identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
44CB08A8AF905080951D93D0BC93B796.text	44CB08A8AF905080951D93D0BC93B796.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nannopus Brady 1880	<div><p>Genus Nannopus Brady, 1880</p><p>Generic diagnosis</p><p>(amended from Fiers and Kotwicki 2013; Kim et al. 2017; Vakati et al. 2016, 2019; Vakati and Lee 2017, 2021; Sciberras et al. 2021). Nannopodidae . Body fusiform, slightly dorsoventrally flattened. Body ornamentation consisting of dorsal denticles and horizontal rows of setules and sensilla. Rostrum bell-shaped, fused to cephalothorax; anterior margin furnished with setules and one pair of sensilla. Cephalothorax anteriorly attenuated in dorsal view; posterior margin serrate, with several sensilla. Prosome 4 - segmented, comprising cephalothorax and three subequal pedigerous somites. The female genital and first abdominal somite fused, forming a genital double-somite. Second abdominal somite with row of spinules on the ventral posterior margin. Operculum well developed, with rows of setules. Antennule 5 - segmented, short in female; 2 nd segment with tri-articulate seta inserted dorsally; 3 rd segment with aesthetasc, and 5 th with acrothek. Male antennule 5–6 - segmented, chirocer. Antenna with one or two abexopodal setae on allobasis; 1 - segmented exopod with three or four setae; endopod with six or seven setae. Mandibular palp 1 - segmented, broad, with 3–5 elements. Maxillule praecoxal arthrite with nine or ten elements at the distal margin. Cephalothorax with / without integumental windows. P 1 – P 4 exopods 3 - segmented; P 1 – P 4 endopods 1-2: 2: 1 - 2: 1 - segmented. P 2 – P 3 exp- 3 with 2-3 – 2 - 3 (♀): 2-4 – 2 - 4 (♂) outer spines. P 3 enp- 2 of male present or absent. P 4 exp- 3 inner subdistal element setiform or pectinate. Female P 5 baseoendopod plate-like, not developed, with three or four setae; exopod with four or five setae. Female P 6 represented by 1 seta. P 6 articulation of males asymmetrically or symmetrically confluent; represented by two or three setae. Female with two egg-sacs, male with one or two spermatophores. Caudal rami cylindrical or trapezoidal, clearly separated from anal somite; with 7 setae; seta V well developed and longest.</p><p>Type species.</p><p>Nannopus palustris Brady, 1880 (by monotypy).</p><p>Other species.</p><p>N. bulbiseta Vakati &amp; W. Lee, 2017, N. canui Sciberras, Cazzaniga &amp; Huys, 2021, N. cylindricus Vakati &amp; Lee, 2021, N. didelphis Fiers &amp; Kotwicki, 2013, N. dimorphicus Vakati &amp; W. Lee, 2017, N. flexibilis (Lilljeborg, 1902), N. ganghwaensis Vakati, Kihara &amp; W. Lee, 2017, N. hirsutus Fiers &amp; Kotwicki, 2013, N. minutus Vakati &amp; W. Lee, 2017, N. parvipilis Kim, J. G., Choi &amp; Yoon, 2017, N. parvus Vakati &amp; W. Lee, 2017, N. perplexus (Sars G. O., 1909), N. procerus Fiers &amp; Kotwicki, 2013, N. robustus Vakati &amp; Lee, 2021, N. scaldicola Fiers &amp; Kotwicki, 2013, N. serratus Vakati &amp; W. Lee, 2017, N. sinusalbi Sciberras, Cazzaniga &amp; Huys, 2021, N. unisegmentatus Shen &amp; Tai, 1964; subspecies N. palustris tiberiadis Por, 1968 .</p></div>	https://treatment.plazi.org/id/44CB08A8AF905080951D93D0BC93B796	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Garlitska, Lesya;Chertoprud, Elena;Mayor, Tatyana;Saburova, Maria;Gevorgyan, Gor;Azovsky, Andrey	Garlitska, Lesya, Chertoprud, Elena, Mayor, Tatyana, Saburova, Maria, Gevorgyan, Gor, Azovsky, Andrey (2026): Morphological redescription and genetic characterization of the enigmatic Nannopus palustris Brady, 1880 (Copepoda, Harpacticoida, Nannopodidae). ZooKeys 1278: 135-161, DOI: 10.3897/zookeys.1278.188797
CD3E47CFEE2F50639AC268549EC3CFD5.text	CD3E47CFEE2F50639AC268549EC3CFD5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nannopus palustris Brady 1880	<div><p>Nannopus palustris Brady, 1880</p><p>Figs 2, 3, 4, 5, 6, 7, 8, 9</p><p>Synonymy.</p><p>Nannopus palustris sensu T. Scott 1902 .</p><p>Type locality.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=1.4736667&amp;materialsCitation.latitude=55.08433" title="Search Plazi for locations around (long 1.4736667/lat 55.08433)">Seaton Sluice</a>, Northumberland, UK (55°05.06'N, 01°28.42'E).</p><p>Material examined.</p><p>• 1 ♀ designated as the neotype, dissected on 4 slides, deposited at the Zoological Museum of SCZHE, reg. no. SCZHE -COP-1–4; • 1 ♀ undissected on 1 slide, same collection data, reg. no. SCZHE -COP-5; • 1 ♂ dissected on 4 slides, same collection data, reg. no. SCZHE -COP-6–9; • 1 ♂ undissected on 1 slide, same collection data, reg. no. SCZHE -COP-10 .</p><p>Other material.</p><p>• 50 ♀♀ and 5 ♂♂, alcohol preserved, same collection data . All specimens were collected from the type locality in August 2013 and September 2023, leg. L. Garlitska.</p><p>Description of the adult female.</p><p>Total body length measured from the tip of the rostrum to the posterior margin of the caudal rami ranged from 618 to 699 µm (mean = 667 µm; n = 10). Habitus fusiform (Figs 2 A, 3). Color of preserved specimens pale yellowish to colorless. Rostrum prominent, fused to the cephalothorax, recurved ventrally; anterior margin with multiple long and slender setules and one pair of sensilla (Fig. 2 B). Cphth bell-shaped, representing 22 % of the total body length; posterior margin serrate. Sensillar pattern on the cephalothorax and body somites are illustrated in Fig. 2 A, B. Posterodorsal margin of each somite serrate except for the anal somite. Somites bearing P 2 – P 4 with pairs of lateral nuchal organs (osmoregulatory integumental windows) (Fig. 2 A). Genital and first abdominal somite fused, forming a GDS; the original segmentation marked by bilateral constriction and by a transverse serrate ridge dorsally. Anal somite cleft medially in ventral view; ventral posterior margin with row of spinules (Fig. 4 A); dorsal surface with two pairs of sensilla (Fig. 4 B). Anal operculum well developed, with rows of setules (Fig. 4 B).</p><p>Caudal ramus (Figs 3 C, 4 B) cylindrical and elongate, dorsal posterior margin with row of spinules, with seven setae. Seta I naked, inserted close to the anterior margin of the ramus. Seta II naked, located dorsolaterally. Seta III naked, inserted approximately halfway down the outer margin. Seta IV bipinnate, longer than ramus (in dorsal view); located at the outer distal corner. Seta V longest (2.6–3 × longer than IV), strongest, and swollen at its base and bipinnate. Seta VI naked; located at inner distal corner. Dorsal seta VII naked; located close to inner margin.</p><p>Antennule (Fig. 2 C – E) compact, short, and 5 - segmented. First, second and third segments with rows of spinules. First segment with one posterior seta and pore. Antennule 2 nd segment with a tri-articulate seta inserted dorsally. Segment 3 with aesthetasc fused basally to long seta. Segment 5 with acrothek. Armature as follows: 1 - [1], 2 - [4 + 5 bipinnate], 3 - [6 + (1 + ae)], 4 - [1], 5 - [8 + acrothek].</p><p>Antenna (Fig. 2 F, G). Relatively short, consisting of allobasis, a 1 - segmented endopod and a 1 - segmented exopod. Allobasis with two abexopodal setae of which one short bipinnate on the middle margin, and one long pinnate on the distal margin. Endopod with a transverse row of spinules in the proximal half; a row of robust spinules near the inner corner and a row of short spinules near the outer corner; armed with six strong spines. Exopod with four elements, one of which is bipinnate.</p><p>Mandible (Fig. 5 A). Gnathobase is well developed; the cutting edge is provided with six rigid, multicuspidate teeth and a setular process. Mandibular palp 1 - segmented with rami completely incorporated into the basis; outer margin and middle part with rows of spinules; armed with four bipinnate setae, one of which (derived from the basis) originates from a subcylindrical pedestal.</p><p>Maxillule (Fig. 5 B). Praecoxal arthrite with two setae on the anterior surface; distal margin with six stout spines, two short pinnate and two long pinnate elements. Syncoxa with cylindrical coxal endite bearing two naked setae. Basis and rami fused; armed with two naked, four bipinnate and two pinnate setae.</p><p>Maxilla (Fig. 5 C). Syncoxa bearing two subequal endites, each with three elements (2 spinulose which are confluent with endites, and 1 slender naked). Allobasis with a row of spinules, a strong claw with one accompanying seta. Endopod incorporated into basis and represent by two naked setae.</p><p>Maxilliped (Fig. 5 D) subchelate, 3 - segmented. Syncoxa shorter than basis, with rows of spinules, unarmed. Basis unarmed; with rows of spinules. Endopod with a strong, curved claw ornamented with spinules in the distal half, and two naked accessory setae.</p><p>Swimming legs (Figs 6, 7) with 3 - segmented exopods and 2 - (P 1 – P 3) or 1 - segmented (P 4) endopods; exopods longer than endopods. P 1 – P 4 with smooth and short concave intercoxal sclerites (not shown). Praecoxae with row of spinules on the anterior surface. Coxae with a row of spinules on the anterior surface and a row of strong spinules along the outer margin. Bases with plumose outer seta; with rows of spinules near the insertion of the exopod and endopod. Exopodal segments (except P 1 exp- 3) with bunches of thin setules on inner margin; first (except P 4), second and third segments of the exopod with row of tiny setules near the outer distal corner (Fig. 6 A, on Figs 6 B, C, 7 marked with arrows); outer margins of exopodal and endopodal segments with spinular ornamentation, except for the P 4 endopod.</p><p>P 1 (Fig. 6 A, B). Basis with row of spinules near the insertion of the inner and outer spines; inner basal spine pinnate. Outer exopodal spine of exp- 1 naked; exp- 2 with outer spine with single tooth and with inner seta; exp- 3 with one naked and one pectinate outer spine, and two terminal elements of which outer bipinnate and inner plumose. Enp- 1 and end- 2 with bunches of tiny setules on inner margin; outer element of enp- 2 naked, terminal element bipinnate, and inner seta short and pinnate.</p><p>P 2 (Fig. 6 C). Basis with a row of long setules along the inner margin. Inner margins of all segments with setular row; outer exopodal spines of exp- 1 and exp- 2 naked. Exp- 3 with three outer spines (of which two proximal ones with a tooth, and distal one pectinate), two apical setae of which outer pinnate and inner plumose, and one inner plumose element. Inner margin of end- 2 with setular row, outer element of enp- 2 spiniform and naked, terminal and inner elements setiform and plumose.</p><p>P 3 (Fig. 7 A). Basis with a row of long setules along the inner margin. Inner margin of all segments (except end- 1) with setular row; outer exopodal spines of exp- 1 and exp- 2 naked. First outer spine of exp- 3 naked, second spine with single tooth and distal spine pectinate; inner and terminal elements plumose. Outer element of enp- 2 spiniform and naked; terminal and inner element long, setiform and plumose.</p><p>P 4 (Fig. 7 B). Inner margin of exopodal segments with row of setules. Outer spine of exp- 1 and exp- 2 naked; first outer spine of exp- 3 naked, second spine with single tooth and distal spine pectinate; inner element of exp- 3 plumose seta; inner subdistal seta shortest and pectinate; inner apical elements plumose. Endopod with long plumose apical seta, and a short (~ 4 × shorter than apical) plumose inner element.</p><p>Setal formula of the swimming legs is as follows:</p><p>P 5 (Fig. 4 C). Baseoendopod transversely elongate, with distal spinules along the plate and near articulation with the exopod; endopodal lobe with four bipinnate elements; outer expansion with bipinnate basal seta. Exopod covered with spinules, articulating with baseoendopod; semicircular and approximately as long as wide; with five pinnate elements.</p><p>P 6 (Fig. 4 A). Closing off paired genital apertures, semi-triangular with protruding outer distal edge bearing one short naked element.</p><p>Description of the adult male.</p><p>Total body length measured from the tip of the rostrum to the posterior margin of the caudal rami ranged from 563 to 610 µm (mean = 588 µm; n = 5). Sexual dimorphism expressed in body size, antennule, P 2, P 3, P 5, P 6, and urosomal segmentation.</p><p>Ornamentation of the body and armature of the caudal ramus like in female (Fig. 8 A, E).</p><p>Antennule (Fig. 8 B – D) short, 6 - segmented; geniculation located between segments 4 and 5. All segments are strongly chitinized. First segment with a row of spinules and a small naked seta. Segment 5 with aesthetasc fused basally to long seta. Segment 6 with acrothek. The armature formula is as follows: 1 - [1], 2 - [5 + 4 bipinnate], 3 - [4], 4 - [6], 5 - [8 + (1 + ae)], 6 - [7 + acrothek].</p><p>Antenna, mouthparts, and P 1 like in female.</p><p>P 2 and P 3 (Fig. 9 A, B). Exp- 1, - 2 and - 3 of both legs with outer spines stronger than in female; the outer terminal spine of exp- 3 with spinules and stronger than in female. P 3 enp- 2 (Fig. 8 B), with the outer spine forming a basally fused, robust apophysis; terminal and inner elements shorter than those in female; inner element naked.</p><p>P 5 (Fig. 9 C) fused with somite. Baseoendopod and exopod fused, forming a common plate with a long row of spinules. Endopodal lobe armed with two pectinate and two bipinnate elements. Outer setophore with bipinnate basal seta. Exopodal lobe with three plumose and two naked elements.</p><p>P 6 (Fig. 9 C) asymmetrical with a functional left leg forming a simple flap; confluent with somite, with spinules along the distal margin and armed with one bipinnate and two plumose elements. Double spermatophore observed underneath the flap (not shown).</p><p>Additional comments.</p><p>The body and appendages of the specimens were densely covered with mud, which could be only partly removed by ultrasonication. The natural population in the type locality exhibited a rather unbalanced sex ratio (in August 2013 and September 2023, females accounted for 90–95 % of adults).</p></div>	https://treatment.plazi.org/id/CD3E47CFEE2F50639AC268549EC3CFD5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Garlitska, Lesya;Chertoprud, Elena;Mayor, Tatyana;Saburova, Maria;Gevorgyan, Gor;Azovsky, Andrey	Garlitska, Lesya, Chertoprud, Elena, Mayor, Tatyana, Saburova, Maria, Gevorgyan, Gor, Azovsky, Andrey (2026): Morphological redescription and genetic characterization of the enigmatic Nannopus palustris Brady, 1880 (Copepoda, Harpacticoida, Nannopodidae). ZooKeys 1278: 135-161, DOI: 10.3897/zookeys.1278.188797
