taxon	materials_examined	Type depository: USNTC: holotype (male) and paratypes (males and females) Type locality: 6 km south of Empalme, Cordillera de Talamanca, Province of San José, Costa Rica	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF804250FF23FEC8FA61FDC9.taxon	materials_examined	Type depositories: ZMB?: holotype (male) (Schulze 1950); USNTC: paratypes (unspecified developmental stages), listed as “ types ” in Keirans & Clifford (1984) Type locality: specimens collected on an Australian host in Berlin Aquarium (Germany) Type host: Diemeniae textilis Current status: a synonym of Amblyomma argentinae in Guglielmone et al. (2021). Comments: Moritz & Fischer (1981) did not list specimens of Amblyomma diemeniae as being held in the ZMB.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF804250FF23FB21FBD8FA69.taxon	materials_examined	Type depository: ZMB: syntypes (males and females) (Moritz & Fischer 1981) Type locality: Mexico Type host: not stated Comments: Camicas et al. (1998) did not recognize Amblyomma dissimile as valid.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF804250FF23FC41FE57FB09.taxon	materials_examined	Type locality: El Paso, Peten, Guatemala Type host: Tayassu pecari ringens Comments: no type specimens of Dermacentor dispar are listed by Keirans & Clifford (1984) as having been deposited in the USNTC.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF804250FF23FA01FD46F8D9.taxon	materials_examined	Type depositories: AMNH?: holotype (female) and paratypes (females and a nymph); USNPC?: a paratype (female), USNTC: paratypes (females and a nymph) (Cooley 1947), MCZ: paratype (a female) Type locality: Las Margaritas, 45 km southeast of Comitán, Chiapas, Mexico Type host: horse Comments: Keirans & Clifford (1984) listed the holotype as deposited in the USNTC, but Hernández et al. (2011) confirmed that the holotype is held in the AMNH. Salley et al. (1978) did not include type specimens deposited in the USNPC. KLC visited the AMNH in 2024 and the paratype nymph was not found there; however, an unreported paratype (labeled as allotype male) was found.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF804250FF23FD61FA83FC29.taxon	materials_examined	Type depository: SAIMR: holotype (female) Type locality: Nightingale Island, Tristan da Cunha, Atlantic Ocean Type host: Diomedea chlororhynchus Comments: the holotype of Ixodes diomedeae was originally identified as Ixodes rothschildi in Zumpt (1952).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF834253FF23FBF5FC2FFA99.taxon	materials_examined	Type depository: USNPC: a syntype (female) in Salley et al. (1978), who designated the specimen as “ type ” Type locality: New Mexico, USA Type host: Procyon lotor Comments: Ixodes diversifossus was described from two females, but only one is preserved in the USNPC (Keirans & Hillyard 1978). Kolonin (2009) treated Ixodes diversifossus as invalid.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF834253FF23FDA9FD67FD59.taxon	description	See Rhipicephalus punctatus Bedford, 1929	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF834252FF23F895FD39FECD.taxon	materials_examined	Type depository: originally deposited in the IEMPVT, but currently held in USNTC: holotype (female) (Keirans & Clifford 1984) Type locality: La Mandraka, Madagascar Type host: Newtonia brunneicauda Comments: Keirans & Clifford (1984) did not specify the stage of the holotype held in the USNTC, but Ixodes domerguei was described from the holotype female.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF824252FF23FE7CFBE5FDC9.taxon	materials_examined	Type locality: Périnet, Moramanga, Madagascar Type host: Nelicurvius nelicourvi Current status: a synonym of Ixodes domerguei (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF824252FF23FD60FBBDFC29.taxon	materials_examined	Type depositories: MHNM: holotype (female); BMNH: a paratype (female) Type locality: Mutuali, Province of Nampula, Mozambique Type host: Sylvicapra grimmia Current status: a synonym of Ixodes neitzi, as discussed in Guglielmone et al. (2009).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF824252FF23FC40FE5BFAD1.taxon	materials_examined	Type depositories: TC: syntypes (a female? nymphs? a larva?), BMNH (nymphs and larvae) (Keirans & Hillyard 2001) Type locality: Scilly Islands, Great Britain Type host: Crocidura cassiteridium Current status: a synonym of Ixodes acuminatus (Keirans & Hillyard 2001). Comments: the developmental stages of specimens deposited in the TC are uncertain. Turk (1948) described this tick from a female, 14 nymphs and a larva collected from six Crocidura cassiteridium, but Keirans & Hillyard (2001) stated that Ixodes dorriensmithi was described from six specimens collected from Crocidura crassiteridium, which contradicts Turk’s (1948) statement above. Additionally, Keirans & Hillyard (2001) listed three larvae as deposited in the BMNH versus the single larva in the original description. Thus, the type data for Ixodes dorriensmithi should be considered provisional.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF8D425DFF23F9B4FBF0F8E5.taxon	materials_examined	Type depository: not stated: syntypes (males) Type locality: Krong-Pha, Vietnam Type host: Bos sondaicus Current status: a synonym of Hyalomma isaaci (Apanaskevich & Horak 2010).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF8D425DFF23FC3CFD2BFAA4.taxon	materials_examined	Type depository: ZMB: syntype (a male) (Moritz & Fischer 1981) Type locality: Bukhara, Uzbekistan Type host: not stated Current status: a synonym of Hyalommma asiaticum and its original name, as discussed in Apanaskevich & Horak (2010), who stated that this tick was described from an unknown number of males, but Schulze & Schlottke (1929) also described the female of Hyalomma asiaticum.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF8D425DFF23FAD4FBD0F985.taxon	materials_examined	Type depository: not stated: syntypes (males and females) Type locality: Canary Islands, Spain Type host: camel Current status: a synonym of Hyalomma dromedarii (Apanaskevich et al. 2008 b).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF8D425DFF23FE81FB92FD95.taxon	materials_examined	Type depositories: OVI: holotype (female) and paratypes (males, females, nymphs and larvae); BMNH and USNTC: paratypes (males, females, nymphs and larvae) Type locality: Giant’s Castle Natural Reserve, Province of KwaZulu-Natal, South Africa	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF8C425CFF23FEA5FAF2FD95.taxon	materials_examined	Type depository: ENV: holotype (female) Type locality: southern Spain? Type host: unknown Comments: the type locality is incorrect because this species is Neotropical (Estrada-Peña et al. 2002).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF8C425CFF23FD84FBB1FCF5.taxon	materials_examined	Type depository: not stated: syntypes (males, females, nymphs and larvae) Type locality: Algeria and Morocco Type host: sheep and cattle Current status: a synonym of Rhipicephalus annulatus (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF8C425CFF23FC65FA92FAD1.taxon	materials_examined	Type depository: BMNH?: holotype (male) Type locality: “ Zambie ” but changed to “ Zambu (Congo indépendant) ” in Neumann (1911 a), while Keirans & Hillyard (2001) named the locality as “ Zambi ” Type host: bovine Comments: Keirans & Hillyard (2001) and Walker et al. (2000) stated that the type depository for this tick was not identified. However, Keirans & Hillyard (2001) listed a male of Rhipicephalus duttoni held in the BMNH as a syntype, although Guglielmone et al. (2014) doubt that this specimen was used to describe Rhipicephalus duttoni. Nevertheless, the data provided by Keirans & Hillyard (2001) agree with those presented by Neumann, who described this species from a single male. Therefore, if the male held in the BMNH is the original specimen used to describe Rhipicephalus duttoni, it should be treated as the holotype, not a syntype of this species. Walker et al. (2000) stated that the type locality is in the Democratic Republic of the Congo, a statement not confirmed here.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF8C425CFF23FA49FD1BF9DD.taxon	materials_examined	Type depository: not stated in Dönitz (1910): syntypes (a male and a female); ZMB: a syntype (male) (Moritz & Fischer 1981) Type locality: Ituri, Democratic Republic of the Congo	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF8F425FFF23FE59FC92FD59.taxon	materials_examined	Type depository: ZMB: syntypes (males) (Moritz & Fischer, 1981) Type locality: Arusha, Tanzania Type host: Varanus saurus Comments: morphological descriptions of Amblyomma eburneum differ among authors (Guglielmone et al. 2020), and further studies would help to clarify the status of this species.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF8F425FFF23F94DFD5CF8D9.taxon	materials_examined	Type depository: BMNH: lectotype (female) and paralectotypes (nymphs) were designated by Keirans & Brewster (1981) Type locality: Umboi Island, Papua New Guinea	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF8E425EFF23FB20FC19FA4D.taxon	materials_examined	Type depository: not stated: holotype? (male) Type localities: Egypt and Senegal Type host: not stated Current status: a synonym of Amblyomma variegatum (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF8E425EFF23F9FCFF0DF8E5.taxon	materials_examined	Type depository: MNHN: syntypes (females) Type locality: Chile Type host: Pudua humilis Current status: Ixodes elegans (preoccupied) is a synonym of Ixodes stilesi and its original name. The current status of Ixodes elegans, as Ixodes stilesi, was provided by Neumann (1911 b), as discussed in Guglielmone & Nava (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF8E425EFF23FD84FDD9FCF5.taxon	description	Dzhaparidze, N. I. (1950) New species of ticks of the family Ixodidae in Georgia. Soobshcheniya Akademy Nauk Gruzinskoy SSSR, 11, 117 – 121. [in Russian] Type depository: IZG: holotype (female) (Filippova 1977) Type locality: Shiraki Plains, Georgia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF894259FF23F928FDC0F83D.taxon	materials_examined	Type depository: GM: syntypes (a male and females) (Hoogstraal 1955) Type locality: Gebel Resas, Tunisia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF894259FF23FC89FB5BFBF1.taxon	materials_examined	Type depositories: not stated in Neumann (1897): syntypes (males and females); BMNH and ENV: syntypes (males and females) (Nuttall & Warburton 1915, Keirans & Hillyard 2001) Type locality: Madagascar Type hosts: Centetes madagascariensis, Centetes ecaudatus, Erinaceus sp. in Neumann (1897)	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF884258FF23FBF4FA95FA4D.taxon	materials_examined	Type depositories: USNTC?: holotype (male) and paratype (unspecified developmental stage); VLM? holotype (male) and paratypes (males, females and nymphs); BMNH?, CZIICT, IMT, MNHN and SAIMR: paratypes (males, females and nymphs) (Santos Dias 1958 b, Keirans & Clifford 1984, Keirans & Hillyard 2001) Type locality: Sábiè, Mozambique Type host: Otolemur crassicaudatus Current status: a synonym of Haemaphysalis spinulosa in Apanaskevich et al. (2007) but not confirmed in Tomlinson et al. (2018). Comments: Keirans & Clifford (1984) listed the holotype and a paratype as held in the USNTC, but Santos Dias (1958 b) stated that the holotype was deposited in the VLM and he did not list any paratype as having been sent to the USNTC. Keirans & Hillyard (2001) did not find paratypes of Haemaphysalis ethiopica held in the BMNH.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF884258FF23F9FDFD0BF949.taxon	materials_examined	Type depositories: ANIC: a paralectotype (life stage undetermined; specimen is in transit); CMC: a lectotype (female) and paralectotypes (females and nymphs) were designated by Heath & Palma (2017) Type locality: Dusky Sound, Fiordland, New Zealand	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF8B425BFF23F9FCFBACF8AD.taxon	materials_examined	Type depository: ZMB: syntypes (males) (Moritz & Fischer 1981) Type locality: Namibia Type host: cattle Current status: a synonym of Rhipicephalus evertsi in Guglielmone & Nava (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF8B425AFF23F8DCFF04FEB5.taxon	materials_examined	Type depository: ZMB?: holotype (male) Type locality: Egypt Type host: not stated Comments: Feldman-Muhsam (1954) redescribed the holotype of Hyalomma excavatum, but this specimen, which is supposed to be in the ZMB, to date has not been found there (Moritz & Fischer 1981, Apanaskevich & Horak 2005).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF8A425AFF23FC3CFE9BFBD5.taxon	materials_examined	Type depository: ZMB: syntypes (males and a female) (Moritz & Fischer 1981) Type locality: Africa	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF8A425AFF23F9B4FDF6F8D9.taxon	materials_examined	Type depository: SDC: holotype: (male) Type locality: Dambacua, Inhambane District, Mozambique Type host: Syncerus caffer caffer Current status: a synonym of Amblyomma sparsum, as discussed in Theiler & Salisbury (1959), who named this tick Amblyomma faifai (a lapsus).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF8A425AFF23FEA5FD0CFD95.taxon	materials_examined	Type depositories: OVI: holotype (male) and paratypes (males, females, nymphs and larvae), BMNH and USNTC: paratypes (males, females, nymphs and larvae) Type locality: farm Kosos 11, Karas region, Namibia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF954245FF23F9FDFD0BF915.taxon	materials_examined	Type depository: BMNH: holotype (female) (Keirans & Hillyard 2001) Type locality: Cranbook, Western Australia, Australia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF944244FF23FEC8FC1FFDB1.taxon	materials_examined	Type depository: ZMB: syntypes (females) (Moritz & Fischer 1981) Type locality: unknown Type host: not stated Current status: a synonym of Dermacentor reticulatus (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF944244FF23FDA9FE28FD65.taxon	materials_examined	Type depository: MRSN: holotype (female) (Gilot & Pérez 1978) Type locality: Derna, Libya	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF944244FF23F929FC46F8D9.taxon	materials_examined	Type depositories: USNTC: holotype (male) and a paratype (female); ZMMO: paratypes (males and a female) Type locality: Buon Luoi, An Khe District, Gia Lai Provnce, Vietnam	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF974247FF23FCADFAD7FBD5.taxon	materials_examined	Type depository: ZMB: holotype (male) (Moritz & Fischer 1981) Type locality: Manila, the Philippines Type host: not stated Comments: Voltzit & Keirans (2002) treated Amblyomma fimbriatum as a synonym of Amblyomma helvolum, but this synonymy was not accepted in Guglielmone et al. (2020). Kaufman (1972) stated that the holotype is deposited in the USNTC, but this species is not listed in the catalog of USNTC types by Keirans & Clifford (1984).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF974247FF23FE59FBA9FCBD.taxon	materials_examined	Type depository: not stated: syntypes? (nymphs and larvae) Type locality: uncertain Type host: marmots Current status: Filippova (1977) and Kolonin (2009) treated Ixodes filippovae as invalid, while the species described in Černý (1961) was considered questionable and provisional in Guglielmone et al. (2014, 2020). Comments: Černý (1961) treated the larva and nymph of Ixodes crenulatus from marmots in Filippova (1958) as a new species that he named Ixodes filippovae, without designation of type specimens.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF974246FF23F905FB64FF7D.taxon	materials_examined	Type depository: MNHN: syntypes (males, females, nymphs and larvae) Type localities: Tokyo, Saga and Sendai, Japan Type hosts: dog, horse, cow and hare Comments: Nuttall & Warburton (1915) stated that specimens used to describe Haemaphysalis flava were deposited in the ENV, but Neumann (1897) listed them as held in the MNHN. Yamaguti et al. (1971) stated that specimens of Haemaphysalis campanulata were included among the syntypes of Haemaphysalis flava.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF914241FF23FCACFF0DFBF1.taxon	materials_examined	Type depository: not stated: holotype (female) Type locality: Retalhuleu, Nicaragua Type host: unknown Current status: a name incertae sedis (Guglielmone & Nava 2014). Comments: the type locality for Amblyomma foreli is erroneously treated as unknown in Guglielmone & Nava (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF914241FF23FB68FCDDF985.taxon	materials_examined	Type depositories: ZMB?: holotype (female) (Schulze 1933 b, Moritz & Fischer 1981); USNTC?: a “ type ” of unspecified developmental stage (Keirans & Clifford 1984) Type locality: Kankau, Taiwan (Moritz & Fischer 1981) and “ Kankan, ” Taiwan, in Keirans & Clifford (1984) Type host: Cyclemis sp. Current status: a probable synonym of Amblyomma geoemydae in Keirans & Clifford (1984). Comments: Schulze (1933 b) named as Amblyomma formosanum a female collected on Cyclemis sp. and figured in Robinson (1926) as Amblyomma cyprium. The holotype of Amblyomma formosanum is probably the specimen listed in Moritz & Fischer (1981), while Keirans & Clifford (1984) may have received another specimen from the same host but not the holotype of Amblyomma formosanum.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF914241FF23F9B5FF3EF83D.taxon	materials_examined	Type depositories: BMNH, ENV and ZMB?: syntypes (males and females) (Nuttall & Warburton 1915, Keirans & Hillyard 2001) Type locality: Kosempo, Taiwan Type host: dog Comments: Moritz & Fischer (1981) did not list specimens of Haemaphysalis formosensis in their catalog of ZMB types.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF914241FF23FE59FEC9FCBD.taxon	materials_examined	Type depository: BMNH: syntypes (males) (Keirans & Hillyard 2001) Type locality: Namibia? South Africa? (Keirans 1985, Walker et al. 2000, Keirans & Hillyard 2001) Type host: probably cattle Comments: Keirans (1985) listed Namibia as the type locality. Keirans & Hillyard (2001) also listed Namibia as the type locality, but they appear unconvinced about its presence there, stating that Rhipicephalus follis is “ known only from the Republic of South Africa and Swaziland. ” Walker et al. (2000, page 185) believe that the type locality is South Africa.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF934243FF23FB1DFCE4F985.taxon	materials_examined	Type depositories: NIAH: holotype (male) and paratypes (males, females, nymphs and larvae) (Kitaoka 1970); USNTC: a paratype (developmental stage not specified) (Keirans & Clifford 1984) Type locality: Shizuoka Prefectural Cattle Breeding Experiment Station, near Fujinomiya, Shizuoka, Japan Type host: Holstein calf Comments: the holotype specimen was obtained from laboratory material derived from an engorged female of Haemaphysalis fujisana collected from a Holstein calf.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF934243FF23FE58FC25FD65.taxon	materials_examined	Type depository: undetermined: holotype? (female) (Guglielmone et al. 2003) Type locality: Germany Type host: Paro maiori Current status: a synonym of Ixodes frontalis and its original name (Guglielmone & Nava 2014). The current status of Acarus frontalis, as Ixodes frontalis, was established by Koch (1844).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF934243FF23F9B5FD81F881.taxon	materials_examined	Type depositories: CCNMNS: holotype (female); CMT and / or CTY: paratypes (females) Type locality: Inukami-gun, Shiga, Japan	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF924242FF23F9FDFD63F915.taxon	materials_examined	Type depository: ZMB: holotype (female) (Labruna et al. 2020) Type locality: “ Casapav ” (Casapava), Brazil	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF9D424DFF23FC18FC61FB41.taxon	materials_examined	Type depository: ZMB: syntypes (a male and females) (Moritz & Fischer 1981) Type locality: Germany Type host: unknown Current status: a synonym of Ixodes ricinus (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF9D424DFF23FD39FD15FC61.taxon	materials_examined	Type depository: LMNH: syntypes (males) Type locality: America? (Neumann 1911 a) Type host: Boa constrictor? Comments: Horak et al. (2002) do not list Amblyomma fuscum as valid. Amblyomma fuscum is a Neotropical species found only in Brazil (Guglielmone et al. 2021, 2023).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF9D424DFF23FE34FAC6FD01.taxon	materials_examined	Type depository: MNHN: syntypes (females) Type locality: South America Type host: Boa constrictor Current status: a synonym of Amblyomma rotundatum (Camicas et al. 1998, Guglielmone et al. 2021).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF9C424CFF23FB68FD16F9F9.taxon	materials_examined	Type depository: MNHN: syntypes (males) (Guglielmone et al. 2003) Type localities: Pará, Brazil, and Darién, Colombia in Neumann (1899), but “ Darién ” is probably currently located in Panama rather than in Colombia Type host: not stated Comments: Santos Dias (1961), Camicas et al. (1998) and others treated Amblyomma geayi as a synonym of Ixodes perpunctatus. It is a nomen dubium in Fairchild et al. (1966) and Guglielmone et al. (2014), but type comparisons may help settle the status of this name (Guglielmone et al. 2021). Guglielmone et al. (2003) stated that a syntype was found in the MCZ, but in fact the specimen in the MCZ corresponds to Ixodes perpunctatus with an added label “ appears to be = Amblyomma geayi ” (http: // mczbase. mcz. harvard. edu / accessed 22 May 2022). This was confirmed during KLC, IR and RGR‘s visit to the MCZ in 2024.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF9C424CFF23F990FB72F83D.taxon	materials_examined	Type depository: ITS: holotype (male) and a paratype (female) Type locality: Minankro, Ivory Coast Type host: cow Current status: a synonym of Rhipicephalus geigyi and its original name (Guglielmone & Nava 2014). The current status of Boophilus geigyi, as Rhipicephalus geigyi, was established by Horak et al. (2002).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF9C424CFF23FE59FBEFFD2D.taxon	materials_examined	Type depositories: VRC: holotype (male) and paratypes (males and females); BMNH, IM and USNTC: paratypes (males and females) Type locality: near Badrinath, Channoli District, Garhwal, Uttar Pradesh, India	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF9F424FFF23FC65FE5CFB6D.taxon	materials_examined	Type depositories: USNTC: holotype (male) and paratypes (males and females); BMNH: paratypes (males and females) Type locality: Andrafiafaly, Ambatomainty-Andranolava Region, Mahajanga Province, Madagascar Type host: Geochelone yniphora Comments: it is probable that Amblyomma geochelone is a synonym of Amblyomma chabaudi (Kushimo 2013, Guglielmone et al. 2020).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF9F424EFF23F8DDFB11FF19.taxon	materials_examined	Type depositories: FMC: holotype (male) and a paratype (female); OVI: paratypes (unspecified developmental stages), ZMB: a paratype (female) (Walker et al. 2000) Type locality: Namaqualand (this territory is currently partly in Namibia and partly in South Africa)	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF9E424EFF23FBF4FE8BFAA5.taxon	materials_examined	Type depository: IM: syntypes (males and females) Type locality: Zoological Garden, Kolkata, State of West Bengal, India Type hosts: various reptiles Current status: a synonym of Amblyomma varanense (Kaufman 1972, Camicas et al. 1998) (under the former genus Aponomma).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF9E424EFF23FAD4FEFAF9DD.taxon	materials_examined	Type depository: ZMM: holotype (male) Type locality: Balochistan (Afghanistan?, Iran?, Pakistan?) Type host: a reptile Current status: a synonym of Amblyomma gervaisi (Kaufman 1972, Camicas et al. 1998), under the former genus Aponomma.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF994249FF23FF79FBA5FEE9.taxon	materials_examined	Type depository: ZIAC: holotype (nymph) and paratypes (nymphs) Type locality: River Akhty-Chai valley, Samur Mountain Range, Daghestan, Russia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF984248FF23FE7DFBDEFD2D.taxon	materials_examined	Type depositories: WAM: holotype (female) and paratypes (males, females, nymphs and larvae); BMNH, MAGNT and USNTC: paratypes (males, females, nymphs and larvae) Type locality: Lachlan Island, Buccaneer Archipelago, Western Australia, Australia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF9B424BFF23FE35FD3FFCF5.taxon	materials_examined	Type depositories: BMNH and ENV: syntypes (males) (Keirans & Hillyard 2001, Guglielmone et al. 2014) Type locality: Pará, Brazil Type host: Myremcophaga tetradactyla Comments: Neumann (1899) described Amblyomma goeldii from seven males and five females, but the alleged females of Amblyomma goeldii were found to be, in fact, females of Amblyomma rotundatum (Keirans 1985, Guglielmone et al. 2021). Therefore, records of females from Bufo agua, one female collected in Demerara (Guyana) and a female allegedly deposited in the ZMB but not found there by Moritz & Fischer (1981), are not treated here as part of the syntype series of Amblyomma goeldii.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF9B424BFF23FC65FB43FBF1.taxon	materials_examined	Type depository: USNTC: holotype (female) and paratypes (females) Type locality: 12 miles east of Edie Creek Road, Bulldog Road, Morobe, Papua New Guinea	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF9B424BFF23FA01FC4FF931.taxon	materials_examined	Type depositories: BMNH and GM: syntypes (females) (Anastos 1950, Keirans & Hillyard 2001) Type localities: Bhamo, Mooleyis, Tarrawaddy and Terinzo, Myanmar	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF9A424AFF23FD5CFDA5FC61.taxon	materials_examined	Type locality: Guernsey, Channel Islands, Great Britain Type host: Arvicola terrestris Current status: a synonym of Ixodes acuminatus (Camicas et al. 1998, Keirans & Hillyard 2001). Comments: Arthur (1955) listed Evotomys glareolus as the type host of Ixodes guernseyensis, but Keirans & Hillyard (2001) found that host to be incorrect.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF9A424AFF23FC19FB8DFB41.taxon	materials_examined	Type depository: USNTC: holotype (female) and paratypes (a female and nymphs) Type locality: Puerto Williams, Isla Navarino, Magallanes y la Antártica Chilena, Chile	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA54275FF23FD5DFC6BFC0D.taxon	materials_examined	Type depositories: USNTC: holotype (male) and paratypes (males, females, nymphs); CFCV, CNCDGS and INTA: paratypes (males, females and nymphs), CDPV: paratypes (larvae) Type locality: Parque Nacional El Rey, Province of Salta, Argentina	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA54275FF23F990FCFBF875.taxon	materials_examined	Type depository: GM: holotype (female) (Walker et al. 2000) Type locality: Yado, Myanmar Type host: not stated Current status: a synonym of Rhipicephalus haemaphysaloides (Camicas et al. 1998) and its original name (Guglielmone & Nava 2014). The current status of Rhipicephalus haemaphysaloides niger, as Rhipicephalus haemaphysaloides, was established by Neumann (1897). See also Rhipicephalus haemaphysaloides for a discussion of the confusion attending the type specimens of this tick.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA44274FF23FE35FB81FD59.taxon	materials_examined	Type depository: IZAS: holotype (male) and a paratype (female) Type locality: Hainan Island, Province of Hainan, China Type host: snake Comments: Kolonin (2009) and others treat Amblyomma hainanense as a synonym of Amblyomma helvolum, but this species is considered provisionally valid in Guglielmone et al. (2014, 2020, 2023).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA44274FF23FA49FDA6F9F9.taxon	materials_examined	Type depositories: USNPC: holotype (male) and paratypes (males and females); FMNH: paratype (an adult); USNTC: paratypes (unspecified developmental stages) (Salley et al. 1978, Keirans & Clifford 1984) Type locality: Kingsville, Texas, USA	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA44274FF23FBD0FC57FAD1.taxon	materials_examined	Type depository: BMNH?: syntypes (males and females) (Tendeiro 1950, as “ types ”) Type locality: Granja Pecuária Experimental de Pessuba, Guinea-Bissau Type hosts: Varanus niloticus and Varanus exanthematicus Current status: a synonym of Amblyomma flavomaculatum (Santos Dias 1993). Comments: Tendeiro (1950) stated that “ types ” would be sent to the BMNH, but in the catalog of BMNH types by Keirans & Hillyard (2001) no specimens of Aponomma halli are listed.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA44274FF23FF78FD18FE05.taxon	materials_examined	Type depository: GM: syntypes (males and females) (Walker et al. 2000) Type locality: Mount Mooleyit, Meteleó, Thagatá, Cagó del Cadú Gianng and north east of Bomó, Myanmar Type host: not stated Current status: a synonym of Rhipicephalus haemaphysaloides (Walker et al. 2000). Comments: see also Rhipicephalus haemaphysaloides.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA74277FF23F9B5FDD2F949.taxon	materials_examined	Type depository: FMNH: holotype (female) Type locality: Mount Mocco, Angola	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA74277FF23FB1DFB76F985.taxon	materials_examined	Type depository: USNTC: a lectotype (male) and paralectotypes (males and a female) were designated by Hoogstraal & Kohls (1968) Type locality: Popa, Myanmar Type host: Melogale personatus personatus Comments: Hoogstraal & Kohls (1968) found specimens which they believed belonged to the syntype series of Haemaphysalis heinrichi. These authors also stated that the vial containing Haemaphysalis heinrichi included one specimen of Haemaphysalis wellingtoni along with larvae and nymphs of Dermacentor sp.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA64276FF23FE58FCB3FD2D.taxon	materials_examined	Type depository: not stated: syntypes (males) Type locality: Motzorongo, State of Veracruz, Mexico Type host: Tapirus bairdi Currents status: a nomen dubium (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA64276FF23FD5DFA83FBB9.taxon	materials_examined	Type depository: BMNH: syntypes? (unspecified developmental stages), holotype? (female) Type locality: England, presumably Type host: hedgehog Comments: on page 25 of Keirans & Hillyard (2001) it is stated under the sub-heading “ TYPE SERIES ” that Ixodes hexagonus was described from an unknown number of syntypes. This is followed by the subheading “ TYPE MATERIAL, ” where the authors state that a presumed holotype (female) of this species is held in the BMNH.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA64276FF23FBD0FC7DFA99.taxon	materials_examined	Type depository: undetermined: holotype? (female) Type localities: Bogdanci and Skopje, North Macedonia Type hosts: cattle and horses Current status: a synonym of Ixodes gibbosus (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA64276FF23FAB0FAEBF985.taxon	materials_examined	Type depository: not stated: syntypes (females) Type locality: Kecskemét, Hungary Type host: fox Current status: a synonym of Ixodes hexagonus (Camicas et al. 1998, under the genus Pholeoixodes).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA64276FF23F9B4FEAEF811.taxon	materials_examined	Type depositories: ENV and USNTC?: syntypes (females) Type localities: England, France and USA Type hosts: dog, fox, badger and squirrel Current status: partly a synonym of Ixodes canisuga and partly a synonym of Ixodes marxi (Guglielmone & Nava 2014). Comments: no specimens of Ixodes hexagonus inchoatus are listed in the USNTC type catalog by Keirans & Clifford (1984).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA14271FF23FBD0FBD8FAFD.taxon	materials_examined	Type depository: ZSH: holotype (male) and a paratype (female) Type locality: Molta, Uttar Pradesh, India Type host: Hemitragus jamlahicus jamlahicus Current status: a synonym of Haemaphysalis sundrai in Guglielmone et al. (2009), but Geevarghese & Mishra (2011) treated both Haemaphysalis himalaya and Haemaphysallis sundrai as valid.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA14271FF23FF78FBEDFE79.taxon	materials_examined	Type depositories: USNPC? and USNTC?: syntypes (males, females and a nymph) Type localities: Texas, Maine and Colorado, USA Type host: otter, Putreolus vison, sheep, cat, fox, squirrel, weasel and porcupine Current status: a synonym of Ixodes cookei (Neumann 1911 a, Keirans & Clifford 1978). Comments: Neumann (1901) listed the USNPC and USNTC as depositories, but no specimens were found in those collections by Salley et al. (1978) and Keirans & Clifford (1984), respectively.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA14271FF23FA6DFF0DF915.taxon	materials_examined	Type depositories: VRC: holotype (female) and paratypes (females, nymphs and larvae); BMNH?, IM and USNTC: paratypes (females, nymphs and larvae) Type locality: Palchan, Kullu District, Himachal Pradesh, India Type host: Mus musculus Comments. no specimens of Ixodes himalayensis are listed in the catalog of BMNH types by Keirans & Hillyard (2001).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA14271FF23FE11FA67FCBD.taxon	materials_examined	Type depositories: FVZ: holotype (male) and paratypes (a female, nymphs and larvae) Type locality: middle of Ebro Valley, La Cartuja Baja, province of Zaragoza, Spain Type host: the holotype specimen was collected from vegetation Comments: Rhipicephalus hibericus was confused with Rhipicephalus turanicus prior to its original description.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA34273FF23FC89FE0EFB9D.taxon	materials_examined	Type depository: BMNH: a lectotype (male) and paralectotypes (males and females) were designated by Keirans & Brewster (1981) Type locality: Banjul, Gambia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA34273FF23F9FCFF13F8D9.taxon	materials_examined	Type depository: BMNH: lectotype (male) and paralectotypes (males and females) were designated by Keirans & Brewster (1981) Type locality: south west shore of Lake Malawi, Malawi Type host: Heterohyrax brucei manningi Current status: a synonym of Haemaphysalis orientalis and its original name (Keirans & Brewster 1981). The current status of Haemaphysalis hoodi orientalis, as Haemaphysalis orientalis, was established by Hoogstraal (1956 b).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA34273FF23FE59FAC4FC91.taxon	materials_examined	Type depositories: BMNH: syntypes (a male and females), USNPC?, ZMB and ZSH: syntypes (females) Type locality: Queensland, and Port Stephens, New South Wales, Australia Type host: dog, sheep and Phascogale penicillata Comments: nymphs from India and the East Indies are included in the original description, but the nymph deposited in the BMNH is not Ixodes holocyclus (Keirans & Hillyard 2001); this is also true for the remaining type nymphs because Ixodes holocyclus is known only from Australia. One nymph deposited in the IP and classified by Neumann (1899) as Ixodes holocyclus from India is, in fact, a specimen of Ixodes petauristae, as discussed in Sénevet (1970). In their catalog of USNPC types, Salley et al. (1978) did not recognize specimens of Ixodes holocyclus.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA34272FF23F871FBFDFEE9.taxon	materials_examined	Type locality: Puerto Princesa, Palawan Island, the Philippines Type host: Mydaus marchei Comments: IR visited the FMNH in 2024 and only one paratype (labeled as allotype female) was found there; however, the paratypes were present in 2011, when the collection was rehoused.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA24272FF23FE81FD7BFDB1.taxon	materials_examined	Type depositories: USNTC: holotype (female) and paratypes (females); BMNH, FMNH, OVI and ZIAC: paratypes (females) (Keirans & Hillyard 2001; Filippova 2008) Type locality: 6 miles south of Ma’bar, Yemen	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFAD427DFF23FF79FE28FEE9.taxon	materials_examined	Type depository: ZMB: syntypes (males) (Moritz & Fischer 1981) Type locality: Bahia, Brazil	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFAD427DFF23FC41FE79FB09.taxon	materials_examined	Type depositories; USNTC: holotype (female) and paratypes (males and females); RCPV: paratypes (females) Type locality: Bamian Mountain, Liye Town, Longshan County, Xiangxi Tujia and Miao Autonomous Prefecture, Hunan Province, China	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFAD427DFF23F9B5FF31F8D9.taxon	materials_examined	Type depositories: BMNH, OVI, SDC and USNTC?: syntypes (males and females) Type locality: Naivasha, Karati Forest, Kenya Type host: buffalo Comments: in their catalog of USNTC types, Keirans & Clifford (1984) did not list specimens of Rhipicephalus hurti.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFAD427CFF23F878FD5FFF7D.taxon	materials_examined	Type depository: IM: syntypes (males and females) Type locality: Akola, State of Maharasthra, India	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFAC427CFF23FD61FCCAFC29.taxon	materials_examined	Type depositories: USNTC: holotype (female) and paratypes (males, females, nymphs and larvae); BMNH: paratypes (a female, nymphs and larvae) Type locality: Bonhote, Muktinah, Mustang District, Nepal	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFAC427FFF23F8E1FC0AFEB5.taxon	materials_examined	Type depositories: ZMB?: holotype (male) (Anastos 1950); USNTC: a paratype (unspecified developmental stage) (Keirans & Clifford 1984) Type locality: Ranau, Sumatra, Indonesia Type host: Hylobates syndactylus Comments: according to the original description, the holotype was to be transferred to the ZMB, but no specimens of Haemaphysalis hylobatis are included in the ZMB type catalog by Moritz & Fischer (1981), which contradicts Anastos (1950), although that author stated that the types had not been seen.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFAF427FFF23FEA5FB75FD95.taxon	materials_examined	Type depositories: USNTC: holotype (male) and paratypes (a female, other stages unknown); BMNH: paratypes (a male and a female), OVI and SAIMR: paratypes (unspecified developmental stages) (Keirans & Hillyard 2001) Type locality: Kogman’s Kloof, Montagu District, Province of Western Cape, South Africa	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFAF427FFF23F9B5FA79F87E.taxon	description	Nakamura, T. & Yajima, A. (1937) On the morphology and distribution of Japanese Ixodidae, specially of the species parasitic on cattle and horses. Report of the Government Experimental Station for Animal Hygiene, 17, 133 – 184. [in Japanese] Type depository: CK (presumed): holotype (male) (Yamaguti et al. 1971) Type locality: Miyake Island, Japan Type host: cow Current status: a synonym of Haemaphysalis cornigera, as presented in Takada (1990). Comments: Haemaphysalis ias was treated as valid in Guglielmone & Nava (2014) and Guglielmone et al. (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFAF427FFF23FB1CFE25F9A1.taxon	materials_examined	Type depository: AI (presumed): holotype (male) and paratypes (males and females) Type locality: Quang Ninh Province, Vietnam Type host: Muntiacus muntjak Current status: a synonym of Haemaphysalis lagrangei in Camicas et al. (1998) and a probable synonym of that species in Kolonin (2009).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFAE427EFF23FE34FDC3FD59.taxon	materials_examined	Type depository: BMNH: a lectotype (male) and paralectotypes (males and a female) were designated by Keirans & Brewster (1981) Type locality: Turrialba, Costa Rica	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFAE427EFF23FF78FB23FE05.taxon	materials_examined	Type depository: undetermined: holotype (female) Type locality: Bórçka, Ibrikli Bay, Turkey Type host: sheep Current status: a synonym of Alloceraea inermis (Guglielmone et al. 2014, under the genus Haemaphysalis). Comments: Haemaphysalis ibrikliensis is treated as provisionally valid by Bursali et al. (2012).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFAE427EFF23FB45FD20F9A1.taxon	description	See also savignyi impeltatum Schulze & Schlottke, 1929, Hyalomma Type depository: USNTC: a neotype (male) was designated by Apanaskevich & Horak (2009) Type locality: Greater Casablanca, Morocco Type host: cattle Comments: Hyalomma impeltatum was originally named Hyalomma savignyi impeltatum, but the neotype was described under the name Hyalomma impeltatum.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFAE4279FF23F8B8FE71FEB5.taxon	materials_examined	Type depository: ZMB: syntype (a male) (Moritz & Fischer 1981) Type locality: Senegal Type host: not stated Comments: Koch (1844) described Hyalomma impressum from at least one male and a female but only a male is preserved in the ZMB.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA94279FF23FD5CFCB0FC60.taxon	materials_examined	Type depository: BMNH: syntypes (males) (Keirans & Hillyard 2001) Type localities: Bolivia, and Cuenca, Ecuador Type host: tapir Comments: one of the males deposited in the BMNH as Amblyomma incisum is, in fact, Amblyomma scalpturatum (Labruna et al. 2005), and this tick was treated as the probable holotype of Amblyomma scalpturatum in Guglielmone et al. (2014), but that species was described from two females.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA94279FF23FA48FC09F931.taxon	materials_examined	Type depositories: BMNH and GM: syntypes (males and females) (Keirans & Hillyard 2001) Type locality: “ Kokarcet ” (“ Tanosserim ” = Tenasserim?), Bhamo, Myanmar Type hosts: Manis aurita and Manis javanica Current status: a synonym of Amblyomma javanense (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA94279FF23F950FE73F852.taxon	materials_examined	Type depositories: VRC: holotype (male) and paratypes (males and females); BMNH, IM and USNTC: paratypes (males and females) Type locality: Mukteswar, Naini Tal District, State of Uttarakhand, India Type host: the holotype specimen was collected from vegetation Comments: some specimens in the original description of Haemaphysalis indoflava were classified as Haemaphysalis flava by Sharif (1928).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA94279FF23FEA4FD73FD2D.taxon	materials_examined	Type depository: USNTC?: syntypes (females) Type locality: Sennar, Sudan Type host: unknown Current status: a synonym of Hyalomma truncatum (Apanaskevich & Horak 2008 a). Comments: Apanaskevich & Horak (2008 a) found specimens that probably belong to the syntype series of Hyalomma impressum luteipes, deposited in the USNTC.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA84278FF23FA24FB8AF915.taxon	materials_examined	Type depository: ZMB: syntypes (a male and a female) (Moritz & Fischer 1981) Type locality: “ Bintang, Hinterindien ”, but Bintan Island, Indonesia, according to Anastos (1950). Type host: not stated Current status: a synonym of Amblyomma testudinarium (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA84278FF23FD14FA98FB9D.taxon	materials_examined	Type depository: ZMB?: syntypes (females and nymphs) (Neumann 1901) Type locality: not stated Type host: not stated Current status: a synonym of Ixodes canisuga, as listed in Camicas et al. (1998 under the genus Pholeoixodes) and Guglielmone & Nava (2014). Comments: in the catalog of ZMB types by Moritz & Fischer (1981), no specimens of Ixodes inermis are listed.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA84278FF23FB8CFB26FA35.taxon	materials_examined	Type depositories: BMNH: a lectotype (female) and paralectotypes (females) were designated by Keirans & Brewster (1981); ENV and ZMB: paralectotypes (females) (Nuttall & Warburton 1915, as “ types ”) Type locality: Belgachia, Kolkata, State of West Bengal, India Type host: Himalayan zebu Current status: a synonym of Alloceraea aponommoides and its original name. The current status of Haemaphysalis inermis aponommoides, as Alloceraea aponommoides, was established by Kelava et al. (2024).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFA8427BFF23F904FE08FEB5.taxon	materials_examined	Type depository: ZMB?: holotype (male) in the original description; USNTC: holotype? (unspecified developmental stage) (Keirans & Clifford 1984) Type locality: Borneo Type host: Sus verrucosus Current status: a synonym of Amblyomma testudinarium (Anastos 1950, Camicas et al. 1998). Comments: Moritz & Fischer (1981) did not recognize the presence of a type specimen held in the ZMB, while Keirans & Clifford (1984) named as “ type ” the specimen deposited in the USNTC, under the name Amblyomma borneense infestum (a lapsus).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFAB427BFF23FEA4FDD6FDC9.taxon	materials_examined	Type depository: ZMB?: holotype (male) (Schulze 1935 b) Type locality: Kosempo, Taiwan Type host: dog Current status: a synonym of Amblyomma testudinarium (Anastos 1950, Camicas et al. 1998). Comments: In their list of ZMB types, Moritz & Fischer (1981) did not recognize the presence of a type specimen of Amblyomma infestum taivanicum.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFAA427AFF23FEA4FD80FE5D.taxon	materials_examined	Type depository: USNTC: holotype (female) and paratypes (a female, nymphs and larvae) Type locality: La Pedraza, Madrid, Spain	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFB54265FF23FBD0FCF9FA99.taxon	materials_examined	Type depositories: USNTC: holotype (male) and paratypes (males and females); BMNH and OVI: paratypes (males and females) Type locality: Kawoko-Masaka, Masaka District, Uganda	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFB44264FF23F990FADEF83D.taxon	materials_examined	Type depository: BMNH: a lectotype (male) and paralectotypes (males) were designated by Keirans & Brewster (1981) Type locality: Shensi, China Type host: Capreolus capreolus Current status: a synonym of Haemaphysalis japonica (Guglielmone & Nava (2014, Nakao & Ito (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFB44264FF23FB1CFDA3F9F9.taxon	materials_examined	Type depository: BMNH: a lectotype (male) and paralectotypes (a male and females) were designated by Keirans & Brewster (1981) Type locality: Kyushu, Japan Type host: Capricornis crispus Comments: the type specimens of Haemaphysalis japonica were found in a vial labeled Haemaphysalis flava that contained Haemaphysalis flava and Haemaphysalis campanulata as well as the types of Haemaphysalis japonica, as discussed in Yamaguti et al. (1971).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFB44264FF23FD39FC8AFC0D.taxon	materials_examined	Type depositories: USNTC: holotype (male) and paratypes (a female and nymphs); CMC: paratypes (a male, a female and a nymph) Type locality: Murray´s Mistake, Banks Peninsula, New Zealand	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFB74267FF23FD60FE67FBB9.taxon	materials_examined	Type depositories: BMNH and GM: syntypes (nymphs) (Anastos 1950, Keirans & Hillyard 2001) Type locality: “ Karkecet ”, (“ Tanosserim ” = Tenasserim?), Myanmar Type host: Manis javanica Current status: a synonym of Amblyomma javanense and its original name (Keirans & Hillyard 2001). The current status of Rhipicephalus javanensis, as Amblyomma javanense, was established by Schulze (1934). Comments: Anastos (1950) stated that the “ type ” is deposited in the GM. Guglielmone et al. (2020) found discrepancies in the morphological descriptions of Amblyomma javanense; further studies of types may aid in better defining this species.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFB74266FF23F8BFFBB5FF51.taxon	materials_examined	Type depositories: USNTC: holotype (female) and paratypes (a male? a female? nymphs and larvae); FMNH: paratypes (a female and a larva) Type locality: southeast of Tabay (La Coromoto), State of Mérida, Venezuela Type host: Thomasomys laniger Comments: it is uncertain which of the above collections contain the adult paratypes.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFB64266FF23FDA9FF43FC0D.taxon	materials_examined	Type depositories: USNTC: holotype (female) and paratypes (males and females); DEEZ and MCZ: paratypes (males and females) Type locality: Panama Type host: the holotype specimen was collected from vegetation. k	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFB64266FF23F9D9FBC8F8E5.taxon	materials_examined	Type depositories: USNTC: holotype (female); AR, BMNH and FMNH?: paratypes (females) Type locality: Borg El Arab, Maryut, Governorate of Alexandria, Egypt Type host: Vulpes vulpes aegyptiaca Comments: IR visited the FMNH in 2024 and the paratypes were not found there.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFB64261FF23F895FC87FF7C.taxon	discussion	Comments: Lindquist et al. (2016) and Schachat et al. (2018) treat Dermacentor kamshadalus as an invalid name, but Apanaskevich & Barker (2021) strongly support its validity.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFB14261FF23FEEDFC7BFE05.taxon	materials_examined	Type depository: AMMS: holotype (female) and paratypes (females and nymphs) Type locality: Kangding, Kangding County, Sichuan Province, China	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFB04260FF23FAD5FC9FF915.taxon	materials_examined	Type depositories: USNTC: lectotype (female) and paralectotype (males and females); MCZ: a paralectotype (female) (Keirans & Clifford 1978, Keirans & Clifford 1984) Type locality: Meeteetse, Wyoming, USA Type host: Taxidea taxus Comments: Bishopp (1911) described the male and female of Ixodes kingi selected from a lot of several adults, nymphs and larvae. A lectotype female was selected by Keirans & Clifford (1978); therefore, the remaining males and female ticks are treated here as paralectotypes. Nevertheless, Keirans & Clifford (1984) stated that “ paralectotypes ” (developmental stages not provided) are deposited in the USNTC.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFB04260FF23FE59FB51FC91.taxon	materials_examined	Type depository: DA: the holotype, paratypes or syntypes are not specifically indicated in the original description (Guglielmone et al. 2020) Type locality: Kerala, India Type host: buffalo Comments: Nosomma keralensis is treated as a name incertae sedis by Schachat et al. (2018). In fact, this name can be considered invalid because it was published after 1999, when the International Code of Zoological Nomenclature began requiring designation of a holotype or expressly indicated syntypes. Nosomma keralensis is treated as provisionally valid in Guglielmone & Nava (2014), Guglielmone et al. (2014, 2023) and here.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFB34263FF23FC18FB2FFB41.taxon	materials_examined	Type depository: not stated: syntypes (males and females) Type localities: Baurú and Jacutinga, State of São Paulo, Brazil Type hosts: Cervus campestris and Cervus rufus Current status: Haemaphysalis kochi, a preoccupied name, was renamed Haemaphysalis kohlsi by Aragão & Fonseca (1951), and both names are synonyms of Haemaphysalis juxtakochi (Guglielmone et al. 2021).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFB34263FF23FAF9FE13FA4D.taxon	materials_examined	Type depositories: ZMB: lectotype (female); BMNH: a paralectotype (male) was designated by Clifford et al. (1983) Type locality: Sadani, Tanzania	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFB34263FF23FD5DFE0DFC61.taxon	materials_examined	Type depository: DA: holotype (male) and paratypes (males and females) Type locality: Chelari Cattle Market, Malappuran District, State of Kerala, India Type host: cow Comments: Haemaphysalus knobigera is a poorly described species considered provisionally valid in Guglielmone et al. (2009, 2014) and others.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFB24262FF23FE10FDBCFBB9.taxon	description	Du, C. H., Sun, Y., Xu, R. M. & Shao, S. T. (2018) Description of Haemaphysalis (Alloceraea) kolonini sp. nov., a new species subgenus Alloceraea Schulze (Ixodidae: Haemaphysalis) in China. Acta Parasitologica, 63, 678 – 691. See kolonini Du, Sun, Xu & Shao, 2018, Haemaphysalis kolonini Du, Sun, Xu & Shao, 2018, Haemaphysalis Du, C. H., Sun, Y., Xu, R. M. & Shao, S. T. (2018) Description of Haemaphysalis (Alloceraea) kolonini sp. nov., a new species subgenus Alloceraea Schulze (Ixodidae: Haemaphysalis) in China. Acta Parasitologica, 63, 678 – 691. Type depository: AMMS: holotype (male) and paratypes (males and females) Type locality: Boshan, Yunnan Province, China Type host: the holotype specimen was collected from vegetation Current status: a synonym of Alloceraea kolonini and its original name, whose current status, as Alloceraea kolonini, was established by Kelava et al. (2024).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFB24262FF23FF79FD97FEE9.taxon	materials_examined	Type depositories: USNTC: holotype (female), paratypes (males and females); BMNH: paratypes (a male and a female) Type locality: Julia Percy Island, Australia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFBD426DFF23FE11FB61FDC9.taxon	description	See kopsteini Oudemans, 1925; Eschatocephalus, as Eschatocephalus ropsteini (a lapsus)	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFBD426DFF23F9D8FF0DF8D9.taxon	materials_examined	Type depository: ZMB?: holotype (female) (Minning 1934) Type locality: Sangir, northwest of Sulawesi Island, Indonesia Type host: cattle Current status: a name incertae sedis (Guglielmone & Nava 2014). Comments: the holotype of Boophilus krijgsmani is not listed in the catalog of ZMB types by Moritz & Fischer (1981).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFBC426CFF23FAD5FF7BF949.taxon	materials_examined	Type depositories: USNTC: holotype (male) and paratypes (males, females, nymphs and larvae); BMNH?, VRC, ZIAC? and ZSI: paratypes (males, females, nymphs and larvae) Type locality: Kyasanur Forest, near Barage, Stattion 3, Shimoga District, State of Karnataka, India Type host: the holotype specimen was collected from vegetation Comments: Keirans & Hillyard (2001) and Filippova (2008) did not recognize type specimens of Haemaphysalis kyasanurensis as having been deposited in the BMNH and ZIAC, respectively. l	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFBC426CFF23FD85FD58FC91.taxon	materials_examined	Type depositories: USNTC: holotype (female) and paratypes (females, nymphs and larvae); BM, BMNH and FMNH: paratypes (females, nymphs and larvae) Type locality: Ho-ping, Tai-chung Hsien, Taiwan	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFBE426EFF23FB45FD73F984.taxon	materials_examined	Type depositories: USNTC: holotype (male) and paratypes (a male? females, nymphs and larvae); paratypes: BMNH (nymphs and larvae), ZIAC (a male, a nymph and a larva), FMNH and VRC (a male?, females, nymphs and larvae) Type locality: Geling, Mustang District, Nepal Type host: Alticola sp. Comments: Camicas et al. (1998) amended the name of the species to lamai. The type depository of the paratype male of Anomalohimalaya lamai is uncertain.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFBE426EFF23FD84FC28FCF5.taxon	materials_examined	Type depository: ZIAC: a lectotype (male) and paralectotypes (females) (Filippova 2008) Type locality: near Babuk-Aul, Adler District, western Caucasus, Russia Type host: Glis glis Current status: a synonym of Ixodes laguri (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFBE426EFF23FC64FC28FBD5.taxon	materials_examined	Type depository: IPCAS: holotype (female) and paratypes (males) Type locality: Dvorniky, Slovakia Type host: the holotype specimen was collected from the nest of Citellus citellus Current status: a synonym of Ixodes laguri (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFB94269FF23FA48FBC2F8AD.taxon	materials_examined	Type depository: BMNH: a lectotype (male) and paralectotypes (a female and a nymph) were designated by Keirans & Brewster (1981) Type locality: Raffles Museum, Singapore, where the type host was captive Type host: Laticauda colubrina Current status: a synonym of Amblyomma nitidum (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFB84268FF23FDA8FBC6FC0D.taxon	materials_examined	Type depository: ZMB: syntypes (males) in Moritz & Fischer (1981) that listed these specimens as the probable syntypes of this tick Type locality: South Africa (Koch 1844) Type host: not stated Current status: Robinson (1926), Guglielmone & Nava (2014) and others treated Hyalomma latum as a synonym of Amblyomma latum, but Neumann (1911 a), Camicas et al. (1998), and Guglielmone et al. (2020) list Hyalomma latum as a synonym of Amblyomma sylvaticum, a position also maintained here.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFB84268FF23FA01FD5BF915.taxon	materials_examined	Type depositories: BM: holotype (female) and paratypes (females and nymphs); USNTC: paratypes (a female and a nymph) (Keirans & Clifford 1978) Type locality: Laysan Island in the Pacific Ocean	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFBB426BFF23FD5CFB02FC0D.taxon	materials_examined	Type depository: OVI: syntypes (males and females) (Apanaskevich et al. 2007) Type locality: Macia, Bilene and Massoane, Mozambique Type host: dog Current status: a synonym of Haemaphysalis elliptica, as discussed in Apanaskevich et al. (2007).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFBB426BFF23FC3CFA6BFB6D.taxon	materials_examined	Type depositories: BMNH and IM: syntypes (females) (Keirans & Hillyard 2001) Type locality: Indian Museum Compound, Kolkata, India Type host: Canis aureus Current status: a synonym of Haemaphysalis indica and its original name, as discussed in Hoogstraal (1970). The current status of Haemaphysalis leachi indica, as Haemaphysalis indica, was established by Santos Dias (1958 a).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFBB426BFF23FEC8FB87FD2D.taxon	materials_examined	Type depository: unknown: “ type ” (male) (Hoogstraal 1958) Type locality: Egypt Type host: unknown Current status: a synonym of Haemaphysalis leachi and its original name (Neumann 1897). The current status of Ixodes leachi, as Haemaphysalis leachi, was also established by Neumann (1897). Comments: see Haemaphsalis leachi for the designation of a neotype under that name.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FFBA426AFF23FEC9FC11FDB1.taxon	materials_examined	Type depository: ZIAC: lectotype (male) and a paralectotype (female) (Filippova 2008) Type locality: environs of Agit-Ma, Kenimekh District, Uzbekistan Type host: hare Comments: Walker et al. (2000) do not list a type depository for this species.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF454295FF23FB1CFF08F985.taxon	materials_examined	Type depository: ZMUC: holotype (male) Type locality: America Type host: unknown Current status: a nomen dubium (Camicas et al. 1998, Guglielmone & Nava 2014). Comments: the type of Acarus lineatus was found labeled as Acarus cajennensis in the ZMUC (Nava et al. 2014 a).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF454295FF23F9B4FBF8F83D.taxon	materials_examined	Type depository: unknown Type locality: Egypt Type host: unknown Current status: a synonym of Rhipicephalus linnaei and its original name, as stated in Šlapeta et al. (2022). Comments: see Rhipicephalus linnaei for a neotype designated under that name.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF444294FF23FD60FD2CFC29.taxon	description	Kolonin, G. V. (1995) Haemaphysalis lobachovi sp. n. (Acarina: Ixodidae) from porcupine (Hystrix cristata) from Ethiopia. Folia Parasitologica, 42, 239. Type depositories: ZMMO: holotype (male); KC: a paratype (male) Type locality: near Awash National Park, Ethiopia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF444294FF23FF79FD45FE79.taxon	materials_examined	Type depository: ESAM: a neotype (male) is to be deposited in this museum (Šlapeta et al. 2022) Type locality: Esna City, Luxor Governorate, Egypt (Šlapeta et al. 2022) Type host: dog (Šlapeta et al. 2022) Comments: this species was originally named Ixodes linnaei, but a neotype was designated under the name Rhipicephalus linnaei by Šlapeta et al. (2022).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF474297FF23FC40FEFDFAA5.taxon	materials_examined	Type depository: BMNH: holotype (female) (Keirans & Hillyard 2001) Type locality: Guyana Type host: host unknown Current status: a synonym of Amblyomma humerale (Camicas et al. 1998, Keirans & Hillyard 2001). Comments: Estrada-Peña & Castellá (1994) listed Amblyomma longirostrum as a synonym of Amblyomma longirostre.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF474297FF23FAD5FD76FA4D.taxon	materials_examined	Type depository: CB: holotype (female), but probably lost (Guglielmone et al. 2003) Type locality: Nogoyá, Entre Ríos, Argentina	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF464296FF23FBF5FB8FFA99.taxon	materials_examined	Type depository: ZIAC: holotype (male) and paratypes (males, females, nymphs and larvae) (Filippova 2008) Type locality: juncture of Obikhingou and Lyulya-Kharvi rivers, Savor Area, Peter the First Mountain Range, Tajikistan Type host: Alticola argentatus Comments: the holotype specimen was reared from larvae collected from the type host.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF464296FF23FD38FA66FC45.taxon	materials_examined	Type depository: BMNH: a lectotype (female) and a paralectotype (female) were designated by Keirans & Brewster (1981) Type locality: Frontera, State of Tabasco, Mexico Type host: Didelphis sp. Current status: a synonym of Ixodes luciae (Keirans & Brewster 1981). Comments: Santos Dias (1954) renamed Ixodes loricatus spinosus (preoccupied) as Ixodes loricatus vogelsangi.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF464296FF23FF79FDE4FEE9.taxon	materials_examined	Type depository: BMNH: holotype (male) Type locality: Kasongo, Malawi	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF464296FF23FAB0FC07F985.taxon	materials_examined	Type depository: not stated: syntypes (males, females and nymphs) Type localities: Cape and Orange Colonies, South Africa Type hosts: horse and cattle Current status: a synonym of Margaropus winthemi (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF464296FF23F9B5FE25F811.taxon	materials_examined	Type depositories: OVI: holotype (male) and paratypes (males, females, nymphs and larvae); USNTC: paratypes (males, females, nymphs and larvae) Type locality: Dordrecht, Province of Eastern Cape, South Africa Type host: not stated Comments: the type specimens of Rhipicephalus lounsburyi were reared by Theiler & Robinson (1953) under the name Rhipicephalus follis.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF414291FF23FD38FD1DFBD5.taxon	materials_examined	Type depositories: SC: holotype (female) and paratypes (females, a nymph and larvae); USNTC: paratypes (a female and a larva) Type locality: Limanowa, Poland Type host: dog Current status: a synonym of Amblyomma sphenodonti, as discussed in Siuda (1982, under the genus Aponomma). Comments: the type locality and host of Aponomma ludovici are wrong, but the valid data are unknown (Siuda 1982). Siuda (1972) stated that a female and a larva were sent to the USNTC, but Keirans & Clifford (1984) listed one paratype (tick stage not provided) deposited there.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF414291FF23FF79FC87FE05.taxon	materials_examined	Type depository: MCZ: syntypes (a female and a male) Type locality: Mount Cholo, Nyasaland, Malawi Type host: Cricetomys gambianus viator Comments: the specimens listed as held in the MCZ are treated here as syntypes, although a holotype and a paratype of Ixodes loveridgei were designated in the original description.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF414291FF23FE35FE66FD01.taxon	materials_examined	Type depository: not stated: holotype? (female), but probably in IPG (Guglielmone et al. 2003) Type locality: Cayenne? [sic], French Guiana Type host: dog? [sic]	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF414291FF23FB45FE24FA99.taxon	materials_examined	Type depository: LMNH: holotype (female) Type locality: Madagascar	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF414291FF23F94DFC40F8E5.taxon	materials_examined	Type depository: BMNH: lectotype (male) and a paralectotype (male) (Keirans & Hillyard 2001); the lectotype was designated by Walker et al. (1988) Type locality: near Lualaba River, Democratic Republic of the Congo	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF404290FF23FE34FBDFFD65.taxon	materials_examined	Type depository: undetermined: syntypes (males and females) Type locality: several northern localities, Algeria Type host: camels Current status: a synonym of Hyalomma lusitanicum (Apanaskevich et al. 2008 a).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF404290FF23FD14FB0AFB9D.taxon	materials_examined	Type depository: not stated: syntypes (males and females); USNTC: a syntype (male) (Apanaskevich et al. 2008 a, named as “ type ”) Type locality: Extremadura, Spain Type host: not stated Current status: a synonym of Hyalomma lusitanicum, as discussed in Apanaskevich et al. (2008 a).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF404290FF23FA6DFE0BF8AD.taxon	materials_examined	Type depository: ZMB? (syntypes); USNTC: a lectotype (female) was designated by Apanaskevich & Schenk (2020) Type locality: Weyland Mountains, Papua, Indonesia Type host: unknown Comments: Schulze (1935 a) stated that the “ type specimens ” were deposited in the ZMB, but they are not listed in the catalogue of type specimens by Moritz & Fischer (1981). Apanaskevich & Schenk (2020) found a female of Ixodes luxuriosus deposited in the USNTC that they considered part of this species’ syntype series and designated that specimen as the lectotype.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF404293FF23F8C4FF4AFEE9.taxon	description	(Ixodoidea: Ixodidae). H. (Kaiseriana) luzonensis sp. n. from Philippine deer and boars. Journal of Parasitology, 54, 402 – 410. Type depositories: USNTC: holotype (male) and paratypes (males, females, nymphs and larvae); BM, BMNH, FMNH and PNM: paratypes (males, females, nymphs and larvae) Type locality: Mount Dumug, Labi, Bongabon, Province of Nueva Ecija, Luzon, Philippines Type host: Cervus philippinus m	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF434292FF23F991FE13FEE9.taxon	materials_examined	Type depository: ZMB: holotype (male) (Moritz & Fischer 1981) Type locality: “ Carolina, ” USA Type host: not stated maculatus Neumann, 1901, Rhipicephalus Neumann, L. G. (1901) Révision de la famille des ixodidés (4 e mémoire). Mémoires de la Société Zoologique de France, 14, 249 – 372. Type depository: ZMB: syntypes (a male and females) (Moritz & Fischer 1981) Type locality: Cameroon? Type host: Platymeris horrida, an insect; and consequently Walker et al. (2000) treated this host as incorrect Comments: Neumann (1901) listed Cameroon as the type locality of Rhipicephalus maculatus, but this tick is not established in that country (Morel 2003, Walker et al. 2000), and both the type locality and host probably resulted from incorrect sample labeling.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF4D429DFF23FE58FC82FD65.taxon	materials_examined	Type depository: unknown: holotype? (female) Type locality: Germany Type host: unknown Current status: a synonym of Dermacentor marginatus and its original name. The current status of Acarus marginata, as Dermacentor marginatus, was established by Brumpt (1913).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF4D429DFF23FF78FC48FE21.taxon	materials_examined	Type locality: northwest of San Antonio, Isla Margarita, State of Nueva Esparta, Venezuela Type host: Pseudoboa neuwiedii Current status: a synonym of Amblyomma dissimile (Guerrero, 1996).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF4D429DFF23FD14FE88FC0D.taxon	materials_examined	Type depository: unknown: holotype? (male) treated as lost (Apanaskevich & Horak 2008 b) Type locality: Italy	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF4D429DFF23FA6CFD3AF8AD.taxon	materials_examined	Type depository: not stated (Schulze & Schlottke 1929): USNTC: syntypes? (males and females) (Apanaskevich & Horak 2008 b) Type localities: Brioni Island, Croatia, and Istria Peninsula, Croatia? Italy? Slovenia? Type host: undetermined Current status: a synonym of Hyalomma marginatum (Apanaskevich & Horak 2008 b). Comments: Apanaskevich & Horak (2008 b) found males and females in the USNTC that may be syntype specimens used to describe Hyalomma marginatum brionicum.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF4D429CFF23F8DCFB8AFF7D.taxon	materials_examined	Type depository: MCNB: holotype (male) Type locality: Santa Galdana, Menorca Island, Spain Type host: undetermined Current status: a synonym of Hyalomma marginatum (Apanaskevich & Horak 2008 b).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF4C429CFF23FBF4FE23FB6D.taxon	materials_examined	Type depository: ZMB: holotype (male) (Moritz & Fischer 1981) Type locality: South Africa	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF4C429CFF23FEECFDE3FD95.taxon	materials_examined	Type depository: ZIAC: lectotype (male) and paralectotypes (males) (Filippova 2008) Type locality: Varzobstroi, Tajikistan Type host: goat Current status: a synonym of Hyalomma turanicum and its original name. The current status of Hyalomma marginatum turanicum, as Hyalomma turanicum, was provided first by Hoogstraal (1956 a), and definitively by Apanaskevich & Horak (2008 b).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF4C429CFF23F9DFFABAF8EE.taxon	materials_examined	Type depository: USNTC: holotype (male) and paratypes (males and females), MCZ: paratype (a female) Type locality: Mayfield, Idaho, USA Type host: Marmota sp. Comments: the holotype was reared from immature stages collected from the type host. According to Keirans & Clifford (1978) the holotype is missing, but Keirans & Clifford (1984) listed the holotype as held in USNTC.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF4C429FFF23F89BFDA2FF51.taxon	materials_examined	Type depositories: USNTC: a lectotype (female), MCZ: a paralectotype (females and nymphs) (Keirans & Clifford 1978) Type locality: Washington, D. C., USA	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF4F429FFF23F9FCFB24F915.taxon	materials_examined	Type depositories: MZD: holotype (male) and a paratype (female); CS: paratypes (males and females) Type locality: near Kotakuji, Nanasawa, Atsugi-machi, Naka-gun, Kanagawa Prefecture, Japan	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF4E4299FF23F8E7FF0DFF7D.taxon	materials_examined	Type depository: GM: holotype? (male or female) (Kohls 1957, who said that the “ type ” is held in the GM) Type locality: Chaco austral, Paraguay Type host: an unknown mammal Current status: a synonym of Rhipicephalus microplus and its original name. The current status of Haemaphysalis micropla, as Rhipicephalus microplus, was provided first by Canestrini (1890) and definitively by Murrell & Barker (2003).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF4E429EFF23FB20FD74FAD1.taxon	materials_examined	Type depository: USNTC: holotype (female) and a paratype (female) Type locality: Tancitaro, Michoacán, México	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF494299FF23FBF4FCB5FB6D.taxon	materials_examined	Type depository: MNHN: syntypes (a male and a female), but apparently lost (Guglielmone et al. 2003) Type locality: Guatemala Type host: Hesperomys sp.? [sic from the original description]	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF494299FF23FB1CFC02FA35.taxon	materials_examined	Type depositories: USNTC: holotype (male) and paratypes (males); BMNH: paratypes (males) Type locality: Nikawewa Camp, near Kantalai, Eastern Province, Sri Lanka	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF494299FF23FD5CFD95FC45.taxon	materials_examined	Type depository: CK: syntypes (males and females) Type locality: “ Nanrai, ” South Keysio, South Korea Type host: cattle Current status: a synonym of Rhipicephalus microplus (Yamaguti et al. 1971, under the genus Boophilus). Comments: Guglielmone & Nava (2014) named the tick described by Kisihida (1936) Boophilus minningi, but that author named it Palpoboophilus minningi.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF484298FF23FE10FE6BFD65.taxon	materials_examined	Type depository: ZMB: lectotype (female) and a paralectotype (male) following designation of a lectotype by Nava et al. (2014 a) Type locality: Mexico	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF484298FF23FBF4FC22FAFD.taxon	materials_examined	Type depository: USNTC: syntypes? (a male and females) (Keirans & Clifford 1984, as “ cotypes ”) Type locality: Mountain Home, Idaho and Prescott, Washington, USA Type host: horse Current status: a synonym of Dermacentor andersoni (Keirans & Clifford 1984, Camicas et al. 1998). Comments: Keirans & Clifford (1984) did not specify the developmental stages of “ cotypes ” held in the USNTC, but Dermacentor modestus was described using a male and female ticks.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF484298FF23FA6CFD5FF9F9.taxon	materials_examined	Type depository: USNTC: holotype (female) and paratypes (males, females, nymphs and larvae) Type locality: Amargosa Valley, California, USA	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF4B429BFF23FAF8FC66F985.taxon	materials_examined	Type depositories: USNTC: holotype (male) and a paratype (female); CNCDGS, CVLPR and IBU: paratypes (males, females and larvae) Type locality: Balneário Pinhal, State of Rio Grande do Sul, Brazil	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF4B429BFF23FF78FBE7FEE9.taxon	materials_examined	Type depository: MZD: holotype (female) Type locality: Mount Lide Series, Kitakambara-gun, Niigata Prefecture, Japan	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF4A429AFF23FCADFCFBFB9D.taxon	materials_examined	Type depositories: USNTC: holotype (male) and paratypes (males, females, nymphs and larvae); ORSTOM: paratypes (males, females, nymphs and larvae) Type locality: Saboya, Department Nioro du Rip, Senegal	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF4A429AFF23FE35FBC8FCBD.taxon	materials_examined	Type depositories: USNTC: holotype (female) and a paratype (female); FMNH? and GML: paratypes (females) Type locality: El Respingo Station, Volcán Barú National Park, Chiriquí, Panama Type host: Odontophorus guttatus Comments: IR visited the FMNH in 2024 and the paratypes were not found there.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF554285FF23FAB1FDF7F9A1.taxon	materials_examined	Type depository: not stated: holotype (male) Type locality: Maroua, Cameroon	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF544284FF23FB1DFDF8FA35.taxon	materials_examined	Type depository: not stated: holotype (female) and a paratype (male) Type locality: Sangalkam, Senegal	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF544284FF23FC65FB5FFB6D.taxon	materials_examined	Type depositories: VLM: holotype (male) but lost; USNTC: a neotype (male) was designated by Tomlinson et al. (2018) Type locality: Calgary Farm, Mashonaland Central, Zimbabwe Type host: Galerella sanguinea Comments: Kolonin (2009) did not include this species in his list of ixodids of the world, and Tomlinson et al. (2018) cautioned that there have been several misidentifications of Haemaphysalis muhsamae.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF514281FF23FAB0FBA2F985.taxon	materials_examined	Type depositories: OVI: holotype (female) and paratypes (males and females); BMNH and USTNC: paratypes (males and females) Type locality: Loskop Dam Nature Reserve, Province of Mpumalanga, South Africa	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF514281FF23FF78FBEBFE05.taxon	materials_examined	Type depository: BMNH: a lectotype (male) and paralectotypes (males and females) were designated by Keirans & Brewster (1981) Type locality: near the mouth of Tasangazi River, Luangwa Valley, Zambia Type host: eland Current status: a synonym of Rhipicephalus kochi (Keirans & Brewster 1981).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF514281FF23FE34FAFBFCBD.taxon	materials_examined	Type depository: BMNH: a lectotype (male) and paralectotypes (males and females) were designated by Keirans & Brewster (1981) Type locality: near Fort Mlangeni, Ncheu District, Malawi Type host: Tragelaphus strepsiceros Current status: a synonym of Rhipicephalus punctatus and its original name (Walker et al. 2000). The current status of Rhipicephalus neavei punctatus, as Rhipicephalus punctatus, was established by Santos Dias (1951).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF504280FF23FF79FDE4FEE9.taxon	materials_examined	Type locality: Leucopani, Nepal	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF504280FF23FA00FE85F8D9.taxon	materials_examined	Type depository: ENV: syntypes (females and nymphs) Type locality: New Zealand Type host: not stated Current status: a synonym of Ixodes eudyptidis (Heath & Palma 2017). Comments: Nuttall & Warburton (1911) regarded the redescription of Ixodes eudyptidis by Neumann (1899) as belonging to a new species that was described under the name Ixodes neumanni. However, that decision was unjustified, as discussed in Heath & Palma (2017), and Ixodes neumanni is currently considered a synonym of Ixodes eudyptidis.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF504280FF23FC64FE08FBB9.taxon	materials_examined	Type depository: not stated: holotype (male) Type locality: Salta, Argentina	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF504283FF23F870FC91FEE9.taxon	materials_examined	Type depositories: OVI: holotype (male) and paratypes (males and females); BMNH and USNTC: paratypes (males and females) Type locality: farm “ Soutdoringvlei ” Bethanien District, Namibia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF534283FF23FC40FC08FB09.taxon	materials_examined	Type depository: not stated in the original description: syntypes (females); IP: a syntype (female) (Morel 1963) Type locality: Kef, Tunisia Type host: the type specimens were collected on stones Current status: a synonym of Haemaphysalis sulcata (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF534282FF23F8E0FECAFEB5.taxon	materials_examined	Type depositories: NIAH: holotype (male) and paratypes (males, females, nymphs and larvae); MZD: paratypes (males, females, nymphs and larvae) (Kitaoka & Saito 1967), ZIAC: paratypes (a male, a female, a nymphal pelt and larvae) (Filippova 2008) Type locality: Kakudayama, Minamikambara-gun, Niigata Prefecture, Japan Type host: Lepus timidus angustidens Comments: the specimens used to describe this species were laboratory reared from a female tick collected from the type host.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF524282FF23FC88FF3EFB09.taxon	materials_examined	Type depositories: BMNH: syntypes (males and females); MNHN: syntypes (males); USNTC?: syntypes (males, females and nymphs), (Neumann 1897, Keirans & Hillyard 2001) Type localities: Maryland (probably), USA; Santo Domingo, Dominican Republic; and Jamaica Type host: horse Comments: Keirans & Clifford (1984) did not list specimens of Dermacentor nitens in their catalog of USNTC types.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF524282FF23FB20FB18FA69.taxon	materials_examined	Type depository: not stated: syntypes (females and a nymph); BMNH: a syntype (female) (Keirans & Hillyard 2001) Type locality: Christmas Island in the Indian Ocean Type host: Rattus macleari Comments: the syntype specimens were collected prior to their host’s extinction (James et al. 2019).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF524282FF23FA00FC72F9DD.taxon	materials_examined	Type depositories: BMNH: a syntype (male); OVI: a syntype (female) (Walker et al. 2000) Type locality: Stellenbosch, Province of Western Cape, South Africa	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF524282FF23FEA5FD94FDB1.taxon	materials_examined	Type depositories: CCNMNS: holotype (female); CMT and / or CTY: paratypes (females) Type locality: Mine City, Yamaguti, Japan	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF5D428DFF23F929FF3EF875.taxon	materials_examined	Type depositories: MNHN, BMNH and USNTC?: syntypes (males, females and nymphs) Type localities: Costa Rica and Brazil Type host: not stated Comments: Keirans & Clifford (1984) did not list specimens of Amblyomma nodosum in their catalog of USNTC types.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF5D428DFF23FCF0FC2FFBB9.taxon	materials_examined	Type depository: undetermined: holotype (female) Type locality: Gotebörg, Sweden Type host: Hypudaeus glareolus Current status: a synonym of Ixodes trianguliceps (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF5D428DFF23FE10FC2FFD59.taxon	materials_examined	Type depository: MRSN: holotype (female) Type locality: Valsavaranche, Italy Type host: Chionomys nivalis Current status: a synonym of Ixodes trianguliceps (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF5D428DFF23FBD0FB4FFB41.taxon	discussion	Comments: Estrada-Peña et al. (2017) and other authors treat Dermacentor niveus as invalid.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF5D428DFF23FAF8FBBEF931.taxon	materials_examined	Type depository: ZIAC: lectotype (male) (Filippova 2008) Type locality: Kizlyar District, Province of Tersk, Daghestan, Russia Type host: not stated Current status: a synonym of Dermacentor niveus (Filippova 1997) Comments: controversy attends the name Dermacentor niveus daghestanicus because some authors, such as Estrada-Peña et al. (2017), treat this name as a synonym of Dermacentor marginatus, while others, such as Kolonin (2009), have treated it as a synonym of Dermacentor daghestanicus. Filippova (2008) listed Dermacentor niveus daghestanicus as a synonym of Dermacentor niveus and her opinion is followed here.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF5C428CFF23FF79FB24FE79.taxon	materials_examined	Type depositories: USNTC: holotype (male) and paratypes (males, females, larvae, and a nymph?); ORSTOM, OVI and VRLH: paratypes (males, females, larvae, and a nymph?) Type locality: Bulawayo, Matebeleland, Zimbabwe Type host: Erinaceus frontalis Comments: it is uncertain where the paratype nymphs of Haemaphysalis norvali are deposited.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF5C428CFF23FE10FE11FCBD.taxon	materials_examined	Type depository: BMNH: a lectotype (male) and paralectotypes (males and females) were designated by Roberts (1963) Type locality: Sattelberg, Huon Gulf, Papua New Guinea Type host: Perameles sp.? [sic]	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF5C428CFF23FCACFC6DFBD5.taxon	materials_examined	Type depository: undetermined: syntypes (males) Type locality: Merauke, New Guinea, Indonesia Type host: wild pig Current status: Haemaphysalis novaeguineae Krijgsman & Ponto, 1932, is a synonym of Haemaphysalis bancrofti but also a preoccupied name. Therefore, Taylor (1944) named it Haemaphysalis meraukensis, while Kohls (1948) altered the Krijgsman & Ponto name to Haemaphysalis krijgsmani.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF5C428CFF23FB44FBD4FA35.taxon	materials_examined	Type depository: BMNH and ENV: syntypes (males and a female) (Nuttall & Warburton 1915, Keirans & Hillyard 2001) Type locality: Tebessa, Algeria Type host: Erinaceus sp. Current status: a synonym of Haemaphysalis erinacei (Keirans & Hillyard 2001).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF5C428CFF23FA24FBADF915.taxon	materials_examined	Type depository: ZIAC: a lectotype (male) and paralectotypes (males and females) (Filippova 2008) Type locality: Biyuk-Lambat, Ukraine Type host: Erinaceus europeus Current status: a synonym of Haemaphysalis erinacei (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF5C428CFF23F904FBADF875.taxon	materials_examined	Type depository: ZIAC: a lectotype (male) and paralectotypes (males and females) (Filippova 2008) Type locality: Vakhsh Village, Tajikistan Type host: hedgehog Current status: a synonym of Haemaphysalis erinacei (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF5F428FFF23F8C4FE58F87E.taxon	materials_examined	Type depository: ZMB: holotype (female) (Moritz & Fischer 1981) Type locality: Pará, Brazil	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF5F428FFF23FC3DFDD1FB09.taxon	materials_examined	Type depository: not stated: holotype (male) Type locality: Sanddown, Zimbabwe	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF5F428FFF23FF79FD89FE21.taxon	materials_examined	Type depositories: BMNH and ZMB: paratypes (males and females) (Moritz & Fischer 1981, Keirans & Hillyard 2001) Type localities: Bagamojo and Dar es-Salaam, Tanzania; Umtali, Zimbabwe Type hosts: land tortoises and Varanus sp.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF5F428FFF23FE59FD68FDC9.taxon	materials_examined	Type depository: ZIAC: lectotype (male) (Filippova 2008) Type locality: River Kobdo Basin, Mongolia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF5E428EFF23FA49FE14F9F9.taxon	materials_examined	Type depository: unknown: syntypes (males and females) Type locality: California, USA	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF5E428EFF23FE58FCF0FCBC.taxon	materials_examined	Type depository: ZSM: holotype (female) Type locality: Funchal, Madeira Island, Portugal Type host: phaisan (Neumann, 1908) Current status: the name Ixodes obscurus was preoccupied and replaced by Ixodes nigricans in Neumann (1908), but the latter name is a synonym of Ixodes ricinus (Camicas et al. 1998). Comments: the type host was not included in the original description, but named in Neumann (1908), where the name Ixodes obscurus was replaced by Ixodes nigricans.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF594289FF23FD5CFBEBFC61.taxon	materials_examined	Type depositories: ZMB and ZHS: syntypes (males and females) Type locality: Zanzíbar, Tanzania Type host: Mabuja striata Current status: a synonym of Amblyomma latum (Guglielmone & Nava 2014). Comments: Santos Dias (1993) treated Aponomma ochraceum as a valid name.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF594289FF23FC19FEA6FA34.taxon	materials_examined	Type depositories: ZMB and ZSH?: syntypes (males and females) (Walker et al. 2000) Type locality: Damaraland, Namibia Type host: Lepus timidus and cow Comments: Neumann (1901) maintained that all specimens from his description of Rhipicephalus oculatus were deposited in the ZMB. Neumann (1901) also stated that one female tick was found at Kilosa, Tanzania, but Keirans et al. (1993) believed that this locality is incorrect for Rhipicephalus oculatus, adding that this specimen, which is lost, may, in fact, be Rhipicephalus pravus. Keirans et al. (1993) also said that from the five specimens Neumann (1901) used to describe this species, only a male and a female exist in the ZMB, numbers confirmed by Moritz & Fischer (1981).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF584288FF23FE58FC1CFCBD.taxon	materials_examined	Type depository: ZMB?: holotype (male) Type locality: “ Berliner Aquarium, ” Germany Type host: Coluber corrais Current status: a name incertae sedis (Guglielmone & Nava 2014, Guglielmone et al. 2017). Comments: Schulze (1932 b) stated that the holotype would be transferred to the ZMB, but no specimen of Aponomma omissum is listed in the ZMB type catalog by Moritz & Fischer (1981). Schulze (1932 b) also indicated that the type host was infested in America, but this is unproven (Guglielmone et al. 2017).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF584288FF23FCACFD18FB9D.taxon	materials_examined	Type depository: JMC: syntypes (males), but lost (Kaufman 1972) Type locality: collected in Bonn, Germany, but this is not the locality of origin Type host: Dipsas sp. Current status: a name incertae sedis (Kaufman 1972).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF58428BFF23F8C3FE4EFEE9.taxon	materials_examined	Type depositories: ZIAC: holotype (female); KC: a paratype (female) Type locality: Shonla, Vietnam, but corrected to Africa in Kolonin (2003) Type host: Python molorus bivittatus Current status: a synonym of Africaniella orlovi and its original name, as discussed in Hornok et al. (2020), who also established the current status of Aponomma orlovi. Comments: this tick was originally treated as found in Vietnam (Oriental Zoogeographic Region), but this was reassessed by Kolonin (2003), who concluded that the type locality is an undetermined locality in Africa (Afrotropical Zoogeographic Region).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF5B428BFF23FA00FB4BF8E5.taxon	materials_examined	Type depositories: ZMA: a lectotype (male) and paralectotypes (males), BMNH: a paralectotype (male) (Santos Dias 1993, Keirans & Hillyard 2001). The lectotype and paralectotypes were designated by Santos Dias (1993) Type locality: Fakfak, New Guinea, Indonesia Type host: Zaglossus bruyni nigroaculeatus Current status: a synonym of Bothriocroton oudemansi and its original name (Beati et al. 2008). The current status of Aponomma oudemansi, as Bothriocroton oudemansi, was also established by Beati et al. (2008). Comments: Keirans & Hillyard (2001) listed the type locality as being in Papua New Guinea.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF5B428BFF23FE81FDC0FDB1.taxon	materials_examined	Type depositories: MCZ: holotype (male) and paratypes (males and a female); USNTC: paratypes (males) Type locality: Doi Angka, Thailand	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF5B428BFF23FB1CFB92FA69.taxon	materials_examined	Type depository: undetermined: syntypes (males and females) Type localities: El-Merg and Tolmetta, Libya Type host: unknown Current status: a synonym of Haemaphysalis parva (Morel (1963, Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF5A428AFF23FE35FE6BFD2D.taxon	materials_examined	Type depository: ZMB: syntypes (males) (Moritz & Fischer 1981) Type locality: Mexico	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF5A428AFF23F8BFFBC0F87E.taxon	materials_examined	Type depository: not stated, but probably in the IOC (Guglielmone et al. 2003): syntypes (a male and females) Type localities: Xerém and Sant’Anna de Macacú, State of Rio de Janeiro, Brazil	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF5A428AFF23FC18FE25FAD1.taxon	materials_examined	Type depository: ENV: syntypes (females) (Yamaguti et al. 1971) Type localities: Akita and Saga, Japan Type hosts: dog, horse and Lepus sp. Comments: Neumann (1899) included in his original description two females and one male collected in Amour and deposited in the ZSH, a depository included in Guglielmone et al. (2014), but Neumann (1911 a) listed only Japan within the range of Ixodes ovatus, and apparently considered ticks from Amour to be, in fact, Ixodes ricinus. Other authors have treated these specimens as probably Ixodes persulcatus or Ixodes nipponensis, as discussed in Guglielmone et al. (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF5A428AFF23FA48FF43F881.taxon	materials_examined	Type depositories: USNTC: holotype (female) and paratypes (males and females); DEP, DEEZ, DEUA and MCZ: paratypes (males and females) Type locality: Huntsville, Arkansas, USA Type host: dog Current status: a synonym of Ixodes scapularis (Keirans & Clifford 1978, Camicas et al. (1998). p	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6542B5FF23F94CFC00F8E5.taxon	materials_examined	Type depository: GM: syntypes (Trapido et al. 1964) Type locality: “ Ramoi, ” New Guinea, most probably an Indonesian locality	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6542B5FF23FD39FCC8FC29.taxon	materials_examined	Type depository: FMNH: holotype (male) and paratypes (males and females); PNM and USNTC: paratypes (males and females) Type locality: Puerto Princesa, Palawan Island, Philippines	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6442B4FF23FE10FAE5FD59.taxon	discussion	Comments: Barker et al. (2024) transferred Amblyomma papuana to the new genus Cryptocroton and treated this species as found only in Australia and Papua New Guinea; however, the type locality is in Indonesia.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6442B4FF23FB8DFDB3FAFD.taxon	materials_examined	Type depositories: USNTC: holotype (male) and paratypes (males and females); MNHN and ORSTOM: paratypes (males and females) Type locality: Nanga-Eboko, Cameroon	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6442B4FF23FCF0FB96FB9D.taxon	materials_examined	Type depositories: USNTC: holotype (female) and paratypes (females and larvae); BM and RBINS: paratypes (females and larvae) Type locality: Ulu Gombak, 27 km north of Kuala Lumpur, Forest Reserve, Selangor, Malaysia Type host: Cheiromeles torquator Current status: a synonym of Ixodes kopsteini, as discussed in Kohls & Clifford (1968).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6742B7FF23FBD1FD91FAFD.taxon	materials_examined	Type depository: IBU: holotype (female) (Labruna et al. 2009) Type locality: Cotia, State of São Pailo, Brazil Type host: Coendou sp. Comments: Amblyomma parkeri was not listed as valid by Horak et al. (2002) and was treated as a doubtful species in Kolonin (2009), but it is a valid species.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6742B7FF23FEA4FE67FD2D.taxon	materials_examined	Type depositories: USNTC: holotype (male) and paratypes (males, females, and a nymphal pelt); BMNH, IM and VRC: paratypes (males and females) Type locality: Hyderabad, State of Telangana, India Type host: Felis chaus Comments: Haemaphysalis paraturturis was originally identified as Haemaphysalis bispinosa intermedia by Nuttall & Warburton (1915).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6642B6FF23F9FCFE6FF915.taxon	materials_examined	Type depository: ZSH: holotype (female) Type locality: Bolivia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6642B6FF23FF78FBC6FE05.taxon	materials_examined	Type depository: not stated: syntypes (females) Type locality: Mesa City, Arizona, USA Type host: jack rabbit Current status: a synonym of Dermacentor parumapertus (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6142B1FF23FE58FB05FD65.taxon	materials_examined	Type depository: MNHN: holotype (male) Type locality: Syria Type host: unknown Current status: a synonym of Haemaphysalis parva and its original name (Morel 1963, Camicas et al. 1998). The current status of Dermacentor parvus, as Haemaphysalis parva, was established by Morel (1963).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6142B1FF23FD14FB8FFBB9.taxon	materials_examined	Type locality: Friedland, Mecklenburg, Germany Type host: Passer montanus Current status: a synonym of Ixodes arboricola, as discussed in Haarløv (1962). Comments: according to the original description, the holotype was to be transferred to the ZMB, but no specimen of Ixodes passericola is listed in the catalog of ZMB types by Moritz & Fischer (1981).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6142B0FF23F928FCBDFF19.taxon	materials_examined	Type depositories: CNCDGS: holotype (male) and paratypes (males and females); USNTC: paratypes (males and a female) Type locality: Villeta, Department of Cundinamarca, Colombia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6042B0FF23FA49FCFDF985.taxon	materials_examined	Type depository: ZIAC: a lectotype (male) and a paralectotype (female) (Filippova 2008) Type locality: “ Khantan Canyon, Province of Syr-Darya ”	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6342B3FF23FD38FD98FC0D.taxon	materials_examined	Type depositories: USNTC: holotype (male) and a paratype (female); MCZ and GML: paratypes (males) (Dunn 1933, Guglielmone et al. 2014) Type locality: Miraflores, Panamá, Panama	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6342B3FF23FF79FBBAFEE9.taxon	materials_examined	Type depository: ZIAC: holotype (female) (Filippova 2008) Type locality: Imanskii Forestry, Dalnerechensky District, Primorsky Territory, Russia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6242B2FF23FE80FDE4FD2D.taxon	materials_examined	Type depository: not stated: syntypes (females) Type locality: Cambridge, England, Great Britain Type host: Syrrhaptes paradoxus Current status: a nomen dubium (Guglielmone & Nava 2014). The original name was corrected to Haemaphysalis peregrina by Neumann (1911 a).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6242B2FF23FAF8FCEDF931.taxon	materials_examined	Type depository: MCZ: syntype (male) Type locality: South America? [sic] Type host: not stated Current status: a nomen dubium (Fairchild et al. 1966, Guglielmone & Nava 2014), but treated as valid, under the name Amblyomma perpunctatum in Santos Dias (1961) and Camicas et al. (1998); type comparison should prove useful in settling this difference (Guglielmone et al. 2021). Comments: the MCZ website notes that the vial label bears the statement “ appears to be = Amblyomma geayi ” (http: // mczbase. mcz. harvard. edu accessed 2 July 2022).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6D42BDFF23FD5CFC2AFC0D.taxon	materials_examined	Type locality: Vardvan Maru River, Kashmir, India Type host: unknown Current status: a synonym of Ixodes kashmiricus and its original name. The current status of Ixodes persulcatus kashmiricus, as Ixodes kashmiricus, was established by Filippova (1969).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6D42BDFF23FC3CFE42FB6D.taxon	materials_examined	Type depository: BMNH: holotype (female) Type locality: Sri Lanka	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6C42BCFF23FE80FB65FCBD.taxon	materials_examined	Type depository: USNTC: holotype (female) and paratypes (males and females) Type locality: Kingsville, Texas, USA Type host: “ rabbit ” Current status: a synonym of Amblyomma inornatum (Cooley & Kohls 1944, Camicas et al. 1998). Comments. Cooley & Kohls (1939) stated that the holotype had been deposited in the USNTC under accession number 14069; however, in their catalog of USNTC types, Keirans & Clifford (1984) listed paratypes but not the holotype of Amblyomma philipi as being in that collection, including the specimen under accession number 14069. One of us (SN) examined the specimen under that number and found that its data agree with those of Cooley & Kohls (1939); accordingly, this tick is treated here as the holotype of Amblyomma philipi.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6C42BCFF23FCADFCF8FC45.taxon	materials_examined	Type depository: USNTC: holotype (female) and a paratype (female) Type locality: Sanganjima Island, Iwate Prefecture, Japan	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6C42BCFF23FAB0FD55FA4D.taxon	materials_examined	Type depository: BMNH: syntypes (males and females) Type locality: South America; and Bahia, Brazil	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6F42BFFF23FC88FCE8FB09.taxon	materials_examined	Type depository: VLM: holotype (male) (Walker et al. 2000) Type locality: “ Lumasse, ” Guijá District, Province of Gaza, Mozambique Type host: Paraxerus cepapi auriventris Current status: a name incertae sedis (Guglielmone & Nava 2014). Comments: Santos Dias (1950) did not mention a type depository for Rhipicephalus piresi, but Walker et al. (2000) stated that the holotype had been deposited in the VLM.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6F42BEFF23F8E1FB51FE05.taxon	description	Neumann, L. G. (1910) Sur quelques espèces d’Ixodidae nouvelles ou insuffisamment connues. Annales des Sciences Naturelles, Zoologie, Série 9, 12, 161 – 176. Type depository: MNHN: syntypes (males and a female) Type locality: Ivindo basin, Cameroon Type host: wild boar Current status: a synonym of Rhipicephalus complanatus and its original name. However, Rhipicephalus planus is a preoccupied name that was subsequently changed to the current name by Neumann (1911 b).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6E42BEFF23FD14FEB4FB9D.taxon	materials_examined	Type depository: BMNH: presumably a syntype (female) (Keirans & Hillyard 2001) Type locality: England, Great Britain (presumed) Type host: Riparia riparia Current status: a synonym of Ixodes lividus (preoccupied) and its original name, which was renamed Ixodes lividus by Koch (1844).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6E42BEFF23FB8CFC36FA35.taxon	materials_examined	Type depository: not stated: syntypes (males, females and nymphs) Type locality: Bayern, Germany Type host: Riparia sp. Current status: a synonym of Ixodes lividus (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6E42BEFF23FA24FC36F8AD.taxon	materials_examined	Type depository: not stated: syntypes (males, females and nymphs) Type locality: Brandenbrug and Mecklenburg, Germany Type host: Riparia sp. Current status: a synonym of Ixodes lividus (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6942B9FF23F9D9FE5EF881.taxon	materials_examined	Type depositories: USNTC: holotype (male) and paratypes (males, females nymphs and larvae); ZMB: a paratype (female) Type locality: Kyrgyzstan	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6942B9FF23FAD4FB50F9A1.taxon	materials_examined	Type locality: Cape of Good Hope, South Africa Type host: Cynhyaena picta Current status: a synonym of Amblyomma hebraeum (Neumann 1911 a, Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6842B8FF23FC40FEE0FAFD.taxon	materials_examined	Type depositories: BMNH, ZMB: syntypes (males and females) (Walker et al. 2000) Type localities: Damaraland, Namibia?; Transvaal, South Africa?, and southern Massai steppe and Mpapua, Tanzania Type hosts: Oryx-Antelopes (Moritz & Fischer 1981) Comments: Walker et al. (2000, page 33) follow Dönitz (1910) in listing Demararaland (Namibia), Transvaal (South Africa), and Massai steppe (Tanzania) as type localities, but in their map 47 (page 357) the type locality is indicated as being in Tanzania, while Namibia and South Africa are treated as countries where Rhipicephalus near pravus is present, but not bona fide Rhipicephalus pravus. The BMNH and ZMB contain type specimens collected in Tanzania.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6842BBFF23F89BFD50FF7D.taxon	description	(Acari: Ixodidae) in the H. (Rhipistoma) spinulosa subgroup, parasites of carnivores and hedgehogs in Africa. Systematic Parasitology, 96, 485 – 509. Type depository: USNTC: holotype (male) and paratypes (males and females) Type locality: Paloich, Upper Nile, South Sudan	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6B42BBFF23FD38FE1EFC0D.taxon	materials_examined	Type depository: USNTC: holotype (female) and paratypes (males and females); MCZ: paratypes (female) Type locality: Kisenyi, Rwanda	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6A42BAFF23FA24FD15F8AD.taxon	materials_examined	Type locality: Shantou, China Type host: Zamenis mucosus Current status: a synonym of Amblyomma pattoni (Kaufman 1972, Camicas et al. 1998). Comments: according to the original description, the holotype was to be transferred to the ZMB, but Aponomma pseudolaeve is not included in the ZMB type catalog by Moritz & Fischer (1981). Teng & Jiang (1991) and Sun et al. (2016) considered Amblyomma pseudolaeve valid.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6A42BAFF23FEC9FC7BFDB1.taxon	materials_examined	Type depositories: USNTC: holotype (male) and paratypes (males, females and nymphs); BM, BMNH and FMNH: paratypes (males and females) Type locality: Mount Kamatis, San Luis, Quezon, Luzon, Philippines	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6A42BAFF23FB8CFEADFA35.taxon	materials_examined	Type depository: IP: holotype (nymph) Type locality: India Type host: red squirrel Current status: a synonym of Ixodes petauristae, as stated in Sénevet (1970). Comments: the holotype was originally classified as Ixodes holocyclus collected in India by Neumann (1899) (Sénevet 1969).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF6A42BAFF23F8C4FB9CF87E.taxon	description	See capensis pseudolongus Santos Dias, 1953, Rhipicephalus Comments: Rhipicephalus pseudolongus is not treated as valid by Camicas et al. (1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7542A5FF23FF79FBF9FE2E.taxon	materials_examined	Type depositories: USNTC: holotype (female); INTA: paratypes (males, females and nymphs), MCN: a paratype (female) Type locality: Rivadavia, Department of Rivadavia, Province of Salta, Argentina	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7542A5FF23FCBDFEF2FB21.taxon	materials_examined	Type depository: OVI?: holotype (female) Type locality: Marion Island, Indian Ocean Type host: Pterodroma mollis Current status: a synonym of Ixodes kerguelenensis (Wilson 1970). Comments: Walker (1991) stated that there is no evidence that the holotype of Ixodes pterodromae was deposited in the OVI.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7542A5FF23FB5FFB69FA43.taxon	materials_examined	Type depository: ZMB: syntypes (males) (Moritz & Fischer 1981) Type localities: Arusha and Uru, Tanzania, and Lake Jipe on the border of Kenya and Tanzania Type host: unknown Current status: a synonym of Rhipicephalus pulchellus and its original name (Neumann, 1897). The current status of Dermacentor pulchellus, as Rhipicephalus pulchellus, also appears in Neumann (1897).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7542A5FF23F9FDFB13F880.taxon	materials_examined	Type depository: MNHN: syntypes (males and females) Type locality: Cayenne, French Guiana Type hosts: Bufo agua, Spilotes variabilis Current status: a synonym of Amblyomma dissimile (Neumann 1911 a, Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7442A4FF23FE34FC95FC29.taxon	materials_examined	Type depositories: QM: holotype (male) and paratype (female); ARI: a paratype (male) Type locality: Rockhampton, Queensland, Australia Type host: Varanus sp. Current status: a synonym of Amblyomma flavomaculatum Kaufman (1972, under the genus Aponomma) (Guglielmone & Nava 2014). Comments: Amblyomma flavomaculatum is an Afrotropical species and Aponomma pulchrum was found in Australia (Australasian Zoogeographic Region). Kaufman (1972) considered the Australian origin of Aponomma pulchrum accidental. Nevertheless, Santos Dias (1993) treated this name as valid, and Camicas et al. (1998) believed it to be a synonym of Amblyomma gervaisi, under the genus Aponomma.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7442A4FF23FB1DFCBFFA4D.taxon	materials_examined	Type depositories: USNTC: holotype (male) and paratypes (males and females); ORSTOM: paratypes (female and males) Type locality: Adiopodoumé, 43 km from Abidjan, Ivory Coast	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7442A4FF23FC41FF1DFB6D.taxon	materials_examined	Type depository: ZMA: holotype (male), identified as “ type 1 ” in the original description Type locality: Maralbashi, Xinjiang Uyghur Autonomous Region, China Type host: not stated Comments: Zahler et al. (1997) considered it probable that Rhipicephalus pumilio is a synonym of Rhipicephalus rossicus.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7742A7FF23FAB0FAD2F931.taxon	materials_examined	Type depository: USNTC: holotype (male) and paratypes (males and females); OVI?: paratypes (a male and a female) Type locality: “ Abudwac ” near Ghelinsor, Somalia Type host: camel Comments: Guglielmone et al. (2014) stated that no paratypes of Hyalomma punt were found in the OVI.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7642A6FF23FA25FB8AF949.taxon	materials_examined	Type depository: IZAS: holotype (male) and paratypes (males and females) Type locality: Huangyuan, Qinghai Province, China Type host: goat Comments: Camicas et al. (1998), Guglielmone et al. (2009, 2014) and Guglielmone & Nava (2014) referred to this tick as Haemaphysalis quinghaiensis, but this is an incorrect spelling (Wen et al. 2016).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7642A6FF23FCF0FC35FBB9.taxon	materials_examined	Type depository: undetermined: holotype? (female) Type locality: Denmark Type host: Procellariae sp. Current status: a synonym of Ixodes uriae, as discussed in Zumpt (1952).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7642A6FF23FF79FF12FE79.taxon	description	See also bursa pusillus Gil Collado, 1936, Rhipicephalus Type depository: USNTC: neotype (male) designated by Estrada-Peña et al. (2018) Type locality: Pina del Ebro, Spain Type host: the neotype specimen was collected from a burrow of Oryctolagus cuniculi Comments: this tick was originally named Rhipicephalus bursa pusillus, but the neotype was named Rhipicephalus pusillus.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7142A1FF23FAD4FB5CF985.taxon	materials_examined	Type depository: MNHN: syntypes (males and females) Type locality: “ Congo ” Type host: Bos brachyceros Current status: a synonym of Amblyomma splendidum (Robinson 1926, Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7142A1FF23F9B4FB6DF811.taxon	materials_examined	Type depository: BMNH: holotype (female) (Keirans & Hillyard 2001) Type locality: Frontera, State of Tabasco, Mexico? Type host: Ateles geoffroyi Current status: a synonym of Amblyomma breviscutatum (Santos Dias 1956 a, 1958 a, Camicas et al. 1998, Guglielmone & Nava 2014), although Keirans & Hillyard (2001) listed Amblyomma quasicyprium as a synonym of Amblyomma cyprium. Comments: the locality label is considered to be incorrect, following Santos Dias (1956 a).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7042A0FF23FC89FC28FBB9.taxon	materials_examined	Type depositories: USNTC: holotype (female) and paratypes (females); BMNH: a paratype (female) Type locality: Kanglatongbi, 19 km from Imphal, State of Manipur, India	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7342A3FF23FCF1FF01FC45.taxon	materials_examined	Type depository: ZIAC: holotype (male) (Filippova 1983) Type locality: Xinjiang, China Type host: not stated Comments: the type locality of Dermacentor raskemensis was uncertain until Filippova (1983) clarified this matter.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7342A3FF23FF79FD23FEE9.taxon	materials_examined	Type depository: USNTC: holotype (female) Type locality: Massif da Tsaratanana, Madagascar	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7342A3FF23FBF4FC2BFAD1.taxon	materials_examined	Type depositories: ENV: syntypes (a male and females) and ZMB? (unknown developmental stage) (Nuttall & Warburton 1911 as “ types ”) Type locality: “ Congo ” Type host: Hyrax sp. Comments: no specimens of Ixodes rasus are listed by Moritz & Fischer (1981) as held in the ZMB. Guglielmone et al. (2020) stated that the morphological and molecular definitions of Ixodes rasus need reevaluation, and redescriptions of the type specimens should prove valuable in this regard.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7342A3FF23FA48FA7FF915.taxon	materials_examined	Type depository: ZMB: syntypes (a male, female and nymphs) (Moritz & Fischer 1981) Type localities: Nyika, Malawi or Zambia; Amani, Nguela and Usambara, Tanzania; Bulwe, Zambia. Type hosts: Bdeogale sp., dog, Cephalophus parvus, Herpestes sp., Potomochoerus sp. and wild boar Current status: a synonym of Ixodes cumulatimpunctatus and its original name (Camicas et al. 1998). The current status of Ixodes rasus cumulatimpunctatus, as Ixodes cumulatimpunctatus, was provided by Lariviéri et al. (1957).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7342A3FF23F90AFAFBF853.taxon	materials_examined	Type depository: ZMB: a syntype (female) (Moritz & Fischer 1981) Type locality: Alen and Rio Benito, Equatorial Guinea Type host: Cricetomys gambianus Current status: a synonym of Ixodes rasus (Camicas et al. 1998). Comments: this tick was described from three females but only one syntype female is held in the ZMB.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7242A2FF23FA6CFD5CF8AD.taxon	materials_examined	Type depository: LC: but lost Type locality: undetermined Type host: cattle and dogs Current status: a synonym of Ixodes ricinus, as proposed by Neumann (1901) and accepted by the International Commission on Zoological Nomenclature with Neumann as first reviser. Comments: the name Acarus reduvius and its synonyms have been used in the past for related and unrelated tick species worldwide (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7242A2FF23FF78FD6BFE21.taxon	materials_examined	Type depositories: USNTC: holotype (male) and paratypes (a male and females); AM and BMNH: paratypes (females) Type locality: Dobodura, Papua New Guinea	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7242A2FF23FBD0FC20FAFD.taxon	materials_examined	Type depository: ZIAC: holotype (female) (Filippova 2008) Type locality: steppes of southern trans-Volga to north of Novaya Kazanka, Russia Type host: Lagurus lagurus Current status: a synonym of Ixodes laguri and its original name (Filippova 2008). The current status of Ixodes redikorzevi lagurae, as Ixodes laguri, was established by Olenev (1931).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7242A2FF23FCF0FAA4FBB8.taxon	materials_examined	Type locality: Talysh, Azerbaijan Type host: Emberiza schoeniclus Current status: a synonym of Ixodes redikorzevi (Filippova 1977). Comments: Camicas et al. (1998) listed Ixodes redikorzevi emberizae as a synonym of Ixodes acuminatus.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7D42ADFF23FB69FB70F9F9.taxon	description	Neumann, L. G. (1897) Révision de la famille des ixodidés (2 e mémoire). Mémoires de la Société Zoologique de France, 10, 324 – 420. Type depository: MNHN: syntypes (males and females) Type locality: 40 km north of Tehran, Iran Type host: sheep and camels Current status: a synonym of Dermacentor niveus and its original name (Camicas et al. 1998). The current status of Dermacentor reticulatus niveus, as Dermacentor niveus, was provided by Yakimov (1923).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7D42ADFF23F991FB4CF83D.taxon	materials_examined	Type depositories: OVI?: holotype (female); unknown depository: a paratype (female) (Arthur 1959) Type locality: Howiesons Poort, Province of Eastern Cape, South Africa Type host: Rhabdomys pumilio Comments: there is no evidence that the holotype specimen was deposited in the OVI (Walker 1991), and Arthur (1959) provided no information about the depository of the paratype of Ixodes rhabdomysae.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7C42ACFF23FD5CFE9FFC45.taxon	materials_examined	Type depository: ZMB: holotype (male) (Moritz & Fischer 1981) Type locality: “ Momberg, ” German East Africa (Burundi? Mozambique? Rwanda? Tanzania?) Type host: undetermined Current status: a synonym of Dermacentor rhinocerinus (Camicas et al. 1998). Comments: Mortiz & Fischer (1981) referred to Dermacentor rhinocerinus schillingsi as Dermacentor rhinocerotis schillingsi, a lapsus.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7C42ACFF23FBF4FB79FAA5.taxon	materials_examined	Type depository: unknown: syntypes (males) Type locality: Cape of Good Hope, South Africa Type host: rhinoceros Current status: a synonym of Amblyomma rhinocerotis and its original name (Camicas et al. 1998). The current status of Acarus rhinocerotis, as Amblyomma rhinocerotis, was established by Koch (1844).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7F42AFFF23FB44FF04FA69.taxon	description	See ricinus Linnaeus, 1758, Acarus Comments: Ixodes ricinus is a Palearctic species, but this name was used for many years for related and unrelated species, and it is still sometimes applied to tick specimens collected outside the Palearctic (Guglielmone et al. 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7F42AFFF23FCACFD60FBD5.taxon	materials_examined	Type depository: LC: (developmental stages undetermined) but lost Type locality: undetermined Type host: cattle and dogs Current status: a synonym of Ixodes ricinus and its original name. The current status of Acarus ricinus, as Ixodes ricinus, was established by Latreille (1804).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7F42AFFF23FA00FDB2F8E5.taxon	materials_examined	Type depositories: IBU: syntypes (males and females) (Barros-Battesti & Knysak 1999); Keirans & Clifford (1984) listed a syntype named as “ cotype ” (unspecified developmental stage) deposited in the USNTC Type locality: Cotia, São Paulo, Brazil Type host: Mazama simplicicornis Current status: a synonym of Ixodes fuscipes, as discussed in Labruna et al. (2020). Comments: Keirans & Clifford (1984) erroneously listed the specimen of Ixodes ricinus aragaoi deposited in the USNTC as a synonym of Ixodes affinis.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7F42AEFF23F894FB30FECD.taxon	materials_examined	Type depository: BMNH: a lectotype (male) and paralectotypes (males and females) were designated by Keirans & Brewster (1981) Type locality: Izmir, Turkey Type host: domestic goat Current status: a synonym of Ixodes gibbosus and its original name, as discussed in Keirans & Brewster (1981). The current status of Ixodes ricinus gibbosus, as Ixodes gibbosus, was established by Schulze (1927).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7E42AEFF23FD5CFBA7FC0D.taxon	materials_examined	Type depositories: LBMA and LIDG: syntypes (males and females), but probably lost Type locality: “ La Sierra, ” Department of Rocha, Uruguay Type host: Bos taurus Current status: a synonym of Ixodes fuscipes, as discussed in Labruna et al. (2020).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7E42AEFF23FE7CFC61FD2D.taxon	materials_examined	Type depository: AC: holotype (female) Type locality: Cluj, Romania Type host: cattle Current status: a synonym of Ixodes ricinus (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7E42AEFF23FC3CFC47FB6D.taxon	materials_examined	Type depositories: USNTC: holotype (female) and paratypes (females); CNCDGS: a paratype (female) Type locality: Alto do Itatiaia, Serra do Itatiaia, Rio de Janeiro, Brazil	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7E42A9FF23F89CFCBFFF51.taxon	materials_examined	Type depository: BMNH: a lectotype (male) and paralectotypes (males and a female) were designated by Keirans & Brewster (1981) Type locality: Komodo Island, Lesser Sunda Islands, Indonesia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7942A9FF23FD14FD2BFC0D.taxon	materials_examined	Type depository: MZNH: syntypes (a male, females and nymphs) (Barros-Battesti et al. 2007) Type localities: Great Falls and Demerara, Guyana	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7942A9FF23F990FECFF811.taxon	materials_examined	Type depository: AISP: syntypes (a male? and a nymph?) Type locality: Nowaya Zemlya Archipelago, Russia Type host: walrus Current status: a nomen nudum in Guglielmone & Nava (2014). Comments: the type depository for Dermacentor rosmari is erroneously treated as unknown in Guglielmone & Nava (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7942A9FF23FAF8FB3BF9F8.taxon	materials_examined	Type depository: ZMB: syntypes (a male and females) (Moritz & Fischer 1981) Type locality: Saint Thomas, West Indies Type host: unknown Current status: a synonym of Rhipicephalus annulatus in Guglielmone & Nava (2014). Comments: Rhipicephalus annulatus is not found in the West Indies, and the above name should be confirmed as a synonym of Rhipicephalus annulatus through inspection of the syntypes of Haemaphysalis rosea.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7842A8FF23FF78FE27FE5D.taxon	materials_examined	Type depository: ZMB?: syntypes (females) (Schulze 1935 a) Type locality: New Zealand Type host: not stated Current status: a nomen nudum (Guglielmone et al. 2017). Comments: in the ZMB type catalog by Moritz & Fischer (1981), no specimens of Ixodes rossianus are listed. Ixodes rossianus in Schulze (1935 a) was treated as a synonym of Ixodes holocyclus by Roberts (1960), Guglielmone & Nava (2014) and others.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7842A8FF23FDCCFDC9FC60.taxon	materials_examined	Type depositories: BMNH: syntypes (a male and a female) (Keirans & Hillyard 2001), ZIAC: a syntype (male) (Filippova 1996, 2008) Type locality: Saratov, Russia Type host: dog Comments: a neotype (male) of Rhipicephalus rossicus was designated by Filippova (1996) and repeated in Filippova (2008) for a tick collected from vegetation in the Province of Volgograd, Russia. However, Keirans & Hillyard (2001) rejected this neotype designation because syntypes of Rhipicephalus rossicus still exist, and this specimen is treated here as a syntype. Zahler et al. (1997) considered it probable that Rhipicephalus pumilio is a synonym of Rhipicephalus rossicus.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7842ABFF23F904FBD3FF7D.taxon	materials_examined	Type depository: ZMB?: holotype (male) Type locality: Singer Island, Indonesia Type host: cattle Current status: a name incertae sedis (Guglielmone & Nava 2014). Comments: Minning (1934) stated that the holotype was deposited in the ZMB, but no specimen of Boophilus rotundiscutatus was included in the list of ZMB types by Moritz & Fischer (1981).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7B42ABFF23FEEDFD2DFD95.taxon	materials_examined	Type depositories: IPP and USNTC?: syntypes (males) (Hoogstraal et al. 1966) Type locality: Cochin China, now “ Tam Ky Province, ” Vietnam (Hoogstraal et al. 1966) Type host: deer Comments: no specimens of Haemaphysalis roubaudi were included in the list of USNTC types by Keirans & Clifford (1984). “ Tam Ky Province ” in Hoogstraal et al. (1966) is probably a lapsus for “ Qu ảng Tín Province, ” currently included in the Province of Qu ảng Nam.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7B42ABFF23FAB1FC1BF9DC.taxon	materials_examined	Type depository: ZMB: a lectotype (male) and a paralectotype (male) (Moritz & Fischer 1981) Type locality: Senegal Type host: not stated Comments: a paralectotype female of Hyalomma rufipes is also listed in Moritz & Fischer (1981), but Apanaskevich & Horak (2008 b) determined that this is a specimen of Hyalomma truncatum.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7B42AAFF23F94CFD0DFF7D.taxon	materials_examined	Type depository: OVI: syntypes or paratypes (males, females, nymphs and larvae) (Apanaskevich & Horak 2006) Type locality: Karoo, South Africa Type host: unknown Current status: a synonym of Hyalomma glabrum and its original name, as discussed in Apanaskevich & Horak (2006). The current status of Hyalomma rufipes glabrum, as Hyalomma glabrum, was established first by Theiler (1956) and definitively by Apanaskevich & Horak (2006). Comments: syntypes or paratypes are from laboratory-reared material based on adult specimens collected in the South African Karoo (Apanaskevich & Horak 2006).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7A42AAFF23FDCDFE45FCBD.taxon	materials_examined	Type depository: USNTC: syntypes (females and nymphs) (Keirans & Clifford 1984, as “ types ”) Type locality: Germany	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7A42AAFF23FEECFC00FE5D.taxon	materials_examined	Type depository: ZMB: syntypes (a male and females) (Moritz & Fischer 1981) Type locality: Germany Type host: not stated Current status: a nomen dubium, as explained in Estrada-Peña et al. (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF7A42D5FF23F8E1FF46FEE9.taxon	materials_examined	Type depository: ZMB: holotype (female) (Moritz & Fischer 1981) Type locality: Damietta, Egypt Type host: unknown Current status: a name incertae sedis in Guglielmone & Nava (2014), but reinstated by Šlapeta et al. (2023), and treated as provisionally valid here. s	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0542D5FF23FB69FA86FAD1.taxon	materials_examined	Type depository: BMNH: holotype (male) (Keirans & Hillyard 2001) Type locality: Chathurangapara, Kottayam District, State of Kerala, India Type host: Cervus unicolor Comments: the holotype of Haemaphysalis sambar was originally identified as Haemaphysalis campanulata.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0542D4FF23F8DCFE7BFF7D.taxon	materials_examined	Type depository: USNTC: neotype (male) (Nava et al. 2018) Type locality: Montpellier, France Type host: the holotype specimen was collected from the wall of a kennel Comments: Rhipicephalus sanguineus was originally named Ixodes sanguineus, but the neotype was described under the former name.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0542D5FF23FA48FF0DF8AD.taxon	materials_examined	Type depository: unknown Type locality: Gallia Type host: unknown Current status: a synonym of Rhipicephalus sanguineus and its original name, as discussed in Nava et al. (2018). The current status of Ixodes sanguineus, as Rhipicephalus sanguineus, was established by Koch (1844). Comments: a neotype of this species was designated under the name Rhipicephalus sanguineus by Nava et al. (2018).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0442D4FF23FD5DFC57FBB8.taxon	materials_examined	Type depository: BMNH?: syntypes (females) Type locality: Ega, Brazil, and Bolivia Type host: Tapirus Comments: Neumann (1906) listed two females deposited in the BMNH, but no specimens of Amblyomma scalpturatum are listed in the catalog of BMNH types by Keirans & Hillyard (2001). Labruna et al. (2005) stated that one of the males of Amblyomma incisum deposited in the BMNH is in fact Amblyomma scalpturatum. This specimen may be the holotype of Amblyomma scalpturatum according to Guglielmone & Nava (2014) and Guglielmone et al. (2014) but Amblyomma scalpturatum was described from two females.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0442D4FF23FEECFCA7FD2D.taxon	materials_examined	Type depository: ZMB?: holotype (male) (Kratz 1940) Type locality: Rio de Oro, Western Sahara Type host: unknown Current status: a synonym of Hyalomma impeltatum and its original name (Apanaskevich & Horak 2009). The current status of Hyalomma savignyi impeltatum, as Hyalomma impeltatum, was established by Kratz (1940). Comments: the holotype specimen is not listed as held in the ZMB (Moritz & Fischer 1981). See Hyalomma impeltatum for the designation of a neotype under that name.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0442D4FF23F973FED3F853.taxon	materials_examined	Type depositories: CZIICT: holotype (male) and paratypes (males, a female and a nymph); BMNH? and OVI: paratype (males) Type locality: Lake Ura, Province of Sofala, Mozambique Type host: Pterodromus tetradactylus Current status: a synonym of Rhipicephalus decoloratus (Keirans & Hillyard 2001, under the genus Boophilus). Comments: Keirans & Hillyard (2001) stated that paratypes of Rhipicephalus scheepersi apparently never arrived at the BMNH.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0742D7FF23FF79FC44FE05.taxon	materials_examined	Type depositories: BMNH and ZMB: syntypes (females) (Keirans & Hillyard 2001) Type locality: German East Africa (Burundi? Mozambique? Rwanda? Tanzania?) Type host: Colobus guereza Comments: Moritz & Fischer (1981) included nymphs in the syntype series of Ixodes schillingsi deposited in the ZMB, but Neumann (1901) described only the female of this species.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0742D7FF23FD14FEE9FB9D.taxon	materials_examined	Type depository: MNHN: holotype (female) Type locality: Algeria Type host: not stated Current status: a synonym of Rhipicephalus annulatus (Camicas et al. 1998, under the genus Boophilus). Comments: Minning (1934) designated as “ type ” a female diagnosed by L. G. Neumann as Rhipicephalus annulatus.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0742D7FF23FB8DFD91FAA5.taxon	materials_examined	Type depository: ZIAC: lectotype (male) and paralectotypes (males and a female) (Filippova 2008) Type locality: Seistan, East Khorasan, Iran	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0642D6FF23FB8DFC60FB41.taxon	materials_examined	Type depository: USNTC: a neotype (male) was designated by Nava et al. (2014 a) Type locality: Alegria Farm, Corumbá, Mato Grosso do Sul, Brazil	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0142D1FF23FD5CFBA1FC0D.taxon	materials_examined	Type depository: CNA: holotype (male) and a paratype (female) Type locality: Finca Germania, Chiapas, Mexico Type host: probably Didelphys mesoamericana tabascensis Current status: a synonym of Ixodes luciae (Jones et al. 1972, Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0142D1FF23FC3DFD35FB09.taxon	materials_examined	Type depository: NTCHU: a lectotype (male) and paralectotypes (males and females) were designated by Mumcuoglu et al. (2022) Type locality: Tirat HaCamel, Haifa District, Israel	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0042D0FF23FAD4FB4AF985.taxon	materials_examined	Type depository: IM: syntypes (females) Type locality: Abbottabad, Hazara District, Province of Khyber Pakhtunkhwa, Pakistan Type host: goat Current status: a synonym of Haemaphysalis sulcata (Filippova (1997, Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0042D0FF23FE58FADFFD59.taxon	materials_examined	Type depository: IOC: syntypes (nymphs) Type localities: Parque Nacional do Itatiaia, State of Rio de Janeiro, and Parque Nacional da Serra da Bocaina, State of São Paulo, Brazil Type hosts: Akodon serrensis, Delomys dorsalis, Delomys sublineatus, Nectomys squamipes and / or Trinomys gratiosus Current status: a nomen nudum (Venzal et al. 2008, Guglielmone et al. 2014, Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0042D3FF23F89CFC33FF7D.taxon	materials_examined	Type depository: USNTC: holotype (female) and paratypes (females, nymphs and larvae) Type locality: Kyangjin Ghyang, Langtang Valley, Rasuwa District, Nepal	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0342D3FF23FDA8FBB0FC91.taxon	materials_examined	Type depository: DZUP: holotype (unspecified developmental stage) Type locality: Bannu, Province of Khyber Pakhtunkhwa, Pakistan Type host: cattle Current status: a synonym of Rhipicephalus microplus (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0342D3FF23FC89FBC9FBB9.taxon	discussion	Comments: Kolonin (2009) treated Haemaphysalis shimoga as an invalid species.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0342D3FF23FA6CFB69F931.taxon	materials_examined	Type depositories: NSM: holotype (larva); NIAH: paratypes (larvae) Type locality: Doi Inthanon, Thailand Type host: Anousorex squamipes Comments: Camicas et al. (1998) and Kolonin (2009) did not treat Ixodes siamensis valid.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0342D3FF23FBD0FB85FAFD.taxon	materials_examined	Type depository: BMNH: syntype (a female) (Keirans & Hillyard 2001) Type locality: Sudan Type host: Hyaena sp. Current status: a synonym of Rhipicephalus praetextatus (Camicas et al. 1998, Walker et al. 2000). Comments: Neumann (1902) described Rhipicephalus shipleyi from several adult ticks allegedly deposited in the BMNH, but only a female specimen is currently held there (Keirans & Hillyard 2001).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0242D2FF23FF78FB0CFE21.taxon	materials_examined	Type depository: not stated: syntypes (females) Type localities: Montnedre, Charente-Maritime Department, and Dax, Landes Department, France Type hosts: Vannellus vannellus and Turdus sp. Current status: a synonym of Ixodes frontalis (Camicas et al. 1998, under the genus Scaphixodes).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0242D2FF23FE59FBE4FD59.taxon	materials_examined	Type depositories: USNTC: holotype (female) and paratypes (females, a nymph and a larva); TTU: paratypes (females and a nymph) Type locality: Parque Nahuelbuta, 25 km west of Angol, Chile Type host: Aconaemys fuscus Comments: Camicas et al. (1998) did not treat Ixodes sigelos as a valid name.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0242D2FF23FCF0FB76FB09.taxon	materials_examined	Type depositories: ZIAC: lectotype (female) and paralectotypes (females) (Filippova 2008), BMNH: a paralectotype (female) (Keirans & Hillyard (2001, as syntype). MCZ: a syntype (lost) (Keirans & Clifford 1978) Type locality: Unalaska Island, Aleutian Islands, Alaska, USA Type host: not stated Comments: only the female of Ixodes signatus was described by Birula (1895). Therefore, a nymph in the ZIAC named as paralectotype by Filippova (2008), and nymphs named as “ types ” in Nuttall & Warburton (1911) and Keirans & Clifford (1978) and deposited in the ZSH, do not qualify as belonging to the type series of Ixodes signatus. KLC, IR and RGR visited the MCZ in 2024 and the syntype was not found there.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0D42DDFF23FC19FBA7FB6D.taxon	materials_examined	Type depositories: USNTC: holotype (male) and a paratype (female); OSAL and OSU: paratypes (males and females) Type locality: Deschutes River State Recreation Area, Wasco County, Oregon, USA	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0D42DDFF23FF79FCCAFECD.taxon	materials_examined	Type depositories: USNTC: holotype (female); INTA: paratypes (females, nymphs and larvae) Type locality: Valle Colorado, Province of Jujuy, Argentina	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0D42DDFF23F9B5FE48F8E5.taxon	materials_examined	Type depositories: BMNH and ENV: syntypes (males and a nymph) (Nuttall & Warburton 1915 named, as “ types, ” Keirans & Hillyard 2001) Type locality: Madagascar Type host: Erinaceus sp.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0D42DCFF23F895FCC2FECD.taxon	materials_examined	Type depository: BMNH: syntypes (females and a nymph) Type localities: Shanghai, China; Kashmir Region (Arthur 1956) Type hosts: Rhinolophus ferrumequinum and Vespertilio sp. Comments: Neumann (1906) included a female collected in Gabon in his original description of Ixodes simplex, but Arthur (1956) treated this record as erroneous, stating that the tick was found in the Kashmir Region (China?, India? Pakistan?). Ixodes simplex has not been recorded from India and Pakistan; consequently, China is considered here to be the only country of origin (Guglielmone et al. 2023).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0C42DCFF23FE7CFBFCFD2D.taxon	materials_examined	Type locality: Mount Menengai, Great Valley Rift, Kenya Type host: unknown Current status: a synonym of Ixodes simplex (Filippova 1977). Comments: IR visited the FMNH in 2024 and the holotype was not found there.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0C42DCFF23FB1CFAD0FA4D.taxon	materials_examined	Type depository: ZMB: holotype (male) (Moritz & Fischer 1981) Type locality: South Africa Type host: not stated Comments: Guglielmone et al. (2020) stated that Rhipicephalus simus is poorly defined morphologically.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0C42DCFF23F9FCFBEBF881.taxon	materials_examined	Type depositories: ZMB: holotype (male) and paratypes (males and females); SAIMR: paratype (unspecified developmental stage); ZSH: paratype (male) (Walker et al. 2000) Type locality: Bismarckburg, Togo (Moritz & Fischer 1981) Type host: Equus caballus Current status: a synonym of Rhipicephalus senegalensis (Walker et al. 2000).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0C42DFFF23F8B8FA84FEB5.taxon	materials_examined	Type localities: Kibonoto, Steppe, Kilimanjaro, Tanzania Type host: Hystrix africaeaustralis Current status: a synonym of Rhipicephalus planus and its original name (Camicas et al. 1998, Walker et al. 2000). The current status of Rhipicephalus simus planus, as Rhipicephalus planus, was established by Morel (1969).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0F42DFFF23F990FB5DF8D9.taxon	materials_examined	Type depository: IAHVS: syntypes (males and females) (Zhang 1981, Keirans & Robbins 1999) Type locality: Fang County, Hubei Province, China Type hosts: cattle and goats Comments: Camicas et al. (1998) did not include this species in their list of ticks of the world.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0F42DFFF23FB44FF13F9F8.taxon	materials_examined	Type depository: not stated: syntypes (males and females); ZMB?: syntypes? (males) (Moritz & Fischer 1981) Type localities: “ Mooanshan ” (Moganshan) in Shangai District, Amoy in Fujian Province, Qingdao in Shandong Province, and Hong Kong, China; Miyazaki, Japan; “ Auping, ” Taiwan; and Tongking, Vietnam Type hosts: cattle, dog, goat, horse and water buffalo Current status: a name incertae sedis in Guglielmone & Nava (2014). Comments: Moritz & Fischer (1981) listed three males of the syntype series of Boophilus sinensis allegedly collected in Japan and held in the ZMB. However, Minning (1934) listed only eight female ticks found in that country; therefore, it is uncertain whether the male specimens deposited in the ZMB are true syntypes of Boophilus sinensis.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0F42DEFF23F877FDD8FF7D.taxon	materials_examined	Type depository: IZAS: holotype (female) and paratypes (males and females) Type locality: Fujian Province, China	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0942D9FF23FE58FD66FD65.taxon	materials_examined	Type depository: MHNH: holotype (male) Type locality: Paris, France Type host: Spilotes variabilis Comments: Amblyomma sparsum is an Afrotropical species, but its holotype was collected in the Palearctic on a Neotropical host (Guglielmone et al. 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0942D9FF23FD14FF0DFB9D.taxon	materials_examined	Type depositories: CI: holotype (male) and paratypes (a nymph and a larva); CG: a paratype (female) Type locality: Stephen Island, New Zealand Type host: Sphenodon punctatus Current status: a synonym of Archaeocroton sphenodonti and its original name (Guglielmone et al. 2020, 2023). The current status of Aponomma sphenodonti, as Archaecroton sphenodonti, was established by Barker & Burger (2018).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0942D9FF23FF78FA6AFE21.taxon	materials_examined	Type depositories: CNC: holotype (female) and paratypes (nymphs and larvae); USNTC?: paratypes (a female, nymph and larvae) Type locality: Harrison Bay, New Westminster District, British Columbia, Canada (Robbins & Keirans 1992) Type host: Sorex sp. Comments: paratype specimens of Ixodes soricis in the USNTC are considered lost in Keirans & Clifford (1978).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0942D9FF23F9FCFAEBF8AD.taxon	materials_examined	Type depository: OVI?: holotype (female) and paratypes (a male and a female) Type locality: Pietersburgh, South Africa Type host: Procavia capensis Comments: there is no evidence that types of Ixodes spinae were deposited in the OVI (Walker 1991).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0842D8FF23FAB1FE33F8D9.taxon	materials_examined	Type depository: BMNH?: lectotype (female) and a paralectotype (female) were designated by Hoogstraal (1964 b) Type locality: Entebbe, Uganda Type host: cane rat? Comments: Neumann (1906) stated that type specimens were deposited in the BMNH, naming Uganda as the type locality but providing no type host in his description of Haemaphysalis spinulosa. The type information above is from Hoogstraal (1964 b), who cited the specimen label in the BMNH, and this museum was confirmed as the depository of “ types ” by Nuttall & Warburton (1915). However, Keirans & Hillyard (2001) stated that there are no indications that specimens of this tick were received by the BMNH, and Tomlinson et al. (2018) were also unable to find the types of Haemaphysalis spinulosa in the BMNH. Tomlinson et al. (2018) cautioned that the morphological definition of Haemaphysalis spinulosa is uncertain, apart from the female specimen redescribed by Hoogstraal (1964 b). Tomlinson et al. (2018) extensively reexamined ticks classified as Haemaphysalis spinulosa that are deposited in the USNTC, finding that none of them represented bona fide specimens of this taxon.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0842DBFF23F878FE60FF7D.taxon	materials_examined	Type depository: undetermined: syntypes (males and females) Type locality: Gabon	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0B42DBFF23FC40FB8AFB09.taxon	materials_examined	Type depository: MCNB: holotype (male) Type locality: Las Palmas, Barcelona, Spain Type host: not stated Current status: a synonym of Hyalomma marginatum (Apanaskevich & Horak 2008 b).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0B42DBFF23FB20FBEAFA69.taxon	materials_examined	Type depository: not stated: syntypes (males and females) Type locality: Biscarosse, 25 km south of Arcachon, France Type host: horse Current status: a synonym of Hyalomma scupense (Apanaskevich et al. 2010).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0A42DAFF23FF78FD18FDB1.taxon	materials_examined	Type depositories: ZMB?: paralectotypes (females and a nymph) (Schulze 1935 a); USNTC: a lectotype (female) and a paralectotype (female) were designated by Apanaskevich & Schenk (2020). Type locality: Weyland Mountains, Province of Papua, Indonesia Type host: Phascogale sp. Comments: Schulze (1935 a) stated that type specimens of Ixodes steini had been deposited in the ZMB, but they are not listed in the catalog of ZMB types, and from the original description, some specimens are thought to be held in the USNTC (Apanaskevich & Schenk, 2020). In the event that type specimens of Ixodes steini are eventually found in the ZMB, they should be considered paralectotypes.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0A42DAFF23FDA8FB1FFC91.taxon	materials_examined	Type depository: undetermined: syntypes (a male and a female) Type localities: Dali, Sumatra and Tengger Mountains, east Java, Indonesia Type hosts: tiger and dog Current status: a synonym of Dermacentor steini (Wassef & Hoogstraal 1986, Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF0A42C5FF23F905FAFBFEB5.taxon	description	Koch, C. L. (1844) Systematische Übersicht über die Ordnung der Zecken. Archiv für Naturgeschichte, 10, 217 – 239. Type depository: ZMB: holotype (male) (Moritz & Fischer 1981) Type locality: Brazil Type host: not stated Current status: a synonym of Amblyomma aureolatum (Aragão & Fonseca 1961, Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1542C5FF23FD38FB6EFC29.taxon	materials_examined	Type depository: ZIAC: a lectotype (nymph) and paralectotypes (larvae) (Filippova 2008) Type locality: Ch-Asku Canyon, Kungei Ala Tau Mountain Range, Tien Shan, Kyrgyzstan	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1542C4FF23F8B9FB24FE21.taxon	materials_examined	Type depositories: USNTC: holotype (male) and paratypes? (males and females); ORSTOM: paratypes? (males and females) (Hoogstraal et al. 1992) Type locality: Ngong, 40 km southwest of Nairobi, Kajiado County, Kenya Type host: Herpestes sanguineus Comments: Tomlinson et al. (2018) found only one paratype (named as allotype) in the USNTC. Tomlinson et al. (2018) stated that there are at least three species among the specimens used to describe Haemaphysalis subterra by Hoogstraal et al. (1992), and only the holotype specimen is bona fide Haemaphysalis subterra.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1742C7FF23FF78FADAFE5D.taxon	materials_examined	Type depositories: MZB: holotype (male) and paratypes (males and females); BMNH? and USNTC: paratypes (males and females) Type locality: Tjabang Pasjik, Lampong District, Sumatra, Indonesia Type host: Sus scrofa vittatus Comments: Hoogstraal et al. (1971) listed the BMNH as the depository of paratypes of Haemaphysalis sumatraensis, but Keirans & Hillyard (2001) stated that there is no indication that the specimens arrived at that museum.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1742C7FF23FDCDFEB7FD59.taxon	materials_examined	Type depository: USNTC: lectotype (female) (Keirans & Clifford 1984); IM: lectotypes (females) (Sharif 1928) Type locality: Bhowali, Nianital District, State of Uttarakhand, India Type host: sheep	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1742C7FF23FA00FB88F949.taxon	materials_examined	Type depository: BMNH?: syntypes (males) (Walker et al. 2000) Type locality: Lualaba River, Democratic Republic of the Congo Type host: horse Comments: Keirans & Hillyard (2001) found no indication that the syntypes had been received by the BMNH. Guglielmone et al. (2020) consider the morphological definition of this species unclear.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1642C6FF23FAF8FF0DF9DD.taxon	materials_examined	Type depositories: QM: holotype (female) and paratypes (females); ANIC: paratypes (females and a larva); ARI: paratypes (females) Type locality: Wovan, Queensland, Australia Type host: Tachyglossus aculeatus Current status: a synonym of Bothriocroton tachyglossi and its original name (Guglielmone et al. 2009). The current status of Aponomma tachyglossi, as Bothriocroton tachyglossi, was also established by Guglielmone et al. (2009).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1642C1FF23F8E0FCF3FF19.taxon	description	Kohls, G. M. (1969) Ixodes taglei n. sp. (Acarina: Ixodidae) a parasite of the deer, Pudu pudu (Wol.), in Chile. Journal of Medical Entomology, 6, 280 – 283. Type depositories: USNTC: holotype (female) and paratypes (a male and females); EMV: paratypes (females) Type locality: Callumapu, 15 km north of Valdivia, Chile	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1142C1FF23FE10FE85FD65.taxon	description	[in Japanese, NAMRU- 3 translation 1791] Type depository: unknown: holotype (male) (Apanaskevich & Apanaskevich 2015) Type locality: Aoshan, Xindien, New Taipei, Taiwan Type host: Sus sp.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1142C1FF23FD14FC6EFBB9.taxon	materials_examined	Type depository: undetermined: syntypes? (males and females) Type locality: Tataka, Taiwan Type host: dog Current status: a synonym of Ixodes ovatus (Yamaguti et al. 1971).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1142C0FF23F8B9FC3CFF19.taxon	materials_examined	Type depository: MZD: holotype (male) and paratypes (females) Type locality: Kurokawamura, Kitakambaragun, Niigata Prefecture, Japan	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1042C0FF23FB44FC9DFA69.taxon	materials_examined	Type depositories: MNHN and ZMB: syntypes (females) Type locality: Tasmania, Australia Type host: undetermined Comments: the female syntype deposited in the ZMB is from an unknown type locality, and two females in the MNHN are from Ile St Pierre (Ponafidin?) [sic] (Neumann 1899).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1042C0FF23FA00FC74F949.taxon	materials_examined	Type depository: ZMUC: holotype (female) Type locality: Victoria, Australia Type host: undetermined Current status: a synonym of Ixodes tasmani (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1342C3FF23F950FDF5F852.taxon	materials_examined	Type depository: ZMB: a lectotype (male) and a paralectotype (male) were designated by Nava et al. (2014 b) Type locality: Mexico Type host: unknown Comments: Moritz & Fischer (1981) recognized the presence of two males in the ZMB but stated that Koch used only one male to describe Amblyomma tenellum, regarding that specimen as the holotype of this species, a view not supported by Nava et al. (2014 b).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1342C3FF23FE7DFDCEFDB1.taxon	materials_examined	Type locality: cave of Méya, Congo	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1342C3FF23FF78FB84FECD.taxon	materials_examined	Type depository: FMNH: holotype (female) and paratypes (females) Type locality: Kirkuk Liwa, Chemchemal Valley, Jarmo, Iraq Type host: Vulpes vulpes Current status: a synonym of Ixodes eldaricus (Filippova 1977, Camicas et al. (1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1342C3FF23FC64FC9AFBD5.taxon	materials_examined	Type depository: ZIAC: holotype (female) (Filippova 2008) Type locality: Olenevka Village, Chernomorskii District, Ukraine Type host: Phalacrocorax aristotelis Current status: a synonym of Ixodes unicavatus (Filippova 2008).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1242C2FF23FCF1FE01FC61.taxon	materials_examined	Type depository: ZMB: holotype (male) (Moritz & Fischer 1981) Type locality: Java, Indonesia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1242C2FF23FE58FF0DFD59.taxon	materials_examined	Type depository: ZMB?: syntypes (females and nymphs) Type localities: Rugen Island, Germany, and Painswick, Gloucestershire, England, Great Britain Type hosts: Arvicola glareolus and Arvicola pratensis Current status: a synonym of Ixodes trianguliceps (Filippova (1997, Camicas et al. 1987). Comments: no specimens of Ixodes tenuirostris were included in the ZMB type catalog by Moritz & Fischer (1981).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1242C2FF23FA6CFCEDF931.taxon	materials_examined	Type depositories: BMNH and GM (presumed): syntypes (females) (Keirans & Hillyard 2001) Type locality: Bhamo, Myanmar Type hosts: Testudo elongata, Nicoria trijuga and Varanus bengalensis Current status: a synonym of Amblyomma supinoi and its original name (preoccupied). The current name and status of Ixodes supinoi were established by Neumann (1905).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1242CDFF23F928FEBBFEE9.taxon	materials_examined	Type depository: NSM: holotype (nymph) and paratypes (nymphs) Type locality: Doi Inthanon, Thailand Type host: Rattus nitidus Comments: a male, female and nymphs of the type series of Rhipicephalus tetracornus were lost in the mail (Kitaoka & Suzuki 1983). Also, it is uncertain whether the larva depicted by these authors in the original description actually belongs to this species. Walker et al. (2000) treat Rhipicephalus tetracornus as a name incertae sedis, while Horak et al. (2002) and Kolonin (2009) did not include this species in their lists of ticks of the world, but Camicas et al. (1998), Guglielmone & Nava (2014) and Guglielmone et al. (2009, 2010, 2014, 2020, 2023) consider Rhipicephalus tetracornus valid.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1D42CDFF23F94DFCC5F875.taxon	materials_examined	Type depository: OVI: syntypes (females) Type localities: Gibeon and Mariental, Namibia; Bothaville, Ficksburg and Onderstepoort, South Africa Type hosts: Creatophora carunculata, Hirudo rustica, migratory swallow, Quelea sanguinirostris lathami and “ vink ” Comments: Walker (1991) and Horak et al. (2018) listed Ixodes theilerae as described in 1958, but it was described in Arthur (1953), under the name Ixodes theileri, a lapsus.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1D42CDFF23FC18FC0BFB41.taxon	materials_examined	Type depository: VLM: holotype (male) Type locality: Magude, Mozambique Type host: Bos taurus Current status: a synonym of Amblyomma hebraeum (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1C42CCFF23FCF1FAF4FC61.taxon	materials_examined	Type depository: MNHN: syntypes (a male and females) Type locality: “ Congo ”; Haut-Oubanghi (undetermined country in Central Africa); Malawi?; Tanzania	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1C42CCFF23FF79FCF7FEE9.taxon	materials_examined	Type depository: OVI: holotype (male) and a paratype (female) (Walker et al. 2000) Type locality: Glen, Province of Free State, South Africa	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1C42CFFF23F94CFCB0FF7D.taxon	materials_examined	Type depository: unknown: holotype? (female) Type locality: Brazil Type host: unknown Current status: Ixodes thoracicus, described in Koch (1844), is considered a nomen dubium in Guglielmone & Nava (2014). Comments: Neumann (1899) treated ticks from birds collected in Chile as identical to Ixodes thoracicus, which Koch (1844) described from specimens found in Brazil. Neumann (1904) later found that the Chilean specimens corresponded to a new species that he named Ixodes auritulus.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1F42CFFF23FB44FE99FAFD.taxon	materials_examined	Type depository: ZMB: holotype (male) (Moritz & Fischer 1981) Type locality: Brazil	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1F42CFFF23FE11FEF1FC91.taxon	materials_examined	Type depositories: BMNH: holotype (male) and paratypes (a male, females and a nymph); USNTC: paratypes (a female and a nymph) Type locality: Yatung, Chumbi Valley, Xizang, China Type host: dog Comments: Kelava et al. (2024) suggest that Haemaphysalis tibetensis is a synonym of Haemaphysalis nepalensis.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1E42CEFF23F973FC7BF853.taxon	materials_examined	Type depositories: BMNH: lectotype (male) and paralectotypes (males and a female); ZMA: paralectotypes (males) (Hoogstraal 1964 a) Type locality: Buru, Maluku Islands, Indonesia Type host: bat Comments: Kolonin (2009) did not include Haemaphysalis toxopei in his list of the ixodid ticks of the world. Hoogstraal (1964 a) named this tick Haemaphysalis papuana toxopei.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1E42CEFF23FAD4FD75F9DD.taxon	materials_examined	Type depositories: USNTC: holotype (female) and paratypes (males, females and nymphs); CT, DEP and MCZ: paratypes (males, females and nymphs) Type locality: Comontoso, Guanajuato, Mexico Type host: hare Comments: no paratypes were found in the MCZ (Keirans & Clifford 1978). KLC, IR and RGR visited the MCZ in 2024 and the paratypes were not found there.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1942C9FF23FF78FB63FE63.taxon	materials_examined	Type depository: ZMB?: holotype (male) and paratypes (males and females) Type locality: Wasior, New Guinea, Indonesia Type host: Varanus sp. Current status: a synonym of Amblyomma fimbriatum, as discussed in Guglielmone & Nava (2014). Comments: although Schulze (1933 b) listed the types as having been deposited in the ZMB, no specimens of Amblyomma trabeatum are included in the list of ZMB types by Moritz & Fischer (1981).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1942C9FF23FE1DFCCDFDD0.taxon	materials_examined	Type depository: LMNH: syntypes (males, females and larvae) (Hoogstraal 1964 a) Type locality: Berhala Island, Riau Archipleago, Indonesia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1942C9FF23F9B8FBA4F88E.taxon	materials_examined	Type depositories: SAIMR: holotype (female); USNTC: a paratype (female) Type locality: Citrusdaal, South Africa Type host: the holotype specimen was collected from a nest of Rattus namaquensis.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1842C8FF23FA24FC6CF83D.taxon	description	Dönitz, W. (1906) Über afrikanische Zecken. Sitzungsberichten der Gesellschaft Naturforschender Freunde zu Berlin, 5, 143 – 148. Type depository: ZMB: a lectotype (female) was designated in Walker et al. (1988) Type locality: Kalahari, Botswana? Namibia? Type host: free-living specimen Comments: Moritz & Fischer (1981) listed a holotype and ten paratypes of Rhipicephalus tricuspis, deposited in the ZMB, but Walker et al. (1988) stated that this tick was described from one male and a female with no holotype designation, also indicating that the original series contains specimens of Rhipicephalus lunulatus and Rhipicephalus tricuspis. Moritz & Fischer (1981) recorded the tick later designated as lectotype as having been found in Namibia, but Walker et al. (1988) believed that it was collected in Botswana.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1B42CBFF23FD38FBABFC45.taxon	materials_examined	Type depositories: QM: holotype (female) and paratypes (males and females); ANIC and ARI: paratypes (males and females) Type locality: Burketown, Queensland, Australia Type host: dog Current status: a synonym of Amblyomma triguttatum (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1B42CBFF23FBF4FBABFA99.taxon	materials_examined	Type depositories: QM: holotype (female) and paratypes (males and females); ANIC and ARI: paratypes (males and females) Type locality: Alice Springs, Northern Territory, Australia Type host: cow Current status: a synonym of Amblyomma triguttatum (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1B42CBFF23FF78FE7AFEE9.taxon	materials_examined	Type depository: ZMB: holotype (female) (Moritz & Fischer 1981) Type locality: Australia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1B42CBFF23FAB1FA79F985.taxon	description	See trimaculatus Lucas, 1878, Ixodes Comments: Guglielmone et al. (2020) stated that the several morphological redescriptions of this tick (under the name Aponomma trimaculatum, a synonym of Amblyomma trimaculatum) fail to define this species with certainty.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1B42CBFF23F9B4FAAFF8D9.taxon	materials_examined	Type depository: MNHN: holotype (female) (Kaufman 1972) Type locality: New Guinea Type host: Varanus chlorostigma Current status: a synonym of Amblyomma trimaculatum and its original name (Guglielmone & Nava 2014). The current status of Ixodes trimaculatus, as Amblyomma trimaculatum, was established by Horak et al. (2002).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1A42CAFF23FF78FC57FE21.taxon	materials_examined	Type depositories: BMNH and TC: syntypes (a female, nymphs and larvae) (Keirans & Hillyard 2001) Type locality: Saint Augustine, Trinidad, Trinidad and Tobago Type host: ground lizard Current status: a synonym of Amblyomma dissimile (Keirans & Hillyard 2001). Comments: Camicas et al. (1998) listed Amblyomma trinitatis as valid.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1A42CAFF23FD38FBB3FC45.taxon	materials_examined	Type depository: ZMB: holotype (female) (Moritz & Fischer 1981) Type locality: Montevideo, Uruguay Type host: unknown Comments: Lado et al. (2018) treated Amblyomma triste as a synonym of Amblyomma maculatum, but Guglielmone et al. (2021) believed that more solid evidence is needed before making that decision.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1A42CAFF23FE58FAEAFD00.taxon	materials_examined	Type depository: IP: holotype (female) (Hoogstraal et al. 1965 b) Type locality: Haut-Chlong, Vietnam Type host: unknown Current status: a synonym of Haemaphysalis hystricis (Hoogstraal et al. 1965 b, Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF1A42CAFF23F9D8FE76F949.taxon	materials_examined	Type depository: ZMB: holotype (male) (Moritz & Fischer 1981) Type locality: Senegal	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2542F5FF23FAB0FA6EF8D9.taxon	description	In: Pomerantzev, B. I., Matikashvili, N. V. & Lotozky, B. V. (1940) An ecological and faunistic outline of Ixodidae ticks occurring in Transcaucasia. Parazitologicheskii Sbornik, 7, 100 – 133. [in Russian, NAMRU- 3 translation 51] Type depository: ZIAC: lectotype (male) and a paralectotype (female) (Filippova 2008) Type locality: Tashkent, Uzbekistan Type host: Ovis aries Comments: Walker et al. (2000) named Pomerantzev (1936) as the authority for Rhipicephalus turanicus. Walker et al. (2000) do not name a type depository for this species, but Rhipicephalus turanicus in Pomerantzev (1936) is a nomen nudum (Filippova 1997). Camicas et al. (1998) gave the authority for this species as “ Pomerantzev, B. I., Matikashvily, N. V. & Lotozky, B. V. ” Guglielmone et al. (2020) treated many records of Rhipicephalus turanicus around the world as speculative and maintained that several descriptions of this tick require confirmation. In that regard, Guglielmone et al. (2023) stated that the only bona fide specimens of this species are the type specimens from Tashkent, Uzbekistan, listed above.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2542F5FF23FEA4FE3CFC0D.taxon	materials_examined	Type depositories: IPCUM or MSNM: syntypes (males) (Tonelli Rondelli 1932), but lost (Hoogstraal & Kaiser 1958); USNTC?: lectotype? (male) and a paralectotype? (male) (Hoogstraal & Kaiser 1958), syntypes? (males) (Apanaskevich et al. 2008 a) Type locality: Tauorga Oasis, Libya Type host: camels and sheep Current status: a synonym of Hyalomma franchinii and its original name (Camicas et al. 1998, Apanaskevich et al. 2008 a). The current status of Hyalomma tunesiacum franchinii, as Haemaphysalis franchinii, was established by Hoogstraal & Kaiser (1958). Comments: Kratz (1940) stated that the “ type ” was in the “ Zoological Museum of Milan, ” probably meaning the MSNM. Hoogstraal & Kaiser (1958) believed that two male specimens deposited in the USNTC were part of the syntype series of Hyalomma tunesianicum franchinii and designated one male as a lectotype and the other male as a paralectotype, while Apanaskevich et al. (2008 a) accepted that the two male ticks in the USNTC are probably part of the syntype series of Hyalomma tunesiacum franchinii but did not confirm the lectotype designation by Hoogstraal & Kaiser (1958).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2542F5FF23FC3CFC00FB6C.taxon	materials_examined	Type depository: IPCUM: holotype (male) (Tonelli Rondelli 1932) and Kratz (1940) Type locality: Agordat, Eritrea Type host: undetermined Current status: a synonym of Hyalomma excavatum (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2442F4FF23FD5CFE42FCF5.taxon	materials_examined	Type depository: ZMB: holotype (male) Type locality: Sri Lanka	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2442F4FF23FB1CFB5EF985.taxon	materials_examined	Type depository: NRS: lectotype (female) and paralectotypes (a male and females) were designated by Morel (1965 a) Type locality: Kibonoto, Kilimanjaro, Tanzania Type host: Genetta pardina Current status: a synonym of Ixodes djaronensis and its original name (Camicas et al. 1998). The current status of Ixodes ugandanus djaronensis, as Ixodes djaronensis, was established by Santos Dias (1958 a).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2742F7FF23FAD4FD9AFA69.taxon	materials_examined	Type depository: EC: syntypes (a female and a nymph) (Nuttall & Warburton 1911) Type locality: Forth, Scotland, Great Britan	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2742F6FF23F8B8FBEAFEB5.taxon	materials_examined	Type depository: undetermined: holotype (male) and paratypes (a male and females) Type locality: Ural Mountains, Russia Type host: Bos taurus Current status: a synonym of Hyalomma scupense (Apanaskevich et al. 2010).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2642F6FF23FBF5FCB7FB41.taxon	materials_examined	Type depositories: ZIAC: holotype (male) and paratypes (males and females); USNTC: paratypes (females) (Filippova 2008) Type locality: Bortagoi area, Chilik River Valley, Kazakhastan	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2642F6FF23FEA4FA81FD2D.taxon	materials_examined	Type depository: undetermined: syntypes (females) Type locality: Arctic regions Type host: Uria troile Comments: Arthur (1965, page 336) stated that a male and a female constituting the types of Ixodes uriae are deposited in the MNHN? [sic]. However, White (1852) described the female but not the male of Ixodes uriae; therefore, it is postulated here that the depository of the type specimens of this species remains undetermined.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2642F6FF23FD5CFAA1FC45.taxon	materials_examined	Type depository: USNTC: holotype (nymph) and paratypes (larvae) (Keirans & Clifford 1984) Type locality: 3 km east of Cardona, Department of Soriano, Uruguay Type host: Scapteromys tumidus Current status: a synonym of Ixodes longiscutatus, as discussed in Venzal et al. (2001). Comments: Keirans & Clifford (1984) did not specify the developmental stages held in the USNTC, but Ixodes uruguayensis was described from a nymph (holotype) and five larvae (paratypes) (Kohls & Clifford 1967).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2642F1FF23F929FA78FF51.taxon	materials_examined	Type depository: ZMB?: holotype (female); undetermined collection: paratypes (males, females, nymphs and larvae) Type locality: Campo, Cameroon Type host: “ mwack, ” a mongoose-like animal (Keirans et al. 1982) Comments: Schulze (1943) stated that the holotype female would be deposited in the ZMB. Schulze (1943) also described the male, nymph and larva of Ixodes vanidicus but did not indicate any depository for them. No specimen of Ixodes vanidicus is listed in the catalog of ZMB types by Moritz & Fischer (1981), and Keirans et al. (1982) stated that the types of Ixodes vanidicus are missing, therefore implying the existence of paratypes for this species.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2142F1FF23FCF0FBA2FBB9.taxon	materials_examined	Type depository: MGH: syntypes (males) Type locality: Brisbane, Australia Type host: Hydrosaurus giganteus Current status: a synonym of Bothriocroton undatum (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2142F1FF23F9D8FBDDF83D.taxon	description	Robinson, L. E. (1912) New species of ticks (Haemaphysalis, Amblyomma). Parasitology, 4, 478 – 484. Type depository: BMNH: a lectotype (male) was designated by Keirans & Brewster (1981) Type locality: Zimbabwe Type host: unknown Current status: a synonym of Amblyomma pomposum (Keirans & Brewster 1981).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2042F0FF23FD38FCC5FBD5.taxon	materials_examined	Type depositories: BMNH and MNHN: syntypes (males) (Neumann 1899, Keirans & Hillyard) Type locality: Havana, Cuba Type host: Cyclura harlani Current status: a synonym of Amblyomma albopictum and its original name (preoccupied). The current name and status of Ixodes variegatus was established by Neumann (1899). Comments: Neumann (1899) listed a male collected from Cercolabes villosus in Brazil in his description of Amblyomma albopictum, but this tick was not considered present in that country by Guglielmone et al. (2021), and Lucas (1852) did not mention any specimens collected outside Cuba in his description of Ixodes variegatus. Keirans & Hillyard (2001) named the type host for Amblyomma albopictum as Cyclura nubila, while ignoring the original preoccupied name of this tick species (Ixodes variegatus).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2042F0FF23FE58FB52FD01.taxon	materials_examined	Type depository: USNPC: syntypes (a male and females) Type locality: State of Washington, USA Type host: Cariacus sp. Current status: a synonym of Dermacentor albipictus (Neumann 1911 a, Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2042F0FF23F8C3FE99F87E.taxon	materials_examined	Type depository: ZMB: holotype (male) (Moritz & Fischer 1981) Type locality: Brazil	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2042F0FF23FF78FB9EFE21.taxon	materials_examined	Type locality: Africa Type host: unknown Current status: a synonym of Amblyomma variegatum and its original name (Neumann 1911 a). The current status of Acarus variegatus, as Amblyomma variegatum, was established by Gerstäcker (1873).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2042F0FF23FB44FC0BFA4D.taxon	materials_examined	Type depository: LMNH?: holotype (male) Type locality: Kamchatka, Russia Type host: Ovis sp. Current status: a synonym of Dermacentor kamshadalus and its original name (Apanaskevich 2013), whose current status was also established by Apanaskevich (2013). Comments: Neumann (1908) stated that the holotype specimen of Dermacentor variegatus kamshadalus had been deposited in the LMNH, but it was not found there by Apanaskevich (2013).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2342F3FF23FA6CFCB7F915.taxon	materials_examined	Type depository: USNTC: “ type ” (syntype? unspecified developmental stage) (Keirans & Clifford 1984) Type locality: Texas and New Mexico, USA (Neumann 1897) Type host: unknown Current status: Dermacentor venustus is a suppressed name, preserving the name Dermacentor andersoni Stiles, 1908, as explained by Keirans (1976). Comments: the “ type ” deposited in the USNTC is from Texas.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2342F3FF23FE35FBE2FD01.taxon	materials_examined	Type depositories: USNTC: holotype (female) and paratypes (females and nymphs); BMNH: a paratype (female) (Keirans & Hillyard 2001) Type locality: Campamento Rafael Rangel, Sierra Maestra, Aragua, Venezuela	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2242F2FF23FAB1FCA6F985.taxon	materials_examined	Type depository: BAC: holotype (larva) Type locality: a fossil species collected from Burmese amber	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2242F2FF23FE35FD57FCBD.taxon	materials_examined	Type depository: VMDFA: syntypes (females) (Hoogstraal 1973) Type locality: Yuncheng, Shanxi, China Type host: dog Comments: the syntypes of Haemaphysalis verticalis were thought to have been deposited in the NSM by Emel’yanova & Hoogstraal (1973) but were not found there; this situation was clarified by Hoogstraal (1973), who stated that the syntypes are held in the VMDFA.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2242F2FF23FCADFE45FC45.taxon	materials_examined	Type depository: ZMB: holotype (female) (Moritz & Fischer 1981) Type locality: Germany	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2D42FDFF23FAD5FB32F9F9.taxon	materials_examined	Type depositories: SAM: holotype (male) and paratypes (males, females, nymphs and larvae); USNTC: paratypes (males, females, nymphs and larvae) Type locality: Warruwarldunha Range, Flinder Ranges near Hawker, South Australia, Australia Type host: Egernia stokesii Comments: type specimens were reared in the laboratory from ticks collected from the type host.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2D42FDFF23F990FC9BF811.taxon	materials_examined	Type depository: ZMB?: holotype (female) Type locality: Bolivia Type host: unknown Current status: a synonym of Amblyomma oblongoguttatum (Neumann 1911 a, Camicas et al. 1998). Comments: Neumann (1899) listed the female holotype of Amblyomma vittatum as held in the ZMB, but it is not included in the catalog of ZMB types by Moritz & Fischer (1981).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2D42FDFF23FC3CFEB0FAA5.taxon	materials_examined	Type depositories: BM: holotype (female); USNTC: a paratype (female) Type locality: 6 km south of Dalat, Vietnam Type host: the holotype specimen was collected from vegetation Current status: a synonym of Alloceraea colasbelcouri, as discussed in Guglielmone et al. (2009, under the genus Haemaphysalis).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2D42FDFF23FF78FDFCFE05.taxon	materials_examined	Type depository: NM: syntypes (males and females) Type locality: Lower Tarwin, Victoria, Australia Type hosts: Macropus nalabatus and Pseudochirus peregrinus Current status: a synonym of Ixodes hirsti and its original name but preoccupied; consequently, Hassall (1931) renamed this species Ixodes hirsti.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2D42FDFF23FE34FDE2FD2D.taxon	materials_examined	Type depository: BMNH: a lectotype (female) was designated by Keirans & Brewster (1981) Type locality: Victoria, Australia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2C42FCFF23FBD1FD61FAA5.taxon	materials_examined	Type depository: USNTC: holotype (male) and paratypes (males and females) Type locality: Athi River, Machakos, Kenya	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2F42FFFF23FD5CFEB9FC45.taxon	description	Izvestiya Akademy Nauk Turkmenskoy, 1, 77 – 80. [in Russian, NAMRU- 3 translation 145] Type depository: ZIAC: neotype (male) (Filippova 2008) Type locality: Yablonevskii cordon, Kopet Dagh Reserve, Central Kopet Dagh, Turkmenistan Type host: Bezoar goat Current status: a synonym of Haemaphysalis kopetdaghica and its original name (Filippova 2008). The current status of Haemaphysalis warburtoni kopetdaghicus, as Haemaphysalis kopetdaghica, was established by Hoogstraal & Wassef (1979).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2F42FFFF23F9B5FD98F881.taxon	materials_examined	Type depository: BMNH: a lectotype (male) and paralectotypes (females and nymphs) were designated by Keirans & Brewster (1981) Type locality: Kuching, Sarawak, Malaysia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2E42FEFF23FE80FD90FDC9.taxon	materials_examined	Type depository: FMNH?: holotype (female) and paratype (a female) Type locality: south slope of Mount Balabag, Mantalingajan Range, Palawan Island, Philippines Type host: Rattus palawanensis Comments: IR visited the FMNH in 2024 and the holotype was not found there; however, the paratype was present in 2011 when the collection was rehoused.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2E42FEFF23FD61FD92FCF5.taxon	materials_examined	Type depositories: BMNH: holotype (male) and paratypes (males and females); LSHTM, OVI, SAIMR, USNTC and VRLKK: paratypes (males and females) (Keirans & Hillyard 2001) Type locality: 24 km west of Isiolo, Kenya	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2E42FEFF23FB69FF0BFA4D.taxon	materials_examined	Type depository: ZMB: holotype (male) (Moritz & Fischer 1981) Type locality: Valparaiso (Karsch 1879), Chile? Type host: not stated Comments: the type locality, Valparaíso, an important Chilean city, has been treated as an indication that this tick was originally collected in the southern Neotropics, but Margaropus winthemi is an Afrotropical species, and the name “ Valparaiso ” is probably a mislabeling. Moritz & Fischer (1981) spelled the type locality as “ Valpareiso ” (a lapsus).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2E42F9FF23F9FDFF41FEE9.taxon	materials_examined	Type depositories: WAM: holotype (female) and paratypes (females and nymphs); ANIC: paratypes (females and nymphs) Type locality: Dryandra Woodland, Western Australia, Australia Type host: Bettongia penicillata x	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2942F9FF23FD38FAF0FBB9.taxon	description	Kishida, K. (1935) Notes on the Acarina – mites and ticks (including Amblyomma yajimai sp. n., on water buffalo) from the Island of Formosa, collected in August, 1935. Lansania, 7 (69), 129 – 144. Type depository: CK: holotype (female) Type locality: Pasicau, Hinan-ku, Taito-choh, Taiwan Type host: cow Current status: a synonym of Amblyomma testudinarium (Yamaguti et al. 1971, Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2942F9FF23FE80FAC0FD2D.taxon	materials_examined	Type depository: IZAS: holotype (male) and paratypes (males and females) Type locality: southern area, Xinjiang, China. Type host: goat Current status: a synonym of Haemaphysalis danieli (Teng & Jiang 1991). Comments: Camicas et al. (1998) and Horak et al. (2002) treated Haemaphysalis xinjiangensis as valid.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2942F9FF23FAD4FBB4F9F9.taxon	materials_examined	Type depository: undetermined: holotype? (male) Type locality: Gevrigcvrik Bay, Turkey Type host: fox Current status: a synonym of Haemaphysalis erinacei (Guglielmone et al. 2014). Comments: Bursali et al. (2012) listed Haemaphysalis yalvaci as provisionally valid.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2942F9FF23F990FBE4F881.taxon	materials_examined	Type depository: IPP: lectotype (female) (Hoogstraal & Trapido 1966 b) Type locality: region of Haut-Chlong, Vietnam (Hoogstraal & Trapido 1966 b)	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2842F8FF23FEA4FF47FD65.taxon	materials_examined	Type depositories: CNCDGS: holotype (male) and paratypes (males, females and nymphs); IBU, INTA and USNTC: paratypes (males, females and nymphs) Type locality: Parque Estadua do Turvo, Derrubadas Municipality, State of Rio Grande do Sul, Brazil Type host: holotype specimen collected from vegetation z	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2842F8FF23FD15FD07FC0D.taxon	materials_examined	Type depositories: AM: holotype (female) and paratypes (a male and females); USNTC: paratypes (females) Type locality: Wissel Lakes, New Guinea, Indonesia	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2842F8FF23F9B5FE9BF875.taxon	materials_examined	Type depositories: OVI: holotype (male) and paratypes (males, females, nymphs and larvae); BMNH, USNTC and VRLH: paratypes (males, females, nymphs and larvae) Type locality: Driehoek Ranch, near West Nicholson, Gwanda District, Zimbabwe Type host: cattle Comments: Rhipicephalus zambeziensis was described from laboratory-reared specimens originating from ticks collected from cattle, whose larvae and nymphs were fed on rabbits, while adult ticks were fed on Friesland steers (Walker et al. 1981).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2B42FBFF23FA6CFC93F931.taxon	materials_examined	Type depositories: SAIMR: holotype (female); unknown: paratypes (females) Type locality: Nightingale Island, Tristan da Cunha, Atlantic Ocean Type host: Puffinus gravis Current status: a synonym of Ixodes kerguelenensis (Wilson 1970, Guglielmone et al. 2009). Comments: Ixodes zumpti is listed as valid in Horak et al. (2002).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2B42FBFF23FF79FBDBFE5D.taxon	materials_examined	Type depository: BMNH: syntypes (females) (Keirans & Hillyard 2001, as Amblyomma zelanicum, lapsus) Type locality: KeLa Oyel, Sri Lanka Type host: unknown Current status: a synonym of Amblyomma clypeolatum (Keirans & Hillyard 2001).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2B42FBFF23F928FC11F811.taxon	materials_examined	Type depository: SDC: holotype (male) Type locality: Govuro, Mozambique Type host: Syncerus caffer Comments: Camicas et al. (1998) do not treat Rhipicephalus zumpti as valid.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2B42FBFF23FDCCFE4AFD01.taxon	materials_examined	Type depository: BMNH: syntypes (a male and a female) Type locality: Cameroon	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2B42FBFF23FBF4FBC8FAFD.taxon	materials_examined	Type depositories: USNTC: holotype (male) and paratypes (males, females, nymphs and larvae); BMNH, FMNH?, MNHN, OVI and SAIMR (males, females, nymphs and larvae) Type locality: Mafikeng, Province of North West, South Africa Type host: Cynictis penicillata Comments: IR visited the FMNH in 2024 and the paratypes were not found there.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2A42FAFF23F9D4FBC9F968.taxon	description	Current status: a synonym of Hyalomma excavatum (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2A42FAFF23FA62FBDEF9BB.taxon	description	Current status: a synonym of Hyalomma anatolicum (Guglielmone & Nava, 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2A42FAFF23FD83FD1FFC91.taxon	description	Current status: a nomen dubium (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2A42FAFF23FC8EFBA8FC4F.taxon	description	Current status: a synonym of Hyalomma truncatum (Apanaskevich & Horak 2008 a).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2A42FAFF23FBE0FEACFB10.taxon	description	Current status: a synonym of Hyalomma impressum (Apanaskevich & Horak 2007), as discussed in Guglielmone & Nava (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF2A42FAFF23FB0FFD1FFACD.taxon	description	Current status: a nomen dubium (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3542E5FF23FF78FCA9FEB5.taxon	description	Current status: a nomen dubium (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3542E5FF23FBF4FCA1FBD5.taxon	description	Current status: a nomen nudum (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3542E5FF23FB44FBDEFA99.taxon	description	Current status: a name incertae sedis, as discussed in Guglielmone & Nava (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3542E5FF23FAB0FBADFA4D.taxon	description	Current status: a synonym of Rhipicephalus annulatus (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3542E5FF23F9FCFB13F915.taxon	description	Current status: this name was treated as a synonym of Rhipicephalus microplus in Guglielmone & Nava (2014), but Margaropus annulatus mexicanus lacks both a description and figures, and is thus a nomen nudum).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3442E4FF23FD5CFBDEFC91.taxon	description	Current status: a name incertae sedis, as discussed in Guglielmone & Nava (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3442E4FF23FEA4FBDDFE79.taxon	description	Current status: a nomen dubium (Camicas et al. 1998, Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3442E4FF23FAB0FD1FFA4D.taxon	description	Current status: a nomen dubium (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3742E7FF23FE10FB90FD2D.taxon	description	Current status: Boophilus capensis was treated as a synonym of Rhipicephalus decoloratus in Guglielmone & Nava (2014), but it is a name lacking both a description and figures, and thus a nomen nudum.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3742E7FF23FB20FE70FA35.taxon	description	Current status: a name incertae sedis (Neumann 1911 a, Guglielmone & Nava 2014). The type locality was regarded as unknown by Guglielmone & Nava (2014), but the specimen was collected in the Canary Islands, Spain.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3742E7FF23FA24FECFF931.taxon	description	Current status: a nomen dubium (Guglielmone & Nava 2014). The type locality for Ixodes cinctus is the Canary Islands, Spain, but this was not recognized in Guglielmone & Nava (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3742E6FF23F89BFB7AFEB5.taxon	description	Current status: a nomen dubium (Camicas et al. 1998, Guglielmone & Nava 2014). cinnabarina punctata musimonis Tonelli Rondelli, 1930, Haemaphysalis Tonelli Rondelli, M. (1930) Ixodoidea del Museo di Milano. Atti della Società Iitaliana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano, 69, 112 – 124. Current status: a synonym of Haemaphysalis sulcata (Filippova 1997, Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3642E6FF23FB1CFC61FA35.taxon	description	Current status: a name incertae sedis (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3642E6FF23FEA4FBDDFE79.taxon	description	Current status: a nomen dubium (Camicas et al. 1998, Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3642E6FF23FD5CFD1EFCF5.taxon	description	Current status: Ixodes colenatyi was treated as a nomen dubium in Guglielmone & Nava (2014), but it is a name without a formal description and thus a nomen nudum.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3642E6FF23FBD0FCA9FB6D.taxon	description	Current status: a nomen dubium (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3642E6FF23F990FC7BF881.taxon	description	Current status: Ixodes crenulatus terecus was treated as a synonym of Ixodes crenulatus in Guglielmone & Nava (2014), but it is a name without description and thus a nomen nudum.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3642E1FF23F8B8FC0AFF7D.taxon	description	Current status: a name incertae sedis (Neumann 1911 a, Guglielmone & Nava 2014). Guglielmone & Nava (2014) stated that Neumann (1897) wrote that the type locality of Dermacentor cruentus was unknown, but that author said that the type locality was “ Orowitza (Banat). ”	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3142E1FF23FE34FC61FDC9.taxon	description	Current status: a name incertae sedis (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3142E1FF23FCACFC05FC61.taxon	description	Current status: a synonym of Hyalomma anatolicum (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3142E1FF23FEECFCFFFE05.taxon	description	Current status: Ixodes cyclurae was treated as a nomen dubium in Guglielmone & Nava (2014), but it is a name without a description or figures and thus a nomen nudum.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3142E1FF23FC18FC2BFBD5.taxon	description	Current status: Ixodes distipes was treated as a nomen nudum in Guglielmone & Nava (2014), but it is a nomen dubium, as already stated in Neumann (1911 a) and Camicas et al. (1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3142E1FF23FB44FC61FA99.taxon	description	Current status: a name incertae sedis (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3042E0FF23FC18FBCFFBD5.taxon	description	Current status: a synonym of Ixodes vespertilionis, as stated in Neumann (1911 a).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3042E0FF23FD38FAE6FC91.taxon	description	Current status: a synonym of Haemaphysalis flava (Nuttall & Warburton 1915, Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3042E3FF23F870FC8DFF7D.taxon	description	Current status: Acarus fuscus is treated here as a nomen dubium.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3342E3FF23FEECFCA1FE05.taxon	description	Current status: a nomen nudum (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3342E3FF23FAF8FAFAFA69.taxon	description	Current status: Aponomma halli in Tendeiro (1948) was treated as a synonym of Amblyomma flavomaculatum in Guglielmone & Nava (2014), but it is a name without a description or figures and thus a nomen nudum.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3342E3FF23F94CFAE2F881.taxon	description	Current status: a synonym of Ixodes hexagonus (Camicas et al. 1998, under the genus Pholeoixodes).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3342E2FF23F8B8FECEFEE9.taxon	description	Current status: a synonym of Ixodes hexagonus (Camicas et al. 1998, under the genus Pholeoixodes). hirudo Fabricius, 1781, Acarus of some authors, but this name is not included in the reference below or in any other studies by Fabricius. Fabricius, J. C. (1781) Species insectorum exhibentes eorum differentias specificas, synonyma auctorum, loca natalia, metamorphosin adiectis observationibus, descriptionibus. Tomus II. Carol Ernest Bohhn, Hamburg and Kiel, 517 pp.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3242E2FF23F9D8FB11F915.taxon	description	Current status: a provisional synonym of Hyalomma albiparmatum in Guglielmone & Nava (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3D42EDFF23FEC8FC45FE21.taxon	description	Current status: a synonym of Ixodes vespertilionis (Neumann 1911 a).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3D42EDFF23FB8CFBDFFB6D.taxon	description	Current status: a name incertae sedis, as explained in Guglielmone & Nava (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3D42EDFF23F8E0FBAFF83D.taxon	description	Current status: a name incertae sedis (Neumann 1911 a, Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3C42ECFF23FE34FC66FDC9.taxon	description	Current status: a name incertae sedis in Guglielmone et al. (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3C42ECFF23FEC8FBDDFE05.taxon	description	Current status: a nomen dubium (Camicas et al. 1998, Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3C42ECFF23F9FCFB86F931.taxon	description	Current status: a synonym of Hyalomma marginatum (Apanaskevich & Horak 2008 b).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3C42ECFF23FAB0FB86FA4D.taxon	description	Current status: a synonym of Hyalomma marginatum (Apanaskevich & Horak 2008 b).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3C42ECFF23FB44FBDEFA99.taxon	description	Current status: a name incertae sedis, as discussed in Guglielmone & Nava (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3C42ECFF23F92EFB50F87E.taxon	description	Current status: a synonym of Hyalomma marginatum (Apanaskevich & Horak 2008 b). marginatus lacteolus Schulze, 1933, Dermacentor Schulze, P. (1933) Die Arten der Zeckengattung Dermacentor s. l. aus Europa, Asien und Neu-Guinea. Zeitschrift für Parasitenkunde, 6, 416 – 431. Current status: a synonym of Dermacentor marginatus (Filippova 1997, Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3F42EFFF23FE10FBE2FD2D.taxon	description	Current status: a synonym of Hyalomma scupense (Apanaskevich et al. 2010).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3F42EFFF23FD5CFBE2FC91.taxon	description	Current status: a synonym of Hyalomma scupense (Apanaskevich et al. 2010).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3F42EFFF23FEA4FBDDFE79.taxon	description	Current status: a nomen dubium (Camicas et al. 1998, Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3E42EEFF23FD5CFB2BFCF4.taxon	description	Current status: a synonym of Amblyomma ovale (Aragão & Fonseca 1961, Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3E42EEFF23FEA4FBA4FE79.taxon	description	Current status: a nomen dubium (Camicas et al. 1998, Guglielmone & Nava, 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3942E9FF23FCACFC10FC0D.taxon	description	Current status: a synonym of Dermacentor reticulatus (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3942E9FF23FB1CFB9FFA35.taxon	description	Current status: Ixodes plumbeus (preoccupied) is a name incertae sedis, which was renamed Ixodes dugesi by Gervais (1844), also a name incertae sedis, as explained in Guglielmone & Nava (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3942E9FF23FB8CFBDFFB6D.taxon	description	Current status: a name incertae sedis, as explained in Guglielmone & Nava (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3942E9FF23F8B8FC8BF875.taxon	description	Current status: a synonym of Ixodes simplex (Beaucournu 1966).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3942E9FF23FD5CFA6EFCBD.taxon	description	Current status: a nomen dubium in Guglielmone & Nava (2014), who treated the year of description as unknown,	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3942E9FF23FC3CFB46FB9D.taxon	description	Current status: a name incertae sedis in Neumann (1911 a) and Guglielmone & Nava (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3842E8FF23FD84FC02FC91.taxon	description	Current status: a synonym of Haemaphysalis sulcata (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3842E8FF23F90BFCA9F87E.taxon	description	putorii Razoumowski, 1789, Acarus Razoumowski, G. (1789) Histoire naturelle de Jurat et de ses environs; et celle des trois lacs de Neuchatel, Morat et Bienne. Libraire J. Mourer, Lausanne, 322 pp. Current status: a nomen dubium (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3B42EBFF23FE10FC00FD01.taxon	description	Current status: a nomen dubium, as explained in Estrada-Peña et al. (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3B42EBFF23FD38FC18FCF5.taxon	description	Current status: a synonym of Dermacentor marginatus (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3B42EBFF23FC64FD33FB9D.taxon	description	Current status: a name without a description or figures and thus a nomen nudum, although Camicas et al. (1998) treat it as a synonym of Dermacentor rhinocerinus.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3B42EBFF23FAF8FBE8FA69.taxon	description	Current status: a synonym of Dermacentor rhinocerinus (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3B42EBFF23FB8CFBA8FB41.taxon	description	Current status: a synonym of Hyalomma truncatum (Apanaskevich & Horak 2008 a).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3B42EBFF23F8DCFBB2F811.taxon	description	Current status: a synonym of Ixodes gibbosus (Saratsiotis 1970, Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3A42EAFF23FAF6FB85F9D9.taxon	description	Current status: a nomen dubium in Camicas et al. (1998) and Guglielmone & Nava (2014). rufipes glabratum Fonseca, Pinto, Colaço, Oliveira, Branco, da Gama, Franco and Lacerda, 1951, Hyalomma. Fonseca, F., Pinto, M. R., Colaço, A. F., Oliveira, J., Branco, F., Marques da Gama, M., Soares Franco, J. S. & Lacerda, M. T. (1951) Febre Q em Portugal. Epidemiologia da doença. Clinica Contemporanea, 5, 1 – 23. Current status: Apanaskevich & Horak (2008 b) treated Hyalomma rufipes glabratum as a synonym of Hyalomma marginatum, but it is a name without descriptions or figures and, thus a nomen nudum.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3A42EAFF23FB83FCA9FB5A.taxon	description	Current status: a nomen dubium (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3A42EAFF23F89EFC05F85E.taxon	description	Current status: a synonym of Hyalomma anatolicum (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FF3A42EAFF23FF78FC1FFEE9.taxon	description	Current status: a nomen nudum, as explained in Guglielmone & Nava (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FEC54315FF23F9FCFB50F931.taxon	description	Current status: a questionable synonym of Hyalomma anatolicum (Apanaskevich et al. 2010).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FEC54315FF23FE10FBCCFD2D.taxon	description	Current status: a synonym of Hyalomma rufipes (Apanaskevich & Horak 2008 b).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FEC54315FF23FD5CFAE9FC91.taxon	description	Current status: a synonym of Hyalomma impressum (Camicas et al. 1998, Apanaskevich et al., 2007).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FEC54315FF23FC88FADCFC45.taxon	description	Current status: a synonym of Hyalomma truncatum (Camicas et al. 1998, Apanaskevich & Horak 2008 a).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FEC54315FF23FF78FC05FEB5.taxon	description	Current status: a synonym of Hyalomma anatolicum (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FEC54315FF23FEA4FAC0FE79.taxon	description	Current status: a synonym of Hyalomma lusitanicum (Camicas et al. 1998, Apanaskevich et al., 2008 a).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FEC44314FF23FF78FCA1FEE9.taxon	description	Current status: a nomen nudum (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FEC44314FF23FBE1FC3CFB3B.taxon	description	Current status: a synonym of Hyalomma aegyptium (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FEC44314FF23FD3FFC13FCFE.taxon	description	Current status: a synonym of Haemaphysalis punctata (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FEC44314FF23FA86FC09F9BA.taxon	description	Current status: a nomen dubium (Dunlop 2011, Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FEC44314FF23F92CFC3CF8E1.taxon	description	Current status: a synonym of Hyalomma aegyptium (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FEC44314FF23F89FFC3CF85E.taxon	description	Current status: a synonym of Hyalomma aegyptium (Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FEC74317FF23FD5CFDE0FC29.taxon	description	Current status: a nomen dubium in Guglielmone & Nava (2014). The type locality of Dermacentor triangulatus was treated as unknown in Guglielmone & Nava (2014), but the locality is Corrientes, Argentina.	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FEC74317FF23F8DCFC61F811.taxon	description	Current status: a name incertae sedis (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FEC74317FF23FA24FBC9F9F9.taxon	description	Current status: a synonym of Hyalomma excavatum (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FEC74317FF23FC40FECFFBD5.taxon	description	Current status: a nomen dubium in Guglielmone & Nava (2014). The type locality for Ixodes trilineatus is the Canary Islands, Spain, but this was not recognized in Guglielmone & Nava (2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FEC64316FF23FEA4FC61FE79.taxon	description	Current status: a name incertae sedis (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FEC64316FF23FE10FBDDFDC9.taxon	description	Current status: a nomen dubium (Camicas et al. 1998, Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FEC64316FF23FD60FC61FCBD.taxon	description	Current status: a name incertae sedis (Guglielmone & Nava 2014).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FEC64316FF23F9D8FBA0F915.taxon	description	Current status: a synonym of Ixodes ricinus (Neumann 1911 a, Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FEC64316FF23FCACFB08FC45.taxon	description	Current status: a suppressed name to preserve Dermacentor andersoni Stiles, 1908 (Keirans 1976).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FEC14311FF23FF78FB1AFEB5.taxon	description	Current status: a synonym of Amblyomma sparsum (Theiler & Salisbury 1959, Camicas et al. 1998).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
8263FE48FEC14311FF23FEA4FC84FE5D.taxon	description	Current status: a synonym of Amblyomma sparsum in Camicas et al. (1998) and Guglielmone & Nava (2014), but it is a nomen nudum, as explained in Guglielmone et al. (2017).	en	Robbins, Richard G., Nava, Santiago, Ronai, Isobel, Chong, Kaylin L., Guglielmone, Alberto A. (2025): Type specimens of the world’s hard tick species (Acari: Ixodida: Ixodidae): collection data and depositories for all valid names and the current status of invalid names. Zootaxa 5606 (1): 1-287, DOI: 10.11646/zootaxa.5606.1.1, URL: https://doi.org/10.11646/zootaxa.5606.1.1
