identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
87368253DA5258988EB5630AA1095A2A.text	87368253DA5258988EB5630AA1095A2A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pycnoscelus aurantius Hanitsch 1935	<div><p>Pycnoscelus aurantius Hanitsch, 1935</p><p>Fig. 7</p><p>Pycnoscelus aurantius Hanitsch, 1935: 18; Bruijning 1947: 210; Roth 1998: 125.</p><p>Pycnoscelis [sic] aurantia: Princis 1964: 275.</p><p>Material examined.</p><p>Malaysia • 1 male; Sabah,  Mt. Trus Madi; 1121 m; 5 Oct. 2015; Gui-Qiang Huang leg.</p><p>Diagnosis.</p><p>General color dark orange, similar to  P. semivitreus, but  P. aurantius can be readily distinguished from  P. semivitreus by its smaller right style and the apical part of sclerite L 2 D blunt without any process.</p><p>Supplementary description of male.</p><p>Tegmina is somewhat bicolored as  P. semivitreus, from proximally orange to distally yellowish, with the boundary indistinct (Fig. 7 E). Male genitalia: right phallomere with distinct bristles, caudal part of sclerite R 1 T nearly rectangular, R 2 curved, R 3 forked, R 4 small; R 5 lies above R 2, irregular, prolonged (Fig. 7 H). The basal part of sclerite L 2 D strongly sclerotized with apical portion less sclerotized and nearly transparent; the apical part of sclerite L 2 D C-shaped, outer margin without any tooth; apical membrane less developed, covered with microtrichia (Fig. 7 I). Sclerite L 3 hook slender, inner curved margin with a tooth at apex; sclerite L 4 U divided into two parts (Fig. 7 J).</p><p>Measurements (mm).</p><p>Body length including tegmen: 21.9; pronotum length × width: 4.6 × 6.5; tegmen length: 19.8.</p><p>Remarks.</p><p>The specimen we examined is generally identical to the description of  P. aurantius provided by Roth (1998). However, Roth (1998) examined the type of  P. aurantius and found its left style present, whereas this structure is absent in the specimen we examined (Fig. 7 G). As only a single specimen is available in our study, it is difficult to determine whether the style is missing or naturally absent. More specimens will be needed in the future to clarify this question, and for now, we have tentatively identified the specimen as  P. aurantius .</p></div>	https://treatment.plazi.org/id/87368253DA5258988EB5630AA1095A2A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wang, Yi-Shu;Dong, Xiao-Jiao;Wang, Zong-Qing;Che, Yan-Li	Wang, Yi-Shu, Dong, Xiao-Jiao, Wang, Zong-Qing, Che, Yan-Li (2025): Taxonomic update of Pycnoscelus Scudder, 1862 (Blattodea, Blaberidae, Pycnoscelinae), with descriptions of two new species from China. ZooKeys 1231: 45-67, DOI: 10.3897/zookeys.1231.141287
72F47432B1A054F590897242613C07D4.text	72F47432B1A054F590897242613C07D4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pycnoscelus indicus (Fabricius 1775)	<div><p>Pycnoscelus indicus (Fabricius, 1775)</p><p>Fig. 2</p><p>Blatta indica Fabricius, 1775: 272.</p><p>Pycnoscelus indicus: Roth 1967: 774; Roth 1998: 99.</p><p>Material examined.</p><p>China • 5 males &amp; 3 females; Fujian Prov., Putian City,  Meizhou Island; 22–23 July 2013; Shun-Hua Gui &amp; Yan Shi leg.  •   2 males; Guangdong Prov., Zhongshan City,  Zimaling Park, 11 Oct. 2018; Ke-Liang Wu leg.  •   1 female; Guangxi Prov., Baise City, Renzhuang Township,  Tengmao Village,  Nongli tun; 9–13 July 2015; Jian-Yue Qiu leg.  •   1 male; Guangxi Prov., Longzhou County;  Nonggang Nature Reserve; 29–30 June 2015; Lu Qiu &amp; Qi-Kun Bai leg.  •   11 males &amp; 2 females; Yunnan Prov.,  Yuxi City, Yuanjiang County; 533 m; 22 May 2018; Lu Qiu, Wen-Bo Deng &amp; Zhi-Wei Dong leg.   Japan • 2 males &amp; 5 females; Okinawa,  Mt. Yonahadake; 222 m; 10 Aug. 2016; Jian-Yue Qiu &amp; Hao Xu leg.</p><p>Diagnosis.</p><p>This species is characterized by its pronotum with a broad anterior and narrow anterolateral yellowish band (Fig. 2 E), a feature also presents in  P. surinamensis . The two species are difficult to distinguish in appearance. The key distinction between  P. indicus and  P. surinamensis lies in their reproductive modes:  P. indicus reproduces bisexually, while  P. surinamensis reproduces parthenogenetically (Roth 1967).</p><p>Supplementary description of male.</p><p>Hind margin of supra-anal plate variable, weakly incised medially, or entire and convexly rounded (Fig. 2 G). Sclerotized level and shape of subgenital plate variable: the inner plate nearly symmetrical or strongly asymmetrical, lateral projection on left posterolateral angle distinct or not (Fig. 2 H – J). Male genitalia: Right phallomere with caudal part of sclerite R 1 T widened; R 2 slightly curved; the projection arising from the junction of R 1 T and R 2 elongate; R 3 forked caudally; R 4 plate-like, close to the caudal part of R 1; R 5 absent (Fig. 2 K, L). Sclerite L 2 D divided into basal and apical parts: the basal part straight; the apical part croissant-shaped with outer margin toothed; apical membrane covered with heavy chaetae (Fig. 2 M). Sclerite L 3 with hook short and robust, sclerite L 4 U divided into two parts (Fig. 2 N).</p><p>Measurements (mm).</p><p>Body length including tegmen: male 15.8–17.2, female 16.4–18.6; pronotum length × width: male 4.0–4.4 × 5.4–5.9, female 4.3–5.1 × 6.0–6.7; tegmen length: male 13.8–15.3, female 12.7–15.0.</p></div>	https://treatment.plazi.org/id/72F47432B1A054F590897242613C07D4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wang, Yi-Shu;Dong, Xiao-Jiao;Wang, Zong-Qing;Che, Yan-Li	Wang, Yi-Shu, Dong, Xiao-Jiao, Wang, Zong-Qing, Che, Yan-Li (2025): Taxonomic update of Pycnoscelus Scudder, 1862 (Blattodea, Blaberidae, Pycnoscelinae), with descriptions of two new species from China. ZooKeys 1231: 45-67, DOI: 10.3897/zookeys.1231.141287
57FF9205CE935FF489DAB55AE590FA9C.text	57FF9205CE935FF489DAB55AE590FA9C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pycnoscelus nigra (Brunner von Wattenwyl 1865)	<div><p>Pycnoscelus nigra (Brunner von Wattenwyl, 1865)</p><p>Fig. 3</p><p>Panchlora nigra Brunner von Wattenwyl, 1865: 280.</p><p>Pycnoscelis [sic] nigra: Princis 1964: 274.</p><p>Pycnoscelus nigra: Princis 1967: 709; Roth, 1998: 103.</p><p>Material examined.</p><p>China • 1 female; Guangdong Prov., Zhongshan City,  Mt. Wuguishan; 5 May 2018; Ke-Liang Wu leg.  •   5 males; Yunnan Prov., Xishuangbanna, Mengla County,  Menglun Town; 25 May 2016; Lu Qiu &amp; Zhi-Wei Qiu leg.  •   1 female; Hainan Prov.,  Mt. Diaoluo shan; 275 m; 24–25 May 2014; Shun-Hua Gui &amp; Xin-Ran Li leg.  •   1 male; Chongqing City, Hechuan District,  Dashi Town; 18 July 2013; Zong-Qing Wang leg.  •   2 males &amp; 4 females; Sichuan Prov., Panzhihua City,  Xinzhuang Village; 1306 m; 15 Oct. 2014; Li He leg.  •   6 males; Yunnan Prov., Menglun Town, Xishuangbanna Tropical Botanical Garden,  Lvshilin (Green stone forest); 25 May 2016; Lu Qiu &amp; Zhi-Wei Qiu leg.  •   1 male; Yunnan Prov., Baoshan City, Mt. Gaoligongshan,  Baihualing,  Hanlong Zhai; 1400–1900 m; 20–23 June 2020; Lu Qiu &amp; Jin-Lin Liu leg.</p><p>Diagnosis.</p><p>This species is characterized by the generally dark coloration (Fig. 3 A – D) and pronotum with narrow yellow area anterolaterally (Fig. 3 E).</p><p>Supplementary description of male.</p><p>Male genitalia: Right phallomere with caudal part of sclerite R 1 T widened distinctly; R 2 nearly straight; the outgrowth arising from the junction of R 1 T and R 2 broad, plate-like; R 3 forked caudally; R 4 plate-like; R 5 absent (Fig. 3 I, J). Sclerite L 2 D with the apical part bifurcated basally; apical membrane covered with heavy chaetae (Fig. 3 K). Sclerite L 3 with hook comparatively slender, sclerite L 4 U divided into two parts (Fig. 3 L).</p><p>Measurements (mm).</p><p>Body length including tegmen: male 19.5–23.8, female 20.3–24.1; pronotum length × width: male 4.5–4.9 × 5.4–6.9, female 5.0–6.0 × 7.5–8.1; tegmen length: male 15.9–19.6, female 16.5–20.8.</p><p>Remarks.</p><p>Roth (1998) recorded a male of  P. nigra from Jingdong (= Kintung), Yunnan, and found its left style present. However, the left style is absent in all our new material from Yunnan (Fig. 3 H).</p></div>	https://treatment.plazi.org/id/57FF9205CE935FF489DAB55AE590FA9C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wang, Yi-Shu;Dong, Xiao-Jiao;Wang, Zong-Qing;Che, Yan-Li	Wang, Yi-Shu, Dong, Xiao-Jiao, Wang, Zong-Qing, Che, Yan-Li (2025): Taxonomic update of Pycnoscelus Scudder, 1862 (Blattodea, Blaberidae, Pycnoscelinae), with descriptions of two new species from China. ZooKeys 1231: 45-67, DOI: 10.3897/zookeys.1231.141287
76574D9C1F145BFDB16761FD879DCEA0.text	76574D9C1F145BFDB16761FD879DCEA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pycnoscelus puteus Wang & Che 2025	<div><p>Pycnoscelus puteus Wang &amp; Che sp. nov.</p><p>Fig. 4</p><p>Type material.</p><p>Holotype. China • male; Yunnan Prov., Xishuangbanna, Mengla County, Menglun Town,  Xishuangbanna Tropical Botanical Garden; 25 May 2016; Lu Qiu &amp; Zhi-Wei Qiu leg.; SWU -B-BB 120101  .  Paratypes. China • 2 males; same collection data as holotype; SWU -B-BB 120102 and 120103 •  1 male; same collection data as holotype; 27 May 2016; SWU -B-BB 120104 •   4 males; Yunnan Prov., Pu’er City,  Meizihu Lake; Lu Qiu &amp; Zhi-Wei Qiu leg.; 20–21 May 2016; SWU -B-BB 120105 to 120109  •   1 male; Yunnan Prov., Xishuangbanna, Jinghong City,  Dadugang; 5 May 2013; Zong-Qing Wang leg.; SWU -B-BB 120110  .</p><p>Differential diagnosis.</p><p>Pycnoscelus puteus sp. nov. is very similar to  P. nigra in general appearance and coloration but differs strongly from the later in interocular distance and male genitalia, especially the sclerotized part associated with sclerite R 1 T and R 2 (compare Fig. 3 F, I and Fig. 4 F, J in present paper). Furthermore, the male genitalia distinguish this new species from all other known species of genus  Pycnoscelus except  P. rothi, but the coloration of the pronotum could separate them easily (compare Fig. 4 E with Anisyutkin 2002: fig. 1).</p><p>Description.</p><p>Male (holotype). General color black (Fig. 4 A, B). Head black except for yellowish brown clypeo-labral area; maxillary palps and antennae brownish black; eyes black, ocellar spots whitish. Pronotum shining black, with two indistinct, narrow yellow borders anterolaterally (Fig. 4 E). Tegmina similar to those of  P. nigra, dark when folded. Abdomen and legs brownish black. Cerci slightly lighter.</p><p>Body slender. Head slightly exposed. Eyes comparatively small; outer margin of ocelli obscure; interocular space wider than the distance between the inner margin of ocellar spots, and smaller than distance between antennal sockets (Fig. 4 F). Pronotum subpentagonal, densely punctured (Fig. 4 E). Tegmina and wings extending beyond the end of abdomen. Front femur Type C 1. Hind metatarsus distinctly longer than other segments combined; four proximal tarsomeres with well-developed pulvilli, the one on the first tarsomere occupying almost the complete length of the segment; claws symmetrical and simple; arolium large. Supra-anal plate transverse, slightly asymmetrical, hind margin with a small medial incision (Fig. 4 G). Paraprocts of blaberid type, asymmetrical. Subgenital plate asymmetrical, with the right posterolateral corner acute and upturned. Left style minute, right one large and robust (Fig. 4 H).</p><p>Male genitalia. Right phallomere with caudal part of sclerite R 1 T rounded; R 2 curved; an outgrowth arising from the junction of R 1 T and R 2, with an additional heavily sclerotized part whose surface is pitted; R 3 forked caudally, R 4 plate-like, R 5 absent (Fig. 4 J, K). Sclerite L 2 D divided into basal and apical parts: the basal part short, apically widened; the apical part strongly sclerotized with posterior margin distinctly toothed; apical membrane well developed, with surface covered by microtrichia (Fig. 4 L). Sclerite L 3 hook with apical incision, inner curved margin with a small tooth; sclerite L 4 U present and divided into two parts (Fig. 4 M, N).</p><p>Variation.</p><p>Body broader, brownish black (Fig. 4 C, D); some individuals lack left style (probably missing, Fig. 4 I); inner curved margin of hook of sclerite L 3 with two small projections (Fig. 4 N).</p><p>Measurements (mm).</p><p>Male, body length including tegmina: 20.1–20.9; pronotum length × width: 3.7–4.0 × 5.0–5.6; tegmen length: 16.1–18.0.</p><p>Etymology.</p><p>Derived from the Latin word puteus, referring to the pitted surface of the sclerotized enlargement associated with sclerites R 1 T and R 2.</p></div>	https://treatment.plazi.org/id/76574D9C1F145BFDB16761FD879DCEA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wang, Yi-Shu;Dong, Xiao-Jiao;Wang, Zong-Qing;Che, Yan-Li	Wang, Yi-Shu, Dong, Xiao-Jiao, Wang, Zong-Qing, Che, Yan-Li (2025): Taxonomic update of Pycnoscelus Scudder, 1862 (Blattodea, Blaberidae, Pycnoscelinae), with descriptions of two new species from China. ZooKeys 1231: 45-67, DOI: 10.3897/zookeys.1231.141287
2BCA4C78EFCA5052B6BCB76A4512BE00.text	2BCA4C78EFCA5052B6BCB76A4512BE00.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pycnoscelus Scudder 1862	<div><p>Genus  Pycnoscelus Scudder, 1862</p><p>Fig. 1 A – C</p><p>Pycnoscelus Scudder, 1862: 421; Hebard 1917: 269; Roth 1998: 94; Anisyutkin 2018: 80. Type species: 
Pycnoscelus surinamensis (Linnaeus, 1758)
= 
Pycnoscelus obscurus Scudder, 1862, nymph, adventive, by monotypy.</p><p>Epilampria Tepper, 1894: 174. Type species: 
Pycnoscelus surinamensis (Linnaeus, 1758)
= 
Epilampria tatei Tepper, by monotypy. Synonymized by Princis 1964: 263.</p><p>Diagnosis.</p><p>Pycnoscelus species can be easily recognized by the shortened fore tibiae, which is thickened distally with strong spines (Fig. 1 A), as well as the right posterolateral corner of male subgenital plate angularly produced and upturned (Fig. 1 C).</p><p>Description.</p><p>Facial part of head and pronotum with distinct punctations. Pronotum with the anterior margin nearly truncate, hind margin convex medially. Anterior margin of fore femur Type C 1. Hind metatarsus longer or nearly equal to the remaining segments combined; armed with large pulvilli, occupying more than half length of the segment (Fig. 1 B). Claws symmetrical and unspecialized, arolium present. Subgenital plate covered by the eighth sternum; right style typically large. Male genitalia: right phallomere with R 3 forked caudally, and the left branch usually wider than the right; R 5 sometimes absent. Apical part of sclerite L 2 D well sclerotized, with the sac-like apical membrane wrapped. Sclerite L 3 hooked with small, curved portion. Sclerite L 4 U distinct and divided into two parts.</p><p>Worldwide checklist and species grouping of  Pycnoscelus</p><p>Two groups of  Pycnoscelus species were established based on the shape of right style (Roth 1998):</p><p>indicus species group: the right style elongate with apex directed posteriorly; includes 12 species:  P. conferta (Walker, 1869: 148);  P. femapterus Roth, 1998: 108;  P. gorochovi Anisyutkin, 2002: 352;  P. indicus (Fabricius, 1775: 272);  P. janetscheki Bey-Bienko, 1968: 60;  P. nigra (Brunner von Wattenwyl, 1865: 280);  P. puteus sp. nov.;  P. rothi Anisyutkin, 2002: 355;  P. schwendingeri Anisyutkin, 2018: 80;  P. surinamensis (Linnaeus, 1758: 424);  P. undulatus sp. nov.;  P. vietnamensis Anisyutkin, 2002: 355 .</p><p>striatus species group: the right style broader, plate-like, with apex directed to the left; includes five species:  P. aurantius Hanitsch, 1935: 18;  P. rufus Bey-Bienko, 1950: 268;  P. semivitreus Princis, 1967: 148;  P. striatus (Kirby, 1903: 378);  P. tenebrigera (Walker, 1868: 31) .</p><p>Note:  P. micropterus Hanitsch, 1931 has not been attributed to any group due to the lack of information on its right style. Additionally, there are six unnamed species:  Pycnoscelus sp. A Roth, 1998: 121;  Pycnoscelus sp. B Roth, 1998: 123;  Pycnoscelus sp. C Roth, 1998: 123;  Pycnoscelus sp. D Lucañas &amp; Lit, 2016: 9;  Pycnoscelus sp. E (Malaysia, Borneo);  Pycnoscelus sp. F (China, Yunnan). The latter two species are included in this work.</p></div>	https://treatment.plazi.org/id/2BCA4C78EFCA5052B6BCB76A4512BE00	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wang, Yi-Shu;Dong, Xiao-Jiao;Wang, Zong-Qing;Che, Yan-Li	Wang, Yi-Shu, Dong, Xiao-Jiao, Wang, Zong-Qing, Che, Yan-Li (2025): Taxonomic update of Pycnoscelus Scudder, 1862 (Blattodea, Blaberidae, Pycnoscelinae), with descriptions of two new species from China. ZooKeys 1231: 45-67, DOI: 10.3897/zookeys.1231.141287
78CD09723BF855BE9D1854EC0DF5219F.text	78CD09723BF855BE9D1854EC0DF5219F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pycnoscelus semivitreus Princis 1967	<div><p>Pycnoscelus semivitreus Princis, 1967</p><p>Fig. 6</p><p>Pycnoscelus semivitreus Princis, 1967: 148; Princis 1971: 1141; Roth 1998: 112.</p><p>Material examined.</p><p>Malaysia • 1 male; Borneo, Mt. Trus Madi,  Jungle Girl Camp; 2–5 Oct. 2015; Ye-Jie Lin leg.  •   1 male; Borneo, Mt. Trus Madi,  Jungle Girl Camp; 3 May 2023; Cai-Xia Yuan leg.</p><p>Diagnosis.</p><p>This species can be easily distinguished from all its congeners by its huge right style, which extends to the left posterolateral angle of subgenital plate (Fig. 6 G).</p><p>Supplementary description of male.</p><p>Male genitalia: right phallomere with caudal part of sclerite R 1 T irregular, inner margin produced at middle; R 2 nearly straight, R 3 forked, R 4 plate-like; R 5 elongated, lying above R 2 (Fig. 6 H). Apical part of sclerite L 2 D elongated, the right side with several sharp teeth; apical membrane well developed, covered with microtrichia (Fig. 6 I, J). Sclerite L 3 hook subquadrate apically, inner margin with a tooth at apex; sclerite L 4 U divided into two parts (Fig. 6 K).</p><p>Measurements (mm).</p><p>Body length including tegmen: 20.7; pronotum length × width: 4.7 × 6.2; tegmen length: 17.0.</p></div>	https://treatment.plazi.org/id/78CD09723BF855BE9D1854EC0DF5219F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wang, Yi-Shu;Dong, Xiao-Jiao;Wang, Zong-Qing;Che, Yan-Li	Wang, Yi-Shu, Dong, Xiao-Jiao, Wang, Zong-Qing, Che, Yan-Li (2025): Taxonomic update of Pycnoscelus Scudder, 1862 (Blattodea, Blaberidae, Pycnoscelinae), with descriptions of two new species from China. ZooKeys 1231: 45-67, DOI: 10.3897/zookeys.1231.141287
CFD40A95E19B54B3AF115671D88A9CD2.text	CFD40A95E19B54B3AF115671D88A9CD2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pycnoscelus striatus (Kirby 1903)	<div><p>Pycnoscelus striatus (Kirby, 1903)</p><p>Fig. 8</p><p>Leucophaea striata Kirby, 1903: 378; Kirby 1904: 151; Hanitsch 1915: 122; Chopard 1919: 358.</p><p>Pycnoscelis [sic] striata: Princis 1964: 274.</p><p>Pycnoscelus striatus: Roth, 1998: 117; Lucañas and Lit 2016: 7; Anisyutkin 2018: 83.</p><p>Material examined.</p><p>Malaysia • 5 males &amp; 1 female; Borneo, Sandakan,  Gomantong cave; 19 Apr. 2024; Wei-Wei Zhang leg.  •   1 male; Borneo, Mt. Trus Madi,  Jungle Girl Camp; 19–25 July 2016; Ren-Zhi Zhang leg.  •   1 male; Borneo, Mt. Trus Madi,  Jungle Girl Camp; 4 May 2023; Cai-Xia Yuan leg.</p><p>Diagnosis.</p><p>This species is close to  P. semivitreus in the structure of male genitalia, but the shape of the right style could distinguish them easily: extending over the middle of subgenital plate in  P. striatus (vs more elongate and extending to the left posterolateral angle of subgenital plate in  P. semivitreus).</p><p>Measurements (mm).</p><p>Macropterous, male, body length including tegmen: 23.6; pronotum length × width: 5.7 × 7.0; tegmen length: 20.8. Brachypterous, body length (from vertex to tip of abdomen): male 14.0–15.0, female 15.0; pronotum length × width: male 3.7–4.1 × 5.4, female 4.2 × 5.6; tegmen length: male 7.4–7.5, female 7.1.</p></div>	https://treatment.plazi.org/id/CFD40A95E19B54B3AF115671D88A9CD2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wang, Yi-Shu;Dong, Xiao-Jiao;Wang, Zong-Qing;Che, Yan-Li	Wang, Yi-Shu, Dong, Xiao-Jiao, Wang, Zong-Qing, Che, Yan-Li (2025): Taxonomic update of Pycnoscelus Scudder, 1862 (Blattodea, Blaberidae, Pycnoscelinae), with descriptions of two new species from China. ZooKeys 1231: 45-67, DOI: 10.3897/zookeys.1231.141287
5727C4BDD4AE521684CBD17B06E59B16.text	5727C4BDD4AE521684CBD17B06E59B16.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pycnoscelus undefined-E	<div><p>Pycnoscelus sp. E</p><p>Fig. 9</p><p>Material examined.</p><p>Malaysia • 1 male; Borneo, Mt. Trus Madi,  Jungle Girl Camp; 2–5 Oct. 2015; Ye-Jie Lin leg.  •   1 male; Borneo, Mt. Trus Madi,  Jungle Girl Camp; 3 May 2023; Cai-Xia Yuan leg.</p><p>Diagnosis.</p><p>This species can be readily distinguished from other  Pycnoscelus species by the color pattern of pronotum and tegmina. The apical part of sclerite L 2 D of this species is also peculiar in this genus (see description below for details).</p><p>Description.</p><p>General color yellowish brown, with pronotum, tegmina, and abdominal sternites partly reddish brown (Fig. 9 A, B). Facial part of head dark brownish, with a three-radial stripe between eyes; other part of head, clypeus, labrum and antennae yellowish brown; eyes black, ocellar spots yellow white. Pronotum dark reddish brown, with a yellow stripe along the anterior margin (Fig. 9 C). Tegmina bicolored, the dark part, principally half anal field reddish brown, remainder yellowish hyaline; the boundary between the two regions extends obliquely from the left posterolateral side of the anal field to half of the radial field; in the dark region, dark brown spots scattered along veins (Fig. 9 E). Abdomen yellowish, with last sternites reddish brown. Legs yellowish brown except for reddish brown tibia and tarsomere. Cerci reddish brown.</p><p>Head slightly exposed. Interocular space narrower than the distance between ocellar spots and antennal sockets (Fig. 9 D). Pronotum subpentagonal, densely punctured. Tegmina and wings fully developed, exceeding the end of abdomen. Front femur Type C 1. Hind metatarsus slightly longer than other segments combined; four proximal tarsomeres with well-developed pulvilli, the one on the first tarsomere occupying almost the whole length of the segment; claws symmetrical and simple; arolium large. Supra-anal plate weakly asymmetrical, covered with bristles; paraprocts of blaberid type, asymmetrical (Fig. 9 F). Subgenital plate asymmetrical, with the right posterolateral corner sharp and upturned. Left style absent, right style plate-like, broadly trigonal (Fig. 9 H).</p><p>Male genitalia. Right phallomere with caudal part of sclerite R 1 T rounded, inner margin produced apically; R 2 nearly straight; R 3 forked caudally, with left branch projected at apex; R 4 plate-like; R 5 lies above R 2 and fused with its distal part (Fig. 9 H). Sclerite L 2 D divided into basal and apical parts, the apical part well sclerotized, irregular, with distal part hairy instead of toothed; apical membrane less developed (Fig. 9 I). Sclerite L 3 hook very short, with the apex widened; sclerite L 4 U present and divided into two parts (Fig. 9 J).</p><p>Measurements (mm).</p><p>Body length including tegmen: 17.7–18.0; pronotum length × width: 3.8–4.0 × 4.4–4.8; tegmen length: 14.7.</p><p>Remarks.</p><p>According to the description of  P. rufus given by Anisyutkin (2004),  Pycnoscelus sp. E closely resembles this species in general color pattern, the shape of right style and right phallomere, but differs in features of pronotum and the apical part of sclerite L 2 D. We refrain from defining  Pycnoscelus sp. E as a new species until we examine the type specimen of  P. rufus .</p></div>	https://treatment.plazi.org/id/5727C4BDD4AE521684CBD17B06E59B16	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wang, Yi-Shu;Dong, Xiao-Jiao;Wang, Zong-Qing;Che, Yan-Li	Wang, Yi-Shu, Dong, Xiao-Jiao, Wang, Zong-Qing, Che, Yan-Li (2025): Taxonomic update of Pycnoscelus Scudder, 1862 (Blattodea, Blaberidae, Pycnoscelinae), with descriptions of two new species from China. ZooKeys 1231: 45-67, DOI: 10.3897/zookeys.1231.141287
D0DBC7514DEB5DAF9DC1293227C5B53A.text	D0DBC7514DEB5DAF9DC1293227C5B53A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pycnoscelus undefined-F	<div><p>Pycnoscelus sp. F</p><p>Fig. 10</p><p>Material examined.</p><p>China • 1 female; Yunnnan Prov., Mengla County, Menglun Town,  Manbian Cun; 30 July 2009; Zong-Qing Wang leg.</p><p>Measurements (mm).</p><p>Body length: 17.4; pronotum length × width: 4.3 × 7.8; tegmen length: 2.5.</p><p>Remarks.</p><p>Pycnoscelus sp. F is currently represented by a single female specimen. This specimen is characterized by its strongly reduced tegmina (lobiform and lateral) and the complete absence of hind wings. Wing reduction is also observed in the females of  P. micropterus Hanitsch, 1931,  P. striatus (Kirby, 1903),  P. femapterus Roth, 1998, and  P. undulatus sp. nov. However,  P. micropterus and  P. striatus differ from  Pycnoscelus sp. F by the tegmina reaching the abdominal terga, and the presence of hind wings. Furthermore,  P. femapterus and  P. undulatus sp. nov. can be distinguished from  Pycnoscelus sp. F by the complete lack of both tegmina and wings.</p><p>It is worth noting that no female specimens of  P. puteus sp. nov. have been recorded, and both  P. puteus sp. nov. and  Pycnoscelus sp. F were collected from Menglun Town, Mengla County, Yunnan Province. It is plausible that  Pycnoscelus sp. F represents the female form of  P. puteus but it needs further study.</p></div>	https://treatment.plazi.org/id/D0DBC7514DEB5DAF9DC1293227C5B53A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wang, Yi-Shu;Dong, Xiao-Jiao;Wang, Zong-Qing;Che, Yan-Li	Wang, Yi-Shu, Dong, Xiao-Jiao, Wang, Zong-Qing, Che, Yan-Li (2025): Taxonomic update of Pycnoscelus Scudder, 1862 (Blattodea, Blaberidae, Pycnoscelinae), with descriptions of two new species from China. ZooKeys 1231: 45-67, DOI: 10.3897/zookeys.1231.141287
152CF0214C7254AA9B34F2FBFD025818.text	152CF0214C7254AA9B34F2FBFD025818.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pycnoscelus undulatus Wang & Che 2025	<div><p>Pycnoscelus undulatus Wang &amp; Che sp. nov.</p><p>Fig. 5</p><p>Type material.</p><p>Holotype. China • male; Guangdong Prov., Guangzhou City,  Longyandong Forest Park; 28 June 2015; Zhi-Wei Qiu &amp; Yong-Quan Zhao leg.; SWU -B-BB 120201  .  Paratypes. China • 1 male; same collection data as holotype; SWU -B-BB 120202 •   1 male; Guangxi Prov., Hepu County,  Shiwan Town; 14 May 2016; Yi-Zhou Liu leg.; SWU -B-BB 120203  •   2 females &amp; 1 nymph; Guangxi Prov., Hepu County,  Shiwan Town; 14 May 2016; Yi-Zhou Liu leg.; SWU -B-BB 120204 to 120206  .</p><p>Differential diagnosis.</p><p>The male of this species is similar to  P. vietnamensis Anisyutkin, 2002 and  P. gorochovi Anisyutkin, 2002 in the shape of supra-anal plate, which has a deep emargination at the right posterolateral side, but  P. undulatus sp. nov. differs strongly from these two species by its dark reddish brown pronotum, light marking on facial part between eyes, and the bulky right style (compare Fig. 5 H with Anisyutkin 2002: figs 15, 16).</p><p>Description.</p><p>Male (holotype). General color brownish yellow (Fig. 5 A, B). Facial part of head dark reddish brown, with an indistinct three-radial stripe between eyes; other part of head, clypeus, labrum and antennae brownish yellow; maxillary palps brownish yellow with reddish brown maculae; eyes black, ocellar spots whitish. Pronotum reddish brown, with two yellow areas anterolaterally (Fig. 5 E). Tegmina brownish yellow. Abdomen yellowish brown to reddish brown. Legs and cerci yellowish brown.</p><p>Head slightly exposed. Ocelli subrectangular; interocular space distinctly less than the distance between ocellar spots and antennal sockets (Fig. 5 F). Pronotum subpentagonal, densely punctured. Tegmina and wings fully developed, exceeding the end of abdomen. Front femur Type C 1. Hind metatarsus not quite as long as other segments combined; four proximal tarsomeres with well-developed pulvilli, the one on the first tarsomere occupying almost the whole length of the segment; claws symmetrical and simple; arolium large. Supra-anal plate asymmetrical with hind margin deeply concave at the right side. Paraprocts of blaberid type, asymmetrical. Cerci stout at base (Fig. 5 G). Subgenital plate asymmetrical, with the right posterolateral corner slightly obtuse and upturned. Left style absent, right style broadly trigonal (Fig. 5 H).</p><p>Male genitalia. Right phallomere with caudal part of sclerite R 1 T irregular and widened; R 2 short and curved; a well-developed projection arising from the junction of R 1 T and R 2; R 3 forked caudally, R 4 plate-like, R 5 absent (Fig. 5 I, J). Sclerite L 2 D divided into basal and apical parts: the basal part forked caudally with the inner margin indented; the apical part well sclerotized and more uniform in width than in other  Pycnoscelus species, posterior margin toothed; apical membrane covered with microtrichia (Fig. 5 K). Sclerite L 3 hook without an apical incision, inner curved margin with a sharp convexity at apex; sclerite L 4 U present and divided into two parts (Fig. 5 L).</p><p>Females. Thoracic segments and abdominal tergum black with minute punctures, abdominal sterna and legs reddish brown. Tegmina and wings absent.</p><p>Measurements (mm).</p><p>Male, body length including tegmen: 22.2–23.0; pronotum length × width: 4.3–4.5 × 6.3–7.1; tegmen length: 17.0–20.1. Female, body length: 16.1–20.6; pronotum length × width: 4.35–4.44 × 7.36–7.37.</p><p>Etymology.</p><p>The species epithet is derived from the Latin word undulatus, which refers to the caudal edge of basal part of sclerite L 2 D undulated.</p></div>	https://treatment.plazi.org/id/152CF0214C7254AA9B34F2FBFD025818	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wang, Yi-Shu;Dong, Xiao-Jiao;Wang, Zong-Qing;Che, Yan-Li	Wang, Yi-Shu, Dong, Xiao-Jiao, Wang, Zong-Qing, Che, Yan-Li (2025): Taxonomic update of Pycnoscelus Scudder, 1862 (Blattodea, Blaberidae, Pycnoscelinae), with descriptions of two new species from China. ZooKeys 1231: 45-67, DOI: 10.3897/zookeys.1231.141287
