taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
E643E79E16E25B38A63F64311C7B2DCD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539530	https://doi.org/10.3897/asp.84.e161376.figure5	Figure 5. SEM images. A A. subpulchellus, head and pronotum, lateral view, ZISP _ ENT 00003767. B A. limbatus, Europe, Head and pronotum, lateral view, ZISP _ ENT 00016011. C A. limbatus, Europe, cuneus and membrane, ZISP _ ENT 00015959. D A. limbatus Europe, head and pronotum, dorsal view, ZISP _ ENT 00015959. E Pretarsus, dorsal view, A. spinolae, ZISP _ ENT 00015686. F Pleura, A. limbatus, Europe, ZISP _ ENT 00016011. G Pleura, A. maackiae, ZISP _ ENT 00016248. H Pretarsus, ventral view, A. spinolae, ZISP _ ENT 00015187. I Scutellum, clavus, corium, A. adustus, ZISP _ ENT 00015920. J Labium, A. subpulchellus, ZISP _ ENT 00015177. K Hind tarsus, A. maackiae, ZISP _ ENT 00016246.	Figure 5. SEM images. A A. subpulchellus, head and pronotum, lateral view, ZISP _ ENT 00003767. B A. limbatus, Europe, Head and pronotum, lateral view, ZISP _ ENT 00016011. C A. limbatus, Europe, cuneus and membrane, ZISP _ ENT 00015959. D A. limbatus Europe, head and pronotum, dorsal view, ZISP _ ENT 00015959. E Pretarsus, dorsal view, A. spinolae, ZISP _ ENT 00015686. F Pleura, A. limbatus, Europe, ZISP _ ENT 00016011. G Pleura, A. maackiae, ZISP _ ENT 00016248. H Pretarsus, ventral view, A. spinolae, ZISP _ ENT 00015187. I Scutellum, clavus, corium, A. adustus, ZISP _ ENT 00015920. J Labium, A. subpulchellus, ZISP _ ENT 00015177. K Hind tarsus, A. maackiae, ZISP _ ENT 00016246.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
E643E79E16E25B38A63F64311C7B2DCD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539531	https://doi.org/10.3897/asp.84.e161376.figure6	Figure 6. Female genitalia. A. adustus, ZISP _ ENT 00016215. A Dorsal labiate plate. B Posterior wall. A. fraxinicola ZISP _ ENT 00016592. C Posterior wall. D Dorsal labiate plate. A. furvus, holotype series, ZISP _ ENT 00016298. E Dorsal labiate plate. F Posterior wall. A. furvus, paratype series, ZISP _ ENT 00016055. G Dorsal labiate plate. H Posterior wall.	Figure 6. Female genitalia. A. adustus, ZISP _ ENT 00016215. A Dorsal labiate plate. B Posterior wall. A. fraxinicola ZISP _ ENT 00016592. C Posterior wall. D Dorsal labiate plate. A. furvus, holotype series, ZISP _ ENT 00016298. E Dorsal labiate plate. F Posterior wall. A. furvus, paratype series, ZISP _ ENT 00016055. G Dorsal labiate plate. H Posterior wall.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
E643E79E16E25B38A63F64311C7B2DCD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539534	https://doi.org/10.3897/asp.84.e161376.figure9	Figure 9. Left parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola. ZISP _ ENT 00014862. С dorsal view. D. posterior view. A. furvus, holotype ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. AK genital capsule. A. gotorum ZISP _ ENT 00016164. I posterior view. A. hilaris ZISP _ ENT 00014891. J dorsal view. K posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00014891. L dorsal view. M posterior view. A. infamis ZISP _ ENT 00014891. N dorsal view. O posterior view. A. maackiae ZISP _ ENT 00014893. P dorsal view. Q posterior view. A. limbatus Europe, ZISP _ ENT 00016012. R dorsal view. S posterior view. A. limbatus Asia, ZISP _ ENT 00014886. T dorsal view. U posterior view. V theca. A. shikotan sp. nov. ZISP _ ENT 00016047 holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	Figure 9. Left parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola. ZISP _ ENT 00014862. С dorsal view. D. posterior view. A. furvus, holotype ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. AK genital capsule. A. gotorum ZISP _ ENT 00016164. I posterior view. A. hilaris ZISP _ ENT 00014891. J dorsal view. K posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00014891. L dorsal view. M posterior view. A. infamis ZISP _ ENT 00014891. N dorsal view. O posterior view. A. maackiae ZISP _ ENT 00014893. P dorsal view. Q posterior view. A. limbatus Europe, ZISP _ ENT 00016012. R dorsal view. S posterior view. A. limbatus Asia, ZISP _ ENT 00014886. T dorsal view. U posterior view. V theca. A. shikotan sp. nov. ZISP _ ENT 00016047 holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
E643E79E16E25B38A63F64311C7B2DCD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539535	https://doi.org/10.3897/asp.84.e161376.figure10	Figure 10. Habitus, dorsal view. A. adustus. A Male, ZISP _ ENT 00016237. B Female, ZISP _ ENT 00015922. A. fraxinicola. C Female, ZISP _ ENT 00016596. D Male, ZISP _ ENT 00015938. A. furvus. E Male, holotype. ZISP _ ENT 00016303. F Female, ZISP _ ENT 00016297. A. gotorum. G Male, ZISP _ ENT 00016232. A. hilaris. H Male, ZISP _ ENT 00015182. I Female, paratype, ZISP _ ENT 00015184. A. infamis. J Male, ZISP _ ENT 00016250, holotype.	Figure 10. Habitus, dorsal view. A. adustus. A Male, ZISP _ ENT 00016237. B Female, ZISP _ ENT 00015922. A. fraxinicola. C Female, ZISP _ ENT 00016596. D Male, ZISP _ ENT 00015938. A. furvus. E Male, holotype. ZISP _ ENT 00016303. F Female, ZISP _ ENT 00016297. A. gotorum. G Male, ZISP _ ENT 00016232. A. hilaris. H Male, ZISP _ ENT 00015182. I Female, paratype, ZISP _ ENT 00015184. A. infamis. J Male, ZISP _ ENT 00016250, holotype.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
E643E79E16E25B38A63F64311C7B2DCD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539540	https://doi.org/10.3897/asp.84.e161376.figure15	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
E643E79E16E25B38A63F64311C7B2DCD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539541	https://doi.org/10.3897/asp.84.e161376.figure16	Figure 16. Vesica. A. adustus, ZISP _ ENT 00014889. A Right lateral view. B Left lateral view. A. sp. nr. furvus (M 199), ZISP _ ENT 00014869. C Right lateral view. D Left lateral view. A. fraxinicola, ZISP _ ENT 00014887. E Right lateral view. F Left lateral view. A. furvus (holotype), ZISP _ ENT 00016303. G Right lateral view. H Left lateral view. A. gotorum, ZISP _ ENT 00016232. I Right lateral view. J Left lateral view. A. furvus (paratype), ZISP _ ENT 00003762. K Right lateral view. L Left lateral view.	Figure 16. Vesica. A. adustus, ZISP _ ENT 00014889. A Right lateral view. B Left lateral view. A. sp. nr. furvus (M 199), ZISP _ ENT 00014869. C Right lateral view. D Left lateral view. A. fraxinicola, ZISP _ ENT 00014887. E Right lateral view. F Left lateral view. A. furvus (holotype), ZISP _ ENT 00016303. G Right lateral view. H Left lateral view. A. gotorum, ZISP _ ENT 00016232. I Right lateral view. J Left lateral view. A. furvus (paratype), ZISP _ ENT 00003762. K Right lateral view. L Left lateral view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
E643E79E16E25B38A63F64311C7B2DCD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539545	https://doi.org/10.3897/asp.84.e161376.figure19	Figure 19. Right parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola ZISP _ ENT 00014862. C dorsal view. D posterior view. A. furvus, holotype, ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. A. gotorum ZISP _ ENT 00016164. I dorsal view, J posterior view. A. hilaris ZISP _ ENT 00014891. K dorsal view. L posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00015180. M dorsal view. N posterior view. A. infamis ZISP _ ENT 00016250. O dorsal view. P posterior view. A. maackiae, ZISP _ ENT 00014893. Q dorsal view. R posterior view. A. limbatus Europe ZISP _ ENT 00016012. S dorsal view. T posterior view. A. limbatus Asia ZISP _ ENT 00014886. U dorsal view. V posterior view. A. shikotan sp. nov. ZISP _ ENT 00016047, holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris, ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. A. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	Figure 19. Right parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola ZISP _ ENT 00014862. C dorsal view. D posterior view. A. furvus, holotype, ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. A. gotorum ZISP _ ENT 00016164. I dorsal view, J posterior view. A. hilaris ZISP _ ENT 00014891. K dorsal view. L posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00015180. M dorsal view. N posterior view. A. infamis ZISP _ ENT 00016250. O dorsal view. P posterior view. A. maackiae, ZISP _ ENT 00014893. Q dorsal view. R posterior view. A. limbatus Europe ZISP _ ENT 00016012. S dorsal view. T posterior view. A. limbatus Asia ZISP _ ENT 00014886. U dorsal view. V posterior view. A. shikotan sp. nov. ZISP _ ENT 00016047, holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris, ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. A. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
1E94B7CA60BB5930A4FBCE072E8AC40B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539531	https://doi.org/10.3897/asp.84.e161376.figure6	Figure 6. Female genitalia. A. adustus, ZISP _ ENT 00016215. A Dorsal labiate plate. B Posterior wall. A. fraxinicola ZISP _ ENT 00016592. C Posterior wall. D Dorsal labiate plate. A. furvus, holotype series, ZISP _ ENT 00016298. E Dorsal labiate plate. F Posterior wall. A. furvus, paratype series, ZISP _ ENT 00016055. G Dorsal labiate plate. H Posterior wall.	Figure 6. Female genitalia. A. adustus, ZISP _ ENT 00016215. A Dorsal labiate plate. B Posterior wall. A. fraxinicola ZISP _ ENT 00016592. C Posterior wall. D Dorsal labiate plate. A. furvus, holotype series, ZISP _ ENT 00016298. E Dorsal labiate plate. F Posterior wall. A. furvus, paratype series, ZISP _ ENT 00016055. G Dorsal labiate plate. H Posterior wall.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
1E94B7CA60BB5930A4FBCE072E8AC40B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539534	https://doi.org/10.3897/asp.84.e161376.figure9	Figure 9. Left parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola. ZISP _ ENT 00014862. С dorsal view. D. posterior view. A. furvus, holotype ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. AK genital capsule. A. gotorum ZISP _ ENT 00016164. I posterior view. A. hilaris ZISP _ ENT 00014891. J dorsal view. K posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00014891. L dorsal view. M posterior view. A. infamis ZISP _ ENT 00014891. N dorsal view. O posterior view. A. maackiae ZISP _ ENT 00014893. P dorsal view. Q posterior view. A. limbatus Europe, ZISP _ ENT 00016012. R dorsal view. S posterior view. A. limbatus Asia, ZISP _ ENT 00014886. T dorsal view. U posterior view. V theca. A. shikotan sp. nov. ZISP _ ENT 00016047 holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	Figure 9. Left parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola. ZISP _ ENT 00014862. С dorsal view. D. posterior view. A. furvus, holotype ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. AK genital capsule. A. gotorum ZISP _ ENT 00016164. I posterior view. A. hilaris ZISP _ ENT 00014891. J dorsal view. K posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00014891. L dorsal view. M posterior view. A. infamis ZISP _ ENT 00014891. N dorsal view. O posterior view. A. maackiae ZISP _ ENT 00014893. P dorsal view. Q posterior view. A. limbatus Europe, ZISP _ ENT 00016012. R dorsal view. S posterior view. A. limbatus Asia, ZISP _ ENT 00014886. T dorsal view. U posterior view. V theca. A. shikotan sp. nov. ZISP _ ENT 00016047 holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
1E94B7CA60BB5930A4FBCE072E8AC40B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539535	https://doi.org/10.3897/asp.84.e161376.figure10	Figure 10. Habitus, dorsal view. A. adustus. A Male, ZISP _ ENT 00016237. B Female, ZISP _ ENT 00015922. A. fraxinicola. C Female, ZISP _ ENT 00016596. D Male, ZISP _ ENT 00015938. A. furvus. E Male, holotype. ZISP _ ENT 00016303. F Female, ZISP _ ENT 00016297. A. gotorum. G Male, ZISP _ ENT 00016232. A. hilaris. H Male, ZISP _ ENT 00015182. I Female, paratype, ZISP _ ENT 00015184. A. infamis. J Male, ZISP _ ENT 00016250, holotype.	Figure 10. Habitus, dorsal view. A. adustus. A Male, ZISP _ ENT 00016237. B Female, ZISP _ ENT 00015922. A. fraxinicola. C Female, ZISP _ ENT 00016596. D Male, ZISP _ ENT 00015938. A. furvus. E Male, holotype. ZISP _ ENT 00016303. F Female, ZISP _ ENT 00016297. A. gotorum. G Male, ZISP _ ENT 00016232. A. hilaris. H Male, ZISP _ ENT 00015182. I Female, paratype, ZISP _ ENT 00015184. A. infamis. J Male, ZISP _ ENT 00016250, holotype.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
1E94B7CA60BB5930A4FBCE072E8AC40B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539540	https://doi.org/10.3897/asp.84.e161376.figure15	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
1E94B7CA60BB5930A4FBCE072E8AC40B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539541	https://doi.org/10.3897/asp.84.e161376.figure16	Figure 16. Vesica. A. adustus, ZISP _ ENT 00014889. A Right lateral view. B Left lateral view. A. sp. nr. furvus (M 199), ZISP _ ENT 00014869. C Right lateral view. D Left lateral view. A. fraxinicola, ZISP _ ENT 00014887. E Right lateral view. F Left lateral view. A. furvus (holotype), ZISP _ ENT 00016303. G Right lateral view. H Left lateral view. A. gotorum, ZISP _ ENT 00016232. I Right lateral view. J Left lateral view. A. furvus (paratype), ZISP _ ENT 00003762. K Right lateral view. L Left lateral view.	Figure 16. Vesica. A. adustus, ZISP _ ENT 00014889. A Right lateral view. B Left lateral view. A. sp. nr. furvus (M 199), ZISP _ ENT 00014869. C Right lateral view. D Left lateral view. A. fraxinicola, ZISP _ ENT 00014887. E Right lateral view. F Left lateral view. A. furvus (holotype), ZISP _ ENT 00016303. G Right lateral view. H Left lateral view. A. gotorum, ZISP _ ENT 00016232. I Right lateral view. J Left lateral view. A. furvus (paratype), ZISP _ ENT 00003762. K Right lateral view. L Left lateral view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
1E94B7CA60BB5930A4FBCE072E8AC40B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539545	https://doi.org/10.3897/asp.84.e161376.figure19	Figure 19. Right parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola ZISP _ ENT 00014862. C dorsal view. D posterior view. A. furvus, holotype, ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. A. gotorum ZISP _ ENT 00016164. I dorsal view, J posterior view. A. hilaris ZISP _ ENT 00014891. K dorsal view. L posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00015180. M dorsal view. N posterior view. A. infamis ZISP _ ENT 00016250. O dorsal view. P posterior view. A. maackiae, ZISP _ ENT 00014893. Q dorsal view. R posterior view. A. limbatus Europe ZISP _ ENT 00016012. S dorsal view. T posterior view. A. limbatus Asia ZISP _ ENT 00014886. U dorsal view. V posterior view. A. shikotan sp. nov. ZISP _ ENT 00016047, holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris, ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. A. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	Figure 19. Right parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola ZISP _ ENT 00014862. C dorsal view. D posterior view. A. furvus, holotype, ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. A. gotorum ZISP _ ENT 00016164. I dorsal view, J posterior view. A. hilaris ZISP _ ENT 00014891. K dorsal view. L posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00015180. M dorsal view. N posterior view. A. infamis ZISP _ ENT 00016250. O dorsal view. P posterior view. A. maackiae, ZISP _ ENT 00014893. Q dorsal view. R posterior view. A. limbatus Europe ZISP _ ENT 00016012. S dorsal view. T posterior view. A. limbatus Asia ZISP _ ENT 00014886. U dorsal view. V posterior view. A. shikotan sp. nov. ZISP _ ENT 00016047, holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris, ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. A. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
0EC4BF09B9FC5EE9B49D8927164D0AB3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539531	https://doi.org/10.3897/asp.84.e161376.figure6	Figure 6. Female genitalia. A. adustus, ZISP _ ENT 00016215. A Dorsal labiate plate. B Posterior wall. A. fraxinicola ZISP _ ENT 00016592. C Posterior wall. D Dorsal labiate plate. A. furvus, holotype series, ZISP _ ENT 00016298. E Dorsal labiate plate. F Posterior wall. A. furvus, paratype series, ZISP _ ENT 00016055. G Dorsal labiate plate. H Posterior wall.	Figure 6. Female genitalia. A. adustus, ZISP _ ENT 00016215. A Dorsal labiate plate. B Posterior wall. A. fraxinicola ZISP _ ENT 00016592. C Posterior wall. D Dorsal labiate plate. A. furvus, holotype series, ZISP _ ENT 00016298. E Dorsal labiate plate. F Posterior wall. A. furvus, paratype series, ZISP _ ENT 00016055. G Dorsal labiate plate. H Posterior wall.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
0EC4BF09B9FC5EE9B49D8927164D0AB3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539534	https://doi.org/10.3897/asp.84.e161376.figure9	Figure 9. Left parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola. ZISP _ ENT 00014862. С dorsal view. D. posterior view. A. furvus, holotype ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. AK genital capsule. A. gotorum ZISP _ ENT 00016164. I posterior view. A. hilaris ZISP _ ENT 00014891. J dorsal view. K posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00014891. L dorsal view. M posterior view. A. infamis ZISP _ ENT 00014891. N dorsal view. O posterior view. A. maackiae ZISP _ ENT 00014893. P dorsal view. Q posterior view. A. limbatus Europe, ZISP _ ENT 00016012. R dorsal view. S posterior view. A. limbatus Asia, ZISP _ ENT 00014886. T dorsal view. U posterior view. V theca. A. shikotan sp. nov. ZISP _ ENT 00016047 holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	Figure 9. Left parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola. ZISP _ ENT 00014862. С dorsal view. D. posterior view. A. furvus, holotype ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. AK genital capsule. A. gotorum ZISP _ ENT 00016164. I posterior view. A. hilaris ZISP _ ENT 00014891. J dorsal view. K posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00014891. L dorsal view. M posterior view. A. infamis ZISP _ ENT 00014891. N dorsal view. O posterior view. A. maackiae ZISP _ ENT 00014893. P dorsal view. Q posterior view. A. limbatus Europe, ZISP _ ENT 00016012. R dorsal view. S posterior view. A. limbatus Asia, ZISP _ ENT 00014886. T dorsal view. U posterior view. V theca. A. shikotan sp. nov. ZISP _ ENT 00016047 holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
0EC4BF09B9FC5EE9B49D8927164D0AB3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539535	https://doi.org/10.3897/asp.84.e161376.figure10	Figure 10. Habitus, dorsal view. A. adustus. A Male, ZISP _ ENT 00016237. B Female, ZISP _ ENT 00015922. A. fraxinicola. C Female, ZISP _ ENT 00016596. D Male, ZISP _ ENT 00015938. A. furvus. E Male, holotype. ZISP _ ENT 00016303. F Female, ZISP _ ENT 00016297. A. gotorum. G Male, ZISP _ ENT 00016232. A. hilaris. H Male, ZISP _ ENT 00015182. I Female, paratype, ZISP _ ENT 00015184. A. infamis. J Male, ZISP _ ENT 00016250, holotype.	Figure 10. Habitus, dorsal view. A. adustus. A Male, ZISP _ ENT 00016237. B Female, ZISP _ ENT 00015922. A. fraxinicola. C Female, ZISP _ ENT 00016596. D Male, ZISP _ ENT 00015938. A. furvus. E Male, holotype. ZISP _ ENT 00016303. F Female, ZISP _ ENT 00016297. A. gotorum. G Male, ZISP _ ENT 00016232. A. hilaris. H Male, ZISP _ ENT 00015182. I Female, paratype, ZISP _ ENT 00015184. A. infamis. J Male, ZISP _ ENT 00016250, holotype.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
0EC4BF09B9FC5EE9B49D8927164D0AB3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539540	https://doi.org/10.3897/asp.84.e161376.figure15	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
0EC4BF09B9FC5EE9B49D8927164D0AB3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539541	https://doi.org/10.3897/asp.84.e161376.figure16	Figure 16. Vesica. A. adustus, ZISP _ ENT 00014889. A Right lateral view. B Left lateral view. A. sp. nr. furvus (M 199), ZISP _ ENT 00014869. C Right lateral view. D Left lateral view. A. fraxinicola, ZISP _ ENT 00014887. E Right lateral view. F Left lateral view. A. furvus (holotype), ZISP _ ENT 00016303. G Right lateral view. H Left lateral view. A. gotorum, ZISP _ ENT 00016232. I Right lateral view. J Left lateral view. A. furvus (paratype), ZISP _ ENT 00003762. K Right lateral view. L Left lateral view.	Figure 16. Vesica. A. adustus, ZISP _ ENT 00014889. A Right lateral view. B Left lateral view. A. sp. nr. furvus (M 199), ZISP _ ENT 00014869. C Right lateral view. D Left lateral view. A. fraxinicola, ZISP _ ENT 00014887. E Right lateral view. F Left lateral view. A. furvus (holotype), ZISP _ ENT 00016303. G Right lateral view. H Left lateral view. A. gotorum, ZISP _ ENT 00016232. I Right lateral view. J Left lateral view. A. furvus (paratype), ZISP _ ENT 00003762. K Right lateral view. L Left lateral view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
0EC4BF09B9FC5EE9B49D8927164D0AB3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539545	https://doi.org/10.3897/asp.84.e161376.figure19	Figure 19. Right parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola ZISP _ ENT 00014862. C dorsal view. D posterior view. A. furvus, holotype, ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. A. gotorum ZISP _ ENT 00016164. I dorsal view, J posterior view. A. hilaris ZISP _ ENT 00014891. K dorsal view. L posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00015180. M dorsal view. N posterior view. A. infamis ZISP _ ENT 00016250. O dorsal view. P posterior view. A. maackiae, ZISP _ ENT 00014893. Q dorsal view. R posterior view. A. limbatus Europe ZISP _ ENT 00016012. S dorsal view. T posterior view. A. limbatus Asia ZISP _ ENT 00014886. U dorsal view. V posterior view. A. shikotan sp. nov. ZISP _ ENT 00016047, holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris, ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. A. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	Figure 19. Right parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola ZISP _ ENT 00014862. C dorsal view. D posterior view. A. furvus, holotype, ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. A. gotorum ZISP _ ENT 00016164. I dorsal view, J posterior view. A. hilaris ZISP _ ENT 00014891. K dorsal view. L posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00015180. M dorsal view. N posterior view. A. infamis ZISP _ ENT 00016250. O dorsal view. P posterior view. A. maackiae, ZISP _ ENT 00014893. Q dorsal view. R posterior view. A. limbatus Europe ZISP _ ENT 00016012. S dorsal view. T posterior view. A. limbatus Asia ZISP _ ENT 00014886. U dorsal view. V posterior view. A. shikotan sp. nov. ZISP _ ENT 00016047, holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris, ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. A. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
0EC4BF09B9FC5EE9B49D8927164D0AB3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539542	https://doi.org/10.3897/asp.84.e161376.figure17	Figure 17. Vesica. A. spinolae. A Right lateral view, ZISP _ ENT 00006623. B Left lateral view, ZISP _ ENT 00005851. C Right lateral view, ZISP _ ENT 00015780. D Anterior view, ZISP _ ENT 00005851. E Lateral and sublateral sclerites, left lateral view, ZISP _ ENT 00005851. A. subpulchellus, ZISP _ ENT 00016047. F Right lateral view. G Left lateral view. A. sp. nr. subhilaris, (M 197) ZISP _ ENT 00014867. H Right lateral view. I Left lateral view. A. subhilaris, ZISP _ ENT 00016193. J Right lateral view. K Left lateral view. A. watajii, ZISP _ ENT 00016193. L Right lateral view, ZISP _ ENT 00016247. M Left lateral view, ZISP _ ENT 00003760.	Figure 17. Vesica. A. spinolae. A Right lateral view, ZISP _ ENT 00006623. B Left lateral view, ZISP _ ENT 00005851. C Right lateral view, ZISP _ ENT 00015780. D Anterior view, ZISP _ ENT 00005851. E Lateral and sublateral sclerites, left lateral view, ZISP _ ENT 00005851. A. subpulchellus, ZISP _ ENT 00016047. F Right lateral view. G Left lateral view. A. sp. nr. subhilaris, (M 197) ZISP _ ENT 00014867. H Right lateral view. I Left lateral view. A. subhilaris, ZISP _ ENT 00016193. J Right lateral view. K Left lateral view. A. watajii, ZISP _ ENT 00016193. L Right lateral view, ZISP _ ENT 00016247. M Left lateral view, ZISP _ ENT 00003760.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
0EC4BF09B9FC5EE9B49D8927164D0AB3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539544	https://doi.org/10.3897/asp.84.e161376.figure18	Figure 18. Vesica. A. hilaris, ZISP _ ENT 00014891. A Right lateral view. B Left lateral view, A. infamis, ZISP _ ENT 00016252. C Right lateral view. D Left lateral view. A. limbatus, Europe, ZISP _ ENT 00016012. E Right lateral view. F Left lateral view. A. maackiae, ZISP _ ENT 00016247. G Right lateral view. H Left lateral view. A. limbatus, Asia, ZISP _ ENT 00014886. I Right lateral view. J Left lateral view. A. subpulchellus, ZISP _ ENT 00003765. K Right lateral view. L Left lateral view.	Figure 18. Vesica. A. hilaris, ZISP _ ENT 00014891. A Right lateral view. B Left lateral view, A. infamis, ZISP _ ENT 00016252. C Right lateral view. D Left lateral view. A. limbatus, Europe, ZISP _ ENT 00016012. E Right lateral view. F Left lateral view. A. maackiae, ZISP _ ENT 00016247. G Right lateral view. H Left lateral view. A. limbatus, Asia, ZISP _ ENT 00014886. I Right lateral view. J Left lateral view. A. subpulchellus, ZISP _ ENT 00003765. K Right lateral view. L Left lateral view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
F52FEC30FA8756379156B55D0B2E70D4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539534	https://doi.org/10.3897/asp.84.e161376.figure9	Figure 9. Left parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola. ZISP _ ENT 00014862. С dorsal view. D. posterior view. A. furvus, holotype ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. AK genital capsule. A. gotorum ZISP _ ENT 00016164. I posterior view. A. hilaris ZISP _ ENT 00014891. J dorsal view. K posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00014891. L dorsal view. M posterior view. A. infamis ZISP _ ENT 00014891. N dorsal view. O posterior view. A. maackiae ZISP _ ENT 00014893. P dorsal view. Q posterior view. A. limbatus Europe, ZISP _ ENT 00016012. R dorsal view. S posterior view. A. limbatus Asia, ZISP _ ENT 00014886. T dorsal view. U posterior view. V theca. A. shikotan sp. nov. ZISP _ ENT 00016047 holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	Figure 9. Left parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola. ZISP _ ENT 00014862. С dorsal view. D. posterior view. A. furvus, holotype ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. AK genital capsule. A. gotorum ZISP _ ENT 00016164. I posterior view. A. hilaris ZISP _ ENT 00014891. J dorsal view. K posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00014891. L dorsal view. M posterior view. A. infamis ZISP _ ENT 00014891. N dorsal view. O posterior view. A. maackiae ZISP _ ENT 00014893. P dorsal view. Q posterior view. A. limbatus Europe, ZISP _ ENT 00016012. R dorsal view. S posterior view. A. limbatus Asia, ZISP _ ENT 00014886. T dorsal view. U posterior view. V theca. A. shikotan sp. nov. ZISP _ ENT 00016047 holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
F52FEC30FA8756379156B55D0B2E70D4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539535	https://doi.org/10.3897/asp.84.e161376.figure10	Figure 10. Habitus, dorsal view. A. adustus. A Male, ZISP _ ENT 00016237. B Female, ZISP _ ENT 00015922. A. fraxinicola. C Female, ZISP _ ENT 00016596. D Male, ZISP _ ENT 00015938. A. furvus. E Male, holotype. ZISP _ ENT 00016303. F Female, ZISP _ ENT 00016297. A. gotorum. G Male, ZISP _ ENT 00016232. A. hilaris. H Male, ZISP _ ENT 00015182. I Female, paratype, ZISP _ ENT 00015184. A. infamis. J Male, ZISP _ ENT 00016250, holotype.	Figure 10. Habitus, dorsal view. A. adustus. A Male, ZISP _ ENT 00016237. B Female, ZISP _ ENT 00015922. A. fraxinicola. C Female, ZISP _ ENT 00016596. D Male, ZISP _ ENT 00015938. A. furvus. E Male, holotype. ZISP _ ENT 00016303. F Female, ZISP _ ENT 00016297. A. gotorum. G Male, ZISP _ ENT 00016232. A. hilaris. H Male, ZISP _ ENT 00015182. I Female, paratype, ZISP _ ENT 00015184. A. infamis. J Male, ZISP _ ENT 00016250, holotype.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
F52FEC30FA8756379156B55D0B2E70D4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539540	https://doi.org/10.3897/asp.84.e161376.figure15	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
F52FEC30FA8756379156B55D0B2E70D4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539541	https://doi.org/10.3897/asp.84.e161376.figure16	Figure 16. Vesica. A. adustus, ZISP _ ENT 00014889. A Right lateral view. B Left lateral view. A. sp. nr. furvus (M 199), ZISP _ ENT 00014869. C Right lateral view. D Left lateral view. A. fraxinicola, ZISP _ ENT 00014887. E Right lateral view. F Left lateral view. A. furvus (holotype), ZISP _ ENT 00016303. G Right lateral view. H Left lateral view. A. gotorum, ZISP _ ENT 00016232. I Right lateral view. J Left lateral view. A. furvus (paratype), ZISP _ ENT 00003762. K Right lateral view. L Left lateral view.	Figure 16. Vesica. A. adustus, ZISP _ ENT 00014889. A Right lateral view. B Left lateral view. A. sp. nr. furvus (M 199), ZISP _ ENT 00014869. C Right lateral view. D Left lateral view. A. fraxinicola, ZISP _ ENT 00014887. E Right lateral view. F Left lateral view. A. furvus (holotype), ZISP _ ENT 00016303. G Right lateral view. H Left lateral view. A. gotorum, ZISP _ ENT 00016232. I Right lateral view. J Left lateral view. A. furvus (paratype), ZISP _ ENT 00003762. K Right lateral view. L Left lateral view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
F52FEC30FA8756379156B55D0B2E70D4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539545	https://doi.org/10.3897/asp.84.e161376.figure19	Figure 19. Right parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola ZISP _ ENT 00014862. C dorsal view. D posterior view. A. furvus, holotype, ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. A. gotorum ZISP _ ENT 00016164. I dorsal view, J posterior view. A. hilaris ZISP _ ENT 00014891. K dorsal view. L posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00015180. M dorsal view. N posterior view. A. infamis ZISP _ ENT 00016250. O dorsal view. P posterior view. A. maackiae, ZISP _ ENT 00014893. Q dorsal view. R posterior view. A. limbatus Europe ZISP _ ENT 00016012. S dorsal view. T posterior view. A. limbatus Asia ZISP _ ENT 00014886. U dorsal view. V posterior view. A. shikotan sp. nov. ZISP _ ENT 00016047, holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris, ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. A. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	Figure 19. Right parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola ZISP _ ENT 00014862. C dorsal view. D posterior view. A. furvus, holotype, ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. A. gotorum ZISP _ ENT 00016164. I dorsal view, J posterior view. A. hilaris ZISP _ ENT 00014891. K dorsal view. L posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00015180. M dorsal view. N posterior view. A. infamis ZISP _ ENT 00016250. O dorsal view. P posterior view. A. maackiae, ZISP _ ENT 00014893. Q dorsal view. R posterior view. A. limbatus Europe ZISP _ ENT 00016012. S dorsal view. T posterior view. A. limbatus Asia ZISP _ ENT 00014886. U dorsal view. V posterior view. A. shikotan sp. nov. ZISP _ ENT 00016047, holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris, ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. A. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
61F87C94DE9153A5B3347D8DFD525997.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539532	https://doi.org/10.3897/asp.84.e161376.figure7	Figure 7. Female genitalia. A. hilaris ZISP _ ENT 00014894. A Dorsal labiate plate. B Posterior wall. A. shikotan sp. nov. ZISP _ ENT 00014895. C Posterior wall. D Dorsal labiate plate. A. limbatus Europe, ZISP _ ENT 00016026. E Dorsal labiate plate. F Posterior wall. A. limbatus Asia. G Posterior wall, ZISP _ ENT 00015935. H Dorsal labiate plate, ZISP _ ENT 00014890. A. subpulchellus, ZISP _ ENT 00003766. I Dorsal labiate plate. G Posterior wall.	Figure 7. Female genitalia. A. hilaris ZISP _ ENT 00014894. A Dorsal labiate plate. B Posterior wall. A. shikotan sp. nov. ZISP _ ENT 00014895. C Posterior wall. D Dorsal labiate plate. A. limbatus Europe, ZISP _ ENT 00016026. E Dorsal labiate plate. F Posterior wall. A. limbatus Asia. G Posterior wall, ZISP _ ENT 00015935. H Dorsal labiate plate, ZISP _ ENT 00014890. A. subpulchellus, ZISP _ ENT 00003766. I Dorsal labiate plate. G Posterior wall.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
61F87C94DE9153A5B3347D8DFD525997.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539534	https://doi.org/10.3897/asp.84.e161376.figure9	Figure 9. Left parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola. ZISP _ ENT 00014862. С dorsal view. D. posterior view. A. furvus, holotype ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. AK genital capsule. A. gotorum ZISP _ ENT 00016164. I posterior view. A. hilaris ZISP _ ENT 00014891. J dorsal view. K posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00014891. L dorsal view. M posterior view. A. infamis ZISP _ ENT 00014891. N dorsal view. O posterior view. A. maackiae ZISP _ ENT 00014893. P dorsal view. Q posterior view. A. limbatus Europe, ZISP _ ENT 00016012. R dorsal view. S posterior view. A. limbatus Asia, ZISP _ ENT 00014886. T dorsal view. U posterior view. V theca. A. shikotan sp. nov. ZISP _ ENT 00016047 holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	Figure 9. Left parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola. ZISP _ ENT 00014862. С dorsal view. D. posterior view. A. furvus, holotype ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. AK genital capsule. A. gotorum ZISP _ ENT 00016164. I posterior view. A. hilaris ZISP _ ENT 00014891. J dorsal view. K posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00014891. L dorsal view. M posterior view. A. infamis ZISP _ ENT 00014891. N dorsal view. O posterior view. A. maackiae ZISP _ ENT 00014893. P dorsal view. Q posterior view. A. limbatus Europe, ZISP _ ENT 00016012. R dorsal view. S posterior view. A. limbatus Asia, ZISP _ ENT 00014886. T dorsal view. U posterior view. V theca. A. shikotan sp. nov. ZISP _ ENT 00016047 holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
61F87C94DE9153A5B3347D8DFD525997.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539535	https://doi.org/10.3897/asp.84.e161376.figure10	Figure 10. Habitus, dorsal view. A. adustus. A Male, ZISP _ ENT 00016237. B Female, ZISP _ ENT 00015922. A. fraxinicola. C Female, ZISP _ ENT 00016596. D Male, ZISP _ ENT 00015938. A. furvus. E Male, holotype. ZISP _ ENT 00016303. F Female, ZISP _ ENT 00016297. A. gotorum. G Male, ZISP _ ENT 00016232. A. hilaris. H Male, ZISP _ ENT 00015182. I Female, paratype, ZISP _ ENT 00015184. A. infamis. J Male, ZISP _ ENT 00016250, holotype.	Figure 10. Habitus, dorsal view. A. adustus. A Male, ZISP _ ENT 00016237. B Female, ZISP _ ENT 00015922. A. fraxinicola. C Female, ZISP _ ENT 00016596. D Male, ZISP _ ENT 00015938. A. furvus. E Male, holotype. ZISP _ ENT 00016303. F Female, ZISP _ ENT 00016297. A. gotorum. G Male, ZISP _ ENT 00016232. A. hilaris. H Male, ZISP _ ENT 00015182. I Female, paratype, ZISP _ ENT 00015184. A. infamis. J Male, ZISP _ ENT 00016250, holotype.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
61F87C94DE9153A5B3347D8DFD525997.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539540	https://doi.org/10.3897/asp.84.e161376.figure15	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
61F87C94DE9153A5B3347D8DFD525997.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539544	https://doi.org/10.3897/asp.84.e161376.figure18	Figure 18. Vesica. A. hilaris, ZISP _ ENT 00014891. A Right lateral view. B Left lateral view, A. infamis, ZISP _ ENT 00016252. C Right lateral view. D Left lateral view. A. limbatus, Europe, ZISP _ ENT 00016012. E Right lateral view. F Left lateral view. A. maackiae, ZISP _ ENT 00016247. G Right lateral view. H Left lateral view. A. limbatus, Asia, ZISP _ ENT 00014886. I Right lateral view. J Left lateral view. A. subpulchellus, ZISP _ ENT 00003765. K Right lateral view. L Left lateral view.	Figure 18. Vesica. A. hilaris, ZISP _ ENT 00014891. A Right lateral view. B Left lateral view, A. infamis, ZISP _ ENT 00016252. C Right lateral view. D Left lateral view. A. limbatus, Europe, ZISP _ ENT 00016012. E Right lateral view. F Left lateral view. A. maackiae, ZISP _ ENT 00016247. G Right lateral view. H Left lateral view. A. limbatus, Asia, ZISP _ ENT 00014886. I Right lateral view. J Left lateral view. A. subpulchellus, ZISP _ ENT 00003765. K Right lateral view. L Left lateral view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
61F87C94DE9153A5B3347D8DFD525997.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539545	https://doi.org/10.3897/asp.84.e161376.figure19	Figure 19. Right parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola ZISP _ ENT 00014862. C dorsal view. D posterior view. A. furvus, holotype, ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. A. gotorum ZISP _ ENT 00016164. I dorsal view, J posterior view. A. hilaris ZISP _ ENT 00014891. K dorsal view. L posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00015180. M dorsal view. N posterior view. A. infamis ZISP _ ENT 00016250. O dorsal view. P posterior view. A. maackiae, ZISP _ ENT 00014893. Q dorsal view. R posterior view. A. limbatus Europe ZISP _ ENT 00016012. S dorsal view. T posterior view. A. limbatus Asia ZISP _ ENT 00014886. U dorsal view. V posterior view. A. shikotan sp. nov. ZISP _ ENT 00016047, holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris, ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. A. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	Figure 19. Right parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola ZISP _ ENT 00014862. C dorsal view. D posterior view. A. furvus, holotype, ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. A. gotorum ZISP _ ENT 00016164. I dorsal view, J posterior view. A. hilaris ZISP _ ENT 00014891. K dorsal view. L posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00015180. M dorsal view. N posterior view. A. infamis ZISP _ ENT 00016250. O dorsal view. P posterior view. A. maackiae, ZISP _ ENT 00014893. Q dorsal view. R posterior view. A. limbatus Europe ZISP _ ENT 00016012. S dorsal view. T posterior view. A. limbatus Asia ZISP _ ENT 00014886. U dorsal view. V posterior view. A. shikotan sp. nov. ZISP _ ENT 00016047, holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris, ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. A. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
61F87C94DE9153A5B3347D8DFD525997.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539542	https://doi.org/10.3897/asp.84.e161376.figure17	Figure 17. Vesica. A. spinolae. A Right lateral view, ZISP _ ENT 00006623. B Left lateral view, ZISP _ ENT 00005851. C Right lateral view, ZISP _ ENT 00015780. D Anterior view, ZISP _ ENT 00005851. E Lateral and sublateral sclerites, left lateral view, ZISP _ ENT 00005851. A. subpulchellus, ZISP _ ENT 00016047. F Right lateral view. G Left lateral view. A. sp. nr. subhilaris, (M 197) ZISP _ ENT 00014867. H Right lateral view. I Left lateral view. A. subhilaris, ZISP _ ENT 00016193. J Right lateral view. K Left lateral view. A. watajii, ZISP _ ENT 00016193. L Right lateral view, ZISP _ ENT 00016247. M Left lateral view, ZISP _ ENT 00003760.	Figure 17. Vesica. A. spinolae. A Right lateral view, ZISP _ ENT 00006623. B Left lateral view, ZISP _ ENT 00005851. C Right lateral view, ZISP _ ENT 00015780. D Anterior view, ZISP _ ENT 00005851. E Lateral and sublateral sclerites, left lateral view, ZISP _ ENT 00005851. A. subpulchellus, ZISP _ ENT 00016047. F Right lateral view. G Left lateral view. A. sp. nr. subhilaris, (M 197) ZISP _ ENT 00014867. H Right lateral view. I Left lateral view. A. subhilaris, ZISP _ ENT 00016193. J Right lateral view. K Left lateral view. A. watajii, ZISP _ ENT 00016193. L Right lateral view, ZISP _ ENT 00016247. M Left lateral view, ZISP _ ENT 00003760.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
61F87C94DE9153A5B3347D8DFD525997.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539541	https://doi.org/10.3897/asp.84.e161376.figure16	Figure 16. Vesica. A. adustus, ZISP _ ENT 00014889. A Right lateral view. B Left lateral view. A. sp. nr. furvus (M 199), ZISP _ ENT 00014869. C Right lateral view. D Left lateral view. A. fraxinicola, ZISP _ ENT 00014887. E Right lateral view. F Left lateral view. A. furvus (holotype), ZISP _ ENT 00016303. G Right lateral view. H Left lateral view. A. gotorum, ZISP _ ENT 00016232. I Right lateral view. J Left lateral view. A. furvus (paratype), ZISP _ ENT 00003762. K Right lateral view. L Left lateral view.	Figure 16. Vesica. A. adustus, ZISP _ ENT 00014889. A Right lateral view. B Left lateral view. A. sp. nr. furvus (M 199), ZISP _ ENT 00014869. C Right lateral view. D Left lateral view. A. fraxinicola, ZISP _ ENT 00014887. E Right lateral view. F Left lateral view. A. furvus (holotype), ZISP _ ENT 00016303. G Right lateral view. H Left lateral view. A. gotorum, ZISP _ ENT 00016232. I Right lateral view. J Left lateral view. A. furvus (paratype), ZISP _ ENT 00003762. K Right lateral view. L Left lateral view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
3C130D67C7495BEC96A47A030686CB24.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539534	https://doi.org/10.3897/asp.84.e161376.figure9	Figure 9. Left parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola. ZISP _ ENT 00014862. С dorsal view. D. posterior view. A. furvus, holotype ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. AK genital capsule. A. gotorum ZISP _ ENT 00016164. I posterior view. A. hilaris ZISP _ ENT 00014891. J dorsal view. K posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00014891. L dorsal view. M posterior view. A. infamis ZISP _ ENT 00014891. N dorsal view. O posterior view. A. maackiae ZISP _ ENT 00014893. P dorsal view. Q posterior view. A. limbatus Europe, ZISP _ ENT 00016012. R dorsal view. S posterior view. A. limbatus Asia, ZISP _ ENT 00014886. T dorsal view. U posterior view. V theca. A. shikotan sp. nov. ZISP _ ENT 00016047 holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	Figure 9. Left parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola. ZISP _ ENT 00014862. С dorsal view. D. posterior view. A. furvus, holotype ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. AK genital capsule. A. gotorum ZISP _ ENT 00016164. I posterior view. A. hilaris ZISP _ ENT 00014891. J dorsal view. K posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00014891. L dorsal view. M posterior view. A. infamis ZISP _ ENT 00014891. N dorsal view. O posterior view. A. maackiae ZISP _ ENT 00014893. P dorsal view. Q posterior view. A. limbatus Europe, ZISP _ ENT 00016012. R dorsal view. S posterior view. A. limbatus Asia, ZISP _ ENT 00014886. T dorsal view. U posterior view. V theca. A. shikotan sp. nov. ZISP _ ENT 00016047 holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
3C130D67C7495BEC96A47A030686CB24.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539535	https://doi.org/10.3897/asp.84.e161376.figure10	Figure 10. Habitus, dorsal view. A. adustus. A Male, ZISP _ ENT 00016237. B Female, ZISP _ ENT 00015922. A. fraxinicola. C Female, ZISP _ ENT 00016596. D Male, ZISP _ ENT 00015938. A. furvus. E Male, holotype. ZISP _ ENT 00016303. F Female, ZISP _ ENT 00016297. A. gotorum. G Male, ZISP _ ENT 00016232. A. hilaris. H Male, ZISP _ ENT 00015182. I Female, paratype, ZISP _ ENT 00015184. A. infamis. J Male, ZISP _ ENT 00016250, holotype.	Figure 10. Habitus, dorsal view. A. adustus. A Male, ZISP _ ENT 00016237. B Female, ZISP _ ENT 00015922. A. fraxinicola. C Female, ZISP _ ENT 00016596. D Male, ZISP _ ENT 00015938. A. furvus. E Male, holotype. ZISP _ ENT 00016303. F Female, ZISP _ ENT 00016297. A. gotorum. G Male, ZISP _ ENT 00016232. A. hilaris. H Male, ZISP _ ENT 00015182. I Female, paratype, ZISP _ ENT 00015184. A. infamis. J Male, ZISP _ ENT 00016250, holotype.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
3C130D67C7495BEC96A47A030686CB24.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539540	https://doi.org/10.3897/asp.84.e161376.figure15	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
3C130D67C7495BEC96A47A030686CB24.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539544	https://doi.org/10.3897/asp.84.e161376.figure18	Figure 18. Vesica. A. hilaris, ZISP _ ENT 00014891. A Right lateral view. B Left lateral view, A. infamis, ZISP _ ENT 00016252. C Right lateral view. D Left lateral view. A. limbatus, Europe, ZISP _ ENT 00016012. E Right lateral view. F Left lateral view. A. maackiae, ZISP _ ENT 00016247. G Right lateral view. H Left lateral view. A. limbatus, Asia, ZISP _ ENT 00014886. I Right lateral view. J Left lateral view. A. subpulchellus, ZISP _ ENT 00003765. K Right lateral view. L Left lateral view.	Figure 18. Vesica. A. hilaris, ZISP _ ENT 00014891. A Right lateral view. B Left lateral view, A. infamis, ZISP _ ENT 00016252. C Right lateral view. D Left lateral view. A. limbatus, Europe, ZISP _ ENT 00016012. E Right lateral view. F Left lateral view. A. maackiae, ZISP _ ENT 00016247. G Right lateral view. H Left lateral view. A. limbatus, Asia, ZISP _ ENT 00014886. I Right lateral view. J Left lateral view. A. subpulchellus, ZISP _ ENT 00003765. K Right lateral view. L Left lateral view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
3C130D67C7495BEC96A47A030686CB24.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539545	https://doi.org/10.3897/asp.84.e161376.figure19	Figure 19. Right parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola ZISP _ ENT 00014862. C dorsal view. D posterior view. A. furvus, holotype, ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. A. gotorum ZISP _ ENT 00016164. I dorsal view, J posterior view. A. hilaris ZISP _ ENT 00014891. K dorsal view. L posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00015180. M dorsal view. N posterior view. A. infamis ZISP _ ENT 00016250. O dorsal view. P posterior view. A. maackiae, ZISP _ ENT 00014893. Q dorsal view. R posterior view. A. limbatus Europe ZISP _ ENT 00016012. S dorsal view. T posterior view. A. limbatus Asia ZISP _ ENT 00014886. U dorsal view. V posterior view. A. shikotan sp. nov. ZISP _ ENT 00016047, holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris, ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. A. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	Figure 19. Right parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola ZISP _ ENT 00014862. C dorsal view. D posterior view. A. furvus, holotype, ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. A. gotorum ZISP _ ENT 00016164. I dorsal view, J posterior view. A. hilaris ZISP _ ENT 00014891. K dorsal view. L posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00015180. M dorsal view. N posterior view. A. infamis ZISP _ ENT 00016250. O dorsal view. P posterior view. A. maackiae, ZISP _ ENT 00014893. Q dorsal view. R posterior view. A. limbatus Europe ZISP _ ENT 00016012. S dorsal view. T posterior view. A. limbatus Asia ZISP _ ENT 00014886. U dorsal view. V posterior view. A. shikotan sp. nov. ZISP _ ENT 00016047, holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris, ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. A. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
3C130D67C7495BEC96A47A030686CB24.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539541	https://doi.org/10.3897/asp.84.e161376.figure16	Figure 16. Vesica. A. adustus, ZISP _ ENT 00014889. A Right lateral view. B Left lateral view. A. sp. nr. furvus (M 199), ZISP _ ENT 00014869. C Right lateral view. D Left lateral view. A. fraxinicola, ZISP _ ENT 00014887. E Right lateral view. F Left lateral view. A. furvus (holotype), ZISP _ ENT 00016303. G Right lateral view. H Left lateral view. A. gotorum, ZISP _ ENT 00016232. I Right lateral view. J Left lateral view. A. furvus (paratype), ZISP _ ENT 00003762. K Right lateral view. L Left lateral view.	Figure 16. Vesica. A. adustus, ZISP _ ENT 00014889. A Right lateral view. B Left lateral view. A. sp. nr. furvus (M 199), ZISP _ ENT 00014869. C Right lateral view. D Left lateral view. A. fraxinicola, ZISP _ ENT 00014887. E Right lateral view. F Left lateral view. A. furvus (holotype), ZISP _ ENT 00016303. G Right lateral view. H Left lateral view. A. gotorum, ZISP _ ENT 00016232. I Right lateral view. J Left lateral view. A. furvus (paratype), ZISP _ ENT 00003762. K Right lateral view. L Left lateral view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
3C130D67C7495BEC96A47A030686CB24.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539542	https://doi.org/10.3897/asp.84.e161376.figure17	Figure 17. Vesica. A. spinolae. A Right lateral view, ZISP _ ENT 00006623. B Left lateral view, ZISP _ ENT 00005851. C Right lateral view, ZISP _ ENT 00015780. D Anterior view, ZISP _ ENT 00005851. E Lateral and sublateral sclerites, left lateral view, ZISP _ ENT 00005851. A. subpulchellus, ZISP _ ENT 00016047. F Right lateral view. G Left lateral view. A. sp. nr. subhilaris, (M 197) ZISP _ ENT 00014867. H Right lateral view. I Left lateral view. A. subhilaris, ZISP _ ENT 00016193. J Right lateral view. K Left lateral view. A. watajii, ZISP _ ENT 00016193. L Right lateral view, ZISP _ ENT 00016247. M Left lateral view, ZISP _ ENT 00003760.	Figure 17. Vesica. A. spinolae. A Right lateral view, ZISP _ ENT 00006623. B Left lateral view, ZISP _ ENT 00005851. C Right lateral view, ZISP _ ENT 00015780. D Anterior view, ZISP _ ENT 00005851. E Lateral and sublateral sclerites, left lateral view, ZISP _ ENT 00005851. A. subpulchellus, ZISP _ ENT 00016047. F Right lateral view. G Left lateral view. A. sp. nr. subhilaris, (M 197) ZISP _ ENT 00014867. H Right lateral view. I Left lateral view. A. subhilaris, ZISP _ ENT 00016193. J Right lateral view. K Left lateral view. A. watajii, ZISP _ ENT 00016193. L Right lateral view, ZISP _ ENT 00016247. M Left lateral view, ZISP _ ENT 00003760.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
C229EB1FAE605FC89A22E14307C9B375.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539530	https://doi.org/10.3897/asp.84.e161376.figure5	Figure 5. SEM images. A A. subpulchellus, head and pronotum, lateral view, ZISP _ ENT 00003767. B A. limbatus, Europe, Head and pronotum, lateral view, ZISP _ ENT 00016011. C A. limbatus, Europe, cuneus and membrane, ZISP _ ENT 00015959. D A. limbatus Europe, head and pronotum, dorsal view, ZISP _ ENT 00015959. E Pretarsus, dorsal view, A. spinolae, ZISP _ ENT 00015686. F Pleura, A. limbatus, Europe, ZISP _ ENT 00016011. G Pleura, A. maackiae, ZISP _ ENT 00016248. H Pretarsus, ventral view, A. spinolae, ZISP _ ENT 00015187. I Scutellum, clavus, corium, A. adustus, ZISP _ ENT 00015920. J Labium, A. subpulchellus, ZISP _ ENT 00015177. K Hind tarsus, A. maackiae, ZISP _ ENT 00016246.	Figure 5. SEM images. A A. subpulchellus, head and pronotum, lateral view, ZISP _ ENT 00003767. B A. limbatus, Europe, Head and pronotum, lateral view, ZISP _ ENT 00016011. C A. limbatus, Europe, cuneus and membrane, ZISP _ ENT 00015959. D A. limbatus Europe, head and pronotum, dorsal view, ZISP _ ENT 00015959. E Pretarsus, dorsal view, A. spinolae, ZISP _ ENT 00015686. F Pleura, A. limbatus, Europe, ZISP _ ENT 00016011. G Pleura, A. maackiae, ZISP _ ENT 00016248. H Pretarsus, ventral view, A. spinolae, ZISP _ ENT 00015187. I Scutellum, clavus, corium, A. adustus, ZISP _ ENT 00015920. J Labium, A. subpulchellus, ZISP _ ENT 00015177. K Hind tarsus, A. maackiae, ZISP _ ENT 00016246.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
C229EB1FAE605FC89A22E14307C9B375.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539532	https://doi.org/10.3897/asp.84.e161376.figure7	Figure 7. Female genitalia. A. hilaris ZISP _ ENT 00014894. A Dorsal labiate plate. B Posterior wall. A. shikotan sp. nov. ZISP _ ENT 00014895. C Posterior wall. D Dorsal labiate plate. A. limbatus Europe, ZISP _ ENT 00016026. E Dorsal labiate plate. F Posterior wall. A. limbatus Asia. G Posterior wall, ZISP _ ENT 00015935. H Dorsal labiate plate, ZISP _ ENT 00014890. A. subpulchellus, ZISP _ ENT 00003766. I Dorsal labiate plate. G Posterior wall.	Figure 7. Female genitalia. A. hilaris ZISP _ ENT 00014894. A Dorsal labiate plate. B Posterior wall. A. shikotan sp. nov. ZISP _ ENT 00014895. C Posterior wall. D Dorsal labiate plate. A. limbatus Europe, ZISP _ ENT 00016026. E Dorsal labiate plate. F Posterior wall. A. limbatus Asia. G Posterior wall, ZISP _ ENT 00015935. H Dorsal labiate plate, ZISP _ ENT 00014890. A. subpulchellus, ZISP _ ENT 00003766. I Dorsal labiate plate. G Posterior wall.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
C229EB1FAE605FC89A22E14307C9B375.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539534	https://doi.org/10.3897/asp.84.e161376.figure9	Figure 9. Left parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola. ZISP _ ENT 00014862. С dorsal view. D. posterior view. A. furvus, holotype ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. AK genital capsule. A. gotorum ZISP _ ENT 00016164. I posterior view. A. hilaris ZISP _ ENT 00014891. J dorsal view. K posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00014891. L dorsal view. M posterior view. A. infamis ZISP _ ENT 00014891. N dorsal view. O posterior view. A. maackiae ZISP _ ENT 00014893. P dorsal view. Q posterior view. A. limbatus Europe, ZISP _ ENT 00016012. R dorsal view. S posterior view. A. limbatus Asia, ZISP _ ENT 00014886. T dorsal view. U posterior view. V theca. A. shikotan sp. nov. ZISP _ ENT 00016047 holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	Figure 9. Left parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola. ZISP _ ENT 00014862. С dorsal view. D. posterior view. A. furvus, holotype ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. AK genital capsule. A. gotorum ZISP _ ENT 00016164. I posterior view. A. hilaris ZISP _ ENT 00014891. J dorsal view. K posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00014891. L dorsal view. M posterior view. A. infamis ZISP _ ENT 00014891. N dorsal view. O posterior view. A. maackiae ZISP _ ENT 00014893. P dorsal view. Q posterior view. A. limbatus Europe, ZISP _ ENT 00016012. R dorsal view. S posterior view. A. limbatus Asia, ZISP _ ENT 00014886. T dorsal view. U posterior view. V theca. A. shikotan sp. nov. ZISP _ ENT 00016047 holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
C229EB1FAE605FC89A22E14307C9B375.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539536	https://doi.org/10.3897/asp.84.e161376.figure11	Figure 11. Habitus, dorsal view. A. limbatus. A Europe, Female, ZISP _ ENT 00015986. B Asia, Male, ZISP _ ENT 00015917. C Europe, Male, ZISP _ ENT 00016005. D Asia, Female, ZISP _ ENT 00015918. A. maackiae. E Male, ZISP _ ENT 00016046. A. shikotan sp. nov. F Male, paratype, ZISP _ ENT 00016048. A. subhilaris. G Male, ZISP _ ENT 00016193. H Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. I Male, ZISP _ ENT 00016162. J Female, ZISP _ ENT 00014874.	Figure 11. Habitus, dorsal view. A. limbatus. A Europe, Female, ZISP _ ENT 00015986. B Asia, Male, ZISP _ ENT 00015917. C Europe, Male, ZISP _ ENT 00016005. D Asia, Female, ZISP _ ENT 00015918. A. maackiae. E Male, ZISP _ ENT 00016046. A. shikotan sp. nov. F Male, paratype, ZISP _ ENT 00016048. A. subhilaris. G Male, ZISP _ ENT 00016193. H Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. I Male, ZISP _ ENT 00016162. J Female, ZISP _ ENT 00014874.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
C229EB1FAE605FC89A22E14307C9B375.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539540	https://doi.org/10.3897/asp.84.e161376.figure15	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
C229EB1FAE605FC89A22E14307C9B375.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539544	https://doi.org/10.3897/asp.84.e161376.figure18	Figure 18. Vesica. A. hilaris, ZISP _ ENT 00014891. A Right lateral view. B Left lateral view, A. infamis, ZISP _ ENT 00016252. C Right lateral view. D Left lateral view. A. limbatus, Europe, ZISP _ ENT 00016012. E Right lateral view. F Left lateral view. A. maackiae, ZISP _ ENT 00016247. G Right lateral view. H Left lateral view. A. limbatus, Asia, ZISP _ ENT 00014886. I Right lateral view. J Left lateral view. A. subpulchellus, ZISP _ ENT 00003765. K Right lateral view. L Left lateral view.	Figure 18. Vesica. A. hilaris, ZISP _ ENT 00014891. A Right lateral view. B Left lateral view, A. infamis, ZISP _ ENT 00016252. C Right lateral view. D Left lateral view. A. limbatus, Europe, ZISP _ ENT 00016012. E Right lateral view. F Left lateral view. A. maackiae, ZISP _ ENT 00016247. G Right lateral view. H Left lateral view. A. limbatus, Asia, ZISP _ ENT 00014886. I Right lateral view. J Left lateral view. A. subpulchellus, ZISP _ ENT 00003765. K Right lateral view. L Left lateral view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
C229EB1FAE605FC89A22E14307C9B375.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539545	https://doi.org/10.3897/asp.84.e161376.figure19	Figure 19. Right parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola ZISP _ ENT 00014862. C dorsal view. D posterior view. A. furvus, holotype, ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. A. gotorum ZISP _ ENT 00016164. I dorsal view, J posterior view. A. hilaris ZISP _ ENT 00014891. K dorsal view. L posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00015180. M dorsal view. N posterior view. A. infamis ZISP _ ENT 00016250. O dorsal view. P posterior view. A. maackiae, ZISP _ ENT 00014893. Q dorsal view. R posterior view. A. limbatus Europe ZISP _ ENT 00016012. S dorsal view. T posterior view. A. limbatus Asia ZISP _ ENT 00014886. U dorsal view. V posterior view. A. shikotan sp. nov. ZISP _ ENT 00016047, holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris, ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. A. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	Figure 19. Right parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola ZISP _ ENT 00014862. C dorsal view. D posterior view. A. furvus, holotype, ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. A. gotorum ZISP _ ENT 00016164. I dorsal view, J posterior view. A. hilaris ZISP _ ENT 00014891. K dorsal view. L posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00015180. M dorsal view. N posterior view. A. infamis ZISP _ ENT 00016250. O dorsal view. P posterior view. A. maackiae, ZISP _ ENT 00014893. Q dorsal view. R posterior view. A. limbatus Europe ZISP _ ENT 00016012. S dorsal view. T posterior view. A. limbatus Asia ZISP _ ENT 00014886. U dorsal view. V posterior view. A. shikotan sp. nov. ZISP _ ENT 00016047, holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris, ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. A. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
2698AB364AF25C97B609AB22053B9E4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539530	https://doi.org/10.3897/asp.84.e161376.figure5	Figure 5. SEM images. A A. subpulchellus, head and pronotum, lateral view, ZISP _ ENT 00003767. B A. limbatus, Europe, Head and pronotum, lateral view, ZISP _ ENT 00016011. C A. limbatus, Europe, cuneus and membrane, ZISP _ ENT 00015959. D A. limbatus Europe, head and pronotum, dorsal view, ZISP _ ENT 00015959. E Pretarsus, dorsal view, A. spinolae, ZISP _ ENT 00015686. F Pleura, A. limbatus, Europe, ZISP _ ENT 00016011. G Pleura, A. maackiae, ZISP _ ENT 00016248. H Pretarsus, ventral view, A. spinolae, ZISP _ ENT 00015187. I Scutellum, clavus, corium, A. adustus, ZISP _ ENT 00015920. J Labium, A. subpulchellus, ZISP _ ENT 00015177. K Hind tarsus, A. maackiae, ZISP _ ENT 00016246.	Figure 5. SEM images. A A. subpulchellus, head and pronotum, lateral view, ZISP _ ENT 00003767. B A. limbatus, Europe, Head and pronotum, lateral view, ZISP _ ENT 00016011. C A. limbatus, Europe, cuneus and membrane, ZISP _ ENT 00015959. D A. limbatus Europe, head and pronotum, dorsal view, ZISP _ ENT 00015959. E Pretarsus, dorsal view, A. spinolae, ZISP _ ENT 00015686. F Pleura, A. limbatus, Europe, ZISP _ ENT 00016011. G Pleura, A. maackiae, ZISP _ ENT 00016248. H Pretarsus, ventral view, A. spinolae, ZISP _ ENT 00015187. I Scutellum, clavus, corium, A. adustus, ZISP _ ENT 00015920. J Labium, A. subpulchellus, ZISP _ ENT 00015177. K Hind tarsus, A. maackiae, ZISP _ ENT 00016246.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
2698AB364AF25C97B609AB22053B9E4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539534	https://doi.org/10.3897/asp.84.e161376.figure9	Figure 9. Left parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola. ZISP _ ENT 00014862. С dorsal view. D. posterior view. A. furvus, holotype ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. AK genital capsule. A. gotorum ZISP _ ENT 00016164. I posterior view. A. hilaris ZISP _ ENT 00014891. J dorsal view. K posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00014891. L dorsal view. M posterior view. A. infamis ZISP _ ENT 00014891. N dorsal view. O posterior view. A. maackiae ZISP _ ENT 00014893. P dorsal view. Q posterior view. A. limbatus Europe, ZISP _ ENT 00016012. R dorsal view. S posterior view. A. limbatus Asia, ZISP _ ENT 00014886. T dorsal view. U posterior view. V theca. A. shikotan sp. nov. ZISP _ ENT 00016047 holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	Figure 9. Left parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola. ZISP _ ENT 00014862. С dorsal view. D. posterior view. A. furvus, holotype ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. AK genital capsule. A. gotorum ZISP _ ENT 00016164. I posterior view. A. hilaris ZISP _ ENT 00014891. J dorsal view. K posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00014891. L dorsal view. M posterior view. A. infamis ZISP _ ENT 00014891. N dorsal view. O posterior view. A. maackiae ZISP _ ENT 00014893. P dorsal view. Q posterior view. A. limbatus Europe, ZISP _ ENT 00016012. R dorsal view. S posterior view. A. limbatus Asia, ZISP _ ENT 00014886. T dorsal view. U posterior view. V theca. A. shikotan sp. nov. ZISP _ ENT 00016047 holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
2698AB364AF25C97B609AB22053B9E4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539536	https://doi.org/10.3897/asp.84.e161376.figure11	Figure 11. Habitus, dorsal view. A. limbatus. A Europe, Female, ZISP _ ENT 00015986. B Asia, Male, ZISP _ ENT 00015917. C Europe, Male, ZISP _ ENT 00016005. D Asia, Female, ZISP _ ENT 00015918. A. maackiae. E Male, ZISP _ ENT 00016046. A. shikotan sp. nov. F Male, paratype, ZISP _ ENT 00016048. A. subhilaris. G Male, ZISP _ ENT 00016193. H Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. I Male, ZISP _ ENT 00016162. J Female, ZISP _ ENT 00014874.	Figure 11. Habitus, dorsal view. A. limbatus. A Europe, Female, ZISP _ ENT 00015986. B Asia, Male, ZISP _ ENT 00015917. C Europe, Male, ZISP _ ENT 00016005. D Asia, Female, ZISP _ ENT 00015918. A. maackiae. E Male, ZISP _ ENT 00016046. A. shikotan sp. nov. F Male, paratype, ZISP _ ENT 00016048. A. subhilaris. G Male, ZISP _ ENT 00016193. H Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. I Male, ZISP _ ENT 00016162. J Female, ZISP _ ENT 00014874.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
2698AB364AF25C97B609AB22053B9E4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539540	https://doi.org/10.3897/asp.84.e161376.figure15	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
2698AB364AF25C97B609AB22053B9E4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539544	https://doi.org/10.3897/asp.84.e161376.figure18	Figure 18. Vesica. A. hilaris, ZISP _ ENT 00014891. A Right lateral view. B Left lateral view, A. infamis, ZISP _ ENT 00016252. C Right lateral view. D Left lateral view. A. limbatus, Europe, ZISP _ ENT 00016012. E Right lateral view. F Left lateral view. A. maackiae, ZISP _ ENT 00016247. G Right lateral view. H Left lateral view. A. limbatus, Asia, ZISP _ ENT 00014886. I Right lateral view. J Left lateral view. A. subpulchellus, ZISP _ ENT 00003765. K Right lateral view. L Left lateral view.	Figure 18. Vesica. A. hilaris, ZISP _ ENT 00014891. A Right lateral view. B Left lateral view, A. infamis, ZISP _ ENT 00016252. C Right lateral view. D Left lateral view. A. limbatus, Europe, ZISP _ ENT 00016012. E Right lateral view. F Left lateral view. A. maackiae, ZISP _ ENT 00016247. G Right lateral view. H Left lateral view. A. limbatus, Asia, ZISP _ ENT 00014886. I Right lateral view. J Left lateral view. A. subpulchellus, ZISP _ ENT 00003765. K Right lateral view. L Left lateral view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
2698AB364AF25C97B609AB22053B9E4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539545	https://doi.org/10.3897/asp.84.e161376.figure19	Figure 19. Right parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola ZISP _ ENT 00014862. C dorsal view. D posterior view. A. furvus, holotype, ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. A. gotorum ZISP _ ENT 00016164. I dorsal view, J posterior view. A. hilaris ZISP _ ENT 00014891. K dorsal view. L posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00015180. M dorsal view. N posterior view. A. infamis ZISP _ ENT 00016250. O dorsal view. P posterior view. A. maackiae, ZISP _ ENT 00014893. Q dorsal view. R posterior view. A. limbatus Europe ZISP _ ENT 00016012. S dorsal view. T posterior view. A. limbatus Asia ZISP _ ENT 00014886. U dorsal view. V posterior view. A. shikotan sp. nov. ZISP _ ENT 00016047, holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris, ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. A. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	Figure 19. Right parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola ZISP _ ENT 00014862. C dorsal view. D posterior view. A. furvus, holotype, ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. A. gotorum ZISP _ ENT 00016164. I dorsal view, J posterior view. A. hilaris ZISP _ ENT 00014891. K dorsal view. L posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00015180. M dorsal view. N posterior view. A. infamis ZISP _ ENT 00016250. O dorsal view. P posterior view. A. maackiae, ZISP _ ENT 00014893. Q dorsal view. R posterior view. A. limbatus Europe ZISP _ ENT 00016012. S dorsal view. T posterior view. A. limbatus Asia ZISP _ ENT 00014886. U dorsal view. V posterior view. A. shikotan sp. nov. ZISP _ ENT 00016047, holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris, ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. A. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
565AF38C49955936BA9E3BFD163D352E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539532	https://doi.org/10.3897/asp.84.e161376.figure7	Figure 7. Female genitalia. A. hilaris ZISP _ ENT 00014894. A Dorsal labiate plate. B Posterior wall. A. shikotan sp. nov. ZISP _ ENT 00014895. C Posterior wall. D Dorsal labiate plate. A. limbatus Europe, ZISP _ ENT 00016026. E Dorsal labiate plate. F Posterior wall. A. limbatus Asia. G Posterior wall, ZISP _ ENT 00015935. H Dorsal labiate plate, ZISP _ ENT 00014890. A. subpulchellus, ZISP _ ENT 00003766. I Dorsal labiate plate. G Posterior wall.	Figure 7. Female genitalia. A. hilaris ZISP _ ENT 00014894. A Dorsal labiate plate. B Posterior wall. A. shikotan sp. nov. ZISP _ ENT 00014895. C Posterior wall. D Dorsal labiate plate. A. limbatus Europe, ZISP _ ENT 00016026. E Dorsal labiate plate. F Posterior wall. A. limbatus Asia. G Posterior wall, ZISP _ ENT 00015935. H Dorsal labiate plate, ZISP _ ENT 00014890. A. subpulchellus, ZISP _ ENT 00003766. I Dorsal labiate plate. G Posterior wall.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
565AF38C49955936BA9E3BFD163D352E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539534	https://doi.org/10.3897/asp.84.e161376.figure9	Figure 9. Left parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola. ZISP _ ENT 00014862. С dorsal view. D. posterior view. A. furvus, holotype ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. AK genital capsule. A. gotorum ZISP _ ENT 00016164. I posterior view. A. hilaris ZISP _ ENT 00014891. J dorsal view. K posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00014891. L dorsal view. M posterior view. A. infamis ZISP _ ENT 00014891. N dorsal view. O posterior view. A. maackiae ZISP _ ENT 00014893. P dorsal view. Q posterior view. A. limbatus Europe, ZISP _ ENT 00016012. R dorsal view. S posterior view. A. limbatus Asia, ZISP _ ENT 00014886. T dorsal view. U posterior view. V theca. A. shikotan sp. nov. ZISP _ ENT 00016047 holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	Figure 9. Left parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola. ZISP _ ENT 00014862. С dorsal view. D. posterior view. A. furvus, holotype ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. AK genital capsule. A. gotorum ZISP _ ENT 00016164. I posterior view. A. hilaris ZISP _ ENT 00014891. J dorsal view. K posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00014891. L dorsal view. M posterior view. A. infamis ZISP _ ENT 00014891. N dorsal view. O posterior view. A. maackiae ZISP _ ENT 00014893. P dorsal view. Q posterior view. A. limbatus Europe, ZISP _ ENT 00016012. R dorsal view. S posterior view. A. limbatus Asia, ZISP _ ENT 00014886. T dorsal view. U posterior view. V theca. A. shikotan sp. nov. ZISP _ ENT 00016047 holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
565AF38C49955936BA9E3BFD163D352E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539536	https://doi.org/10.3897/asp.84.e161376.figure11	Figure 11. Habitus, dorsal view. A. limbatus. A Europe, Female, ZISP _ ENT 00015986. B Asia, Male, ZISP _ ENT 00015917. C Europe, Male, ZISP _ ENT 00016005. D Asia, Female, ZISP _ ENT 00015918. A. maackiae. E Male, ZISP _ ENT 00016046. A. shikotan sp. nov. F Male, paratype, ZISP _ ENT 00016048. A. subhilaris. G Male, ZISP _ ENT 00016193. H Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. I Male, ZISP _ ENT 00016162. J Female, ZISP _ ENT 00014874.	Figure 11. Habitus, dorsal view. A. limbatus. A Europe, Female, ZISP _ ENT 00015986. B Asia, Male, ZISP _ ENT 00015917. C Europe, Male, ZISP _ ENT 00016005. D Asia, Female, ZISP _ ENT 00015918. A. maackiae. E Male, ZISP _ ENT 00016046. A. shikotan sp. nov. F Male, paratype, ZISP _ ENT 00016048. A. subhilaris. G Male, ZISP _ ENT 00016193. H Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. I Male, ZISP _ ENT 00016162. J Female, ZISP _ ENT 00014874.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
565AF38C49955936BA9E3BFD163D352E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539540	https://doi.org/10.3897/asp.84.e161376.figure15	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
565AF38C49955936BA9E3BFD163D352E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539542	https://doi.org/10.3897/asp.84.e161376.figure17	Figure 17. Vesica. A. spinolae. A Right lateral view, ZISP _ ENT 00006623. B Left lateral view, ZISP _ ENT 00005851. C Right lateral view, ZISP _ ENT 00015780. D Anterior view, ZISP _ ENT 00005851. E Lateral and sublateral sclerites, left lateral view, ZISP _ ENT 00005851. A. subpulchellus, ZISP _ ENT 00016047. F Right lateral view. G Left lateral view. A. sp. nr. subhilaris, (M 197) ZISP _ ENT 00014867. H Right lateral view. I Left lateral view. A. subhilaris, ZISP _ ENT 00016193. J Right lateral view. K Left lateral view. A. watajii, ZISP _ ENT 00016193. L Right lateral view, ZISP _ ENT 00016247. M Left lateral view, ZISP _ ENT 00003760.	Figure 17. Vesica. A. spinolae. A Right lateral view, ZISP _ ENT 00006623. B Left lateral view, ZISP _ ENT 00005851. C Right lateral view, ZISP _ ENT 00015780. D Anterior view, ZISP _ ENT 00005851. E Lateral and sublateral sclerites, left lateral view, ZISP _ ENT 00005851. A. subpulchellus, ZISP _ ENT 00016047. F Right lateral view. G Left lateral view. A. sp. nr. subhilaris, (M 197) ZISP _ ENT 00014867. H Right lateral view. I Left lateral view. A. subhilaris, ZISP _ ENT 00016193. J Right lateral view. K Left lateral view. A. watajii, ZISP _ ENT 00016193. L Right lateral view, ZISP _ ENT 00016247. M Left lateral view, ZISP _ ENT 00003760.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
565AF38C49955936BA9E3BFD163D352E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539545	https://doi.org/10.3897/asp.84.e161376.figure19	Figure 19. Right parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola ZISP _ ENT 00014862. C dorsal view. D posterior view. A. furvus, holotype, ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. A. gotorum ZISP _ ENT 00016164. I dorsal view, J posterior view. A. hilaris ZISP _ ENT 00014891. K dorsal view. L posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00015180. M dorsal view. N posterior view. A. infamis ZISP _ ENT 00016250. O dorsal view. P posterior view. A. maackiae, ZISP _ ENT 00014893. Q dorsal view. R posterior view. A. limbatus Europe ZISP _ ENT 00016012. S dorsal view. T posterior view. A. limbatus Asia ZISP _ ENT 00014886. U dorsal view. V posterior view. A. shikotan sp. nov. ZISP _ ENT 00016047, holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris, ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. A. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	Figure 19. Right parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola ZISP _ ENT 00014862. C dorsal view. D posterior view. A. furvus, holotype, ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. A. gotorum ZISP _ ENT 00016164. I dorsal view, J posterior view. A. hilaris ZISP _ ENT 00014891. K dorsal view. L posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00015180. M dorsal view. N posterior view. A. infamis ZISP _ ENT 00016250. O dorsal view. P posterior view. A. maackiae, ZISP _ ENT 00014893. Q dorsal view. R posterior view. A. limbatus Europe ZISP _ ENT 00016012. S dorsal view. T posterior view. A. limbatus Asia ZISP _ ENT 00014886. U dorsal view. V posterior view. A. shikotan sp. nov. ZISP _ ENT 00016047, holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris, ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. A. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
565AF38C49955936BA9E3BFD163D352E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539541	https://doi.org/10.3897/asp.84.e161376.figure16	Figure 16. Vesica. A. adustus, ZISP _ ENT 00014889. A Right lateral view. B Left lateral view. A. sp. nr. furvus (M 199), ZISP _ ENT 00014869. C Right lateral view. D Left lateral view. A. fraxinicola, ZISP _ ENT 00014887. E Right lateral view. F Left lateral view. A. furvus (holotype), ZISP _ ENT 00016303. G Right lateral view. H Left lateral view. A. gotorum, ZISP _ ENT 00016232. I Right lateral view. J Left lateral view. A. furvus (paratype), ZISP _ ENT 00003762. K Right lateral view. L Left lateral view.	Figure 16. Vesica. A. adustus, ZISP _ ENT 00014889. A Right lateral view. B Left lateral view. A. sp. nr. furvus (M 199), ZISP _ ENT 00014869. C Right lateral view. D Left lateral view. A. fraxinicola, ZISP _ ENT 00014887. E Right lateral view. F Left lateral view. A. furvus (holotype), ZISP _ ENT 00016303. G Right lateral view. H Left lateral view. A. gotorum, ZISP _ ENT 00016232. I Right lateral view. J Left lateral view. A. furvus (paratype), ZISP _ ENT 00003762. K Right lateral view. L Left lateral view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
A47CF3C0B1E35703ABC5097668878ECE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539530	https://doi.org/10.3897/asp.84.e161376.figure5	Figure 5. SEM images. A A. subpulchellus, head and pronotum, lateral view, ZISP _ ENT 00003767. B A. limbatus, Europe, Head and pronotum, lateral view, ZISP _ ENT 00016011. C A. limbatus, Europe, cuneus and membrane, ZISP _ ENT 00015959. D A. limbatus Europe, head and pronotum, dorsal view, ZISP _ ENT 00015959. E Pretarsus, dorsal view, A. spinolae, ZISP _ ENT 00015686. F Pleura, A. limbatus, Europe, ZISP _ ENT 00016011. G Pleura, A. maackiae, ZISP _ ENT 00016248. H Pretarsus, ventral view, A. spinolae, ZISP _ ENT 00015187. I Scutellum, clavus, corium, A. adustus, ZISP _ ENT 00015920. J Labium, A. subpulchellus, ZISP _ ENT 00015177. K Hind tarsus, A. maackiae, ZISP _ ENT 00016246.	Figure 5. SEM images. A A. subpulchellus, head and pronotum, lateral view, ZISP _ ENT 00003767. B A. limbatus, Europe, Head and pronotum, lateral view, ZISP _ ENT 00016011. C A. limbatus, Europe, cuneus and membrane, ZISP _ ENT 00015959. D A. limbatus Europe, head and pronotum, dorsal view, ZISP _ ENT 00015959. E Pretarsus, dorsal view, A. spinolae, ZISP _ ENT 00015686. F Pleura, A. limbatus, Europe, ZISP _ ENT 00016011. G Pleura, A. maackiae, ZISP _ ENT 00016248. H Pretarsus, ventral view, A. spinolae, ZISP _ ENT 00015187. I Scutellum, clavus, corium, A. adustus, ZISP _ ENT 00015920. J Labium, A. subpulchellus, ZISP _ ENT 00015177. K Hind tarsus, A. maackiae, ZISP _ ENT 00016246.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
A47CF3C0B1E35703ABC5097668878ECE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539533	https://doi.org/10.3897/asp.84.e161376.figure8	Figure 8. Female genitalia. A. spinolae. A Dorsal labiate plate, ZISP _ ENT 00014490. B Posterior wall, ZISP _ ENT 00015712. A. subhilaris ZISP _ ENT 000116157. C Posterior wall. D Dorsal labiate plate. A. sp. nr. subhilaris, ZISP _ ENT 00014784. E Dorsal labiate plate. F Posterior wall. A. watajii, ZISP _ ENT 0006039. G Dorsal labiate plate. H Posterior wall. I Ventral wall.	Figure 8. Female genitalia. A. spinolae. A Dorsal labiate plate, ZISP _ ENT 00014490. B Posterior wall, ZISP _ ENT 00015712. A. subhilaris ZISP _ ENT 000116157. C Posterior wall. D Dorsal labiate plate. A. sp. nr. subhilaris, ZISP _ ENT 00014784. E Dorsal labiate plate. F Posterior wall. A. watajii, ZISP _ ENT 0006039. G Dorsal labiate plate. H Posterior wall. I Ventral wall.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
A47CF3C0B1E35703ABC5097668878ECE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539534	https://doi.org/10.3897/asp.84.e161376.figure9	Figure 9. Left parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola. ZISP _ ENT 00014862. С dorsal view. D. posterior view. A. furvus, holotype ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. AK genital capsule. A. gotorum ZISP _ ENT 00016164. I posterior view. A. hilaris ZISP _ ENT 00014891. J dorsal view. K posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00014891. L dorsal view. M posterior view. A. infamis ZISP _ ENT 00014891. N dorsal view. O posterior view. A. maackiae ZISP _ ENT 00014893. P dorsal view. Q posterior view. A. limbatus Europe, ZISP _ ENT 00016012. R dorsal view. S posterior view. A. limbatus Asia, ZISP _ ENT 00014886. T dorsal view. U posterior view. V theca. A. shikotan sp. nov. ZISP _ ENT 00016047 holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	Figure 9. Left parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola. ZISP _ ENT 00014862. С dorsal view. D. posterior view. A. furvus, holotype ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. AK genital capsule. A. gotorum ZISP _ ENT 00016164. I posterior view. A. hilaris ZISP _ ENT 00014891. J dorsal view. K posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00014891. L dorsal view. M posterior view. A. infamis ZISP _ ENT 00014891. N dorsal view. O posterior view. A. maackiae ZISP _ ENT 00014893. P dorsal view. Q posterior view. A. limbatus Europe, ZISP _ ENT 00016012. R dorsal view. S posterior view. A. limbatus Asia, ZISP _ ENT 00014886. T dorsal view. U posterior view. V theca. A. shikotan sp. nov. ZISP _ ENT 00016047 holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
A47CF3C0B1E35703ABC5097668878ECE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539537	https://doi.org/10.3897/asp.84.e161376.figure12	Figure 12. Habitus, dorsal view. A. spinolae. Specimens initially identified as A. nigronasutus. A Male, ZISP _ ENT 00014451. B Female, ZISP _ ENT 00014477. Specimens initially identified as A. nigrovirens. C Male, ZISP _ ENT 00014590. D Male, ZISP _ ENT 00014604. E Female, ZISP _ ENT 00014618. F Female, ZISP _ ENT 00014738. Specimens initially identified as A. spinolae. G Male, ZISP _ ENT 00014775. H Female, ZISP _ ENT 00014766. Specimens initially identified as A. malaisei. I Female, ZISP _ ENT 00015747. J Male, ZISP _ ENT 00015764. K Male, ZISP _ ENT 00015656. Specimens initially identified as A. lucorum. L Male, ZISP _ ENT 00006565. M Female, ZISP _ ENT 00006918.	Figure 12. Habitus, dorsal view. A. spinolae. Specimens initially identified as A. nigronasutus. A Male, ZISP _ ENT 00014451. B Female, ZISP _ ENT 00014477. Specimens initially identified as A. nigrovirens. C Male, ZISP _ ENT 00014590. D Male, ZISP _ ENT 00014604. E Female, ZISP _ ENT 00014618. F Female, ZISP _ ENT 00014738. Specimens initially identified as A. spinolae. G Male, ZISP _ ENT 00014775. H Female, ZISP _ ENT 00014766. Specimens initially identified as A. malaisei. I Female, ZISP _ ENT 00015747. J Male, ZISP _ ENT 00015764. K Male, ZISP _ ENT 00015656. Specimens initially identified as A. lucorum. L Male, ZISP _ ENT 00006565. M Female, ZISP _ ENT 00006918.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
A47CF3C0B1E35703ABC5097668878ECE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539539	https://doi.org/10.3897/asp.84.e161376.figure14	Figure 14. The digital images of paralectotypes preserved at the American Museum of Natural History, New York. Capsus lucorum Meyer-Dür, 1843 (AMNH _ IZC 00396564), female. A Dorsal view. B Frontal view. C Labels. D Left lateral view. Capsus spinolae Meyer-Dür, 1841 (AMNH _ IZC 00396565), female. E Dorsal view. F Frontal view. G Labels. H Left lateral view.	Figure 14. The digital images of paralectotypes preserved at the American Museum of Natural History, New York. Capsus lucorum Meyer-Dür, 1843 (AMNH _ IZC 00396564), female. A Dorsal view. B Frontal view. C Labels. D Left lateral view. Capsus spinolae Meyer-Dür, 1841 (AMNH _ IZC 00396565), female. E Dorsal view. F Frontal view. G Labels. H Left lateral view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
A47CF3C0B1E35703ABC5097668878ECE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539540	https://doi.org/10.3897/asp.84.e161376.figure15	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
A47CF3C0B1E35703ABC5097668878ECE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539542	https://doi.org/10.3897/asp.84.e161376.figure17	Figure 17. Vesica. A. spinolae. A Right lateral view, ZISP _ ENT 00006623. B Left lateral view, ZISP _ ENT 00005851. C Right lateral view, ZISP _ ENT 00015780. D Anterior view, ZISP _ ENT 00005851. E Lateral and sublateral sclerites, left lateral view, ZISP _ ENT 00005851. A. subpulchellus, ZISP _ ENT 00016047. F Right lateral view. G Left lateral view. A. sp. nr. subhilaris, (M 197) ZISP _ ENT 00014867. H Right lateral view. I Left lateral view. A. subhilaris, ZISP _ ENT 00016193. J Right lateral view. K Left lateral view. A. watajii, ZISP _ ENT 00016193. L Right lateral view, ZISP _ ENT 00016247. M Left lateral view, ZISP _ ENT 00003760.	Figure 17. Vesica. A. spinolae. A Right lateral view, ZISP _ ENT 00006623. B Left lateral view, ZISP _ ENT 00005851. C Right lateral view, ZISP _ ENT 00015780. D Anterior view, ZISP _ ENT 00005851. E Lateral and sublateral sclerites, left lateral view, ZISP _ ENT 00005851. A. subpulchellus, ZISP _ ENT 00016047. F Right lateral view. G Left lateral view. A. sp. nr. subhilaris, (M 197) ZISP _ ENT 00014867. H Right lateral view. I Left lateral view. A. subhilaris, ZISP _ ENT 00016193. J Right lateral view. K Left lateral view. A. watajii, ZISP _ ENT 00016193. L Right lateral view, ZISP _ ENT 00016247. M Left lateral view, ZISP _ ENT 00003760.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
A47CF3C0B1E35703ABC5097668878ECE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539545	https://doi.org/10.3897/asp.84.e161376.figure19	Figure 19. Right parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola ZISP _ ENT 00014862. C dorsal view. D posterior view. A. furvus, holotype, ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. A. gotorum ZISP _ ENT 00016164. I dorsal view, J posterior view. A. hilaris ZISP _ ENT 00014891. K dorsal view. L posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00015180. M dorsal view. N posterior view. A. infamis ZISP _ ENT 00016250. O dorsal view. P posterior view. A. maackiae, ZISP _ ENT 00014893. Q dorsal view. R posterior view. A. limbatus Europe ZISP _ ENT 00016012. S dorsal view. T posterior view. A. limbatus Asia ZISP _ ENT 00014886. U dorsal view. V posterior view. A. shikotan sp. nov. ZISP _ ENT 00016047, holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris, ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. A. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	Figure 19. Right parameres. A. adustus ZISP _ ENT 00014862. A dorsal view. B posterior view. A. fraxinicola ZISP _ ENT 00014862. C dorsal view. D posterior view. A. furvus, holotype, ZISP _ ENT 00016303. E dorsal view. F posterior view. A. furvus, paratype ZISP _ ENT 00015924. G dorsal view. H posterior view. A. gotorum ZISP _ ENT 00016164. I dorsal view, J posterior view. A. hilaris ZISP _ ENT 00014891. K dorsal view. L posterior view. A. hilaris, holotype of A. syringae ZISP _ ENT 00015180. M dorsal view. N posterior view. A. infamis ZISP _ ENT 00016250. O dorsal view. P posterior view. A. maackiae, ZISP _ ENT 00014893. Q dorsal view. R posterior view. A. limbatus Europe ZISP _ ENT 00016012. S dorsal view. T posterior view. A. limbatus Asia ZISP _ ENT 00014886. U dorsal view. V posterior view. A. shikotan sp. nov. ZISP _ ENT 00016047, holotype. W dorsal view. X posterior view. A. spinolae ZISP _ ENT 00005858. Y dorsal view. Z posterior view. A. subhilaris, ZISP _ ENT 00016582. AA dorsal view. AB posterior view. A. sp. nr. A. subhilaris ZISP _ ENT 00014867. AC dorsal view. AD posterior view. A. subpulchellus ZISP _ ENT 00003765. AE dorsal view. AF posterior view. A. watajii ZISP _ ENT 00003760. AG dorsal view. AH posterior view. A. sp. nr. furvus ZISP _ ENT 00014869. AI dorsal view. AJ posterior view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
3ADC97F7451359C69A5D2253926DC68E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539533	https://doi.org/10.3897/asp.84.e161376.figure8	Figure 8. Female genitalia. A. spinolae. A Dorsal labiate plate, ZISP _ ENT 00014490. B Posterior wall, ZISP _ ENT 00015712. A. subhilaris ZISP _ ENT 000116157. C Posterior wall. D Dorsal labiate plate. A. sp. nr. subhilaris, ZISP _ ENT 00014784. E Dorsal labiate plate. F Posterior wall. A. watajii, ZISP _ ENT 0006039. G Dorsal labiate plate. H Posterior wall. I Ventral wall.	Figure 8. Female genitalia. A. spinolae. A Dorsal labiate plate, ZISP _ ENT 00014490. B Posterior wall, ZISP _ ENT 00015712. A. subhilaris ZISP _ ENT 000116157. C Posterior wall. D Dorsal labiate plate. A. sp. nr. subhilaris, ZISP _ ENT 00014784. E Dorsal labiate plate. F Posterior wall. A. watajii, ZISP _ ENT 0006039. G Dorsal labiate plate. H Posterior wall. I Ventral wall.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
3ADC97F7451359C69A5D2253926DC68E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539536	https://doi.org/10.3897/asp.84.e161376.figure11	Figure 11. Habitus, dorsal view. A. limbatus. A Europe, Female, ZISP _ ENT 00015986. B Asia, Male, ZISP _ ENT 00015917. C Europe, Male, ZISP _ ENT 00016005. D Asia, Female, ZISP _ ENT 00015918. A. maackiae. E Male, ZISP _ ENT 00016046. A. shikotan sp. nov. F Male, paratype, ZISP _ ENT 00016048. A. subhilaris. G Male, ZISP _ ENT 00016193. H Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. I Male, ZISP _ ENT 00016162. J Female, ZISP _ ENT 00014874.	Figure 11. Habitus, dorsal view. A. limbatus. A Europe, Female, ZISP _ ENT 00015986. B Asia, Male, ZISP _ ENT 00015917. C Europe, Male, ZISP _ ENT 00016005. D Asia, Female, ZISP _ ENT 00015918. A. maackiae. E Male, ZISP _ ENT 00016046. A. shikotan sp. nov. F Male, paratype, ZISP _ ENT 00016048. A. subhilaris. G Male, ZISP _ ENT 00016193. H Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. I Male, ZISP _ ENT 00016162. J Female, ZISP _ ENT 00014874.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
3ADC97F7451359C69A5D2253926DC68E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539540	https://doi.org/10.3897/asp.84.e161376.figure15	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
3ADC97F7451359C69A5D2253926DC68E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539542	https://doi.org/10.3897/asp.84.e161376.figure17	Figure 17. Vesica. A. spinolae. A Right lateral view, ZISP _ ENT 00006623. B Left lateral view, ZISP _ ENT 00005851. C Right lateral view, ZISP _ ENT 00015780. D Anterior view, ZISP _ ENT 00005851. E Lateral and sublateral sclerites, left lateral view, ZISP _ ENT 00005851. A. subpulchellus, ZISP _ ENT 00016047. F Right lateral view. G Left lateral view. A. sp. nr. subhilaris, (M 197) ZISP _ ENT 00014867. H Right lateral view. I Left lateral view. A. subhilaris, ZISP _ ENT 00016193. J Right lateral view. K Left lateral view. A. watajii, ZISP _ ENT 00016193. L Right lateral view, ZISP _ ENT 00016247. M Left lateral view, ZISP _ ENT 00003760.	Figure 17. Vesica. A. spinolae. A Right lateral view, ZISP _ ENT 00006623. B Left lateral view, ZISP _ ENT 00005851. C Right lateral view, ZISP _ ENT 00015780. D Anterior view, ZISP _ ENT 00005851. E Lateral and sublateral sclerites, left lateral view, ZISP _ ENT 00005851. A. subpulchellus, ZISP _ ENT 00016047. F Right lateral view. G Left lateral view. A. sp. nr. subhilaris, (M 197) ZISP _ ENT 00014867. H Right lateral view. I Left lateral view. A. subhilaris, ZISP _ ENT 00016193. J Right lateral view. K Left lateral view. A. watajii, ZISP _ ENT 00016193. L Right lateral view, ZISP _ ENT 00016247. M Left lateral view, ZISP _ ENT 00003760.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
3ADC97F7451359C69A5D2253926DC68E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539544	https://doi.org/10.3897/asp.84.e161376.figure18	Figure 18. Vesica. A. hilaris, ZISP _ ENT 00014891. A Right lateral view. B Left lateral view, A. infamis, ZISP _ ENT 00016252. C Right lateral view. D Left lateral view. A. limbatus, Europe, ZISP _ ENT 00016012. E Right lateral view. F Left lateral view. A. maackiae, ZISP _ ENT 00016247. G Right lateral view. H Left lateral view. A. limbatus, Asia, ZISP _ ENT 00014886. I Right lateral view. J Left lateral view. A. subpulchellus, ZISP _ ENT 00003765. K Right lateral view. L Left lateral view.	Figure 18. Vesica. A. hilaris, ZISP _ ENT 00014891. A Right lateral view. B Left lateral view, A. infamis, ZISP _ ENT 00016252. C Right lateral view. D Left lateral view. A. limbatus, Europe, ZISP _ ENT 00016012. E Right lateral view. F Left lateral view. A. maackiae, ZISP _ ENT 00016247. G Right lateral view. H Left lateral view. A. limbatus, Asia, ZISP _ ENT 00014886. I Right lateral view. J Left lateral view. A. subpulchellus, ZISP _ ENT 00003765. K Right lateral view. L Left lateral view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
5F64AA50314050FB9D98D7148A80D155.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539530	https://doi.org/10.3897/asp.84.e161376.figure5	Figure 5. SEM images. A A. subpulchellus, head and pronotum, lateral view, ZISP _ ENT 00003767. B A. limbatus, Europe, Head and pronotum, lateral view, ZISP _ ENT 00016011. C A. limbatus, Europe, cuneus and membrane, ZISP _ ENT 00015959. D A. limbatus Europe, head and pronotum, dorsal view, ZISP _ ENT 00015959. E Pretarsus, dorsal view, A. spinolae, ZISP _ ENT 00015686. F Pleura, A. limbatus, Europe, ZISP _ ENT 00016011. G Pleura, A. maackiae, ZISP _ ENT 00016248. H Pretarsus, ventral view, A. spinolae, ZISP _ ENT 00015187. I Scutellum, clavus, corium, A. adustus, ZISP _ ENT 00015920. J Labium, A. subpulchellus, ZISP _ ENT 00015177. K Hind tarsus, A. maackiae, ZISP _ ENT 00016246.	Figure 5. SEM images. A A. subpulchellus, head and pronotum, lateral view, ZISP _ ENT 00003767. B A. limbatus, Europe, Head and pronotum, lateral view, ZISP _ ENT 00016011. C A. limbatus, Europe, cuneus and membrane, ZISP _ ENT 00015959. D A. limbatus Europe, head and pronotum, dorsal view, ZISP _ ENT 00015959. E Pretarsus, dorsal view, A. spinolae, ZISP _ ENT 00015686. F Pleura, A. limbatus, Europe, ZISP _ ENT 00016011. G Pleura, A. maackiae, ZISP _ ENT 00016248. H Pretarsus, ventral view, A. spinolae, ZISP _ ENT 00015187. I Scutellum, clavus, corium, A. adustus, ZISP _ ENT 00015920. J Labium, A. subpulchellus, ZISP _ ENT 00015177. K Hind tarsus, A. maackiae, ZISP _ ENT 00016246.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
5F64AA50314050FB9D98D7148A80D155.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539532	https://doi.org/10.3897/asp.84.e161376.figure7	Figure 7. Female genitalia. A. hilaris ZISP _ ENT 00014894. A Dorsal labiate plate. B Posterior wall. A. shikotan sp. nov. ZISP _ ENT 00014895. C Posterior wall. D Dorsal labiate plate. A. limbatus Europe, ZISP _ ENT 00016026. E Dorsal labiate plate. F Posterior wall. A. limbatus Asia. G Posterior wall, ZISP _ ENT 00015935. H Dorsal labiate plate, ZISP _ ENT 00014890. A. subpulchellus, ZISP _ ENT 00003766. I Dorsal labiate plate. G Posterior wall.	Figure 7. Female genitalia. A. hilaris ZISP _ ENT 00014894. A Dorsal labiate plate. B Posterior wall. A. shikotan sp. nov. ZISP _ ENT 00014895. C Posterior wall. D Dorsal labiate plate. A. limbatus Europe, ZISP _ ENT 00016026. E Dorsal labiate plate. F Posterior wall. A. limbatus Asia. G Posterior wall, ZISP _ ENT 00015935. H Dorsal labiate plate, ZISP _ ENT 00014890. A. subpulchellus, ZISP _ ENT 00003766. I Dorsal labiate plate. G Posterior wall.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
5F64AA50314050FB9D98D7148A80D155.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539538	https://doi.org/10.3897/asp.84.e161376.figure13	Figure 13. Habitus, dorsal view. A. subpulchellus. A Male, ZISP _ ENT 00016042, paralectotype, B Female, ZISP _ ENT 00016044, paralectotypeю C Female, ZISP _ ENT 00016045, paralectotype. A. watajii. D Male, paratype, ZISP _ ENT 00016241. E Female, ZISP _ ENT 00016243. F Female, ZISP _ ENT 00016244.	Figure 13. Habitus, dorsal view. A. subpulchellus. A Male, ZISP _ ENT 00016042, paralectotype, B Female, ZISP _ ENT 00016044, paralectotypeю C Female, ZISP _ ENT 00016045, paralectotype. A. watajii. D Male, paratype, ZISP _ ENT 00016241. E Female, ZISP _ ENT 00016243. F Female, ZISP _ ENT 00016244.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
5F64AA50314050FB9D98D7148A80D155.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539540	https://doi.org/10.3897/asp.84.e161376.figure15	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
5F64AA50314050FB9D98D7148A80D155.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539544	https://doi.org/10.3897/asp.84.e161376.figure18	Figure 18. Vesica. A. hilaris, ZISP _ ENT 00014891. A Right lateral view. B Left lateral view, A. infamis, ZISP _ ENT 00016252. C Right lateral view. D Left lateral view. A. limbatus, Europe, ZISP _ ENT 00016012. E Right lateral view. F Left lateral view. A. maackiae, ZISP _ ENT 00016247. G Right lateral view. H Left lateral view. A. limbatus, Asia, ZISP _ ENT 00014886. I Right lateral view. J Left lateral view. A. subpulchellus, ZISP _ ENT 00003765. K Right lateral view. L Left lateral view.	Figure 18. Vesica. A. hilaris, ZISP _ ENT 00014891. A Right lateral view. B Left lateral view, A. infamis, ZISP _ ENT 00016252. C Right lateral view. D Left lateral view. A. limbatus, Europe, ZISP _ ENT 00016012. E Right lateral view. F Left lateral view. A. maackiae, ZISP _ ENT 00016247. G Right lateral view. H Left lateral view. A. limbatus, Asia, ZISP _ ENT 00014886. I Right lateral view. J Left lateral view. A. subpulchellus, ZISP _ ENT 00003765. K Right lateral view. L Left lateral view.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
D6794063B383586ABF98F45C00CEE127.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539533	https://doi.org/10.3897/asp.84.e161376.figure8	Figure 8. Female genitalia. A. spinolae. A Dorsal labiate plate, ZISP _ ENT 00014490. B Posterior wall, ZISP _ ENT 00015712. A. subhilaris ZISP _ ENT 000116157. C Posterior wall. D Dorsal labiate plate. A. sp. nr. subhilaris, ZISP _ ENT 00014784. E Dorsal labiate plate. F Posterior wall. A. watajii, ZISP _ ENT 0006039. G Dorsal labiate plate. H Posterior wall. I Ventral wall.	Figure 8. Female genitalia. A. spinolae. A Dorsal labiate plate, ZISP _ ENT 00014490. B Posterior wall, ZISP _ ENT 00015712. A. subhilaris ZISP _ ENT 000116157. C Posterior wall. D Dorsal labiate plate. A. sp. nr. subhilaris, ZISP _ ENT 00014784. E Dorsal labiate plate. F Posterior wall. A. watajii, ZISP _ ENT 0006039. G Dorsal labiate plate. H Posterior wall. I Ventral wall.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
D6794063B383586ABF98F45C00CEE127.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539538	https://doi.org/10.3897/asp.84.e161376.figure13	Figure 13. Habitus, dorsal view. A. subpulchellus. A Male, ZISP _ ENT 00016042, paralectotype, B Female, ZISP _ ENT 00016044, paralectotypeю C Female, ZISP _ ENT 00016045, paralectotype. A. watajii. D Male, paratype, ZISP _ ENT 00016241. E Female, ZISP _ ENT 00016243. F Female, ZISP _ ENT 00016244.	Figure 13. Habitus, dorsal view. A. subpulchellus. A Male, ZISP _ ENT 00016042, paralectotype, B Female, ZISP _ ENT 00016044, paralectotypeю C Female, ZISP _ ENT 00016045, paralectotype. A. watajii. D Male, paratype, ZISP _ ENT 00016241. E Female, ZISP _ ENT 00016243. F Female, ZISP _ ENT 00016244.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
D6794063B383586ABF98F45C00CEE127.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539540	https://doi.org/10.3897/asp.84.e161376.figure15	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	Figure 15. Head, frontal view. A. adustus. A Male, ZISP _ ENT 00016237. A. fraxinicola. B Female, ZISP _ ENT 00016596. A. furvus. C Male, ZISP _ ENT 00016303, holotype. A. gotorum. D Male, ZISP _ ENT 00016232. A. hilaris. E Female, paratype, ZISP _ ENT 00015184. A. infamis. F Male, paratype, ZISP _ ENT 00016250, holotype. A. limbatus. G Asia, Male, ZISP _ ENT 00015917. H Europe, Male, ZISP _ ENT 00015986. A. maackiae. I Male, ZISP _ ENT 00016046. A. shikotan sp. nov. J Male, holotype, ZISP _ ENT 00016047 K Male, paratype, ZISP _ ENT 00016047. A. subhilaris. L Female, ZISP _ ENT 00016158. A. sp. nr. subhilaris. M Female, ZISP _ ENT 00014874. A. subpulchellus. N Male, ZISP _ ENT 00016042, paralectotype. A. watajii. O Male, paratype, ZISP _ ENT 00016241. P Female, ZISP _ ENT 00016243. A. spinolae. Q Specimen initially identified as A. lucorum, Male, ZISP _ ENT 00006565. R Specimen initially identified as A. spinolae, Female, ZISP _ ENT 00014766. S Specimen initially identified as A. nigronasutus, Male, ZISP _ ENT 00014551. T Specimen initially identified as A. nigrovirens, Male, ZISP _ ENT 00014590. U Specimen initially identified as A. nigrovirens, Female, ZISP _ ENT 00014618. V Specimen initially identified as A. malaisei, Female, ZISP _ ENT 00015747. W Specimen initially identified as A. malaisei Male, ZISP _ ENT 00015764.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
D6794063B383586ABF98F45C00CEE127.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1539542	https://doi.org/10.3897/asp.84.e161376.figure17	Figure 17. Vesica. A. spinolae. A Right lateral view, ZISP _ ENT 00006623. B Left lateral view, ZISP _ ENT 00005851. C Right lateral view, ZISP _ ENT 00015780. D Anterior view, ZISP _ ENT 00005851. E Lateral and sublateral sclerites, left lateral view, ZISP _ ENT 00005851. A. subpulchellus, ZISP _ ENT 00016047. F Right lateral view. G Left lateral view. A. sp. nr. subhilaris, (M 197) ZISP _ ENT 00014867. H Right lateral view. I Left lateral view. A. subhilaris, ZISP _ ENT 00016193. J Right lateral view. K Left lateral view. A. watajii, ZISP _ ENT 00016193. L Right lateral view, ZISP _ ENT 00016247. M Left lateral view, ZISP _ ENT 00003760.	Figure 17. Vesica. A. spinolae. A Right lateral view, ZISP _ ENT 00006623. B Left lateral view, ZISP _ ENT 00005851. C Right lateral view, ZISP _ ENT 00015780. D Anterior view, ZISP _ ENT 00005851. E Lateral and sublateral sclerites, left lateral view, ZISP _ ENT 00005851. A. subpulchellus, ZISP _ ENT 00016047. F Right lateral view. G Left lateral view. A. sp. nr. subhilaris, (M 197) ZISP _ ENT 00014867. H Right lateral view. I Left lateral view. A. subhilaris, ZISP _ ENT 00016193. J Right lateral view. K Left lateral view. A. watajii, ZISP _ ENT 00016193. L Right lateral view, ZISP _ ENT 00016247. M Left lateral view, ZISP _ ENT 00003760.	2026-02-13	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.		Zenodo	biologists	Namyatova, Anna A.;Dzhelali, Polina A.;Bolshakova, Darya S.			
