taxonID	type	description	language	source
7C4C87C2FF8BFFBCA3D556B6FBFCF8CC.taxon	description	urn: lsid: zoobank. org: act: 145934 A 3 - 4 BA 2 - 4 A 8 A-AC 15 - 294 C 0150 D 5 CB Figs 1, 3 A – C, 4 A – D, 6 – 7	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF8BFFBCA3D556B6FBFCF8CC.taxon	diagnosis	Diagnosis Larger-sized (TL: 4.9 – 5.8 mm), oblong-oval species. Elytra with moderately broad, irregularly shaped, transverse basal yellowish brown band not reaching the suture. Each elytron with six complete discal striae and one submarginal stria. However, the new species can be undoubtedly recognized only based on the shape of male genitalia: median lobe sickle-shaped in lateral view, strongly curved, broadest at mid-length (Fig. 3 A); turning right in apical eighth of its length in ventral view (Fig. 3 B).	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF8BFFBCA3D556B6FBFCF8CC.taxon	etymology	Etymology The species is named after Jinling (ẁK), the ancient name of its type locality, Nanjing. The epithet is an adjective in the nominative singular.	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF8BFFBCA3D556B6FBFCF8CC.taxon	vernacular_names	Vernacular name The Chinese common name of C. jinlingensis Jiang, Jia & Wang sp. nov. is ‘ ẁKIJĖAEǡ ’.	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF8BFFBCA3D556B6FBFCF8CC.taxon	materials_examined	Type material Holotype CHINA – Jiangsu • ♂; “ 江Ā ẘ 南京市 £ 武区 ẍẁ 山KDZdz局 ḣȕ (临Ú ṁ 水坑) ” [Jiangsu Province, Nanjing, Xuanwu District, Zijinshan Mt., near Lingyuan Post Office (temporary small puddle)]; 32.0512 ° N, 118.8499 ° E; 48.1 m a. s. l.; 1 Apr. 2024; “ a 卓Dz, n 以 ¶ ” [Z. Jiang and Y. Zheng leg.]; SYSU. Paratypes CHINA – Jiangsu • 7 ♂♂, 17 ♀♀; same data as for holotype; SYSU • 5 ♂♂, 3 ♀♀; same data as for holotype; ZJCQ • 3 ♂♂; “ 江Ā ẘ 南京市 £ 武区 ẍẁ 山KDZdz局 ḣȕ (水塘) ” [Jiangsu Province, Nanjing, Xuanwu District, Zijinshan Mt., near Lingyuan Post Office (pond)]; 32.0478 ° N, 118.8493 ° E; 26.5 m a. s. l.; 29 Mar. 2024; “ a 卓Dz, n 以 ¶ ” [Z. Jiang and Y. Zheng leg.]; ZJCQ. Type locality China, Jiangsu Province, Nanjing, Xuanwu District, Zijinshan Mt., near Lingyuan Post Office; ca 32.0512 ° N, 118.8499 ° E; 48.1 m a. s. l.	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF8BFFBCA3D556B6FBFCF8CC.taxon	description	Description MEASUREMENTS. TL: 4.9 – 5.8 mm (mean value: 5.4 ± 0.2 mm); holotype: 5.7 mm. TL-h: 4.4 – 5.2 mm (mean value: 4.8 ± 0.2 mm); holotype: 5.2 mm. MW: 2.4 – 2.9 mm (mean value: 2.7 ± 0.1 mm); holotype: 2.9 mm. VARIABILITY. All specimens of the type series are rather uniform, except for sexual dimorphism. There is only slight variability in dorsal surface colouration, paler or darker than the holotype. Minor variability can be seen in number and length of strioles presenting laterally on pronotum. Male (holotype) HABITUS (Fig. 1 A). Oblong-oval in shape, with continuous outline, broadest at one-third of elytral length, slightly convex. Dorsal surface shiny. COLOURATION. Head brown, with reddish brown clypeus and two oval spots on vertex; pronotum dark brown with broadly orange sides; elytra brown, laterally yellowish brown, with moderately broad, irregularly shaped, transverse basal yellowish brown band not reaching suture; appendages yellowish brown; ventral side reddish brown to brown. HEAD. Moderately broad, ca 0.61 × as wide as pronotum, trapezoidal. Anterior margin of clypeus indistinctly concave. Antenna with antennomeres long and slender. Reticulation consisting of moderately deeply impressed polygonal isodiametric meshes. Punctation double, consisting of fine punctures and coarse setigerous punctures; fine punctures spread sparsely on clypeus and slightly larger and more densely on vertex; row of coarse punctures present around inner margin of eyes, several punctures present at frontal level of eyes, and anterolaterally to eyes in fronto-clypeal depressions. PRONOTUM. Strongly transverse (width / length ratio = 2.85), broadest between posterior angles, lateral margins moderately curved. Sides with lateral beading very thin and indistinct. Reticulation similar to that of head. Punctation double; rows of coarse setigerous punctures present along anterior margin, laterally close to sides, several punctures present also in shallow basolateral depressions along basal margin; fine punctures smaller and sparser than on head, spread throughout whole surface. Pronotum basolaterally and especially laterally with numerous, irregularly distributed short longitudinal or oblique strioles. Centre of disc with shallowly impressed medial longitudinal smooth line. ELYTRA. Base of elytra as broad as pronotal base; lateral margins of elytra slightly diverging in basal third, then distinctly narrowing to apex. Six discal and one submarginal longitudinal striae present on each elytron: stria 1 beginning slightly posteriorly to base, striae 2 – 6 beginning at base; stria 1 longest, ending close to apex; striae 2 – 5 somewhat shorter, ending subapically; stria 6 ending at apical fourth; submarginal stria long, beginning before elytral mid-length and ending subapically, at same level as dorsal striae 2 – 5. Reticulation similar to that of head and pronotum, but less impressed. Punctation consisting of coarse setigerous punctures and very fine sparse punctures; coarse punctures present along elytral striae and lateral margins of elytra. LEGS. Protibia modified, angled near base, distinctly broadened anteriorly, club-shaped. Pro- and mesotarsomeres 1 – 3 distinctly broadened, with four rows of adhesive setae on their ventral side; claws simple. VENTRAL SIDE (Fig. 1 B). Prosternum sinuate anteriorly, obtusely keeled medially. Prosternal process shortly lanceolate, in cross-section convex, apex obtuse; process distinctly bordered laterally; reticulation not perceptible. Metaventrite with microsculpture consisting of polygonal meshes; lateral parts of metaventrite (“ metasternal wings ”) tongue-shaped, slender. Metacoxal lines nearly complete, absent only very close to metaventrite. Metacoxal plates covered with long, longitudinal strioles and some transverse wrinkles; reticulation consisting of elongate, oblique polygonal meshes. Metacoxal processes rounded at posterior margin. Abdominal ventrites I – II with longitudinal strioles; ventrites III – IV with oblique strioles laterally. Abdominal reticulation consisting of elongate polygonal meshes, longitudinal on ventrites I – II, oblique on ventrite III and transverse on ventrites IV – VI. Punctation consisting of fine, sparsely distributed punctures; corase setigerous punctures present medially on ventrites III – V, and mediolaterally on ventrite VI. MALE GENITALIA. Median lobe of aedeagus in lateral view simple, sickle-shaped, strongly curved; broadest at mid-length; in apical third gradually narrowing to pointed apex (Fig. 3 A). In ventral view, median lobe turned right at a right angle in apical eighth of its length (Fig. 3 B). Parameres slender, C-shaped, broadest at mid-length; apical lobes long, club-shaped (Fig. 3 C). Female Identical to male in habitus. Protibia simple, not angled basally and only slightly broadened distally; pro- and mesotarsomeres not broadened, without adhesive setae. Females dimorphic; smooth form differs from male in more developed striolation laterally on pronotum (Fig. 1 C); striolate form submatt, with long striolae covering pronotum and elytra except for apical fourth (Fig. 1 D).	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF8BFFBCA3D556B6FBFCF8CC.taxon	diagnosis	Differential diagnosis Based on the presence of six dorsal striae and a submarginal stria on each elytron, the new species can be assigned to the Copelatus irinus species group sensu Guignot (1961). Oblong-oval habitus, elytral striation consisting of six dorsal and one submarginal stria with sutural stria complete, and the median lobe of aedeagus without a distinct process place Copelatus jinlingensis Jiang, Jia & Wang sp. nov. in the Copelatus japonicus complex. The male genitalia of new species are most similar to Copelatus tengchongensis Hájek, Jiang & Jia, 2022, but the median lobe in lateral view of the new species is more curved (see Fig. 3 A and Jiang et al. 2022: fig. 63). The male genitalia of new species are also similar to Copelatus diversistriatus Jiang, Hájek & Jia, 2022, but the lateral carina of the median lobe is without small spines in apical half which is more curved in the new species (see Fig. 3 A and Jiang et al. 2022: fig. 37). It is difficult to distinguish the new species from the coexisting Copelatus japonicus Sharp, 1884 based on the external appearance. Males can be distinguished by the genitalia: in lateral view, the median lobe slender in apical half in C. japonicus (Fig. 3 D) but moderately broad in C. jinlingensis (Fig. 3 A); in ventral view, the apical part of the median lobe is turned right in C. jinlingensis (Fig. 3 B) but bent downward in C. japonicus (Fig. 3 E). Females of C. jinlingensis are dimorphic (Fig. 1 C – D) but monomorphic in C. japonicus. However, smooth form females of C. jinlingensis sp. nov. are almost indistinguishable from the females of C. japonicus in morphology. The females in the paratype series were identified because all the males found simultaneously in the temporary small puddles at the type locality are the same species, C. jinlingensis.	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF8BFFBCA3D556B6FBFCF8CC.taxon	biology_ecology	Ecology Three male specimens of Copelatus jinlingensis sp. nov. were collected from a large pond with duckweed (Fig. 4 A) at Zijinshan Mt. They were found in syntopy with Copelatus japonicus and Austrelatus parallelus at the pond’s edge (Fig. 4 B). A large number of C. jinlingensis were also found in several temporary small puddles near the pond (Fig. 4 C – D).	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF8BFFBCA3D556B6FBFCF8CC.taxon	distribution	Distribution Only known from the type locality in southwestern Jiangsu so far (Figs 6, 7 A – B).	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF8FFFB3A3DA550AFBCDFC8D.taxon	description	urn: lsid: zoobank. org: act: F 11775 FA- 2346 - 408 D-A 494 - 80 A 2 C 43 BDDAE Figs 2, 3 F – H, 5 A, 6, 8	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF8FFFB3A3DA550AFBCDFC8D.taxon	diagnosis	Diagnosis Medium-sized (TL: 4.7 – 5.2 mm), oblong-oval species. Elytra with moderately broad, irregularly shaped, transverse basal yellowish brown band not reaching the suture. Each elytron with six complete discal striae and one submarginal stria. However, the new species can be unequivocally recognized only based on the shape of the male genitalia: median lobe sickle-shaped in lateral view, moderately curved, basal third almost straight, slightly swollen near apex (Fig. 3 F); sinuate in apical half, turned right in apical eighth of its length in ventral view (Fig. 3 G).	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF8FFFB3A3DA550AFBCDFC8D.taxon	etymology	Etymology The species is named after its type locality, Lankeshan Mt. It is renowned as the “ Go Fairyland ”, famed for the legendary story of Wang Zhi from the Jin Dynasty who watched a Go game until his axe handle rotted away (= “ Lanke ” in Chinese Pinyin). The specific epithet is an adjective in the nominative singular.	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF8FFFB3A3DA550AFBCDFC8D.taxon	vernacular_names	Vernacular name The Chinese common name of C. lankeensis Jiang, Jia & Wang sp. nov. is ‘ 烂柯IJĖAEǡ ’.	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF8FFFB3A3DA550AFBCDFC8D.taxon	materials_examined	Type material Holotype CHINA – Zhejiang • ♂; “ ť 江 ẘae 州市柯城区烂柯山 ” [Zhejiang Province, Quzhou, Kecheng District, Lankeshan Mt.]; 28.8781 ° N, 118.9199 ° E; 118.1 m a. s. l.; 5 Feb. 2024; “ a 卓Dz, LJ 巍峰 ” [Z. Jiang and W. Yan leg.]; SYSU. Paratypes CHINA – Zhejiang • 3 ♂♂, 2 ♀♀; same data as for holotype; SYSU • 3 ♂♂, 1 ♀; same data as for holotype; ZJCQ. Type locality China, Zhejiang Province, Quzhou, Kecheng District, Lankeshan Mt.; ca 28.8781 ° N, 118.9199 ° E; 118.1 m a. s. l.	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF8FFFB3A3DA550AFBCDFC8D.taxon	description	Description MEASUREMENTS. TL: 4.7 – 5.2 mm (mean value: 5.0 ± 0.2 mm); holotype: 5.2 mm. TL-h: 4.1 – 4.6 mm (mean value: 4.4 ± 0.2 mm); holotype: 4.6 mm. MW: 2.3 – 2.6 mm (mean value: 2.5 ± 0.1 mm); holotype: 2.6 mm. VARIABILITY. All specimens of the type series are rather uniform, except for sexual dimorphism. There is only slight variability in dorsal surface colouration, paler or darker than holotype. Minor variability can be seen in number and length of strioles presenting laterally on pronotum. Male (holotype) HABITUS (Fig. 2 A). Oblong-oval in shape, with continuous outline, broadest at one-third of elytral length, slightly convex. Dorsal surface shiny. COLOURATION. Head dark brown, with orange clypeus; pronotum dark brown with broadly orange sides; elytra brown, laterally yellowish brown, with moderately broad, irregularly shaped, transverse basal yellowish brown band not reaching suture; appendages yellowish brown; ventral side reddish brown to brown. HEAD. Moderately broad, ca 0.61 × as wide as pronotum, trapezoidal. Anterior margin of clypeus indistinctly concave. Antenna with antennomeres long and slender. Reticulation consisting of moderately deeply impressed polygonal isodiametric meshes. Punctation double, consisting of fine punctures and coarse setigerous punctures; fine punctures spread sparsely on clypeus and slightly larger and more densely on vertex; row of coarse punctures present around inner margin of eyes, several punctures present at frontal level of eyes, and anterolaterally to eyes in fronto-clypeal depressions. PRONOTUM. Strongly transverse (width / length ratio = 2.87), broadest between posterior angles, lateral margins moderately curved. Sides with lateral beading very thin and indistinct. Reticulation similar to that of head. Punctation double; rows of coarse setigerous punctures present along anterior margin, laterally close to sides, several punctures present also in shallow basolateral depressions along basal margin; fine punctures smaller and sparser than on head, spread throughout whole surface. Pronotum basolaterally and especially laterally with numerous, irregularly distributed short longitudinal or oblique strioles. Centre of disc with shallowly impressed medial longitudinal smooth line. ELYTRA. Base of elytra as broad as pronotal base; lateral margins of elytra slightly diverging in basal third, then distinctly narrowing to apex. Six discal and one submarginal longitudinal striae present on each elytron: stria 1 beginning slightly posteriorly to base, striae 2 – 6 beginning at base; stria 1 ending close to apex; striae 2 – 5 almost equal in length, ending subapically; stria 6 ending at apical fourth; submarginal stria long, beginning before elytral mid-length and ending subapically, at same level as dorsal striae 2 – 5. Reticulation similar to that of head and pronotum, but less impressed. Punctation consisting of coarse setigerous punctures and very fine sparse punctures; coarse punctures present along elytral striae and lateral margins of elytra. LEGS. Protibia modified, angled near base, distinctly broadened anteriorly, club-shaped. Pro- and mesotarsomeres 1 – 3 distinctly broadened, with four rows of adhesive setae on their ventral side; claws simple. VENTRAL SIDE (Fig. 2 B). Prosternum sinuate anteriorly, obtusely keeled medially. Prosternal process shortly lanceolate, in cross-section convex, apex obtuse; process distinctly bordered laterally; reticulation not perceptible. Metaventrite with microsculpture consisting of polygonal meshes; lateral parts of metaventrite (“ metasternal wings ”) tongue-shaped, slender. Metacoxal lines nearly complete, absent only very close to metaventrite. Metacoxal plates covered with long, longitudinal strioles and some transverse wrinkles; reticulation consisting of elongate, oblique polygonal meshes. Metacoxal processes rounded at posterior margin. Abdominal ventrites I – II with longitudinal strioles; ventrites III – IV with oblique strioles laterally. Abdominal reticulation consisting of elongate polygonal meshes, longitudinal on ventrites I – II, oblique on ventrite III and transverse on ventrites IV – VI. Punctation consisting of fine, sparsely distributed punctures; corase setigerous punctures present medially on ventrites III – V, and mediolaterally on ventrite VI. MALE GENITALIA. Median lobe of aedeagus in lateral view simple, sickle-shaped, moderately curved; basal third almost straight; slightly swollen near apex; apex bluntly pointed (Fig. 3 F). In ventral view, median lobe sinuate in apical half, turned right in apical eighth of its length (Fig. 3 G). Parameres slender, C-shaped, broadest in basal third; apical lobes long, club-shaped (Fig. 3 H). Female Identical to male in habitus. Protibia simple, not angled basally and only slightly broadened distally; pro- and mesotarsomeres not broadened, without adhesive setae. Females dimorphic; smooth form differs from male in more developed striolation laterally on pronotum (Fig. 2 C); striolate form submatt, with several short oblique striolae covering posterior part of head, and long striolae covering pronotum and elytra except for apical fourth (Fig. 2 D).	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF8FFFB3A3DA550AFBCDFC8D.taxon	diagnosis	Differential diagnosis Based on the presence of six dorsal striae and a submarginal stria on each elytron, the new species can be assigned to the Copelatus irinus species group sensu Guignot (1961). Oblong-oval habitus, elytral striation consisting of six dorsal and one submarginal stria with sutural stria complete, and the median lobe of the aedeagus without a distinct process place Copelatus lankeensis Jiang, Jia & Wang sp. nov. in the Copelatus japonicus complex. The male genitalia of the new species are not very similar to any other Chinese species by its unique shape, especially the swelling near the apex in lateral view and the apical half sinuate in ventral view. Although the new species can be distinguished from the coexisting Copelatus chinensis Régimbart, 1899 by its broader habitus and darker coloration, they can be reliably separated only by the male genitalia: in lateral view, the median lobe almost straight in basal third, slightly swollen near apex, apex bluntly pointed in C. lankeensis (Fig. 3 F) but evenly curved, not swollen near apex, apex skewed in C. chinensis (Fig. 3 I); in ventral view, median lobe sinuate in apical half, turned right apically in C. lankeensis (Fig. 3 G) but almost straight, slightly turned left apically in C. chinensis (Fig. 3 J). Females of C. lankeensis dimorphic (Fig. 2 C – D) but monomorphic in C. chinensis.	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF8FFFB3A3DA550AFBCDFC8D.taxon	biology_ecology	Ecology At Lankeshan Mt., Copelatus lankeensis Jiang, Jia & Wang sp. nov. was collected syntopically with Copelatus chinensis in a temporary long puddle with decaying leaves in the bamboo forest (Fig. 5 A).	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF8FFFB3A3DA550AFBCDFC8D.taxon	distribution	Distribution Only known from the type locality in western Zhejiang so far (Figs 6, 8 A – B).	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF82FFB2A01B557BFE71FA1D.taxon	description	Figs 3 I – J, 5 A – B, D, 6 Literature records from China Régimbart 1899: 298 (Hubei, original description); Falkenström 1932: 192 (Sichuan or Chongqing); Falkenström 1933: 14 (Sichuan or Chongqing); Jiang et al. 2022: 253 (Hunan, Shandong, Zhejiang).	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF82FFB2A01B557BFE71FA1D.taxon	materials_examined	Material examined (first province record) First records CHINA – Jiangsu • 4 ♂♂, 1 ♀; “ 江Ā ẘ 南京市浦口区 # 山Þ ḢDZ ” [Jiangsu Province, Nanjing, Pukou District, Laoshan Mt., Bainiaoyuan]; 32.1059 ° N, 118.6002 ° E; 100.4 m a. s. l.; 20 Mar. 2024; “ a 卓Dz, n 以 ¶, Ẹk ” [Z. Jiang, Y. Zheng and Z. Deng leg.]; ZJCQ • 2 ♂♂; “ 江Ā ẘ 淮 ẋ 市ƃ ḃ 县M山 ḦMũ 桥 ḣȕ ” [Jiangsu Province, Huaian, Xuyi County, Tieshansi, near Tiesuoqiao]; 32.7311 ° N, 118.4621 ° E; 110.9 m a. s. l.; 12 Apr. 2025; “ a 卓Dz, n 以 ¶, Ā ṪNJ, 卢琮Ý ” [Z. Jiang, Y. Zheng, F. Su and C. Lu leg.]; ZJCQ. Additional records CHINA – Zhejiang • 2 ♂♂; “ ť 江 ẘae 州市柯城区烂柯山 ” [Zhejiang Province, Quzhou, Kecheng District, Lankeshan Mt.]; 28.8781 ° N, 118.9199 ° E; 118.1 m a. s. l.; 5 Feb. 2024; “ a 卓Dz, LJ 巍峰 ” [Z. Jiang and W. Yan leg.]; ZJCQ.	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF82FFB2A01B557BFE71FA1D.taxon	biology_ecology	Ecology Copelatus chinensis was found co-occurring with Copelatus lankeensis Jiang, Jia & Wang sp. nov. in a long, temporary puddle with decaying leaves under a bamboo forest at Lankeshan Mt. (Fig. 5 A), whereas at Laoshan Mt., it was collected at the edge of a muddy reservoir (Fig. 5 B, D).	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF82FFB2A01B557BFE71FA1D.taxon	distribution	Distribution Endemic in China. Central and eastern China: Hubei, Hunan, Jiangsu (Fig. 6), Shandong, Sichuan or Chongqing, and Zhejiang.	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF82FFB7A07551C9FB82FEC2.taxon	description	Figs 3 D – E, 4 A, 6 Literature records from China Jiang et al. 2022: 260 (Shanghai); Jiang et al. 2025: 49 (Hubei).	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF82FFB7A07551C9FB82FEC2.taxon	materials_examined	Material examined (first province record) First records CHINA – Jiangsu • 3 ♂♂; “ 江Ā ẘ 南京市 £ 武区 ẍẁ 山KDZdz局 ḣȕ (水塘) ” [Jiangsu Province, Nanjing, Xuanwu District, Zijinshan Mt., near Lingyuan Post Office (pond)]; 32.0478 ° N, 118.8493 ° E; 26.5 m a. s. l.; 29 Mar. 2024; “ a 卓Dz, n 以 ¶ ” [Z. Jiang and Y. Zheng leg.]; ZJCQ • 1 ♀; “ 江Ā ẘ 南 京市江Ť区ŴAE堰水库 ” [Jiangsu Province, Nanjing, Jiangning District, Huanglongyan Reservoir]; 31.9684 ° N, 118.9576 ° E; 79.5 m a. s. l.; 24 Apr. 2024; “ a 卓Dz, n 以 ¶ ” [Z. Jiang and Y. Zheng leg.]; ZJCQ • 1 ♂; “ 江Ā ẘ 南京市浦口区 # 山狮 + 岭AEí ” [Jiangsu Province, Nanjing, Pukou District, Laoshan Mt., Shiziling, Yaotou]; 32.0467 ° N, 118.5518 ° E; 81.7 m a. s. l.; 2 Oct. 2024; “ a 卓Dz, n 以 ¶, Ẹk ” [Z. Jiang, Y. Zheng and Z. Deng leg.]; ZJCQ • 1 ♂, 1 ♀; “ 江Ā ẘ 淮 ẋ 市ƃ ḃ 县M山 ḦMũ 桥 ḣȕ ” [Jiangsu Province, Huaian, Xuyi County, Tieshansi, near Tiesuoqiao]; 32.7311 ° N, 118.4621 ° E; 110.9 m a. s. l.; 12 Apr. 2025; “ a 卓Dz, n 以 ¶, Ā ṪNJ, 卢琮Ý ” [Z. Jiang, Y. Zheng, F. Su and C. Lu leg.]; ZJCQ.	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF82FFB7A07551C9FB82FEC2.taxon	biology_ecology	Ecology At Zijinshan Mt., Copelatus japonicus was found syntopically with Copelatus jinlingensis Jiang, Jia & Wang sp. nov. and Austrelatus parallelus in a large pond with duckweed (Fig. 4 A).	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF82FFB7A07551C9FB82FEC2.taxon	distribution	Distribution Eastern China (Hubei, Jiangsu (Fig. 6), Shanghai), Japan, South Korea.	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF87FFB7A037558DFD2AFB36.taxon	description	Figs 5 C, E, 6 Literature records from China Balfour-Browne 1947: 440 (Nei Mongol, original description); Li 1992: 35 (Liaoning); Nilsson 1995: 42 (Hebei); Jiang et al. 2022: 274 (Liaoning, Shanghai).	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF87FFB7A037558DFD2AFB36.taxon	materials_examined	Material examined (first province record) First records CHINA – Heilongjiang • 2 ♀♀; “ ḙAE 江 ẘ 牡丹江市东 ẋ 区牡丹峰DÊ森林公DZ ḣȕ ” [Heilongjiang Province, Mudanjiang, Dong’an District, near Mudanfeng National Forest Park]; 44.4783 ° N, 129.7355 ° E; 314.0 m a. s. l.; 11 Aug. 2025; “ n 以 ¶ ” [Y. Zheng leg.]; ZJCQ. – Jilin • 6 ♂♂, 4 ♀♀; “ 吉林 ẘ 吉林市丰 ¡ 区 Ẅṝ 公DZ ” [Jilin Province, Jilin, Fengman District, Emerald Park]; 43.8036 ° N, 126.5965 ° E; 235.5 m a. s. l.; 17 Aug. 2024; “ n 以 ¶ ” [Y. Zheng leg.]; ZJCQ.	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF87FFB7A037558DFD2AFB36.taxon	biology_ecology	Ecology In Emerald Park (Jilin), Copelatus weymarni was collected from temporary small puddles under the grass (Fig. 5 C, E). The specimens from Heilongjiang were collected by light traps.	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF87FFB7A037558DFD2AFB36.taxon	distribution	Distribution Northeastern China (Hebei, Heilongjiang, Jilin (Fig. 6), Liaoning, Nei Mongol, Shanghai), Far East of Russia, Japan, North Korea, South Korea.	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF87FFB6A03250E1FB34FEC2.taxon	description	Figs 4 A – B, E, 6 Literature records from China Jiang et al. 2022: 266 (Hebei, Hubei, Hunan, Shanghai).	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF87FFB6A03250E1FB34FEC2.taxon	materials_examined	Material examined (first province record) First records CHINA – Jiangsu • 2 ♂♂, 1 ♀; “ 江Ā ẘ 南京市 £ 武区 ẍẁ 山KDZdz局 ḣȕ (水塘) ” [Jiangsu Province, Nanjing, Xuanwu District, Zijinshan Mt., near Lingyuan Post Office (pond)]; 32.0478 ° N, 118.8493 ° E; 26.5 m a. s. l.; 29 Mar. 2024; “ a 卓Dz, n 以 ¶ ” [Z. Jiang and Y. Zheng leg.]; ZJCQ • 1 ♀; “ 江Ā ẘ 南京市江Ť区ŴAE堰水库 ” [Jiangsu Province, Nanjing, Jiangning District, Huanglongyan Reservoir]; 31.9684 ° N, 118.9576 ° E; 79.5 m a. s. l.; 24 Apr. 2024; “ a 卓Dz, n 以 ¶ ” [Z. Jiang and Y. Zheng leg.]; ZJCQ.	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF87FFB6A03250E1FB34FEC2.taxon	biology_ecology	Ecology At Zijinshan Mt., Austrelatus parallelus was found in a large duckweed pond (Fig. 4 A), where it was collected in syntopy with Copelatus jinlingensis Jiang, Jia & Wang sp. nov. and Copelatus japonicus at the edge (Fig. 4 B, E).	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
7C4C87C2FF87FFB6A03250E1FB34FEC2.taxon	distribution	Distribution Eastern China (Hebei, Hubei, Hunan, Jiangsu (Fig. 6), Shanghai), Japan, South Korea.	en	Jiang, Zhuo-yin, Zheng, Yi-li, Deng, Zhen, Jia, Feng-long, Wang, Bei-xin (2026): Two new species of Copelatus Erichson, 1832 from urban low mountains of eastern China, with new province records of the other Copelatinae in China (Coleoptera: Dytiscidae). European Journal of Taxonomy 1052: 105-124, DOI: 10.5852/ejt.2026.1052.3271, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3271/14446
