taxonID	type	description	language	source
76CBCD16D4005DDA87CC45A4A9D12D95.taxon	description	Figs 17, 18, 19, 20, 21	en	Samraoui, Boudjéma, Vuataz, Laurent, Sartori, Michel (2025): Endemism and hidden diversity of Habrophlebia, Eaton, 1881 in Algeria’s Northeastern Mountains (Ephemeroptera, Leptophlebiidae). Alpine Entomology 9: 113-151, DOI: 10.3897/alpento.9.171490
76CBCD16D4005DDA87CC45A4A9D12D95.taxon	etymology	Etymology. The name Annaba derives from the Arabic ʿannāb (jujube), in reference to the jujube tree (Ziziphus jujuba), once abundant in the region. In antiquity, the city was known as Hippo Regius, a major centre of Roman North Africa and the episcopal See of Saint Augustine (354 – 430 CE). Later, it was renamed Bouna in Arabic and was known as Bône during the French period.	en	Samraoui, Boudjéma, Vuataz, Laurent, Sartori, Michel (2025): Endemism and hidden diversity of Habrophlebia, Eaton, 1881 in Algeria’s Northeastern Mountains (Ephemeroptera, Leptophlebiidae). Alpine Entomology 9: 113-151, DOI: 10.3897/alpento.9.171490
76CBCD16D4005DDA87CC45A4A9D12D95.taxon	description	Descriptions. Nymph. Body length of final instar, excluding caudal filaments, up to 6 mm for male and up to 7 mm for female. Cerci as long as body length. Coloration. General coloration medium brown; dark brown between ocelli, light brown in front and on the clypeus, especially in young nymphs. Upper portion of male eyes red-brick to reddish brown (Fig. 17 B). Antenna with pedicel dark brown, scape greyish brown and filament light brown, more or less entirely whitish in young nymphs. Pronotum medium brown, washed with dark brown in middle and anteriorly, lateral margins whitish; mesonotum medium brown, with dark brown maculae; lateroparapsidal sutures dark brown. Legs light to medium brown; femora dark brown in mature nymphs; tibiae medium brown; tarsi light brown to yellowish. Abdominal tergites dark brown to greyish brown, with two lighter maculae medio-anteriorly, especially well visible on segments I – VII (Fig. 17 A). Sternites medium brown in the middle, posterior corner underlined with dark brown laterally, thicker in mature nymphs, nervous ganglia well visible. Cerci and paracercus light brown at base, whitish distally, without dark banding. Head. Labrum rectangular, ca. 1.55 × wider than long (Fig. 17 C); dorsal surface covered distally with scattered stout setae; anterior margin with a row of stout, long and spatulate setae medially; emargination flared, U-shaped with flat denticles; ventral surface with two bunches of stout setae medially. Mandibles similar to other Habrophlebia species, incisor with 3 teeth, the lowest of left mandible with a clear indentation at base (Fig. 17 D), kinetodontium with 3 teeth, 10 – 11 long and thin setae below mola of right mandible (Fig. 17 E). Maxilla stocky, subapical row of 7 or 8 pectinate setae (Fig. 17 F); maxillary palp with three segments, segment 1 and 2 subequal in length, and longer than segment 3; segment 3 triangular 1.42 – 1.70 × longer than wide at base; all setae on palp segments stout and entire. Hypopharynx with highly developed superlinguae terminated by a membranous digitation. Labium with rhomboid glossae, outer margin and apex covered by short, broad setae; paraglossae enlarged laterally, covered with fine and long setae on the dorsal surface; with long, thick setae on the outer margin; labial palp with three segments, inner margin of segment 1 highly dilated near the middle, about 1.35 × longer than the maximum width, segments 2 slightly longer than segment 3, both ca. 0.65 – 0.70 × length of segment 1; segment 3 ca. 1.41 – 1.54 × longer than wide at base, conical shape and with 8 – 10 stout and long, simple setae on dorsal surface. Submentum with numerous long, stout and pointed setae laterally; ventral surface with numerous long, stout and pointed setae (Fig. 17 G). Thorax. Pronotum with anterior corners bearing few long and pointed setae laterally, and 5 – 7 stout and strong setae dorsally. Anterior margin of pronotum with numerous stout setae (Fig. 18 A). Fore legs femora elongated, ca. 2.45 × longer than wide, outer margin and upper surface covered with long, entire and pointed setae (Fig. 18 B); fore tibiae subequal in length to femora, outer margin with few thin and long setae, inner margin with several rows of long stout and entire setae; tarsi 0.5 × length of tibiae, outer margin with few long and thin setae, inner margin with long and pointed entire setae. Middle legs similar to fore legs, femora ca. 2.6 × longer than wide, dorsal surface of femora with more numerous and slightly longer stout and pointed setae; tibiae and femora of subequal length; tarsi 0.5 × length of tibiae. Hind legs with femora 2.8 × longer than wide (Fig. 18 C), dorsal surface covered with stout, long, pointed or blunt and entire (non-feathered) setae (Fig. 18 D); hind tibiae as long as hind femora, outer margin with numerous stout, pointed setae; inner margin with stout, pointed and entire setae; tarsi 0.5 × length of tibiae, outer margin with long and thin setae, inner margin with long, stout, pointed and entire setae. Claws of all legs slightly hooked, with 15 – 17 denticles that decrease in size from the apex to the tarsus (Fig. 18 E). Abdomen. Posterolateral expansions only on segments VIII and IX. Posterior margin of tergite IX with triangular, broad, pointed spines, ca. 1.5 – 2 × longer than wide at base (Fig. 19 A); tergite VIII with triangular, broad spines 1.5 – 2 × longer than wide (Fig. 19 B); tergite VII with slightly shorter spines than tergite VIII (Fig. 19 C); tergite VI, with short and broad spines (Fig. 19 D), and tergite V with minute but broad spines (Fig. 19 E); tergites I – IV with barely visible spines. Gills present on segments I – VII; all gills elongated with long filaments; dorsal lamella bearing 8 – 10 filaments, ventral lamella with 3 – 6 filaments (Fig. 19 F). Imagos. Male imago. Size: body length: 5.5 – 6.0 mm; forewings missing; cerci and terminal filament broken. Head dark brown; basal portion of compound eyes blackish, upper portion orange brown, scape and pedicel dark brown, flagellum broken. Thorax. Pronotum dark brown; meso- and metanotum uniformly dark brown, pleurae, coxae, and trochanters greyish brown, washed with dark brown; fore femora and fore tibiae dark brown, tarsi light brown, apex of each segment dark brown; mid- and hind legs with femora light brown, dark brown distally, tibiae medium brown, dark brown at base, and tarsi medium brown, except the last segment light brown. Femur / tibia / tarsi ratio in fore leg: 1 / 1.4 / 0.5 / 0.5 / 0.3 / 0.2; mid leg: 1 / 1.2 / 0.1 / 0.07 / 0.05 / 0.1; hind leg: 1 / 1.2 / 0.1 / 0.05 / 0.05 / 0.15. Fore claws similar, paddle-shaped, mid- and hind claws dissimilar, one paddle-like and one hooked. Hindwing with rounded costal process approximately in the middle of the wing; vein Sc long and well separated from the costal vein, reaching the costal margin almost at the tip of the wing (Fig. 20 C). Abdomen. Tergites and sternites medium brown, posterior margin of each segment dark brown. Styliger plate medium brown, dark brown along the margins, gonopods broken. Posterior margin of the styliger plate straight and clearly protruding in the middle, median incision U-shaped (Fig. 20 D). Penis lobes rounded and tight together, ventral spine long, thin and straight, almost reaching the base of the styliger plate. Cerci and terminal filament broken. Female imago. Size: body length: 6.5 – 7.0 mm; forewing length: 7.0 – 7.5 mm. Forewing transparent, slightly tinted in brown in subcostal field; longitudinal and transversal veins dark brown; pterostigmatic area with ca. 10 oblique and simple transversal veins. MA and MP forks asymmetrical, cubital field with two long and two short intercalary veins (Fig. 20 A). Hindwing with rounded costal process approximately in the middle of the wing; vein Sc long and close to the costal vein, reaching the costal margin before the tip of the wing (Fig. 20 B). Eggs ovoid, ca. 180 – 195 µm × 80 – 100 µm, with longitudinal ribs non-punctuated, long and entire, most of them running from one pole to the other (Fig. 21 A). Ribs relatively broad, ca. 4.5 – 5.0 µm in width. In some specimens, chorion folded. Micropyle in equatorial area (Fig. 21 B).	en	Samraoui, Boudjéma, Vuataz, Laurent, Sartori, Michel (2025): Endemism and hidden diversity of Habrophlebia, Eaton, 1881 in Algeria’s Northeastern Mountains (Ephemeroptera, Leptophlebiidae). Alpine Entomology 9: 113-151, DOI: 10.3897/alpento.9.171490
499CE1AB98475D2481076080AF5C9829.taxon	description	Figs 12, 13, 14, 15, 16	en	Samraoui, Boudjéma, Vuataz, Laurent, Sartori, Michel (2025): Endemism and hidden diversity of Habrophlebia, Eaton, 1881 in Algeria’s Northeastern Mountains (Ephemeroptera, Leptophlebiidae). Alpine Entomology 9: 113-151, DOI: 10.3897/alpento.9.171490
499CE1AB98475D2481076080AF5C9829.taxon	etymology	Etymology. The name of El Kala derives from the Arabic al-Qalʿa (“ the fortress ”), a reference to its strategic role through history. During the French period it was known as La Calle, while in antiquity it was recorded as Thinisa in Numidia. Descriptions. Nymph. Body length of final instar, excluding caudal filaments, up to 6 mm for male and up to 7 mm for female. Cerci as long as body length. Coloration. General coloration medium brown; dark brown between ocelli, whitish in front of the eyes, light brown in front and on the clypeus. Upper portion of male eyes light brown to orange. Antenna with pedicel greyish brown, scape light brown and filament yellowish. Pro- and mesonotum dark brown, with light brown maculae, on medium and lateral margins; lateroparapsidal sutures dark brown. Legs light to medium brown; dorsal surface of fore femora dark brown, fore tibiae greyish brown; mid- and hind legs light brown to yellowish, except at apex of femora grayish brown; tarsi light brown to yellowish. Abdominal tergites medium brown with three pairs of lighter maculae, especially well visible on segments I – VI (Fig. 12 A); posterolateral angles and posterior margin of tergites I – VIII dark brown. Sternites medium brown to greyish brown, nervous ganglia well visible (Fig. 12 B). Cerci and paracercus medium brown at base, lighter distally, dark banding slightly visible every two segments proximally. Head. Labrum rectangular, ca. 1.85 × wider than long (Fig. 12 C); dorsal surface covered distally with scattered stout setae; anterior margin with a row of stout, long and spatulate setae medially; emargination narrow, U-shaped with flat denticles; ventral surface with two bunches of stout setae medially. Mandibles similar to other Habrophlebia species, incisor with 3 teeth, the lowest of the left mandible with a clear indentation (Fig. 12 D), kinetodontium with 3 teeth, 14 – 15 long and thin setae below mola of right mandible (Fig. 12 E). Maxilla stocky, subapical row of 5 or 6 pectinate setae (Fig. 12 F); maxillary palp with three segments, segment 1 and 2 subequal in length, and longer than segment 3; segment 3 triangular 1.51 – 1.71 × longer than wide at base; all setae on palp segments stout and entire. Hypopharynx with highly developed superlinguae terminated by a membranous digitation (Fig. 12 G). Labium with rhomboid glossae, outer margin and apex covered by short, broad setae; paraglossae enlarged laterally, covered with fine and long setae on the dorsal surface; with long, thick setae on the outer margin; labial palp with three segments, inner margin of segment 1 highly dilated near the middle, about 1.15 – 1.20 × longer than the maximum width, segments 2 0.85 × length of segment 1, segment 3, ca. 0.75 × length of segment 1; segment 3 ca. 1.7 – 1.8 × longer than wide at base, conical shape and with 5 – 7 stout and long, simple setae on dorsal surface. Submentum with numerous long, stout and pointed setae laterally and on ventral surface (Fig. 12 H). Thorax. Pronotum with anterior corners bearing numerous long and pointed setae laterally, and dorsally. Anterior margin of pronotum with a row of stout setae (Fig. 13 A). Fore legs femora stocky, ca. 2.1 × longer than wide, outer margin and upper surface covered with long, entire and pointed setae (Fig. 13 B); fore tibiae slightly shorter than femora, outer margin with thin and long setae, inner margin with long stout and feathered or entire setae especially near apex; tarsi 0.55 × length of tibiae, outer margin with long and thin setae, inner margin with long and pointed entire setae. Middle legs similar to fore legs, femora ca. 2.5 × longer than wide, dorsal surface of femora with numerous and long, stout and pointed setae; tibiae and femora of subequal length; tarsi 0.5 × length of tibiae. Hind legs with femora 2.8 – 3.0 × longer than wide (Fig. 13 C), dorsal surface covered with stout, long, pointed and entire setae (Fig. 13 D); hind tibiae as long as hind femora, outer margin with a row of stout, pointed setae and few thin and long setae; inner margin with stout, pointed and entire setae (Fig. 13 E); tarsi 0.4 × length of tibiae, outer margin with few long and thin setae, inner margin with long, stout, pointed and entire setae. Claws of all legs slightly hooked, with 17 – 21 denticles subequal in size (Fig. 13 F). Abdomen. Posterolateral expansions only on segments VIII and IX. Posterior margin of tergite IX with triangular, broad, pointed spines, ca. 1.5 – 2 × longer than wide at base (Fig. 14 A); tergite VIII with triangular, narrow spines 2 × longer than wide (Fig. 14 B); from tergite VII to tergite V, spines becoming smaller, needle-shaped (Fig. 14 C – E); tergites I – IV with barely visible spines. Gills present on segments I – VII; all gills elongated with long filaments; dorsal lamella bearing 7 – 8 filaments, ventral lamella with 4 – 5 filaments (Fig. 14 F). Imagos. Male imago in poor condition, only mid- and hind legs and abdomen preserved. Mid and hind legs coloration as in nymphs, whitish with apex of femur brownish. Femur / tibia / tarsi ratio in mid leg: 1 / 1.3 / 0.06 / 0.05 / 0.05 / 0.1; hind leg: 1 / 1.3 / 0.1 / 0.08 / 0.06 / 0.1. Mid- and hind claws dissimilar, one paddle-like and one hooked. Abdomen. Coloration similar to male nymphs. Styliger plate medium brown, dark brown along the margins, first segment of the gonopods medium brown, segments 2 and 3 yellowish brown. Posterior margin of the styliger plate concave, median incision U-shaped (Fig. 15 C); segment 1 slightly longer than segments 2 and 3 combined; inner margin of segment 1 with a broad base, and with bulge on the outer margin. Penis lobes rounded and tight together, ventral spine long, thin and slightly curved outwards, reaching the base of the styliger plate (Fig. 15 D). Cerci and terminal filament broken. Female imago: body length: 4.0 – 4.5 mm; forewing length: 4.5 – 5.0 mm. Forewing transparent, slightly tinted in brown in subcostal field; longitudinal and transversal veins dark brown; pterostigmatic area with ca. 10 oblique and simple transversal veins. MA and MP forks asymmetrical, cubital field with two long and three short intercalary veins (Fig. 15 A). Hindwing with rounded costal process approximately in the middle of the wing; vein Sc long, reaching the costal margin almost at the tip of the wing (Fig. 15 B). Eggs ovoid, ca. 200 µm × 90 µm, with longitudinal ribs non-punctuated, the majority running from one pole to the other (Fig. 16 A); chorion covered with thin membranous filaments perpendicular to pole axis, sometimes longitudinal. Micropyle in equatorial area (Fig. 16 B).	en	Samraoui, Boudjéma, Vuataz, Laurent, Sartori, Michel (2025): Endemism and hidden diversity of Habrophlebia, Eaton, 1881 in Algeria’s Northeastern Mountains (Ephemeroptera, Leptophlebiidae). Alpine Entomology 9: 113-151, DOI: 10.3897/alpento.9.171490
ECABEE96AA6E53318E794500392EB387.taxon	description	Figs 22, 23, 24, 25	en	Samraoui, Boudjéma, Vuataz, Laurent, Sartori, Michel (2025): Endemism and hidden diversity of Habrophlebia, Eaton, 1881 in Algeria’s Northeastern Mountains (Ephemeroptera, Leptophlebiidae). Alpine Entomology 9: 113-151, DOI: 10.3897/alpento.9.171490
ECABEE96AA6E53318E794500392EB387.taxon	etymology	Etymology. The name Edough (Djebel Edough), designating the prominent mountain massif overlooking the city of Annaba, is thought to derive from undocumented Berber (Amazigh) roots. In antiquity, it was referred to as Mons Pappua by Procopius who describes it as a Numidian stronghold, and as Mons Giddaba by Saint Augustine.	en	Samraoui, Boudjéma, Vuataz, Laurent, Sartori, Michel (2025): Endemism and hidden diversity of Habrophlebia, Eaton, 1881 in Algeria’s Northeastern Mountains (Ephemeroptera, Leptophlebiidae). Alpine Entomology 9: 113-151, DOI: 10.3897/alpento.9.171490
ECABEE96AA6E53318E794500392EB387.taxon	description	Descriptions. Nymph. Body length of final instar, excluding caudal filaments, up to 7 mm for female; male nymphs unknown. Cerci as long as body length. Coloration. General coloration medium brown; dark brown between ocelli, light brown in front and on the clypeus (Fig. 22 A), especially in young nymphs (Fig. 22 B). Antenna with pedicel dark brown, scape greyish brown and filament light brown, more or less entirely whitish in young nymphs. Pronotum medium brown, washed with dark brown sub-laterally and with a dark brown M-shaped macula in middle, lateral margins whitish; mesonotum medium brown, with dark brown maculae; lateroparapsidal sutures dark brown. Legs light to medium brown; femora dark brown in mature nymphs; tibiae medium brown, with dark brown apex in mature nymphs; tarsi light brown to yellowish. Abdominal tergites dark brown to greyish brown, with two lighter maculae medio-anteriorly, especially well visible on segments I – VII in young nymphs (Fig. 22 B); maculae less visible in mature nymphs (Fig. 22 A). Sternites medium brown in the middle, posterior margin underlined with dark brown laterally, nervous ganglia hardly visible. Cerci and paracercus light brown at base, whitish distally, without dark banding. Head. Labrum rectangular, 1.75 – 1.79 × wider than long (Fig. 22 C); dorsal surface covered distally with scattered stout setae; anterior margin with a row of stout, long and spatulate setae medially; emargination flared, U-shaped with flat denticles; ventral surface with two bunches of stout setae medially. Mandibles similar to other Habrophlebia species, incisor with 3 teeth, the lowest of left mandible with a clear indentation at base (Fig. 22 D), kinetodontium with 3 teeth, 17 – 22 long and thin setae below mola of right mandible (Fig. 22 E). Maxilla stocky, subapical row of 7 or 8 pectinate setae (Fig. 22 F); maxillary palp with three segments, segment 1 and 2 subequal in length, and longer than segment 3; segment 3 triangular 1.41 – 1.75 × longer than wide at base; setae on the dorsal surface of segment 2 feathered (Fig. 23 B). Hypopharynx with highly developed superlinguae terminated by a membranous digitation (Fig. 23 A). Labium with rhomboid glossae, outer margin and apex covered by short, broad setae; paraglossae enlarged laterally, covered with fine and long setae on dorsal surface; with long, thick setae on the outer margin; labial palp with three segments, inner margin of segment 1 highly dilated near the middle, about 1.35 × longer than maximum width, segments 2 slightly shorter than segment 3, segment 2 ca. 0.67 × length of segment 1, and segment 3 ca. 0.74 × length of segment 1; segment 3 ca. 1.8 – 2.0 × longer than wide at base, conical shape and with 4 – 7 stout and long, simple setae on dorsal surface (Fig. 22 G). Submentum with numerous long, stout and pointed setae laterally; ventral surface with few long, stout and pointed setae (Fig. 22 G). Thorax. Pronotum with anterior corners bearing few long and pointed setae laterally, and 4 – 6 stout and strong setae dorsally. Anterior margin of pronotum with numerous stout setae (Fig. 23 C). Fore legs femora elongated, ca. 2.75 × longer than wide, outer margin and upper surface covered with long, entire and pointed setae (Fig. 23 D); fore tibiae length 0.95 × femora length, outer margin without or with few thin and long setae, inner margin with several rows of long stout, entire or feathered setae; tarsi 0.6 × length of tibiae, outer margin with few long and thin setae, inner margin with long, pointed and feathered setae. Middle legs similar to fore legs, femora ca. 3.2 × longer than wide (Fig. 23 E), dorsal surface of femora with more numerous and slightly longer stout, entire or slightly feathered, pointed setae; tibiae length 0.92 × femora length; tarsi 0.45 × length of tibiae. Hind legs with femora 3.5 × longer than wide (Fig. 23 F), dorsal surface covered with stout, long, pointed, feathered setae (Fig. 23 G); tibiae length 0.94 × femora length, outer margin with numerous stout, pointed and entire setae; inner margin with stout, pointed and feathered setae; tarsi 0.42 × length of tibiae, outer margin with long and thin setae, inner margin with long, stout, pointed and entire setae. Claws of all legs slightly hooked, with 15 – 17 denticles that decrease in size from the apex to the tarsus (Fig. 23 H). Abdomen. Posterolateral expansions only on segments VIII and IX. Posterior margin of tergite IX with triangular, narrow, pointed spines, ca. 2.5 – 3 × longer than wide at base (Fig. 24 A); tergite VIII with triangular, narrow spines 1.5 – 2 × longer than wide (Fig. 24 B); tergite VII and VI with a mix of short, narrow spines and minute, needle-shaped spines (Fig. 24 C, D); tergite V with minute needle-shaped spines (Fig. 24 E); tergites I – IV with barely visible spines. Gills present on segments I – VII; all gills elongated with long filaments; dorsal lamella bearing 8 – 10 filaments, ventral lamella with 4 – 5 filaments. Imagos. unknown. Eggs (extracted from mature female nymphs) ovoid, ca. 175 – 185 µm × 70 – 80 µm, with longitudinal ribs non-punctuated, long and entire, most of them not running from one pole to the other (Fig. 25 A). Ribs relatively narrow, ca. 2.5 – 3.0 µm in width. Micropyle in equatorial area (Fig. 25 B).	en	Samraoui, Boudjéma, Vuataz, Laurent, Sartori, Michel (2025): Endemism and hidden diversity of Habrophlebia, Eaton, 1881 in Algeria’s Northeastern Mountains (Ephemeroptera, Leptophlebiidae). Alpine Entomology 9: 113-151, DOI: 10.3897/alpento.9.171490
7E42BD861B6E56D88325BF7B444E0626.taxon	description	Figs 2, 3, 4, 5, 6	en	Samraoui, Boudjéma, Vuataz, Laurent, Sartori, Michel (2025): Endemism and hidden diversity of Habrophlebia, Eaton, 1881 in Algeria’s Northeastern Mountains (Ephemeroptera, Leptophlebiidae). Alpine Entomology 9: 113-151, DOI: 10.3897/alpento.9.171490
7E42BD861B6E56D88325BF7B444E0626.taxon	etymology	Etymology. The name “ Ghora ” is derived from the Coot (Fulica atra), known locally as “ Ghor ”, whose conspicuous white frontal shield evokes the image of a snow-capped peak. Mount Ghora (Djebel Ghora), situated on the Algerian – Tunisian border, takes its name from this association and forms part of the Kroumiria massif.	en	Samraoui, Boudjéma, Vuataz, Laurent, Sartori, Michel (2025): Endemism and hidden diversity of Habrophlebia, Eaton, 1881 in Algeria’s Northeastern Mountains (Ephemeroptera, Leptophlebiidae). Alpine Entomology 9: 113-151, DOI: 10.3897/alpento.9.171490
7E42BD861B6E56D88325BF7B444E0626.taxon	description	Descriptions. Nymph. Body length of final instar, excluding caudal filaments, up to 5 mm for male and up to 6 mm for female. Cerci as long as body length. Coloration. General coloration medium brown; light brown between compound eyes and antennae. Upper portion of male eyes light brown to orange (Fig. 2 A). Antenna with scape and pedicel greyish brown, filament yellowish. Pro- and mesonotum medium brown, with light brown maculae, on medium; lateroparapsidal sutures dark brown. Legs light to medium brown; dorsal surface of femora darker; tibiae and tarsi light brown to yellowish. Abdominal tergites medium brown with a sagittal line paler; distal half of each tergite darker, sometimes with two elongated light maculae medio-anteriorly, on segments I – VI, especially in younger nymphs. Sternites light to medium brown, nervous ganglia well visible. Cerci and paracercus medium brown at base, lighter distally, rather uniform in color, without apparent banding. Head. Labrum rectangular, ca. 1.6 × wider than long (Fig. 2 B); dorsal surface covered distally with scattered stout setae; anterior margin with a row of stout, long and spatulate setae medially; emargination smooth, U-shaped, flared, with indistinct flat denticles; ventral surface with two bunches of stout setae medially. Mandibles similar to other Habrophlebia species, incisor with three teeth, the lowest on left mandible characterised by distinct basal indentation (Fig. 2 C), kinetodontium with 3 teeth, 11 – 15 long and thin setae below mola of right mandible (Fig. 2 D). Maxilla stocky, subapical row of 6 – 7 pectinate setae (Fig. 2 E); maxillary palp with three segments, segment 1 and 2 subequal in length, and longer than segment 3; segment 3 triangular 1.11 – 1.15 × longer than wide at base; all setae on palp segments stout and entire. Hypopharynx with highly developed superlinguae terminated by a membranous digitation (Fig. 2 F). Labium with rhomboid glossae, outer margin and apex covered by short, broad setae; paraglossae enlarged laterally, covered with fine and long setae on the dorsal surface; with long, thick setae on outer margin; labial palp with three segments, inner margin of segment 1 highly dilated near the middle, about 1.20 × longer than the maximum width, segment 2 as long as segment 3, ca. 0.8 × length of segment 1; segment 3 ca. 1.75 – 1.80 × longer than wide at base, conical shape and with 7 – 8 stout and long, simple setae on dorsal surface. Submentum with numerous long, stout and pointed setae laterally; ventral surface with few long, stout and pointed setae (Fig. 2 G). Thorax. Pronotum with anterior corners bearing few long and pointed setae laterally, and 6 – 8 stout and strong setae dorsally. Anterior margin of pronotum with 3 – 5 stout setae laterally (Fig. 3 A). Fore legs femora elongated, ca. 2.65 × longer than wide, outer margin and upper surface covered with long, entire and pointed setae (Fig. 3 B); tibiae slightly shorter than femora, outer margin with sparse thin and long setae, inner margin with long stout and feathered or entire setae more numerous near apex; tarsi 0.55 × length of tibiae, outer margin with sparse long and thin setae, inner margin with long and pointed entire setae. Middle legs similar to fore legs, femora ca. 2.9 × longer than wide, dorsal surface of femora with more numerous and slightly longer stout and pointed setae (Fig. 3 C); tibiae slightly shorter than femora; tarsi 0.5 × length of tibiae. Hind legs with femora 3.4 × longer than wide (Fig. 3 D), dorsal surface covered with stout, long and short, pointed or rounded and entire (non-feathered) setae more or less in rows (Fig. 3 E); hind tibiae as long as hind femora, outer margin with scattered stout, pointed setae; inner margin with stout, pointed and entire setae (Fig. 3 F); tarsi 0.45 × length of tibiae, outer margin with few long and thin setae, inner margin with long, stout, pointed and entire setae. Claws of all legs slightly hooked, with 12 – 14 denticles that decrease in size from the apex to the tarsus (Fig. 3 G). Abdomen. Posterolateral expansions only on segments VIII and IX. Posterior margin of tergite IX with triangular, broad and narrow, pointed spines, ca. 1.5 – 2.5 × longer than wide at base (Fig. 4 A); tergite VIII with small triangular, narrow spines 1.5 – 2 × longer than wide (Fig. 4 B); tergite VII with tiny needle-shaped spines (Fig. 4 C); tergites VI and above with barely visible spines. Gills present on segments I – VII; all gills elongated with long filaments; dorsal lamella bearing 4 – 5 filaments, ventral lamella with 2 – 3 filaments (Fig. 4 D). Imago. Male imago. Size: body length: ca. 5.5 mm; forewing length: 5.0 – 5.5 mm; cerci and terminal filament broken. Head dark brown; basal portion of compound eyes greyish, upper portion light brown, scape dark brown, pedicel medium brown, flagellum light brown (Fig. 5 A). Thorax. Pronotum dark brown; meso- and metanotum uniformly dark brown, pleurae, coxae, and trochanters medium brown, washed with dark brown; fore femora dark brown, fore tibiae medium brown, tarsi light brown; mid- and hind legs with femora medium brown, dark brown distally, tibiae and tarsi medium brown. Femur / tibia / tarsi ratio in fore leg: 1 / 1.15 / 0.6 / 0.4 / 0.3 / 0.15; mid leg: 1 / 1.15 / 0.05 / 0.08 / 0.08 / 0.2; hind leg: 1 / 1.25 / 0.1 / 0.1 / 0.1 / 0.15. Fore claws similar, paddle-shaped, mid- and hind claws dissimilar, one paddle-like and one hooked. Forewing transparent, pterostigmatic area milky with 6 – 7 oblique and simple transversal veins, longitudinal veins medium brown, transversal veins light brown. MA and MP forks asymmetrical, cubital field with two long and sometimes one short intercalary veins (Fig. 5 B). Hindwing with rounded costal process approximately in the middle of the wing; vein Sc long, very close to the costal one and reaching it almost at the tip of the wing (Fig. 5 C). Abdomen. Tergites greyish brown, posterior margin of each segment dark brown; sternites light brown, nervous ganglia tinted with medium brown. Styliger plate medium brown, dark brown along the margins, first segment of the gonopods medium brown at base, lighter at apex, segments 2 and 3 yellowish brown. Posterior margin of the styliger plate almost straight in the middle, median incision V-shaped (Fig. 5 D); segment 1 subequal in length to segments 2 and 3 combined; inner margin of segment 1 with a broad base, and without bulge on outer margin. Penis lobes rounded and tight together, ventral spine long, thick and curved outwards, almost reaching the base of the styliger plate (Fig. 5 E). Cerci and terminal filament broken. Female imago. Size: body length: 4.5 – 5.0 mm; forewing length: 5.0 – 5.5 mm. Forewing transparent, longitudinal and transversal veins dark brown; pterostigmatic area with 7 – 9 oblique and simple transversal veins. MA and MP forks asymmetrical, cubital field with two long intercalary veins. Hindwing with rounded costal process approximately in the middle of the wing; vein Sc reaching the costal margin at mid length to the tip of the wing. Eggs ovoid, ca. 225 µm × 95 µm, with longitudinal ribs punctuated or not punctuated, bearing transversal small outgrowths, making the whole structure resemble barbed wire (Figs 6 A, B). Micropyle in equatorial area (Figs 6 C, D).	en	Samraoui, Boudjéma, Vuataz, Laurent, Sartori, Michel (2025): Endemism and hidden diversity of Habrophlebia, Eaton, 1881 in Algeria’s Northeastern Mountains (Ephemeroptera, Leptophlebiidae). Alpine Entomology 9: 113-151, DOI: 10.3897/alpento.9.171490
50EA07024EA9590093CBE490288061A8.taxon	description	Figs 7, 8, 9, 10, 11	en	Samraoui, Boudjéma, Vuataz, Laurent, Sartori, Michel (2025): Endemism and hidden diversity of Habrophlebia, Eaton, 1881 in Algeria’s Northeastern Mountains (Ephemeroptera, Leptophlebiidae). Alpine Entomology 9: 113-151, DOI: 10.3897/alpento.9.171490
50EA07024EA9590093CBE490288061A8.taxon	etymology	Etymology. The name Ubus derives from pre-Roman, possibly Punic or Amazigh (Berber), roots. In ancient sources, Ubus referred to the river now known as Oued Seybouse, one of Algeria’s major waterways. The river originates in the Hauts Plateaux and flows northwards to the Mediterranean Sea near Annaba.	en	Samraoui, Boudjéma, Vuataz, Laurent, Sartori, Michel (2025): Endemism and hidden diversity of Habrophlebia, Eaton, 1881 in Algeria’s Northeastern Mountains (Ephemeroptera, Leptophlebiidae). Alpine Entomology 9: 113-151, DOI: 10.3897/alpento.9.171490
50EA07024EA9590093CBE490288061A8.taxon	description	Descriptions. Nymph. Body length of final instar, excluding caudal filaments, up to 7 mm for male and up to 8 mm for female. Cerci as long as body length. Coloration. General coloration medium brown; dark brown between ocelli, light brown in front and on the clypeus. Upper portion of male eyes light brown to orange (Fig. 7 A). Antenna with pedicel greyish brown, scape light brown and filament yellowish. Pro- and mesonotum medium brown, with light brown maculae, on medium and lateral margins; lateroparapsidal sutures dark brown. Legs light to medium brown; dorsal surface of fore femora dark brown in mature nymphs; tibiae light brown to yellowish, except at base with the femur grayish brown; tarsi light brown to yellowish. Abdominal tergites dark brown with a sagittal line paler and two elongated light maculae medio-anteriorly, especially well visible on segments I – VI. Sternites light to medium brown, nervous ganglia well visible. Cerci and paracercus medium brown at base, lighter distally, dark banding slightly visible every two segments proximally. Head. Labrum rectangular, ca. 1.75 × wider than long (Fig. 7 B); dorsal surface covered distally with scattered stout setae; anterior margin with a row of stout, long and spatulate setae medially; emargination narrow, U – shaped without flat denticles; ventral surface with two bunches of stout setae medially. Mandibles similar to other Habrophlebia species, incisor with 3 teeth, kinetodontium with 3 teeth (Fig. 7 C), 13 – 14 long and thin setae below mola of right mandible (Fig. 7 D). Maxilla stocky, subapical row of 6 – 7 pectinate setae (Fig. 7 E); maxillary palp with three segments, segment 1 and 2 subequal in length, and longer than segment 3; segment 3 triangular 1.21 – 1.45 × longer than wide at base; all setae on palp segments stout and entire. Hypopharynx with highly developed superlinguae terminated by a membranous digitation (Fig. 7 F). Labium with rhomboid glossae, outer margin and apex covered by short, broad setae; paraglossae enlarged laterally, covered with fine and long setae on dorsal surface; with long, thick setae on outer margin; labial palp with three segments, inner margin of segment 1 highly dilated near the middle, about 1.25 – 1.30 × longer than maximum width, segment 2 as long as segment 3, ca. 0.7 – 0.8 × length of segment 1; segment 3 ca. 1.8 – 2.0 × longer than wide at base, conical shape and with 6 – 7 stout and long, simple setae on dorsal surface. Submentum with numerous long, stout and pointed setae laterally; ventral surface with few long, stout and pointed setae (Fig. 7 G). Thorax. Pronotum with anterior corners bearing few long and pointed setae laterally, and 1 – 3 stout and strong setae dorsally. Anterior margin of pronotum without or with rarely 1 – 2 stout setae (Fig. 8 A). Fore legs femora elongated, ca. 2.5 × longer than wide, outer margin and upper surface covered with long, entire and pointed setae (Fig. 8 B); fore tibiae subequal in length to femora, outer margin with thin and long setae, inner margin with long stout and feathered or entire setae especially near apex; tarsi 0.5 × length of tibiae, outer margin with long and thin setae, inner margin with long and pointed entire setae. Middle legs similar to fore legs, femora ca. 3 × longer than wide, dorsal surface of femora with more numerous and slightly longer stout and pointed setae (Fig. 8 C); tibiae and femora of subequal length; tarsi 0.5 × length of tibiae. Hind legs with femora 3.5 × longer than wide (Fig. 8 D), dorsal surface covered with stout, long, pointed and entire (non-feathered) setae (Fig. 8 E); hind tibiae as long as hind femora, outer margin with scattered stout, pointed setae; inner margin with stout, pointed and entire setae (Fig. 8 F); tarsi 0.5 × length of tibiae, outer margin with long and thin setae, inner margin with long, stout, pointed and entire setae. Claws of all legs slightly hooked, with 14 – 16 denticles that decrease in size from the apex to the tarsus (Fig. 8 G). Abdomen. Posterolateral expansions only on segments VIII and IX. Posterior margin of tergite IX with triangular, broad, pointed spines, ca. 1.5 – 2 × longer than wide at base (Fig. 9 A); tergite VIII with triangular, narrow spines 2 × longer than wide (Fig. 9 B); from tergite VII to tergite V, spines becoming smaller, needle-shaped (Figs 9 C, D); tergites I – IV with barely visible spines. Gills present on segments I – VII; all gills elongated with long filaments; dorsal lamella bearing 5 – 6 filaments, ventral lamella with 3 – 4 filaments (Fig. 9 E). Imagos. Male imago. Size: body length: 5.5 – 6.0 mm; forewing length: 5.3 – 5.6 mm; cerci and terminal filament broken. Head medium brown, dark brown between ocelli; basal portion of compound eyes greyish, upper portion orange brown, scape medium brown, pedicel dark brown, flagellum light brown. Thorax. Pronotum greyish brown, washed with dark brown; meso- and metanotum uniformly dark brown, pleurae, coxae, and trochanters greyish brown, washed with dark brown; fore femora dark brown, fore tibiae medium brown, tarsi light brown; mid- and hind legs with femora greyish brown, dark brown distally, tibiae medium brown and tarsi medium brown, except the last segment light brown. Femur / tibia / tarsi ratio in fore leg: 1 / 1.2 / 0.4 / 0.4 / 0.3 / 0.1; mid leg: 1 / 1.2 / 0.1 / 0.06 / 0.06 / 0.1; hind leg: 1 / 1.05 / 0.1 / 0.05 / 0.05 / 0.15. Fore claws similar, paddle-shaped, mid- and hind claws dissimilar, one paddle-like and one hooked. Forewing transparent, pterostigmatic area milky with 6 – 8 oblique and simple transversal veins, longitudinal veins medium brown, transversal veins light brown. MA and MP forks asymmetrical, cubital field with two long intercalary veins (Fig. 10 A). Hindwing with rounded costal process approximately in the middle of the wing; vein Sc short, reaching the costal margin at mid-length between costal process and the tip of the wing (Fig. 10 C). Abdomen. Tergites and sternites medium brown, posterior third or half of each segment dark brown. Styliger plate medium brown, dark brown along the margins, first segment of the gonopods medium brown, segments 2 and 3 yellowish brown. Posterior margin of the styliger plate almost straight in the middle, median incision V-shaped (Fig. 10 D); segment 1 subequal in length to segments 2 and 3 combined; inner margin of segment 1 with a broad base, and without bulge on the outer margin. Penis lobes rounded and tight together, ventral spine long, thin and curved outwards, almost reaching the base of the styliger plate (Fig. 10 E). Cerci and terminal filament broken. Female imago. Size: body length: 7.0 – 7.5 mm; forewing length: 7.5 – 8.0 mm. Forewing transparent, slightly tinted in brown in subcostal field; longitudinal and transversal veins dark brown; pterostigmatic area with ca. 7 – 8 oblique and simple transversal veins. MA and MP forks asymmetrical, cubital field with two long intercalary veins. Hindwing with rounded costal process approximately in the middle of the wing; vein Sc long, reaching the costal margin almost at the tip of the wing (Fig. 10 B). Eggs ovoid, ca. 245 µm × 100 µm, with longitudinal ribs punctuated, bearing transversal small outgrowths, making the whole structure resemble barbed wire (Fig. 11 A). Micropyle in equatorial area (Fig. 11 B).	en	Samraoui, Boudjéma, Vuataz, Laurent, Sartori, Michel (2025): Endemism and hidden diversity of Habrophlebia, Eaton, 1881 in Algeria’s Northeastern Mountains (Ephemeroptera, Leptophlebiidae). Alpine Entomology 9: 113-151, DOI: 10.3897/alpento.9.171490
