taxonID	type	description	language	source
E881AB7106EF53F589CB6436ABF171BD.taxon	description	Figs 1, 4	en	Yin, Zi-Wei, Feng, Ting, Zhou, De-Yao (2025): First report of subfamily Scydmaeninae (Coleoptera, Staphylinidae) from Shanghai, with description of two new species. ZooKeys 1231: 133-143, DOI: 10.3897/zookeys.1231.142538
E881AB7106EF53F589CB6436ABF171BD.taxon	diagnosis	Diagnosis. Male. Body length 1.4 – 1.7 mm. Dorsum of body finely punctate. Thick bristles present on tempora and sides of pronotum, especially dense on tempora. Anterior margin of clypeus (Fig. 1 B) angularly prominent at middle. Antenna elongate, club loosely formed by apical four moderately enlarged antennomeres, occupying about half of antennal length. Pronotum bell-shaped, with two asetose basolateral pits connected by shallow transverse impression. Each elytron with two closely-placed basal pits. Abdomen greatly modified (Fig. 1 C), sternite 4 (VI) and 5 (VII) each with two posterolateral nodules directed posteromedially, area between nodules on sternite 4 filled with peg-like granules distributed roughly in two transverse rows. Aedeagus (Fig. 1 D, E) with apical projection much longer than parameres; endophallus armature composed of group of symmetric sclerites; parameres each greatly broadened, bearing two long setae near apex and three similar setae along lateral margin. Female (Fig. 1 A). External morphology similar to male. Abdomen unmodified. Spermatheca (Fig. 1 F) spherical; spermathecal duct broadened at base.	en	Yin, Zi-Wei, Feng, Ting, Zhou, De-Yao (2025): First report of subfamily Scydmaeninae (Coleoptera, Staphylinidae) from Shanghai, with description of two new species. ZooKeys 1231: 133-143, DOI: 10.3897/zookeys.1231.142538
E881AB7106EF53F589CB6436ABF171BD.taxon	description	Description. See Vit (2006) and Jałoszyński (2022). Measurement for Shanghai population: male body length 1.48 mm; length / width of head 0.35 mm / 0.34 mm, pronotum 0.38 mm / 0.37 mm, elytra 0.78 mm / 0.63 mm, length of antenna 0.81 mm, club 0.42 mm, length of aedeagus 0.27 mm; female body length 1.49 – 1.53 mm; length / width of head 0.37 – 0.39 mm / 0.34 – 0.38 mm, pronotum 0.41 – 0.43 mm / 0.39 – 0.41 mm, elytra 0.86 – 0.88 mm / 0.67 – 0.71 mm, length of antenna 0.77 – 0.80 mm, club 0.36 – 0.38 mm, maximum diameter of spermatheca 0.27 mm.	en	Yin, Zi-Wei, Feng, Ting, Zhou, De-Yao (2025): First report of subfamily Scydmaeninae (Coleoptera, Staphylinidae) from Shanghai, with description of two new species. ZooKeys 1231: 133-143, DOI: 10.3897/zookeys.1231.142538
E881AB7106EF53F589CB6436ABF171BD.taxon	distribution	Distribution. East China: Anhui, Shanghai (Minhang, Songjiang, Jiading) (Fig. 4 A); Japan: Honshu, Kyushu; South Korea: Jeju. New record for Shanghai.	en	Yin, Zi-Wei, Feng, Ting, Zhou, De-Yao (2025): First report of subfamily Scydmaeninae (Coleoptera, Staphylinidae) from Shanghai, with description of two new species. ZooKeys 1231: 133-143, DOI: 10.3897/zookeys.1231.142538
B9BF696E0AF65B24B48D6068CC860011.taxon	description	Figs 2, 4	en	Yin, Zi-Wei, Feng, Ting, Zhou, De-Yao (2025): First report of subfamily Scydmaeninae (Coleoptera, Staphylinidae) from Shanghai, with description of two new species. ZooKeys 1231: 133-143, DOI: 10.3897/zookeys.1231.142538
B9BF696E0AF65B24B48D6068CC860011.taxon	diagnosis	Diagnosis. Male. Habitus elongate; body length approximately 1.7 mm. Head and elytra finely punctate, subglabrous, with sparse long setae; punctation and setae of pronotal disc similar to those of head and elytra, lateral margins densely setose and with numerous thick bristles. Head subspherical, eyes anteriorly situated, tempora much longer than eyes. Antennae elongate, antennomeres elongate, clubs loosely formed by apical four enlarged antennomeres. Pronotum lacking antebasal pits, transverse impression, or sublateral carinae; broadest slightly posterior to middle. Tarsomere 1 of protarsus modified, ventrally protruding to form apically truncate projection. Aedeagus moderately elongate, dorso-ventrally symmetric; compressor plate in ventral view with two lateral lobes; apical projection of median lobe broad at base and narrowing apically; parameres broadened before apices, each paramere with three macrosetae at apex. Female. Unknown.	en	Yin, Zi-Wei, Feng, Ting, Zhou, De-Yao (2025): First report of subfamily Scydmaeninae (Coleoptera, Staphylinidae) from Shanghai, with description of two new species. ZooKeys 1231: 133-143, DOI: 10.3897/zookeys.1231.142538
B9BF696E0AF65B24B48D6068CC860011.taxon	description	Description. Male. Body (Fig. 2 A) length 1.66 mm; body uniformly reddish-brown, mouthparts and tarsi paler in color. Setae long and suberect, sparse on head and pronotal and elytral discs, sides of pronotum with dense, thick bristles. Dorsum of body finely and sparsely punctate, almost glabrous. Head subrounded, as long as wide, broadest at eyes, length and width 0.36 mm; vertex and frons confluent, weakly convex; supraantennal tubercles barely prominent; eyes relatively small, barely convex and finely faceted. Punctures on vertex and frons fine; setae long and sparse, suberect, tempora much longer than eyes, lacking bristles. Antenna elongate, length 0.93 mm, club 0.48 mm; antennomeres 1 – 7 each elongate, 2 and 7 longest, 8 – 11 broader than 2 – 7, enlarged, form loose club. Pronotum in dorsal view slightly longer than wide, subglobose, broadest slightly posterior to middle and narrowing anteriorly and posteriorly, length 0.4 width 0.41 mm; lacking lateral antebasal pits, transverse antebasal groove and sublateral carinae. Punctures on pronotal disc rather fine, almost glabrous; setae sparse and long, laterally obscured by dense, long and thick bristles. Elytra suboval and slightly flattened, broadest approximately at middle, length 0.94 mm, width 0.73 mm, length / width 1.29; basal impressions shallow, with four small, asetose basal pits, humeral calli weak; apices of elytra separately rounded. Punctures on elytral disc fine and shallow; sparse setae long and suberect. Metathoracic wings fully developed, functional. Meso- and metaventrite fused. Mesoventral intercoxal process posteriorly extending far beyond level of posterior margin of mesocoxae. Metaventral intercoxal process broad, emarginate at middle. Legs long and slender; protarsus with tarsomere 1 projecting ventrally, forming short, apically truncate lobe (Fig. 2 B). Aedeagus (Fig. 2 C – F) moderately elongate, dorso-ventrally almost symmetric, length 0.34 mm, in ventral view median lobe with long, apically narrowing projection greatly curved dorsally, with two rows of fine setae along dorsal wall of projection; compressor plate broadened in dorso-ventral view, with pair of short and broad lateral lobes, plate inclined to apically-projected median lobe in lateral view and curved dorsally; endophallus armature composed of pairs of elongate and sclerotized plates and membranous structures; parameres elongate, narrow, extending just beyond base of apical projection of median lobe, areas before apices greatly broadened and then abruptly narrowing apically, each paramere with three long macrosetae at apex. Female. Unknown.	en	Yin, Zi-Wei, Feng, Ting, Zhou, De-Yao (2025): First report of subfamily Scydmaeninae (Coleoptera, Staphylinidae) from Shanghai, with description of two new species. ZooKeys 1231: 133-143, DOI: 10.3897/zookeys.1231.142538
B9BF696E0AF65B24B48D6068CC860011.taxon	distribution	Distribution. East China: Shanghai (Jiading) (Fig. 4 A).	en	Yin, Zi-Wei, Feng, Ting, Zhou, De-Yao (2025): First report of subfamily Scydmaeninae (Coleoptera, Staphylinidae) from Shanghai, with description of two new species. ZooKeys 1231: 133-143, DOI: 10.3897/zookeys.1231.142538
B9BF696E0AF65B24B48D6068CC860011.taxon	etymology	Etymology. The specific epithet is derived from a combination of E. impar, a closely related species, and the Latin suffix “ - itus (- a, - um) ”, denoting an affinity between these two species.	en	Yin, Zi-Wei, Feng, Ting, Zhou, De-Yao (2025): First report of subfamily Scydmaeninae (Coleoptera, Staphylinidae) from Shanghai, with description of two new species. ZooKeys 1231: 133-143, DOI: 10.3897/zookeys.1231.142538
72F9430C540054B7A61651F1AA31E520.taxon	description	Figs 3, 4	en	Yin, Zi-Wei, Feng, Ting, Zhou, De-Yao (2025): First report of subfamily Scydmaeninae (Coleoptera, Staphylinidae) from Shanghai, with description of two new species. ZooKeys 1231: 133-143, DOI: 10.3897/zookeys.1231.142538
72F9430C540054B7A61651F1AA31E520.taxon	diagnosis	Diagnosis. Male. Body length approximately 1.5 mm. Eyes greatly prominent, approximately 1.4 × as long as tempora. Terminal four antennomeres greatly enlarged and forming distinct club, occupying approximately 5.5 / 10 of antennal length. Sides of elytra distinctly narrowing posteriorly from broadest point. Aedeagus with compressor plate elongate and subfusiform in ventral view; apical projection of median lobe rounded and greatly protruding in ventral view, curved dorsally in apical portion in lateral view; median lobe with pair of apically rounded lateral projections, and transversely rhomboidal plate on ventral wall; broad and elongate parameres narrowing from bases toward apices, each with two long setae at apex and three similar long setae along apical 2 / 5. Female. Unknown.	en	Yin, Zi-Wei, Feng, Ting, Zhou, De-Yao (2025): First report of subfamily Scydmaeninae (Coleoptera, Staphylinidae) from Shanghai, with description of two new species. ZooKeys 1231: 133-143, DOI: 10.3897/zookeys.1231.142538
72F9430C540054B7A61651F1AA31E520.taxon	description	Description. Male. Body (Fig. 3 A) length 1.52 mm; body uniformly reddish-brown, mouthparts and tarsi paler in color. Setae long and suberect, tempora of head and sides of pronotum with dense, thick bristles. Dorsum of body finely and sparsely punctate. Head roundly rhomboidal, as long as wide, broadest at eyes, length and width 0.31 mm; vertex and frons confluent, weakly convex; supraantennal tubercles barely prominent; eyes large, strongly convex and coarsely faceted. Punctures on vertex and frons fine; setae long and sparse, suberect, additionally tempora with long bristles directed posteriorly. Antenna moderately short, length 0.56 mm, club 0.31 mm; antennomeres 1 and 2 subcylindrical, elongate, 3 – 7 compact, gradually larger, 8 – 11 greatly enlarged, conical, 11 largest, distinctly shorter than 9 and 10 combined. Pronotum in dorsal view subtrapezoidal, broadest at base and strongly narrowing anteriorly, length 0.35 mm, width 0.38 mm; lateral antebasal pits small but distinct, asetose, connected by transverse antebasal groove. Punctures on pronotal disc fine; setae long, obscured by dense, long and thick bristles especially on sides. Elytra suboval and slightly flattened, broadest approximately at middle, length 0.89 mm, width 0.63 mm, length / width 1.39; basal impressions shallow but distinct, with four small, asetose basal pits, humeral calli elongate; apices of elytra separately rounded. Punctures on elytral disc fine and shallow; setae long, sparse and suberect. Metathoracic wings fully developed, functional. Meso- and metaventrite fused. Sides of mesoventral intercoxal process posteriorly divergent, form pair of ridges, similar to condition in E. maklinii (Mannerheim) (Jałoszyński 2021: fig. 8). Metaventral intercoxal process relatively narrow. Legs long and slender; unmodified. Aedeagus (Fig. 3 B – E) moderately elongate, dorso-ventrally almost symmetric, length 0.31 mm, in ventral view median lobe with abruptly delimited and long, broad apical projection greatly curved dorsally, rounded at apex; compressor plate relatively narrow and subfusiform in dorso-ventral view, with narrowed anterior and posterior margins, connected in parallel to median lobe in lateral view; lateral projections broad and partially sclerotized, curved dorsally, with round apices; endophallus armature composed of pairs of complex, symmetric sclerotized plates and large, transversely rhomboidal plate, its apical margin with two admesal roundly acute projections; parameres broad and elongate, narrowing from bases toward apices, each with two long setae at apex, and three similar long setae along apical 2 / 5. Female. Unknown.	en	Yin, Zi-Wei, Feng, Ting, Zhou, De-Yao (2025): First report of subfamily Scydmaeninae (Coleoptera, Staphylinidae) from Shanghai, with description of two new species. ZooKeys 1231: 133-143, DOI: 10.3897/zookeys.1231.142538
72F9430C540054B7A61651F1AA31E520.taxon	distribution	Distribution. East China: Shanghai (Hongqiao) (Fig. 4 A).	en	Yin, Zi-Wei, Feng, Ting, Zhou, De-Yao (2025): First report of subfamily Scydmaeninae (Coleoptera, Staphylinidae) from Shanghai, with description of two new species. ZooKeys 1231: 133-143, DOI: 10.3897/zookeys.1231.142538
72F9430C540054B7A61651F1AA31E520.taxon	etymology	Etymology. The name is a combination of the Latin adjective “ magnus (great, large) ” and noun “ oculus (eye) ”, referring to the large eyes of this species.	en	Yin, Zi-Wei, Feng, Ting, Zhou, De-Yao (2025): First report of subfamily Scydmaeninae (Coleoptera, Staphylinidae) from Shanghai, with description of two new species. ZooKeys 1231: 133-143, DOI: 10.3897/zookeys.1231.142538
