identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
6CBEA4590CE4529E88195D917325A862.text	6CBEA4590CE4529E88195D917325A862.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amathimysis sarbui Bacescu 1991	<div><p>Amathimysis sarbui Băcescu, 1991</p><p>Fig. 2 E, F</p><p>Amathimysis sarbui Băcescu, 1991: 6–8, fig. 2 except for the erroneous legend (original description); Wittmann 2024 a: 145 (in key).</p><p>Amathimysis illifei Băcescu, 1991: legend of fig. 2 (nomen nudum, species name confused with that of Mysidium iliffei Băcescu, 1991, described in the same publication).</p><p>Note.</p><p>Only the ♀ holotype known. Băcescu (1991) claimed deposition of slides with registry number 247 at the MINGA. However, there is neither a respective entry in the inventory booklet nor such material in the type collection (Iorgu Petrescu, pers. comm. in 2025).</p><p>Short diagnosis.</p><p>Based on adult female holotype: Amathimysis Brattegard, 1969, with very large eyes, larger than the small antennal scale; antennal scale stout, only twice as long as maximum width, terminal segment 1 / 4–1 / 3 of total scale length, lateral margin with one tooth marking end of bare portion; endopod of uropods without spine; telson subtriangular, roughly heart-shaped, terminal margin narrowly truncate with two densely set large spines, each lateral margin with only one small subterminal spine.</p><p>Distribution</p><p>(Fig. 3 B). Băcescu (1991) described the holotype from a submarine cave at the Abaco Islands in the Bahamas (ca 26°21'N, 77°09'W). Unless additional records become available, the very large eyes point to a troglophile rather than troglobiont status.</p></div>	https://treatment.plazi.org/id/6CBEA4590CE4529E88195D917325A862	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
7199468F0D495F5EBCFF64CBE322B329.text	7199468F0D495F5EBCFF64CBE322B329.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anchialina typica subsp. typica (Kroyer 1861)	<div><p>Anchialina typica typica (Krøyer, 1861)</p><p>Fig. 2 A – D</p><p>Anchialus typicus Krøyer, 1861: 53–59, fig. 7 a – l in pl. 2 (original description, tropical Atlantic).</p><p>Anchialina typica: Norman and Scott 1906: 24 (replacement name Anchialina for preoccupied genus name Anchialus); Ii 1964: 188 –195, figs 48, 49 (records, description); Brattegard 1970 a: 24 –28, fig. 6 (description, Bahamas); Meland et al. 2015: 23 (biogeography).</p><p>Anchialina typica typica: Nouvel 1971: 327, fig. 1 (diagnosis, Atlantic distribution); Fukuoka and Murano 2002: 100 (in key); Wittmann et al. 2009: 144 (records from Canary and Selvagens islands).</p><p>Anchialina tipica: Băcescu 1991: 6 (misspelling of species name, record, Bahamas).</p><p>Material examined.</p><p>Leeward Antilles — Curaçao • 1 imm. (BL = 1.5 mm) during daytime extracted from a large empty shell of Strombus sp. at Sun Reef (Station 17) .</p><p>Short diagnosis at species level.</p><p>Anchialina Norman &amp; Scott, 1906, with trapezoid rostrum with concave terminal margin, rostrum covering base of eyestalks; antennal scale short, not extending beyond median segment of antennular trunk; eyes normal-sized, cornea calotte-shaped, in dorsal view weakly bean-shaped; pleomere 1 of female and pleomeres 1–5 of male with well-developed pleural plates; pleopods with unilobed pseudobranchiae; exopod of male pleopod 3 with 11–13 segments, distal segment with three setae; exopod of uropod with 16–20 spines along distal 65–75 %; telson trapezoid with short apical cleft, distal half of each lateral margin with dense series of 25–35 spines; distolateral edges of telson each with large spine mesially accompanied by a half as long spine; cleft 1 / 6 telson length, its margin completely armed with laminae.</p><p>Differential diagnosis at subspecies level.</p><p>The nominotypical Atlantic-Mediterranean subspecies A. typica typica (Krøyer, 1861) differs from the Indian Ocean and Pacific A. typica orientalis H. Nouvel, 1971, by exopod of male pleopod 3 (Fig. 2 C) with modified bent setae at fourth to sixth segments from tip and by disto-lateral angle of third segment from tip not clearly projecting laterally vs modified bent setae at fifth to seventh or eighth segments, and disto-lateral angle of third segment strongly projecting laterally.</p><p>Distribution</p><p>(Fig. 3 A). This species shows an essentially epipelagic, nearly circumtropical distribution (Meland et al. 2015). The subspecies A. typica typica occurs in the western and eastern Atlantic and in the Mediterranean Sea (Wittmann et al. 2009). Only one ♀ was reported by Băcescu (1991) from a submarine cave at the Abaco Islands in the Bahamas (ca 26°21'N, 77°09'W). The present record from the interior of a large gastropod shell at Curaçao (Station 17) confirms a somewhat sciaphilic habitat preference. Nonetheless, this species is considered clearly trogloxene due to the great dominance of pelagic records.</p></div>	https://treatment.plazi.org/id/7199468F0D495F5EBCFF64CBE322B329	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
B1C7E09B8BA25C678255232EFB4ADD2F.text	B1C7E09B8BA25C678255232EFB4ADD2F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Antromysis cenotensis Creaser 1936	<div><p>Antromysis cenotensis Creaser, 1936</p><p>Fig. 4 A, B</p><p>Antromysis cenotensis Creaser, 1936: 121–123, figs 13–24 (original description, Balamcanché Cave in Yucatán); Bowman 1977: figs 1–4 (records, supplemental description); Reddell 1981: 105, Table 2, fig. 13 (records in Yucatán); Pesce et al. 1994: 115 (stygobiont status, endemic to Yucatán); Wittmann and Ariani 2019: fig. 2, suppl. (vaterite statoliths, biogeography); Price 2020: 978, figs 23.181 C, 23.183 F (in key, neotropical fauna); Daneliya and Wittmann 2021: table 10.1 (endangerment status); Liévano-Beltrán and Simões 2021: 1–13, Table 1, figs 1–6 (biogeography, records, bioindicator); Mejía-Ortíz et al. 2022: tables 1, 3 (freshwater subterranean species, vulnerability of subterranean waters); Wittmann 2023: table SS 1 (sensory organs).</p><p>Antromysis (Antromysis) cenotensis: Bowman 1977: 27 –31; Reddell 1981: 103; Iliffe 1992 b: 201, 202 (in checklist, distribution); Pesce and Iliffe 2002: 275, 276, figs 14–21 (record, morphological variability); Chávez-Solís et al. 2018: 2, 4 (micro-detritivorous, prey of decapods).</p><p>Antromysis coenotensis: Băcescu and Orghidan 1971: 228 (misspelling in key).</p><p>Material examined.</p><p>Yucatán • 5 ♀♀ ad. (BL = 2.6–3.0 mm), 4 ♂♂ ad. (BL also 2.6–3.0 mm) from anchialine cave in the Grutas de Tzab-Nah (Station 27) .</p><p>Short diagnosis.</p><p>Antromysis with eyestalks closely set together; exopod of male pleopod 3 two-segmented, basal segment without seta; telson slightly longer than maximum width near basis, with bare, convex lateral margins; telson terminally truncate, terminal margin slightly concave, terminal width 0.3–0.5 × maximum width; each disto-lateral edge with a comparatively strong spine; the pair of disto-lateral spines flanking 1–3 (mostly 2) small spines in median or paramedian position.</p><p>Distribution</p><p>(Fig. 3 C). In freshwater and anchialine cenotes of Quintana Roo and the northern Yucatán Peninsula, including Caribbean and Gulf of Mexico coasts, 18– 21°N, 87– 90°W, 1–37 m depth (Bowman 1977; Reddell 1981; Pesce and Iliffe 2002; Liévano-Beltrán and Simões 2021).</p></div>	https://treatment.plazi.org/id/B1C7E09B8BA25C678255232EFB4ADD2F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
23FB3B3394A55C4BAE2EA4D28F2B2E32.text	23FB3B3394A55C4BAE2EA4D28F2B2E32.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Antromysis Creaser 1936	<div><p>Genus Antromysis Creaser, 1936</p><p>Figs 4, 5</p><p>Antromysis Creaser, 1936: 121 (original description from cave in Yucatán); W. M. Tattersall 1951: 229, 230 (revised diagnosis); Mauchline 1980: 33, fig. 8.26 (in key to genera); Henderson and Bamber 1983: 143 (removal of invalid subgenus Parvimysis); Bamber and Henderson 1990: 398 (removal of invalid subgenus Surinamysis); Pesce et al. 1994: 115 (in list of hypogean species); Fukuoka 2006: fig. 1 (distribution); Price 2020: 974, 975 (in key).</p><p>Antromysis (Antromysis): Bowman 1977: 36, 37 (correct but meanwhile obsolete subgeneric assignment); Bowman 1982: 201.</p><p>nec Antromysis (Anophelina): Bowman 1977: 36, 37 (synonymous subgenus); Bowman 1982: 201.</p><p>nec Antromysis (Surinamysis) Bowman 1977: 37 (outdated combination); Bowman 1980: 208; Bowman 1981 a: 27; Bowman 1982: 201.</p><p>nec Antromysis (Parvimysis): Bowman 1977: 37 (invalid combination); Bowman 1981 a: 27; Bowman 1982: 201.</p><p>nec Antromysis: Mauchline 1980: 36, fig. 10.5 (invalid assignment of Parvimysis species in key to genera).</p><p>Short diagnosis.</p><p>Diamysini with separate subovate eyestalks, no cornea, no pigment (except for Antromysis anophelinae W. M. Tattersall, 1951, showing fused eyestalks, distally with pigmented band of rudimentary ommatidia); antennular trunk without accessory flagellum; antennal scale setose except for a short bare basal portion of the lateral margin; thoracic endopods 3–7 with 2 - or 3 - segmented carpopropodus; female pleopods forming a setose, uniramous, unsegmented bilobate plate; male pleopods biramous, endopod also forming an unsegmented bilobate plate, exopod 2 - or 3 - segmented, apically with a single strong seta; uropods without spine; telson small, stout, length &lt;1.3 × maximum width, lateral margins each with 0–4 spines.</p><p>Species inventory.</p><p>Six species are acknowledged at a world-wide scale; bold italics highlight species recorded from Caribbean caves: Antromysis cenotensis Creaser, 1936 (type species), A. anophelinae W. M. Tattersall, 1951, A. cubanica Băcescu &amp; Orghidan, 1971, A. juberthiei Băcescu &amp; Orghidan, 1977, A. peckorum Bowman, 1977, and A. reddelli Bowman, 1977 . See also key to Caribbean species below.</p></div>	https://treatment.plazi.org/id/23FB3B3394A55C4BAE2EA4D28F2B2E32	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
02078E7A7A625EE699FC8320465AC618.text	02078E7A7A625EE699FC8320465AC618.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Antromysis cubanica Bacescu & Orghidan 1971	<div><p>Antromysis cubanica Băcescu &amp; Orghidan, 1971</p><p>Figs 4 D, 4 E, 5</p><p>Antromysis cubanica Băcescu &amp; Orghidan, 1971: 226–228, fig. 1 (original description, Cuba, anchialine cave); Pesce et al. 1994: 115 (stygobiont status, endemic to Cuba); Petrescu and Wittmann 2009: 62 (types in museum collection); Ortiz and Lalana 2018: 74, fig. 16 B (Cuban archipelago, in key); Price 2020: 979, figs 23.181 F, 23.183 G (in key, neotropical fauna); Daneliya and Wittmann 2021: table 10.1 (endangerment status); Wittmann 2023: table SS 1 (sensory organs).</p><p>Types examined.</p><p>Cuba • holotype, ♂ ad. (BL = 3.0 mm; MINGA 49070) from the anchialine Cueva Juanelo Piedra (Station 24) • paratype, ♀ with 4 eggs (BL = 3.3 mm; MINGA MYS 455), sampling data as for holotype .</p><p>Short diagnosis.</p><p>Eyestalks set widely apart; exopod of male pleopod 3 two-segmented, basal segment without seta; telson slightly longer than maximum width near basis, with bare, convex lateral margins; telson with transverse, weakly convex terminal margin flanked by a pair of thin spines at distolateral edge, no additional spines.</p><p>Notes on male holotype</p><p>(Fig. 5). Distal segment of antennular trunk ventrally with small brush of setae, probably homologous with the appendix masculina otherwise not detected there. Eyes dorsoventrally flattened by a factor of 2.0, no cornea, no pigment in the ethanol-preserved specimen. Eyes with single-plane dorsal face and weakly convex ventral face. Anterior 1 / 5 of eye obliquely truncate in the transversal plane (i. e. in lateral view). Penes stout, length 6 % BL, telson length 8 % BL.</p><p>Distribution</p><p>(Fig. 3 D). At species level so far reported only from a brackish-water lake in the anchialine Cueva Juanelo Piedra at the Caribbean coast of Cuba (Station 24: type locality). It cannot be excluded that “ Antromysis ” recorded by Bolívar y Pieltain (1943) from the Cueva de Quíntanal in the municipality of Alquízar in Cuba belongs to this species (locality details not found).</p></div>	https://treatment.plazi.org/id/02078E7A7A625EE699FC8320465AC618	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
08350BBE00925F58939E34EB723A3D6E.text	08350BBE00925F58939E34EB723A3D6E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Antromysis juberthiei Bacescu & Orghidan. Modified 1977	<div><p>Antromysis juberthiei Băcescu &amp; Orghidan, 1977</p><p>Fig. 4 F</p><p>Antromysis juberthiei Băcescu &amp; Orghidan, 1977: 263–265, fig. 1 (original description); Pesce et al. 1994: 115 (stygobiont status, endemic to Cuba); Petrescu and Wittmann 2009: 62 (types in museum collection); Ortiz and Lalana 2018: 74, fig. 16 A (Cuban archipelago, in key); Price 2020: 979, fig. 23.181E (in key, neotropical fauna); Daneliya and Wittmann 2021: table 10.1 (endangerment status).</p><p>Type examined.</p><p>Cuba • holotype, ♀ ad. (BL = 3.8 mm; MINGA 49161) from the Cueva de los Murciélagos (= Bat Cave; Station 22) .</p><p>Short diagnosis.</p><p>Antromysis with eyestalks set well apart; exopod of male pleopod 3 two-segmented, basal segment distally with comparatively large barbed seta; telson slightly shorter than maximum width near basis; telson terminally broadly rounded, all around with bare margin.</p><p>Distribution</p><p>(Fig. 3 E). So far known only from brackish waters in the Cueva de los Murciélagos (Station 22: type locality) 1500 m landwards from the coast of Isla de Pinos, also called Isla de Juvendud; this island off the Caribbean coast of Cuba.</p></div>	https://treatment.plazi.org/id/08350BBE00925F58939E34EB723A3D6E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
52DE87ED7BB257BABEB55C2EF8A8C9AB.text	52DE87ED7BB257BABEB55C2EF8A8C9AB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Antromysis peckorum Bowman 1977	<div><p>Antromysis peckorum Bowman, 1977</p><p>Fig. 4 C</p><p>Antromysis peckorum Bowman, 1977: 31–33, figs 5, 6 (original description); Pesce et al. 1994: 116 (stygobiont status, endemic to Jamaica); Wittmann and Ariani 2019: fig. 2, suppl. (vaterite statoliths, biogeography); Price 2020: 979, fig. 23.181 D (in key, neotropical fauna); Wittmann 2023: table SS 1 (sensory organs).</p><p>Material examined.</p><p>Jamaica • 2 ♀♀ subad. (BL = 2.4–2.8 mm, ZMH -K-066768) from Jackson Bay Cave (Station 20) • 2 ♀♀ ad. (BL = 2.7–2.9 mm, NHMW -ZOO-CR-31423), sampling data as for preceding .</p><p>Short diagnosis.</p><p>Antromysis with eyestalks closely set together; exopod of male pleopod 3 two-segmented, basal segment without seta; telson slightly shorter than maximum width near basis; telson with bare, convex lateral margins; telson terminally truncate, terminal width 0.2 × maximum width; distolateral corners with pair of long slender spines flanking 0–2 (mostly 0) (para) median spines.</p><p>Distribution</p><p>(Fig. 3 F). So far known only from the system of the anchialine Jackson Bay Cave (type locality) at the southern coast of Jamaica (Bowman 1977). This also includes the present new records from Station 20.</p></div>	https://treatment.plazi.org/id/52DE87ED7BB257BABEB55C2EF8A8C9AB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
9C4048EBBF7557A8BA9C4F77C8C10E7D.text	9C4048EBBF7557A8BA9C4F77C8C10E7D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bermudamysis Bacescu & Iliffe 1986	<div><p>Genus Bermudamysis Băcescu &amp; Iliffe, 1986</p><p>Figs 33, 34, 35, 36, 37, 38, 39, 40, 41, 42</p><p>Bermudamysis Băcescu &amp; Iliffe, 1986 a: 95, 96 (definition, short description); Iliffe 1992 a: 623 (in list of anchialine taxa); Pesce et al. 1994: 117 (in list of subterranean taxa); Nouvel et al. 1999: 79 (morphology, taxonomy, in list of genera); San Vicente and Monniot 2014: 333, 341 (taxonomy, in key); Wittmann et al. 2014: 324, 341 (taxonomy, biogeography, transfer to Mysidetini); Daneliya 2021: 6 (taxonomy, claimed transfer to Heteromysini).</p><p>Type species.</p><p>Bermudamysis speluncola Băcescu &amp; Iliffe, 1986, by monotypy.</p><p>Species inventory.</p><p>Bermudamysis speluncola from Bermuda and B. caribbaea sp. nov. from islands of the Lesser Antilles are described below and given in key to species.</p><p>Revised diagnosis.</p><p>Designed for inclusion of the below-described B. caribbaea sp. nov. and underlining differences from the genus Mysidetes Holt &amp; Tattersall, 1906: Mysidetini with well-developed eyes; antennal scale setose all around, with small apical segment, no spines, no teeth; thoracic endopod 3 normal, non-prehensile, its carpopropodus with 2–4 segments, endopods 4–8 with 3 - to 6 - segmented carpopropodus; female sympod 6 with vestigial oostegite; marsupial chamber formed by oostegites from sympods 7–8; penes tube-like, &lt;1 / 4 BL; pleopods non-dimorphic, reduced to unsegmented setose plates with inconspicuous rudiments of pseudobranchial lobes, no spines, no teeth; distal portion of pleopods 1–4 not longer than basal portion marked by the pseudobranchial rudiments; uropods setose all around, no spines; telson terminally incised; cleft with laminae (plus one small spine subterminally on each side in the type species); lateral and terminal margins of telson with spines, no setae; nauplioid larvae with smooth cuticle all around, no cercopods.</p><p>Remarks.</p><p>As already discussed above for Platyops, Wittmann et al. (2014) placed Bermudamysis within the tribe Mysidetini mainly based on the non-prehensile structure of thoracic endopod 3. Daneliya (2021) transferred Bermudamysis to the Heteromysini by valuating a 2 - segmented carpopropodus of thoracic endopod 3 rather than the (more complex) prehensile structure in Heteromysini as decisive. However, Băcescu and Iliffe (1986 a) had indicated a 2 - segmented carpopropodus 3 only in drawing (fig. 1 G) but a 3 - segmented carpopropodus (expressed as “ 4 - segmented tarsus ”) in text (p. 97) upon original description of the type species B. speluncola . The different numbers are due to individual variability as confirmed by present findings of 2 - to 3 - segmented carpopropodites 3 in the type specimens. In addition, the here-described B. caribbaea sp. nov. has 3 - to 4 - segmented carpopropodites 3. This indicates that segmental numbers of carpopropodites are more variable (as expected) in serial pediform endopods compared with stand-alone prehensile endopods. Consequently, Bermudamysis is here retransferred to the Mysidetini .</p><p>Băcescu and Iliffe (1986 a) provided a short diagnosis of the genus Bermudamysis, regarding the marsupium by stating “ Two marsupial laminae with 2 or 4 eggs, .... ” and stating for the type species B. speluncola “ It is provided with only 1 pair of oostegites and bears 2 or 4 genital produces ”. The counts of oostegites are revised by the present findings (Fig. 36 B – D) of two pairs of fully functional oostegites (as in most Mysidae) plus one pair of vestigial oostegites in the type specimens.</p><p>According to the above-revised diagnosis the genus Bermudamysis differs from the genus Mysidetes by shorter pleopods 1–4 in both sexes, nauplioid larvae without cercopods, and by the combination of shorter penes, carpopropodus of thoracic endopods 3–8 with fewer segments and endopod of uropods without spines.</p></div>	https://treatment.plazi.org/id/9C4048EBBF7557A8BA9C4F77C8C10E7D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
7E0C1A664EF4521E895DAA0284FBA20A.text	7E0C1A664EF4521E895DAA0284FBA20A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bermudamysis caribbaea Wittmann & Chevaldonné 2026	<div><p>Bermudamysis caribbaea sp. nov.</p><p>Figs 37, 38, 39, 40, 41, 42</p><p>Type material.</p><p>All types from Lesser Antilles — Guadeloupe, Barracuda Cave (Station 11) • holotype, ♂ ad. (BL = 2.6 mm; NHMW -ZOO-CR-31410) • allotype, ♀ ad. (BL = 3.2 mm; MNHN -IU-2025-2825) • paratypes, 1 ♀ ad., 1 ♂ ad. (MNHN -IU-2025-2826) • paratypes, 1 ♀ ad., 1 imm. (NHMW -ZOO-CR-31411) • paratypes, 1 ♀ ad., 1 ♂ ad. (ZMH -K-066758) • paratypes, 3 ♀♀ ad., 1 ♂ ad. (MINGA MYS 456) .</p><p>Diagnosis.</p><p>Bermudamysis with triangular rostrum, distally narrowly rounded to acute, short, not reaching to half-length of basal segment of antennular trunk; antennal scale reaching to half-length of distal segment of antennular trunk, up to 1 / 10 its length beyond this segment; scale reaching &lt;1 / 10 length beyond antennal peduncle; thoracic endopods 3–8 long and slender, endopod 7 the longest, 0.4–0.6 × BL; carpopropodus of endopods 3–8 with 3 or 4, 3 or 4, 3 or 4, 4 or 5, 5 or 6 and 4 or 5 segments, respectively; penes tubular, 1 / 6 BL and 1.2–1.5 × length of ischium 8; each lateral margin of telson with six or seven spines only on distal 1 / 2–2 / 3; spines weakly increasing in length distally; U-shaped terminal cleft penetrating 24–35 % telson length, margins of cleft with nine or ten laminae only on proximal half; distolateral lobes of telson transversely truncate, each with large spine mesially accompanied by a &lt;1 / 2 as long spine; telson with total of 16–18 spines.</p><p>Description.</p><p>♀♀ ad. with BL = 2.6–3.4 mm (n = 11) and ♂♂ ad. with BL = 2.6–2.8 mm (n = 5). Cephalothorax comprises 33–38 % BL, pleon (without telson) 48–54 %, telson 10–14 %, carapace (without rostrum) 28–31 %, and rostrum 3–5 %. Thoracic sternites without mid-ventral processes in both sexes. Pleomeres 1–5 showing 0.7–1.0, 0.6–0.9, 0.7–0.9, 0.6–0.8, and 0.7–0.8 × the length of pleomere 6, respectively.</p><p>Carapace (Fig. 38 D – F). Rostrum as long as 0.5–0.8 × length of terminal segment of antennular trunk. Posterior margin of carapace leaves 0.5–1 posterior thoracic somite mid-dorsally exposed. Cervical sulcus very short but distinct. Cervical pore group (Fig. 38 E) formed by 11–16 pores with varying diameter (1–3 μm) arranged closely in front of the cervical sulcus. Another pore group (Fig. 38 F) closely in front of the mid-posterior margin of carapace; it consists of 10–12 pores ≈ 1.4 μm in diameter, arranged in subgroups around a larger but indistinct, rounded structure.</p><p>Eyes (Fig. 37). Eye length 0.23–0.30 mm (n = 11), this is 7–10 % BL. OB completely internal, at basis of eyestalk, diameter 34–59 µm (n = 4). Cornea calotte-shaped in dorsal view, diameter of calotte 0.20–0.27 mm, length 0.14–0.18 mm, this is 0.6–0.9 × length of eyestalk (cornea not included); ovoid in lateral view, length 1.2–1.5 × maximum width. Pigment to varying degrees centripetally retracted from crystalline bodies of eye (Fig. 37 C), thus feigning a smaller cornea upon low-power microscopy (Fig. 37 B, D). According to a review by Land (2004), retreated pigment is indicative of light adaption by pigment migration (a slow process) in crustaceans with superposition eyes.</p><p>Antennulae (Fig. 38 A, B). Antennular trunk of both sexes with setose apophyses and flagella, no spines, no teeth. Trunk extends 36–57 % its length beyond eyes. Measured along dorsal midline, the basal segment is 47–51 % trunk length, median segment 13–20 %, and terminal segment 31–39 %. Basal segment on basal half of its outer face with three or four small, barbed setae. Its dorsal apophysis with two or three simple barbed setae and 1–3 smooth whip setae. The slender lateral apophysis (lobe) reaching to distal margin of median segment or shortly beyond; its transverse terminal margin with three or four barbed setae and none or one smooth seta. Median segment dorsally with bifid apophysis bearing four or five barbed setae; mesial branch of this apophysis longer in males than in females. Terminal segment 0.9–1.1 × as long as wide; mesial half with three or four large plumose setae in females only. Mid-dorsal apophysis with 3–5 barbed setae. In both sexes outer antennular flagellum is thicker than inner one. Male lobe well setose, somewhat longer than wide, 1 / 3 as long as terminal segment of trunk, inserts ventrally close to terminal margin of antennular trunk.</p><p>Antennae (Fig. 38 C). Antennal sympod with two longer than basally wide, anteriorly projecting lobes between scale and peduncle. Antennal scale length is 3.1–4.3 × maximum width; apical segment contributes 12–16 % to total length; scale reaching 3–10 % of its length beyond antennal peduncle. Basal segment contributes 18–21 % to total length of peduncle, median segment 40–43 %, and terminal segment 38–42 %.</p><p>Primary mouthparts (Figs 38 G, 38 H, 39 A – C). Labrum (Fig. 38 G) not produced into a spiniform process; caudally weakly rugged and with small, stiff bristles; fields of setae on caudal and oral faces. Left and right mandibles (Fig. 39 A – C) essentially alike. Proximal segment of palp 11–18 % palp length, no setae. Length of median segment 2.1–2.6 × its maximum width and 63–70 % palp length. Mesial margin of median segment with 4–6 mostly sparsely barbed whip setae in near-median position, lateral margin almost all along with 6–9 smooth whip setae, terminal margin with two smooth whip setae. Terminal segment distally well setose, 19–23 % palp length. Pars molaris with well-developed grinding surface. Pars incisiva with 2–4 teeth, digitus mobilis with two or three teeth, and pars centralis with three or four spines bearing stiff bristles. Labium (Fig. 38 H) normal, comprising two hairy lobes with dense set of stiff bristles on distal third of inner face, no spines.</p><p>Foregut (Fig. 40). Gross structure as described by Wittmann and Abed-Navandi (2019) for Heteromysis domusmaris . Lateralia anteriorly with apically coronate spines (Fig. 40 B) and apically pronged spines (Fig. 40 C), each with loose series of small denticles along distal 1 / 5–1 / 3. Lateralia more caudally with separate group of three unilaterally serrated spines (Fig. 40 D). Dorsolateral infoldings with spine group formed by a small apically toothed spine together with two long spines, the latter furnished with unequal-sized teeth along distal 2 / 3 of shaft (Fig. 40 E). Two of three dissected foreguts were almost empty, the third one filled to ≈ 20 % with copepod remains, mostly unidentified organic material and few mineral particles.</p><p>Maxillula (Fig. 38 I) normal; distal segment terminally with 10–12 mostly weakly serrated spines; subterminally with four whip setae barbed on their distal 2 / 3; no pores detected. Endite of maxillula terminally with three strong, distally spiny setae accompanied by two barbed whip setae; inner and outer margins with numerous shorter, barbed whip setae.</p><p>Maxilla (Fig. 39 D) normal; terminal segment of endopod 1.7–1.9 × as long as maximum width. Lateral margin of exopod all along with 10–13 plumose setae, the two apical setae 2–3 × longer than the lateral setae.</p><p>Thoracopods (general, Figs 37 D, 41). Both sexes with flagellum of thoracic exopod 1 with eight segments, flagella 2–8 with nine segments. Basal plates expanded, 1.9–2.5 × as long as wide in both sexes, distolateral corner well rounded (Fig. 41 C). First thoracopods with leaf-like, smooth epipod. A large plumose seta present at the intersegmental joint connecting sympod 2 with the corresponding thoracic sternite; no such setae detected in remaining thoracopods. Total length of endopods, including length and slenderness of ischium, increase in series of endopods 1–7 and then decrease to endopod 8. Endopod 3 (Fig. 41 D) measures 1 / 3 BL and shows somewhat thicker and shorter ischium compared with endopods 4–8. Endopods 3, 4, partly also endopod 5, with one basally spiny (Fig. 41 H) and several smooth paradactylar whip setae. Endopod 3 with additional basally spiny setae (Fig. 41 F) on merus and carpopropodus, none on praeischium and ischium. Endopods 4, 5 only with smooth setae from praeischium to merus; endopods 6–8 all along with smooth setae. Endopod 1 with smooth, slender, weakly bent nail (Fig. 41 B), endopod 2 without nail, endopods 3–8 with smooth, even more slender, weakly bent to straight nail (Fig. 41 E). Nail 3 longer than nail 1, length then decreasing in series of nails 3–8. When stretched anteriorly, endopod 8 reaches to antennae, when stretched posteriorly to pleopod 5.</p><p>Maxillipeds (Fig. 41 A – C). Coxa of first maxilliped (thoracic endopod 1, Fig. 41 A, B) with small endite bearing one barbed seta at tip. Basis with large, prominent endite that is densely setose on mesial margin. Ischium and merus each with one smaller, mesially setose endite. Basis of second maxilliped (thoracic endopod 2, Fig. 41 C) with mesially projecting endite bearing one or two large plumose seta. In both sexes, combined praeischium plus ischium 0.5–0.7 × length of merus, carpopropodus plus dactylus 1.0–1.2 × merus. Dactylus very large, with dense brush formed by large numbers of normal setae and 7–9 modified setae, the latter apically bent, bearing two symmetrical series of stiff cils on either side in subbasal to median portions.</p><p>Marsupium (Figs 37 B, 39 E, 39 F). Gross structure of marsupium as described above for B. speluncola . The vestigial oostegite 1 (Fig. 39 E) with only one seta penetrating into marsupial chamber, this seta ‘ hispid’ by bilateral series of minute cils along subbasal to subapical portions. Caudal third of oostegite 2 (Fig. 39 F) with several hispid setae, rostral third with bilaterally well-barbed setae, central third with hispid as well as barbed setae. Oostegite 3 with only few hispid setae near conjunction with sympod; most margins of the (folded) plate with numerous setae barbed at least along distal 2 / 3. No setae on outer face of marsupium.</p><p>Penes (Figs 37 D, 41 K) 1.1–1.4 × length of ischium 7, when stretched anteriorly extending up to thoracic sympods 2–4. Tip trilobate. Penes subapically with one barbed seta, subbasally with group of three shorter barbed setae.</p><p>Pleopods (Fig. 42). Structure as in the above genus definition. Pleopod 1 shorter than the subequal pleopods 2–4, pleopod 5 longest. Male pleopods slightly longer than corresponding female pleopods. No additional differences between sexes.</p><p>Tail fan (Fig. 39 G – I). Scutellum paracaudale triangular, tip acute (Fig. 39 G) to narrowly rounded. Exopod of uropods 1.1–1.2 × length of endopod (Fig. 39 H). Exopod extends by 4–9 % its length beyond endopod and 29–33 % beyond telson; endopod 20–24 % its length beyond telson. Diameter of fluorite statoliths 55–71 μm (n = 6). Statolith formula 2 + 3 + (2–3) + 3 + 2 = 12–13 or 2 + 3 + (2–3) + 5 = 12–13 (n = 4 statoliths from 3 specimens). Latero-apical spines of telson (Fig. 39 I) 12–15 % telson length; medio-apical spines 0.3–0.4 × the length of latero-apical spines. Telson length 1.3–1.7 × its maximum width or 1.1–1.4 × length of sixth pleomere.</p><p>DNA data.</p><p>Only one 629 bp mitochondrial COI sequence obtained from a single individual at Barracuda Cave (GenBank PV 990330).</p><p>Derivatio nominis.</p><p>The species name is a Latinized adjective with feminine ending, derived from a Kalinago language noun, referring to occurrence in the Caribbean.</p><p>Distribution and habitat</p><p>(Fig. 10 A). Known only from the type locality in the marine sublittoral Barracuda Cave (Station 11), Guadeloupe. The mysids were observed occurring in small groups hovering above the sediment of semi-dark parts in the cave. They were reddish in situ and co-occurred with Parvimysis laminata . The observed retreated eye pigment (Fig. 37 C) correlates with occurrence in light rather than dark parts. The species is classified as a troglophile based on its occurrence in somewhat lighter rather than very dark cave parts in combination with large functional eyes.</p><p>Remarks.</p><p>The new species differs from its only congener B. speluncola by carpopropodus of thoracic endopods 6–8 with 4–6 segments (vs three segments), carpopropodus of endopod 3 with three or four segments (vs two or three segments), and penes 1.2–1.5 × (vs 0.4–0.6 ×) as long as ischium 8.</p></div>	https://treatment.plazi.org/id/7E0C1A664EF4521E895DAA0284FBA20A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
223566FA0C695F5C9B50FD2A88ADE1C8.text	223566FA0C695F5C9B50FD2A88ADE1C8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bermudamysis speluncola Bacescu & Iliffe 1986	<div><p>Bermudamysis speluncola Băcescu &amp; Iliffe, 1986</p><p>Figs 33, 34, 35, 36</p><p>Bermudamysis speluncola Băcescu &amp; Iliffe, 1986 a: 96–99, fig. 1 A – J (original description); Iliffe 1996: e. T 2766 A 9478950: 1–3 (Red List assessment as critically endangered); Wittmann and Ariani 2019: suppl. (statolith composition, biogeography); Kou et al. 2020: Table 1, figs 7–9 (molecular analysis, phylogenetic analysis); Iliffe and Calderón-Gutiérrez 2021: Table 1, fig. 3 C (stygobiont species, Bermuda); Wittmann 2023: table SS 1 (troglophile, non-visual eyestalk organs).</p><p>Note.</p><p>Redescription of this non-Caribbean species necessary for differentiation from the below described B. caribbaea sp. nov.</p><p>Type material.</p><p>Bermuda • Here designated lectotype, ♀ (BL = 2.6 mm, TL = 3.3 mm, MINGA MYS 447, Fig. 33 A) carrying nauplioid larvae from Green Bay Cave (Station 30) • paralectotypes, 33 ♀♀ ad. (BL = 2.2–2.9 mm, MINGA MYS 446), sampling data as for lectotype • paralectotypes, 2 ♀♀ ad. (BL = 2.2–2.5 mm), 9 ♂♂ ad. (BL = 2.3–2.5 mm), 9 imm., 8 juv. (MINGA MYS 452) from Palm Cave (Station 32) • paralectotypes, 5 ♀♀ ad. (BL = 2.3–2.9 mm), 3 ♂♂ ad. (BL = 2.4–2.7 mm) (MINGA MYS 450) from Cherry Pit Cave (Station 33) .</p><p>Other material.</p><p>Bermuda • 3 ♀♀ ad. (BL = 3.2–4.5 mm), 1 ♂ ad. (BL = 3.0 mm), 6 imm., 10 juv. from Green Bay Cave (Station 31) • 1 ♂ ad. (BL = 3.1 mm; ZMH -K-066761) from Deep Blue Cave (Station 34) • 7 ♀♀ ad., 10 ♂♂ ad., 13 imm. (MNHN -IU-2025-2827) from Grenadier Pool (Station 35) • 8 ♀♀ ad., 9 ♂♂ ad., 12 imm., 1 juv. (NHMW -ZOO-CR-31421), sampling data as for preceding • 5 ♀♀ ad., 10 ♂♂ ad., 14 imm., 1 juv. (ZMH -K-066762), sampling data as for preceding • 11 ♀♀ ad., 2 ♀♀ subad., 11 ♂♂ ad., 6 imm. (MNHN -IU-2025-2828), sampling data as for preceding • 5 ♀♀ ad., 1 ♀ subad., 11 ♂♂ ad., 2 ♂♂ subad., 11 imm. (NHMW -ZOO-CR-31422), sampling data as for preceding • 13 ♀♀ ad., 10 ♂♂ ad., 7 imm. (ZMH -K-066763), sampling data as for preceding .</p><p>Identification of the types.</p><p>Just recently discovered and identified as types by Iorgu Petrescu (pers. comm. in 15 Mar. 2024) in vials with hand-written labels by Mihai Băcescu.</p><p>Type locality and distribution.</p><p>Type locality not stated originally, here defined as Bermuda, Green Bay Cave (Station 30). Upon original description, Băcescu and Iliffe (1986 a) listed seven cave localities from Bermuda, five of which are represented in the present material (types plus non-types). The mysids were recorded up to now only from polyhaline to euhaline cave waters in Bermuda, depth range 0.5–18 m, where they tend to gather in swarms hovering close to the sediment surface or in the water column (Băcescu and Iliffe 1986 a).</p><p>Revised diagnosis.</p><p>Bermudamysis with triangular rostrum with rounded apex reaching to at most half-length of basal segment of antennular trunk; antennal scale reaching 0.3–0.5 × its length beyond antennular trunk and 0.3–0.4 × beyond antennal peduncle; thoracic endopod 3 non-prehensile, endopods 4–8 longer and more slender than endopod 3, endopod 7 the longest, ≈ 1 / 2 BL; carpopropodus of endopods 3–8 with 2 or 3, 3, 3, 3, 3 and 3 segments, respectively; penes tubular, &lt;1 / 10 BL and 0.4–0.6 × length of ischium 8; each lateral margin of telson with 4–6 (mostly 4) subequal spines positioned only on distal half; U-shaped terminal cleft penetrating 15–30 % telson length, its margins bearing seven to ten long, slender laminae on proximal 2 / 3; distolateral lobes of telson each with narrow apex ending in one large spine, one additional &lt;1 / 2 as long spine positioned nearby on each side shortly inside cleft; telson with total of 12–16 spines.</p><p>Short description of the lectotype</p><p>(Fig. 33 A). Only external features checked: cephalothorax 29 % BL, rostrum 6 %, carapace without rostrum 25 %, pleon without telson 53 %, and telson 12 %. Antennal scale length 4.7 × maximum width. Scale extends 1 / 5 its length beyond antennular trunk. Eye length 7 % BL. Eyes dorsoventrally compressed by a factor of 1.7. Eye pigment slightly retracted from cornea. Thoracic endopod 3 non-prehensile, its carpopropodus with two segments, carpopropodi 4–6 with three segments. Marsupial chamber formed by large oostegites from thoracopods 7–8, inside with total of four nauplioid larvae. Paramedian pairs of muscle structures, visible as near-pentagonal images (arrows in Fig. 36 E), are located in pleomeres 1–3 between neuronal cord and intestine. Each distolateral lobe of telson with large terminal spine mesially inside cleft accompanied by a shorter subterminal spine; each lateral margin with four spines on distal half. Cleft penetrates 1 / 4 telson length.</p><p>Non-type supplements to the original description.</p><p>Dissected parts included: eye length 0.24–0.29 mm (n = 10 adults), this is 6–11 % BL. OB completely internal, close to cornea, diameter 28–71 µm (n = 6). Setose mesial lobe dorsally at median segment of antennular trunk with length not exceeding width at basis (Fig. 34 A) in both sexes. Antennal sympod with two lobes emerging between scale and peduncle, both lobes shorter than basally wide (Fig. 34 B). Antennal scale length 3.8–5.1 × maximum width. Thoracic epipod 1 leaf-like, with subbasal seta. Thoracic endopod 3 with 2 - to 3 - segmented carpopropodus, endopod length 1 / 3 BL, distinctly less slender (Fig. 35 A) than endopods 4–8 (Fig. 35 C – E). Endopod 3 with smooth paradactylar whip setae except one whip seta with spiny handle and smooth flagellum (Fig. 35 B). Merus and carpopropodus of endopod 3 with additional basally spiny whip setae plus several smooth whip setae. Endopods 4–8 with only smooth setae all along (Fig. 35 C – E), not considering setae of the sympod. Basal plates of thoracic exopods 1–8 are ≈ 3 × longer than wide, distolateral corner rounded; flagellum with 7, 8, 8, 8, 8, 8, 8 and 7 segments, respectively. Penes terminally trilobate with barbed seta (Fig. 35 F). Paramedian pairs of muscle structures, visible as near-pentagonal images (arrows in Fig. 36 E), are located in pleomeres 1–3 between neuronal cord and intestine.</p><p>Marsupium (Fig. 36 B – D). As in most Mysidae, the marsupium is formed by two pairs of fully functional oostegites emerging from thoracic sympods 7, 8. Length of chamber ≈ 1 / 3 BL, maximum height and width each 1 / 6–1 / 4 BL, the values varying with brood volume by adjustment of the non-locked, motile oostegites. The main large oostegite 3 (Fig. 36 D) from sympod 8 forms a deep, rostrally converging bowl, rostrally enclosing part of oostegite 2 and caudally containing, if any, one egg or larva; rostral half of mesial margin with series of sparsely barbed setae. Oostegite 2 (Fig. 36 C) from sympod 7 forms a less deep bowl with, if any, again only one egg or larva; rostral margin obliquely truncate, bearing a short series of barbed setae. Outer face of marsupium without setae, no spines. The left and right pairs of oostegites together forming a main chamber with four subchambers (bowls), each providing space for only one offspring. In addition to the main oostegites, a vestigial oostegite 1 is present on sympod 6. This vestige represents a small flat lobe, distally obliquely truncate with two setae on tip. Both setae with minute cils along distal half (only marginally in focus in Fig. 36 B).</p></div>	https://treatment.plazi.org/id/223566FA0C695F5C9B50FD2A88ADE1C8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
BE061DF64581580083CC3D20EAA82428.text	BE061DF64581580083CC3D20EAA82428.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chelitrapezura dennisi (Bowman 1985) Wittmann & Chevaldonné 2026	<div><p>Chelitrapezura dennisi (Bowman, 1985) comb. nov.</p><p>Fig. 43</p><p>Heteromysoides dennisi Bowman, 1985; Juberthie-Jupeau and Iliffe 1994: Table 1 (stygobiont, Blue Holes in Bahama Islands); Hanamura and Kase 2001 b: 70 (in key); Daneliya 2021: 4 (taxonomy).</p><p>Platyops dennisi: Daneliya 2021: 4, 45–46 (revised combination).</p><p>Distribution</p><p>(Fig. 10 B). Known only from the type locality at Grand Bahama Island, Cemetery Cave, ca 100 m off the coast in a large blue hole, in 17 m depth, 330 m from entrance (as Heteromysoides dennisi in Bowman 1985, and in Juberthie-Jupeau and Iliffe 1994).</p><p>Short diagnosis.</p><p>Rostrum evenly rounded; distolateral edges of eyestalks anteriorly extended into a tooth-like projection, penes with long apical seta; endopod and exopod of uropods (sub) equal in length, no spines; telson slightly longer than maximum width near basis, distal ≤ 1 / 2 of each lateral margin with four spines increasing in length distally, terminal margin with pair of large disto-lateral spines flanking two half as long paramedian spines.</p></div>	https://treatment.plazi.org/id/BE061DF64581580083CC3D20EAA82428	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
C45B178767DA5B259FBAAFD2028E2D7B.text	C45B178767DA5B259FBAAFD2028E2D7B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chelitrapezura Wittmann & Chevaldonné 2026	<div><p>Chelitrapezura gen. nov.</p><p>Fig. 43</p><p>Heteromysoides: Bowman 1985: 945 (partim; first description of stygobiont species).</p><p>Platyops: Daneliya 2021: 4 (partim; reassignment of Heteromysoides species).</p><p>Type species.</p><p>Heteromysoides dennisi Bowman, 1985, by present designation. The combination Platyops dennisi (Bowman, 1985) emend. Daneliya 2021, here recombined as Chelitrapezura dennisi (Bowman, 1985), comb. nov. This species so far known only from the type locality at Grand Bahama Island, Cemetery Cave.</p><p>Derivatio nominis.</p><p>The genus name is a noun with feminine ending, formed by fusion of the Greek nouns χηλή (claw), τραπέζιο (trapeze) and ουρά (tail), referring to the subchelate thoracic endopod 3 in combination with the trapezoid telson as the main diagnostic characters.</p><p>Diagnosis.</p><p>Heteromysini with large, subquadrangular eyestalks, no eye papilla, small cornea implanted laterally; antennular trunk with normal setae, no flagellate spines, no backward-directed modified setae; antennal scale, pleopods and uropods setose, no spine, no teeth; thoracic endopod 3 forming a stout subchela with thick fused carpopropodus mesially armed with spines; thoracic endopods 4–8 normal, long, slender, with 2 - to 4 - segmented carpopropodus, praeischium without distal projection, ischium and merus without flagellate spines; penes tube-like; pleopods reduced to unsegmented setose plates with inconspicuous rudiments of pseudobranchial lobes; uropods normal, both rami undivided; telson trapezoid with bare proximal half, only distal ≤ 1 / 2 of lateral margins and terminal margin with spines.</p><p>Remarks.</p><p>Due to its prehensile thoracic endopod 3 with swollen unsegmented carpopropodus mesially armed with spines, Platyops dennisi (Bowman, 1985) or (as originally) Heteromysoides dennisi Bowman, 1985, respectively, cannot remain in the genus Platyops . The combination of a strong prehensile endopod 3 and trapezoid telson does not fit with any currently known genus of the subfamily Heteromysinae . This combination points to morphologically based paraphyly that arose from the integration of Heteromysoides dennisi into Platyops based on failed assumptions (as shown above) about the morphology of the type species P. sterreri by Daneliya (2021). This problem is addressed here by installing the new genus Chelitrapezura with Heteromysoides dennisi Bowman, 1985, as type species, this taxon then recombined as Chelitrapezura dennisi (Bowman, 1985), comb. nov. The new genus differs from the four other genera of the tribe Heteromysini and the single genus of the tribe Harmelinellini (namely Harmelinella Ledoyer, 1989) by a terminally truncate telson without cleft (Fig. 43 D). It differs from all seven currently known genera of Mysidetini by the subchelate thoracic endopod 3 with fused, mesially spinose carpopropodus (Fig. 43 B). Considering the modification of endopod 3 as a more complex structure than telson shape, the genus Chelitrapezura is here affiliated to the Heteromysini .</p></div>	https://treatment.plazi.org/id/C45B178767DA5B259FBAAFD2028E2D7B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
4EDF50A52DAA5E9A93DA1CFF40CE9DAD.text	4EDF50A52DAA5E9A93DA1CFF40CE9DAD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gironomysis lalanai Ortiz, Garcia-Debras & Perez 1997	<div><p>Gironomysis lalanai Ortiz, García-Debrás &amp; Pérez, 1997</p><p>Fig. 11</p><p>Gironomysis lalanai Ortiz et al., 1997: 22, 27, figs 1–25 (diagnosis, original description); Ortiz and Lalana 2018: 67, fig. 6 (in list, Cuban archipelago).</p><p>Short diagnosis.</p><p>Gironomysis with 3 - segmented mesial antennular flagellum; long, well-developed lateral flagellum of antennula with four or five apically blunt aesthetascs on basal segment; rudimentary antennal scale distally with two medium-sized setae, no spines, no teeth; male pleopod 4 with long smooth apical seta; lateral margin of exopod of uropods with only two setae on distal half.</p><p>Note.</p><p>Ortiz et al. (1997) misinterpreted the “ widened setae ” (aesthetascs) at the basal segment of the lateral antennular flagellum as representing the male lobe (see below remarks on the same subject in Palaumysis species).</p><p>Distribution</p><p>(Fig. 10 C). Known only from the Casimba Laguna Larga, a brackish well in the cave system of Playa Girón (near 22°04.54'N, 81°03.37'W) at the Caribbean coast of the province Matanzas, Cuba (Ortiz et al. 1997).</p></div>	https://treatment.plazi.org/id/4EDF50A52DAA5E9A93DA1CFF40CE9DAD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
BDE491CFF94F50C687E15D5454926F55.text	BDE491CFF94F50C687E15D5454926F55.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gironomysis Ortiz, Garcia-Debras & Perez 1997	<div><p>Genus Gironomysis Ortiz, García-Debrás &amp; Pérez, 1997</p><p>Gironomysis Ortiz et al., 1997: 22 (diagnosis, original description); Ortiz et al. 2012: 988, Table 1, fig. 12 (in key to genera, in species list, Caribbean); Wittmann et al. 2014: 340 (taxonomy); Ortiz and Lalana 2018: 67 (in taxa list, in key).</p><p>Short diagnosis.</p><p>Palaumysinae with normal-sized carapace leaving 1–2 ultimate thoracomeres mid-dorsally exposed; thoracic endopods 3–8 with unsegmented or 2 - segmented carpopropodus; endopod of uropods setose all around, longer than exopod, no spines; lateral margin of exopod with only few setae on distal half.</p></div>	https://treatment.plazi.org/id/BDE491CFF94F50C687E15D5454926F55	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
0A3B6FAA9DA75092A563A97F10BB8FDF.text	0A3B6FAA9DA75092A563A97F10BB8FDF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heteromysis (Heteromysis) cyanogoleus Bamber 2000	<div><p>Heteromysis (Heteromysis) cyanogoleus Bamber, 2000</p><p>Fig. 44 D, E</p><p>Heteromysis cyanogoleus Bamber, 2000: 129–133, figs 1–3 (original description, Bahamas); Hanamura and Kase 2001 a: 17 (discussion on subgeneric assignment); Fukuoka 2005: 1363, table III (habitat, distribution).</p><p>Diagnosis.</p><p>Modified after Bamber (2000). Heteromysis with pointed triangular rostrum extending beyond basal segment of antennular trunk; disto-mesial edge of antennular trunk in addition to normal setae with one blunt seta, mesially finely barbed and with distal pair of minute flagella (arrow in Fig. 44 D); eyes normal, cornea large, calotte-shaped, distally attached on the eyestalk; merus and carpus of the strong, subchelate thoracic endopod 3 mesially with flagellate spines; pleopods reduced to simple setose plates in both sexes, no teeth, no spines; endopod of uropods with only one spine mesially located beneath statocyst; telson trapezoid with incised terminal margin, lateral margins all along with nine or ten spines, disto-lateral lobes narrowly truncate, each armed with large spine mesially accompanied by a very small spine; triangular cleft armed all along with 18 acute laminae.</p><p>Affiliation.</p><p>Pleopods without spines together with modified seta at the disto-mesial corner of the antennular trunk bearing small apical flagella rather than a spine with subapical flagellum point to the subgenus Heteromysis S. I. Smith, 1873.</p><p>Distribution</p><p>(Fig. 10 E). Sandy’s Cave, east end of Grand Bahama Island, ca 27°N, 78°W (Bamber 2000). The species provisionally considered troglophile.</p></div>	https://treatment.plazi.org/id/0A3B6FAA9DA75092A563A97F10BB8FDF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
ECA752DC8A5C544AB455714EEE02884C.text	ECA752DC8A5C544AB455714EEE02884C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heteromysis (Olivemysis) bermudensis G. O. Sars 1885	<div><p>Heteromysis (Olivemysis) bermudensis G. O. Sars, 1885</p><p>Fig. 44 F – H</p><p>Heteromysis bermudensis G. O. Sars, 1885: 216, figs 1–7 in pl. XXXVIII (original description, Bermuda); Clarke 1955: 5, fig. 7 (description, Bermuda); Brattegard 1973: 51–53, fig. 20 (description, Caribbean); Modlin 1987 a: 116 (records, Caribbean); Fukuoka 2005: table III (habitat, distribution); Price and Heard 2008: Table 1 (morphology); Wittmann 2023: table SS 1 (records, trogloxene status).</p><p>Heteromysis bermudensis bermudensis: Bowman 1981 b: 458 –461, fig. 1 (first description of male).</p><p>Heteromysis (Olivaemysis) bermudensis: Băcescu and Iliffe 1986 a: 99, 100, fig. 2 M – R (description of male, misspelling of subgeneric name).</p><p>Heteromysis (Olivemysis) bermudensis: Price et al. 2002: 47, figs 3, 4 (records, in key); Ortiz et al. 2017 c: 49 (record, Yucatán); Wittmann and Abed-Navandi 2021: 137, 141 (morphology, in key).</p><p>Material examined.</p><p>Leeward Antilles — Curaçao • 1 ♂ subad. (BL = 3.1 mm) inside marine cave at Playa Lagun (Station 19) .</p><p>Diagnosis.</p><p>Heteromysis with triangular rostrum covering only basal part of eyestalks; eyes normal, cornea large, calotte-shaped, terminally attached to the eyestalk; disto-mesial edge of antennular trunk with modified spine characterized by a flattened handle with thin, subapically inserting flagellum; antennal scale with small distal segment, not reaching beyond distal margin of antennular trunk; thoracic endopod 3 subchelate, strong, with six or seven strong flagellate spines on mesial margin of fused carpopropodus; pleopods reduced to simple setose plates in both sexes, only male pleopod 4 distally with 27–55 densely set, small flagellate spines (modified setae); endopod of uropods not curved, its mesial margin with 10–16 subequal spines between statocyst and terminus; telson trapezoid with incised terminal margin; proximal 3 / 5 with bare lateral margins, distal 2 / 5 of lateral margins with 6–8 spines distally weakly increasing in length; disto-lateral lobes narrowly truncate, each armed with large spine mesially accompanied by a 0.6–0.8 × as long spine; U-shaped telson cleft with margin furnished with 11–20 laminae along basal half.</p><p>Distribution</p><p>(Fig. 10 F). Known from Bermuda, Gulf of Mexico, Belize, Caribbean coast of Mexico (Yucatán), Cuba, Caribbean coasts of Colombia, Cayman Islands, Turks and Caicos Islands and Antilles, total range 16– 32°N, 63– 88°W, depth 0.1–18 m. Found in marine caves, among algae, in sponges and empty shells, in the spaces between stones, over detritus bottoms in shallow water, also in association with sea anemones (Brattegard 1973; Băcescu and Iliffe 1986 a; Modlin 1987 a; Price et al. 2002; Fukuoka 2005; Ortiz et al. 2017 c; Wittmann 2023). Wittmann (2023) considers this species as trogloxene due to the great dominance of non-cave records.</p></div>	https://treatment.plazi.org/id/ECA752DC8A5C544AB455714EEE02884C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
F13D1BE004F254408F564B4BF5B3B385.text	F13D1BE004F254408F564B4BF5B3B385.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heteromysis (Olivemysis) floridensis Brattegard. Modified 1969	<div><p>Heteromysis (Olivemysis) floridensis Brattegard, 1969</p><p>Fig. 44 L – N</p><p>Heteromysis floridensis Brattegard 1969: 99–103, Table 1, figs 33, 34 (original description, Florida); Modlin 1987 b: 299 –301, fig. 2 (first record of male, Bahamas); Fukuoka 2005: table III (habitat, distribution).</p><p>Heteromysis (Olivemysis) floridensis: Ortiz et al. 2017 c: 49 (record, Yucatán); Ortiz and Lalana 2018: 67, 75, fig. 18 B (in key, Cuban records).</p><p>Diagnosis.</p><p>Modified and pooled after Brattegard (1969), Modlin (1987 b) and Ortiz and Lalana (2018). Heteromysis with triangular, distally right-angled, narrowly blunt rostrum covering basal part of eyestalks; eyes normal, cornea of moderate size, calotte-shaped, disto-laterally attached to eyestalk, no tooth-like process on anterior margin of stalk; disto-mesial edge of antennular trunk with flagellate spine characterized by a flattened handle with subapically inserting flagellum; antennal scale unsegmented, short, reaching to half-length of terminal segment of antennular trunk; thoracic endopod 3 forming a stout subchela; the thick fused carpopropodus mesially armed with five or six flagellate spines along its distal 2 / 3 (not every spine well visualized in Fig. 44 L due to arrangement of the distal four spines in pairs); pleopods reduced to simple setose plates in both sexes, only male pleopod 4 distally with 26 densely set, small flagellate spines (modified setae); uropods setose all around, endopod with 13–19 spines along mesial margin between statocyst and terminus, endopod curved mesially; telson linguiform, terminally incised, with bare proximal half of lateral margins, distal half of each margin with 6–8 spines increasing in length distally; disto-lateral lobes narrowly truncate, each armed with large spine mesially accompanied by a slightly longer spine; U-shaped cleft penetrates 0.4 × telson length, its margin furnished with 9–11 laminae on proximal quarter.</p><p>Distribution and habitat</p><p>(Fig. 10 G). Type locality is Key Biscayne, Florida, 25°42'11"N, 80°11'W, depth 2 m (Brattegard 1969). Known from the Caribbean coast of Mexico (Quintana Roo), Cuba, Florida and Grand Bahama Island, total range 21– 27°N, 78– 87°W, depth 2–4 m, on sandy bottoms, in sponges and near entrance of Cemetery Cave (Brattegard 1969; Modlin 1987 c; Fukuoka 2005; Ortiz and Lalana 2018).</p></div>	https://treatment.plazi.org/id/F13D1BE004F254408F564B4BF5B3B385	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
DE23E4D5FFFA5E4D9276B06D43EB9FAD.text	DE23E4D5FFFA5E4D9276B06D43EB9FAD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heteromysis S. I. Smith 1873	<div><p>Genus Heteromysis S. I. Smith, 1873</p><p>Figs 44, 45, 46, 47, 48, 49, 50, 51</p><p>Heteromysis S. I. Smith, 1873: 553, 554 (original description); G. O. Sars 1883: 55 (synonymy); Daneliya 2021: 6 –8 (revision, key to subgenera).</p><p>Chiromysis G. O. Sars, 1877: 56, 57 (original description of junior synonym); G. O. Sars 1883: 55 (in synonymy).</p><p>nec Heteromysis Czerniavsky, 1882 a: 57 (homonym of Heteromysis S. I. Smith, diagnosis in key); Czerniavsky 1882 b: 33 (diagnosis).</p><p>Short diagnosis.</p><p>Heteromysini with cornea normal or reduced to various degrees; disto-mesial corner of terminal segment of antennular trunk with or without flagellate spine opposed by a large smooth seta, no backward-oriented modified setae at this corner; thoracic endopod 3 prehensile, mostly stout, with swollen 1 - to 2 - segmented carpopropodus armed with spines on mesial (= inner) margin; praeischium of thoracic endopod 8 without spiniform extension in both sexes; penes tubular; pleopods reduced to simple setose plates in both sexes, with or without flagellate spines in males; endopod of uropods armed with spines in most species; telson with well-developed cleft at least basally armed with spine-like laminae; lateral margins of telson armed with spines at least in distal portions, terminal margin of disto-lateral lobes with at least one large spine and mostly with additional spines.</p><p>Species inventory.</p><p>Five species from Caribbean caves are here treated and given in below key to species: H. (Heteromysis) cyanogoleus Bamber, 2000, H. (Olivemysis) bermudensis G. O. Sars, 1885, H. (O.) floridensis Brattegard, 1969, H. spongicola (Băcescu, 1968), and H. troglophila sp. nov. Additional 103 species are acknowledged at world-wide scale. Type species is Heteromysis formosa S. I. Smith, 1873.</p><p>Remarks.</p><p>Băcescu (1968) did not explicitly designate a type species upon installing the genus-group taxon Heteromysoides . He added two species to this genus by using the wordings “ Heteromysoides spongicola n. g. n. sp. ” (in abstract and in legend of fig. 5) and “ Heteromysoides cotti (Calman, 1932) ” (on p. 231). From this, Daneliya (2021) argued that it would be impossible to determine which of the two species could be the type species and so the name ‘ Heteromysoides ’ would be unavailable by not matching Art. 13.3. of the Nomenclatorial Code (ICZN 1999). However, another article, namely Art. 13.4. is very well matched by inclusion of a single new species marked by “ n. g. n. sp. ”, thus deeming to confer availability. In conclusion, the type species was fixed from the very beginning as Heteromysoides spongicola Băcescu, 1968 .</p></div>	https://treatment.plazi.org/id/DE23E4D5FFFA5E4D9276B06D43EB9FAD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
F9825443A1755D0281B03E4FEC8C56C0.text	F9825443A1755D0281B03E4FEC8C56C0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heteromysis spongicola (Bacescu 1968)	<div><p>Heteromysis spongicola (Băcescu, 1968)</p><p>Figs 44 A – C, 45</p><p>Heteromysoides spongicola Băcescu, 1968: 231–233, fig. 5 (original description from Cuba); Pesce 1982: 113, 115 (in list of cave-dwelling taxa); Escobar-Briones and Soto 1991: Table 1 (biogeography); Hanamura and Kase 2004: 2146, 2147, fig. 1 (first record from Caribbean cave, redescription); Petrescu and Wittmann 2009: 58 (in type collection); Daneliya 2021: 3, 4 (type species, synonymy).</p><p>Heteromysis spongicola: Daneliya 2021: 4 (revised combination); Wittmann et al. 2021: 516 (taxonomy).</p><p>Type material examined.</p><p>Cuba • holotype, ♂ ad. (BL ≈ 3.5 mm, TL ≈ 4.2 mm; MINGA 49009) broken in two damaged pieces; from sponges in stands of corals ( Station 25: type locality) • paratype, damaged ♀ ad. (BL = 2.9 mm, TL = 3.5 mm; MINGA MYS 454; Fig. 45); sampling data as for holotype . The holotype listed as female in Petrescu and Wittmann (2009).</p><p>Diagnosis.</p><p>Heteromysis with ovoid, dorsally flattened eyestalks and with small, laterally implanted cornea; distal 1 / 5 of the stalks obliquely truncate in the sagittal plane, rendering the eye anteriorly chisel-shaped in lateral view, not so in dorsal view; disto-mesial corner of the antennular trunk with normal setae only, no strongly modified seta; pleopods reduced to simple setose plates in both sexes, no teeth, no spines; endopod of uropods with only one spine mesially implanted below statocyst; telson trapezoid with incised terminal margin; distal half of lateral margins with four spines increasing in length distally, proximal half bare; disto-lateral lobes narrowly truncate, each armed with large spine mesially accompanied by a very small spine; triangular cleft armed all along its margin with nine or ten acute laminae.</p><p>Descriptive notes.</p><p>Female paratype (Fig. 45). No ventral hump on distal segment of antennular trunk. Eyestalk subrectangular with rounded distal margin, 1.5 × longer than broad in dorsal view. Eyestalk dorsoventrally compressed by a factor of 1.8, dorsally more strongly flattened than ventrally. Thoracic endopods 1–8 and most pleopods broken. Thoracic sympod 3 thicker than sympods 2, 4–8 (Fig. 45), most likely supporting a thick, potentially prehensile endopod in analogy to other species of Heteromysis . Terminal cleft penetrates 1 / 5 telson length. Each lateral margin of telson with four spines increasing in length distally.</p><p>Male holotype. Antennular trunk extends 0.3 × its length beyond antennal scale. Distal segment of trunk ventrally with short papillary hump, possibly homologous with the otherwise missing appendix masculina. Antennal scale 3 × longer than broad. Eye dorsoventrally compressed by a factor of 2.0 in lateral view, 1.4 × longer than broad in dorsal view. Penes tube-like with 11 % BL, no apical stylet (seta). Pleopod 3 without setae measures 1.5 % BL, so reaching to half-length of pleomere 4; pleopod 4 is 1.8 % BL, reaching to proximal 1 / 4 of the longer pleomere 5.</p><p>Distribution</p><p>(Fig. 10 D). So far recorded from sponges in stands of corals at the northern coast of Cuba (type locality) and from a submarine cave off the western coast of Grand Cayman Island (Caribbean), total range 19– 23°N, 81– 82°W, 9–26 m depth (Băcescu 1968; Hanamura and Kase 2004). Both records are from cryptic habitats.</p><p>Remarks.</p><p>Upon synonymization of the genus Heteromysoides Băcescu, 1968, Daneliya (2021) proposed as a provisory to recombine the poorly known type species Heteromysoides spongicola Băcescu, 1968, as Heteromysis spongicola (Băcescu, 1968) . Our external inspection of the strongly damaged female paratype of Heteromysoides spongicola at the MINGA yielded a previously unknown thickening of thoracic sympod 3 (Fig. 45) pointing to a strong (broken) endopod 3. This would exclude attribution to the genus Mysidetes Holt &amp; Tattersall, 1906, thus indirectly (though weakly) supporting the recombination of this species taxon with the genus Heteromysis S. I. Smith, 1873, by Daneliya (2021).</p></div>	https://treatment.plazi.org/id/F9825443A1755D0281B03E4FEC8C56C0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
24B7E7BA206E50E59BA6B41D3757363D.text	24B7E7BA206E50E59BA6B41D3757363D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heteromysis troglophila Wittmann & Chevaldonné 2026	<div><p>Heteromysis troglophila sp. nov.</p><p>Figs 44 I – K, 46 –51</p><p>Heteromysis dispar: Wittmann 2023: table SS 1 (eyestalk organs).</p><p>Type material.</p><p>Lesser Antilles — Martinique • holotype, ♀ ad. (BL = 4.7 mm; NHMW -ZOO-CR-31424) from Anse Noire Cave (Station 1) • paratype, 1 imm. (BL = 2.4 mm), sampling data as for holotype — Guadeloupe • allotype, ♂ ad. (BL = 2.9 mm; NHMW -ZOO-CR-31788), paratypes, 1 imm. (BL = 1.9 mm; NHMW -ZOO-CR-31789), 1 ♀ ad. (BL = 5.1 mm; ZMHK-K-066767) from Amedien Cave (Station 12) .</p><p>Diagnosis.</p><p>Based on adults of both sexes. Heteromysis with wide-angled broadly rounded rostrum 0.2–1.0 × as long as terminal segment of antennular trunk; eyes normal, cornea calotte-shaped with diameter 0.5–0.7 × the length of antennular trunk; disto-mesial corner of trunk with flagellate spine characterized by a thick handle with subapically inserting flagellum, no additional modified spines (setae); appendix masculina well-developed; antennal scale with small distal segment, scale reaching to terminus of antennular trunk and 0.2–0.4 × its length beyond antennal peduncle; thoracic endopod 2 without nail; carpopropodus 3, 4 with two segments and carpopropodus 5–8 with three segments; thoracic endopod 3 non-dimorphic, subchelate, with carpopropodus 3–4 × longer than wide, carpus not fused with propodus, carpus furnished with three flagellate spines along distal 1 / 7–1 / 5 of its mesial margin; penes tubular, 1 / 2 as long as ischium 8, distally with four lobes around ejaculatory opening; pleopods reduced to simple setose plates, no spines in both sexes; uropods setose all around, no spines, endopod not curved; telson linguiform with incised terminal margin; 13–18 spines on convex distal half of each lateral margin, proximal half bare; disto-lateral lobes narrowly truncate with transverse terminal margin armed with large spine mesially accompanied by a much smaller spine and laterally by none or one medium-sized spine; V-shaped cleft penetrates 1 / 3 telson length, basal 1 / 2–3 / 4 of its margin furnished with 26–40 laminae; nauplioid larvae normal, with smooth cuticle all around.</p><p>Etymology.</p><p>The species name is a Latinized adjective with feminine ending referring to the troglophilic habit.</p><p>Description.</p><p>Body length of adult females 4.7–5.1 mm (n = 2), the only adult male 2.9 mm; female holotype and male allotype dissected. Cephalothorax comprises 29–34 % BL, pleon (without telson) 50–56 %, telson 13–14 %, carapace (without rostrum) 24–28 %, and rostrum 1–4 %. Male thoracic sternites 2, 8 with small, acute triangular, ventral processes (Figs 47 C, 49 G). Potential processes on male sternites 3–7 not clearly identified. No such processes in females. Pleomeres 1–5 show 0.6–0.9, 0.6–0.9, 0.7–0.9, 0.7–0.9, and 0.8–0.9 × the length of pleomere 6 in both sexes. Posterior margin of carapace leaves the ultimate thoracic somite mid-dorsally exposed. Cervical sulcus indistinct. Penes (Fig. 50 I) treated in Diagnosis.</p><p>Eyes (Fig. 46 A, C, D). Eyes very large, length 0.38–0.71 mm (n = 6), this is 11–15 % BL. Cornea calotte-shaped in dorsal view, diameter of calotte 0.36–0.52 mm, length 0.27–0.38 mm, this is 0.8–1.3 × length of eyestalk (cornea not included); spherical in lateral view. Cornea strongly black pigmented. Eyestalk without lobes, no spiniform processes.</p><p>Antennulae (Fig. 48 A – D). Antennular trunk with setose apophyses and flagella, no teeth. Trunk extends 0.3–0.5 × its length beyond eyes. Measured along dorsal midline, the basal segment is 41–48 % trunk length, median segment 13–20 %, and terminal segment 35–41 %. Basal segment on basal half of its outer face with three or four small, barbed setae. Its dorsal apophysis with six barbed setae. The subrectangular lateral apophysis (lobe) reaching to the distal extremity of the median segment in dorsal view but not so in ventral view; its transverse terminal margin with 3–5 barbed setae. Median segment with dorsal apophysis bearing four barbed setae. Terminal segment 1.1–1.3 × as long as wide, its mesial half with three or four large plumose setae. Shaft of the flagellate spine at disto-mesial corner with shorter and more acute distal portion in adult male (Fig. 48 D) compared with that in both adult females available (Fig. 48 A, B). Mid-dorsal apophysis with three or four barbed setae. Appendix masculina (dashed line in Fig. 48 C) is 1 / 5 length of terminal segment. Appendix with brush of smooth setae. Outer antennular flagellum thicker than inner one.</p><p>Antennae (Fig. 48 E). Antennal sympod with shorter than basally wide, anteriorly projecting lobe (dashed line in Fig. 48 E) above peduncle; lateral margin with small spiniform projection at some distance behind the scale. Scale length is 3–4 × maximum width; apical segment is 1 / 10 total length. Basal segment contributes 23–26 % to total length of peduncle, median segment 34–35 %, and terminal segment 41–43 % (n = 3).</p><p>Primary mouthparts (Fig. 48 F – J). Labrum (Fig. 48 F) rostrally produced into a short distally rounded triangular process; caudally with small, stiff bristles; fields of setae on caudal and oral faces. Proximal segment of mandibular palp (Fig. 48 G) contributing 8–10 % palp length, no setae. Length of median segment 3 × maximum width and 60–64 % palp length. Lateral margin of median segment with two or three smooth whip setae in median position; rostral face of proximal half with 2–4 smaller whip setae with barbed shaft; mesial margin almost all along with 6–9 smooth whip setae; terminal margin with one or two setae. Terminal segment distally well setose, 27–30 % palp length. Pars molaris with moderately strong grinding surface in both mandibles. Other parts strongly differing between left and right mandibles (Fig. 48 H, I). Left mandible (Fig. 48 H) with pars incisiva and digitus mobilis each with four blunt teeth, pars centralis with seven or eight comparatively small spines bearing stiff bristles. Pars incisiva of right mandible (Fig. 48 I) with three blunt teeth, digitus mobilis representing a large triangular tooth, and pars centralis with four large acute spines bearing stiff bristles. Labium (Fig. 48 J) normal, composed by two lobes with stiff bristles and setae on distal fourth, no spines.</p><p>Foregut (Fig. 49 A – D). Gross structure as described by Wittmann and Abed-Navandi (2019) for Heteromysis domusmaris . Lateralia anteriorly with apically coronate spines (Fig. 49 B) and apically pronged spines (Fig. 49 A), each with loose series of small denticles along distal 1 / 3–1 / 2. Lateralia more caudally with separate cluster of 6–8 bilaterally serrated spines (Fig. 49 C). Dorsolateral infoldings with spine group formed by a large spine serrated by dense series of small teeth and additional one or two smaller spines with fewer, larger, loosely set teeth (Fig. 49 D). Both dissected foreguts were empty.</p><p>Maxillula (Fig. 49 E) normal; distal segment terminally with 9–12 strong spines, at least three of which armed with two or three minute subapical teeth; this segment subterminally with two setae furnished with long barbs on their proximal half; no pores detected. Endite terminally with five strong setae bearing stiff bristles; inner margin with one and outer margin with two whip setae, both margins also armed with stiff bristles.</p><p>Maxilla (Fig. 49 F) normal; terminal segment of endopod 1.8–2.1 × as long as maximum width. Lateral margin of exopod all along with eight or nine plumose setae, not counting the apical seta and its mesially adjoining seta which are 1 / 3 longer than average lateral setae.</p><p>Thoracopods (general; Figs 49 G – J, 50 A – H, 51 H). Flagellum of thoracic exopod 1 with eight segments (Fig. 49 G), exopods 2–8 with nine segments (Fig. 51 H). Basal plates expanded, 1.5–2.3 × as long as wide, distolateral corner well rounded (Figs 49 G, 51 H). Thoracopod 1 with leaf-like epipod bearing none or one seta. Total length of endopods, including length of ischium, increases in series of endopods 1–6 and then decreases weakly to endopod 8. Endopod 3 (Fig. 50 A) measures 1 / 4 BL and shows somewhat thicker and shorter ischium compared with endopods 4–8. Endopods 4–8 only with smooth paradactylar setae, none on endopod 3. Endopod 1 with strong, weakly bent nail bilaterally lined with stiff bristles (Fig. 49 H, I) along part of distal half; endopod 2 without nail, endopods 4–8 with smooth, more slender, weakly bent nail (Fig. 50 F, H). Nail 3 approx. as long as nail 1 but 1.4–1.6 × as thick at basis, nails 4–8 are 4–15 % shorter and only 1 / 2 as thick as nail 3. When stretched anteriorly, endopod 8 reaches to labrum or clypeus, when stretched posteriorly to pleomeres 3 or 4.</p><p>Maxillipeds (Fig. 49 G – J). Coxa of first maxilliped (thoracic endopod 1, Fig. 49 G – I) with small endite bearing one barbed seta at tip. Basis with comparatively small endite (dashed line in Fig. 49 G) that is densely setose on mesial and terminal margins. Ischium to dactylus without endite. Intersegmental joint between thoracic sternite 2 and sympod 2 with large barbed seta as long as 1.2 × combined length of praeischium plus ischium of endopod 2. This seta accompanied by a small whip seta with many long barbs (Fig. 49 G). Basis of second maxilliped (thoracic endopod 2, Fig. 49 J) with mesially projecting endite bearing 1–3 large plumose setae. Combined praeischium plus ischium 0.7 × length of merus, carpopropodus plus dactylus 1.2–1.3 × merus. Dactylus large, with dense brush formed by large numbers of normal setae and 6–9 modified setae, the latter apically bent, bearing two symmetrical series of stiff cils on either side between 1 / 5–2 / 3 length from basis.</p><p>Gnathopods (Fig. 50 A – D). Thoracic endopod 3 non-dimorphic, forming a strong subchela. Basis with weakly distinct endite bearing one or two barbed setae. Ischium and merus strong, as normal in gnathopods. Ischium 2.7–3.3 × as long as wide; merus 4.4–4.6 × as long as wide and 1.6–1.7 × length of ischium. Carpopropodus 0.8–0.9 × the length of merus and 1.2–1.5 × ischium. Length of carpus 2.7–2.9 × maximum width. Carpus with two flagellate spines at disto-mesial corner and a single flagellate spine at 1 / 7–1 / 5 carpus length proximally from this corner on the mesial margin. Propodus 1 / 6–1 / 5 carpus length, mesially with two smooth setae and one bilaterally barbed seta. Dactylus 1 / 10 carpus length, distally with three long and one or two short smooth setae flanking the nail.</p><p>Marsupium (Figs 46 A, 50 G, 51 H). The vestigial oostegite 1 (Fig. 50 G) without seta. Oostegite 2 with four setae near conjunction with sympod 7, these setae all along hispid by minute stiff barbs; caudal fourth of lower margin with four smooth setae, rostral 3 / 4 with series of numerous well-barbed setae, most of which unilaterally barbed, only few setae bilaterally barbed. Lower margin of oostegite 3 (Fig. 51 H) most caudally with three hispid setae, rostrally followed by dense series of unilaterally barbed setae until rostral fourth of the margin with transition from unilaterally to bilaterally barbed. No setae on outer face of marsupium.</p><p>Pleopods (Fig. 51 A – G) non-dimorphic, representing setose plates in both sexes, no spines, no teeth. Length increases continuously by an increment of ≈ 1 / 15 upon each step from pleopod 1 to 4, whereas 9 / 10 from 4 to 5. Pleopod 5 (without setae) measures 7 % BL in both sexes. Absolute size differences of pleopods related to different BL are compensated in Fig. 51 by using inversely proportional scale length for male with BL = 2.9 mm (panels A – E) vs female with 4.7 mm (panels F, G).</p><p>Tail fan (Figs 44 J, 44 K, 46 B). Scutellum paracaudale right-angled triangular with blunt tip (Fig. 46 B). Exopod of uropods 1.2–1.4 × length of endopod (Fig. 44 J). Exopod extends by 20–27 % its length beyond endopod and 31–38 % beyond telson; endopod 9–17 % its length beyond telson. Diameter of fluorite statoliths 77–126 μm (n = 6). Statolith formula (2–3) + (1–2) + (0–1) + (5–6) + (4–7) = 13–18 (n = 4). Telson length 1.8–2.0 × its maximum width and 1.2–1.4 × length of sixth pleomere. Spines at lateral telson margins increase continuously in length distally up to 1 / 2 length of the spine series and then decrease weakly again in continuous series; the pattern is analogous for the laminae in the cleft (Fig. 46 B). The large terminal spines of the distal lobes of the telson (Figs 44 K, 46 B) measure 15–17 % telson length; the disto-mesial spines 0.3–0.4 × the length of the large apical spines.</p><p>Larvae.</p><p>The female with BL 4.7 mm carried five nauplioid larvae at substage N 2, the female with BL 5.1 mm three larvae at substage N 1. Nauplioids with normal gross morphology, body and antennae with smooth cuticle all around, no caudal furca, no spines, no setae.</p><p>DNA data.</p><p>A 623 bp mitochondrial COI sequence was obtained from a single specimen from Amedien Cave (GenBank PV 990328). The 801 bp sequence of nuclear 18 S (GenBank PV 990201) of H. troglophila displays highest similarities with other 18 S sequences available for Heteromysis species in GenBank, closest to an unidentified Japanese species (GenBank AM 422515) from which it differs by 2 %.</p><p>Distribution and habitat</p><p>(Fig. 10 H). Type locality is Martinique, semi-dark area of the Anse Noire Cave, 14°31.70'N, 61°05.30'W, 4 m depth (Station 1), where the mysids occurred in small groups. In Guadeloupe they dwelled solitarily in the dark zone of the Amedien Cave (Station 12), displaying a somewhat reddish color in situ. Total distribution range 15– 17°N, 61°W, depth 4–6 m.</p><p>Remarks.</p><p>Arguments for affiliation with the genus Heteromysis arise from a single flagellate spine at the disto-mesial corner of the antennular trunk, subchelate thoracic endopod 3, normal endopod 8 and reduced setose pleopods in both sexes. Affiliation at subgenus level is unclear.</p><p>The new species appears closely related to H. dispar Brattegard, 1970, from the Florida Keys based on telson structure and on penis with four lobes. The penis structure of H. dispar was first described by Wittmann (2013: fig. 4 e). That species differs from the new species by triangular vs broadly rounded rostrum; antennal peduncle reaching halfway vs 2 / 3–3 / 4 along antennal scale; thoracic endopod 2 with short nail vs no nail developed; each lateral margin of telson along distal half with 7–11 vs 13–18 spines.</p><p>A very similar telson is also found in H. gymnura W. M. Tattersall, 1922, from the northern Indian Ocean (W. M. Tattersall 1922; Wittmann and Ariani 2019). The distribution is discussed in Wittmann (2013) and the pleopod structure in Wittmann and Abed-Navandi (2021). Heteromysis gymnura differs from the new species by larger eyes and triangular rostrum; unsegmented antennal scale vs presence of a small distal segment; carpopropodus of thoracic endopod 4 with three vs two segments; telson cleft with ≈ 50 vs 26–40 laminae.</p><p>A similar telson structure is also found in H. komaii Fukuoka, 2005, from coastal waters of Japan (Fukuoka 2005). It differs from the new species by antennal peduncle reaching to distal 3 / 7 of antennal scale; carpopropodus of thoracic endopod 3 with two vs three flagellate spines; nearly straight vs distinctly convex distal half of lateral margins of telson; each lateral margin with 10–12 vs 13–18 spines; distally more widely open telson cleft; penes distally with two vs four lobes.</p></div>	https://treatment.plazi.org/id/24B7E7BA206E50E59BA6B41D3757363D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
4E61B99CA52853D1A968C72B698748B8.text	4E61B99CA52853D1A968C72B698748B8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mysidetini Holt & Tattersall 1906	<div><p>Tribe Mysidetini Holt &amp; Tattersall, 1906</p><p>Mysidetinae Holt &amp; Tattersall, 1906: 39 (new subfamily, definition, type genus defined); Hansen 1910: 4 (unaccepted); W. M. Tattersall 1923: 277 (withdrawal in favor of Leptomysini).</p><p>Mysidetini: Wittmann et al. 2014: 341, 346 (re-establishment at tribe level, diagnosis, in key); San Vicente 2017: 166 (distribution); Kou et al. 2020: Table 1 (molecular analysis); Wittmann and Chevaldonné 2021: 166, Table 1 (morphology, sensory structures); Daneliya 2023: 415 (short diagnosis).</p><p>Type genus.</p><p>Mysidetes Holt &amp; Tattersall, 1906 .</p><p>Diagnosis.</p><p>Slightly modified after Wittmann et al. (2014): Heteromysinae with appendix masculina well-developed to reduced; thoracic endopods 3–8 normal, pediform; endopod 3 not prehensile, not conspicuously swollen, carpopropodus with two or more (in some species many) segments; penes mostly long, tubular, stiff or changeable in size and form; pleopods reduced to simple, unsegmented setose plates in both sexes, no modified setae or spines; telson entire or with apical cleft, lateral margins with spines.</p><p>Seven genera included.</p><p>Bold italics highlight two genera with species recorded from Caribbean caves: Bermudamysis Băcescu &amp; Iliffe, 1986, Burrimysis Jaume &amp; Garcia, 1993, Deltamysis Bowman &amp; Orsi, 1992, Mysidetes Holt &amp; Tattersall, 1906, Mysifaun Wittmann, 1996, Platyops Băcescu &amp; Iliffe, 1986, and Pseudomysidetes W. M. Tattersall, 1936 .</p></div>	https://treatment.plazi.org/id/4E61B99CA52853D1A968C72B698748B8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
DF6678D53A825F81BB5F4CBFE35F1602.text	DF6678D53A825F81BB5F4CBFE35F1602.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mysidium (Mysidium) cubanense Bacescu & Ortiz 1984	<div><p>Mysidium (Mysidium) cubanense Băcescu &amp; Ortiz, 1984</p><p>Fig. 9 D, E</p><p>Mysidium cubanense Băcescu &amp; Ortiz, 1984: 18–20, fig. 2 A – K (original description, record from small cave in Cuba); Harrison and Bowman 1987: 678 (in comparison); Băcescu 1991: 3, 4 (records, Jamaica, Bahamas); Price and Heard 2009: 938 (distribution); Ortiz et al. 2017 b: 72 –74, fig. 4 (distribution, diagnosis); Wittmann 2023: table SS 1 (trogloxene status, sensory organs).</p><p>Mysidium (Mysidium) cubanense: Wittmann and Wirtz 2019: 12 –14, tables 2, 3, fig. 3 (revised diagnosis, in key, cave record).</p><p>Mysidium cubanensis: Harrison and Bowman 1987: 674 (gender inconsistency).</p><p>Types examined.</p><p>Cuba • holotype, ♂ ad. (BL = 4.5 mm; previously partly dissected; MINGA 49346) from entrance of small cave ( Station 26 — type locality) • allotype, ♀ ad. broken in two partly dissected pieces (MINGA 49347) from same sample as for holotype .</p><p>Non-types examined.</p><p>Leeward Antilles — Curaçao • 1 ♀ ad. (BL = 3.8 mm), 1 ♂ ad. (BL = 5.0 mm) from rock recesses at Sun Reef (Station 16) • 15 ♀♀ ad., 3 ♂♂ ad., 10 subad. (ZMH K-55259) from small marine cave at Playa Lagun (Station 18) .</p><p>Short diagnosis.</p><p>Mysidium with large cornea, diameter&gt; 2.5 × as long as terminal segment of antennular trunk; only males with anterior margin of antennular trunk dorsally with rounded, shield-like, mediodistal extension; thoracic endopod 3 with 3 - segmented carpopropodus; male pleopod 3 with medial widening at ≈ 1 / 2 length; male pleopod 4 with 4 - segmented exopod, endite of sympod reduced to weak medial hump or missing; telson linguiform to subrectangular or trapezoid, terminally not emarginated.</p><p>Distribution</p><p>(Fig. 7 A). Widely distributed in coastal waters of the Gulf of Mexico, Caribbean and Bahamas in the range of 12– 26°N, 68– 82°W, 0–26 m depth (Băcescu 1991; Price and Heard 2009; Ortiz et al. 2017 b; Wittmann and Wirtz 2019; Wittmann 2023). Present records (Stations 16, 18) from the coast of Curaçao. The species mainly forming swarms between corals, also found in caves and rock recesses.</p></div>	https://treatment.plazi.org/id/DF6678D53A825F81BB5F4CBFE35F1602	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
8F7F45BAC5D050D999109BC66103A617.text	8F7F45BAC5D050D999109BC66103A617.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mysidium (Mysidium) gracile (Dana 1852)	<div><p>Mysidium (Mysidium) gracile (Dana, 1852)</p><p>Fig. 9 H</p><p>Macromysis gracilis Dana, 1852: 653–655 (original description, combination with generic junior synonym).</p><p>Mysidium gracile: Czerniavsky 1887: 85 –87 (revised combination, description); Price and Heard 2004: 155, fig. 3 K (distribution, morphological remarks); Price and Heard 2009: 938 (distribution); Miyashita and Calliari 2014: 9, Table 1 (distribution); Ortiz et al. 2017 b: 74, 75, fig. 5 (distribution, diagnosis); Ortiz and Lalana 2018: 67, 72, fig. 11 (in species list for Cuba, in key).</p><p>Mysidia gracilis: Zimmer 1915 b: 215 (combination with generic junior homonym).</p><p>Mysidium (Mysidium) gracile: Wittmann and Ariani 2019: suppl. (fluorite statoliths, biogeography); Wittmann and Wirtz 2019: 6 –8, Table 1, fig. 1 (revised diagnosis, in key).</p><p>Material examined.</p><p>Lesser Antilles — St. Vincent • 1 ♂ ad. (BL = 4.1 mm), 2 subad., 1 juv. from entrance of Bat Cave (Station 3) — Martinique • 1 ♂ ad. (BL = 6.6 mm) from entrance of Zeb Cave (Station 2) — Saint-Martin • 10 imm. (NHMW -ZOO-CR-31415) from small holes ( Station 8) • 5 ♀♀ ad. (BL = 5.2–6.8 mm), 1 ♂ ad. (BL = 5.4 mm) from small holes ( Station 7) .</p><p>DNA data.</p><p>Eleven individuals of M. gracile were successfully amplified (548 and 658 bp fragments), ten from Basse Espagnole (Station 8), Saint-Martin, and one from Bat Cave (Station 3) in St. Vincent (GenBank PV 990300 – PV 990310). 453 bp aligned together with M. columbiae and M. triangulare (Fig. 8), the Saint-Martin M. gracile displayed six closely-related (divergence 0.2–0.4 %) different haplotypes. The St. Vincent sequence is quite divergent from the latter (16 %), which no doubt reflects the great geographical distance (≈ 560 km) and natural habitat fragmentation between the two sites. Only one of the concerned substitutions is non-synonymous.</p><p>Short diagnosis.</p><p>Mysidium with large cornea, diameter 2–3 × as long as terminal segment of antennular trunk; only males with anterior margin of antennular trunk dorsally with rounded, shield-like, mediodistal extension; antennal scale clearly reaching beyond antennular trunk; thoracic endopod 3 with 3 - segmented carpopropodus; male pleopod 3 with medial widening at ≈ 1 / 2 length; male pleopod 4 with 4 - segmented exopod, endite of sympod reduced to weak medial hump or missing. Telson subrectangular, terminally with shallow, rounded indentation.</p><p>Distribution and habitat</p><p>(Fig. 7 B). Widely distributed, mainly in sublittoral, also in littoral waters of the western Atlantic, ranges 32°N to 23°S, 42– 68°W, 3–35 m depth, including Bermuda, Gulf of Mexico, Caribbean and coast of Brazil (Price and Heard 2004; Ortiz et al. 2017 b; Wittmann and Wirtz 2019). Type locality is the harbor of Rio de Janeiro, ca 22°54'S, 43°10'W (Dana 1852). Found in a great variety of marine to metahaline habitats. Swarms hovering during the daytime over the sea floor, also found between corals and associated with sea urchins and fish (Randall et al. 1964; Emery 1968). Present records in the Lesser Antilles are from small swarms at the entrance of caves (Stations 2, 3) or in small holes (Stations 7, 8), also recovered as a single specimen from a large swarm of M. columbiae in the entrance of Zeb Cave (Station 2). The great dominance of non-cave records classifies this species as trogloxene.</p></div>	https://treatment.plazi.org/id/8F7F45BAC5D050D999109BC66103A617	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
07F6ECDEE0A55D4F88CCB24B28C37A59.text	07F6ECDEE0A55D4F88CCB24B28C37A59.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mysidium (Mysidium) integrum W. M. Tattersall 1951	<div><p>Mysidium (Mysidium) integrum W. M. Tattersall, 1951</p><p>Fig. 9 A – C</p><p>Mysidium integrum W. M. Tattersall, 1951: 223–225, fig. 96 (original description); Brattegard 1970 b: 128 (distribution); Brattegard 1973: 50, 51 (taxonomy, Caribbean coast); Modlin 1987 a: 115 (records, Caribbean, Belize); Price and Heard 2004: 155, fig. 3 J (distribution, morphological remarks); Price and Heard 2009: 938 (distribution); Ortiz et al. 2017 b: 76, 77, fig. 7 (diagnosis, in key, distribution); Wittmann 2023: table SS 1 (trogloxene status, sensory organs).</p><p>Mysidium (Mysidium) integrum: Wittmann and Ariani 2019: suppl. (fluorite statoliths, biogeography); Wittmann and Wirtz 2019: 9 –12, tables 2, 3, fig. 2 (revised definition, in key, cave records).</p><p>Material examined.</p><p>Lesser Antilles — Martinique • 1 ♀ ad. (BL = 4.9 mm) from Zeb Cave (Station 2) — Saint-Martin • 1 ♀ ad. (BL = 3.9 mm), 1 ♂ ad. (BL = 4.4 mm) from small holes at Basse Espagnole (Station 8) — Guadeloupe • 1 ♀ ad. (BL = 5.1 mm) from entrance of Cathedral Cave (Station 10) . Leeward Antilles — Curaçao • 6 ♂♂ ad. (BL = 3.4–4.9 mm), 3 ♀♀ ad. (BL = 3.3–4.3 mm), 4 ♀♀ subad., 3 ♂♂ subad., 4 imm. (NHMW -ZOO-CR-31420) from Blue Chamber (Station 15) • 19 ♀♀ ad., 19 ♂♂ ad., 1 ♀ subad., 1 juv. (ZMH -K-066772), sampling data as for preceding • 9 ♀♀ ad., 15 ♂♂ ad., 5 ♀ subad., 1 imm. (ZMH -K-066771) from entrance of small caves at Sun Reef (Station 17) • 10 ♀♀ ad., 16 ♂♂ ad., 2 ♀♀ subad., 2 ♂♂ subad. (MNHN -IU-2025-2831), sampling data as for preceding .</p><p>Short diagnosis.</p><p>Mysidium with cornea diameter &lt;2.5 × as long as terminal segment of antennular trunk; only males with anterior margin of antennular trunk dorsally with a small rounded, mediodistal extension; thoracic endopod 3 with 3 - segmented carpopropodus; male pleopod 3 with medial widening at ≈ 1 / 2 length; male pleopod 4 with 4 - segmented exopod, endite of sympod reduced to weak medial hump or missing; telson linguiform to subrectangular, with roughly straight or with convex, continuously rounded terminal margin.</p><p>Distribution and habitat</p><p>(Fig. 7 C). Distribution most similar to that of M. gracile, namely widely distributed in marine and occasionally also in metahaline waters of the western Atlantic from Bermuda to Bahamas, Caribbean, Gulf of Mexico and coast of Brazil, range 32°N to 23°S, 42– 69°W, 0–35 m depth (Brattegard 1970 b, 1973; Ortiz et al. 2017 b; Wittmann and Wirtz 2019). Type locality is tropical NW-Atlantic, Cruz Bay, St. John, Virgin Islands, ca 18°19.93'N, W 64°47.80'W (W. M. Tattersall 1951; Wittmann and Wirtz 2019). The present records (Stations 2, 8, 10, 15, 17) from cave entrances and small holes are within the already known range. In swarms close to the bottom of diverse coral reef and mangrove habitats, also found associated with sea urchins (Zoppi de Roa and Alonso 1997). The mysids were abundant at the entrance of caves and holes during the sampling in Curaçao (Stations 15, 17), but could be recovered only as rare individuals in Martinique (Station 2), Guadeloupe (Station 10) and Saint-Martin (Station 8), within swarms of other Mysidium species ( M. columbiae, M. gracile).</p></div>	https://treatment.plazi.org/id/07F6ECDEE0A55D4F88CCB24B28C37A59	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
C8C1DB381FC2522582F81384150E355E.text	C8C1DB381FC2522582F81384150E355E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mysidium (Mysidium) triangulare Wittmann 2019	<div><p>Mysidium (Mysidium) triangulare Wittmann in Wittmann &amp; Wirtz, 2019</p><p>Fig. 9 F, G</p><p>Mysidium (Mysidium) triangulare Wittmann in Wittmann &amp; Wirtz, 2019: 16–23, table 3, figs 5–8 (original description, in key, cave records); Wittmann and Ariani 2019: suppl. (fluorite statoliths, biogeography).</p><p>Mysidium triangulare: Hendrickx et al. 2023: 204 (in comparison); Wittmann 2023: table SS 1 (trogloxene status, sensory organs).</p><p>Material examined.</p><p>Lesser Antilles — Saint-Martin • 3 ♀♀ ad. (BL = 4.7–6.1 mm), 5 ♂♂ ad. (BL = 4.4–4.9 mm) from small holes ( Station 8) . Leeward Antilles — Curaçao • 3 ♀♀ ad. (BL = 4.0– 4.3 mm), 2 ♂♂ ad. (BL = 4.5–4.8 mm) (NHMW -ZOO-CR-31416) from Blue Chamber (Station 15) • 1 ♂ ad. (BL = 4.1 mm; ZMH -K-066773), sampling data as for preceding • 2 ♀♀ ad. (BL = 3.7–3.8 mm), 1 ♂ ad. (BL = 3.1 mm) (MNHN -IU-2025-2832) from entrance of small caves ( Station 17) .</p><p>DNA data.</p><p>Only two individuals were successfully amplified at the COI locus (615 and 658 bp fragments), both from Basse Espagnole (Station 8), Saint-Martin (GenBank PV 990311 and PV 990312). Over the aligned portion, they represented two different haplotypes that differed only by one (synonymous) substitution (0.16 % divergence). In relation to the other Mysidium species (Fig. 8), they appeared as a well-separated species, equally distinct from M. columbiae and M. gracile (≈ 25 % divergence).</p><p>Short diagnosis.</p><p>Mysidium with cornea diameter ≤ 2.5 × length of terminal segment of antennular trunk; only males with anterior margin of antennular trunk dorsally with rounded, shield-like, mediodistal extension; thoracic endopod 3 with 3 - segmented carpopropodus; male pleopod 3 with medial widening at ≈ 1 / 2 length; male pleopod 4 with 4 - segmented exopod; telson spatulate, its terminal portion triangular with rounded tip.</p><p>Distribution and habitat</p><p>(Fig. 7 D). Type locality is Caribbean, Curaçao, Playa Lagun, 12°19.09'N, 69°09.07'W, diverse habitats in 4–6 m depth (Wittmann and Wirtz 2019). This species previously known only from three sublittoral localities including an intertidal cave in euhaline waters off the island of Curaçao. The present findings from Curaçao (Stations 15, 17) and Saint-Martin (Station 8) extend the known geographic range to 12– 18°N, 63– 69°W, 3–28 m depth. By including these records, M. triangulare appears to be a troglophile rather than trogloxene as previously suggested by Wittmann (2023). Previously known from mostly large swarms hovering during daytime around and between corals. The present records always yielded small numbers, mostly at the entrance of caves or holes, only one specimen inside an intertidal cave. At Saint-Martin the mysids were captured from small swarms of M. gracile .</p></div>	https://treatment.plazi.org/id/C8C1DB381FC2522582F81384150E355E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
7BFD586AB7CB5152A5C16937F51E0A02.text	7BFD586AB7CB5152A5C16937F51E0A02.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mysidium (Orientomysidium) antillarum Wittmann 2019	<div><p>Mysidium (Orientomysidium) antillarum Wittmann, 2019</p><p>Fig. 9 J, K</p><p>Mysidium (Orientomysidium) antillarum Wittmann in Wittmann &amp; Wirtz, 2019: 30–37, table 3, figs 10–14 (original description, in key, cave records); Wittmann and Ariani 2019: suppl. (fluorite statoliths, biogeography).</p><p>Mysidium antillarum: Hendrickx et al. 2023: 203 (in citation, distribution).</p><p>Material examined.</p><p>Lesser Antilles — Bequia • 2 ♂♂ ad. (BL = 7.5–7.6 mm) from Full Moon Cave (Station 6) . Leeward Antilles — Curaçao • 1 ♀ ad., 4 ♀♀ subad., 3 ♂♂ subad., 12 imm. (MNHN -IU-2025-2829) from entrance of small caves ( Station 17) • 2 ♀♀ ad. (BL = 6.3–7.4 mm), 2 ♂♂ ad. (BL = 7.1–7.2 mm) (NHMW -ZOO-CR-31417), sampling data as for preceding • 1 ♂ ad., 5 ♀♀ subad., 4 ♂♂ subad., 10 imm. (ZMH -K-066769), sampling data as for preceding .</p><p>Short diagnosis.</p><p>Mysidium with cornea diameter 1.4–1.7 × as long as terminal segment of antennular trunk; only males with anterior margin of antennular trunk dorsally with rounded, shield-like, mediodistal extension; antennal scale extends clearly beyond antennular trunk; thoracic endopod 3 with 3 - segmented carpopropodus; male pleopod 4 with 3 - segmented exopod, endite of sympod reduced to indistinct projection; telson subrectangular with distinctly concave lateral margins and with distinct apical cleft separating two broadly rounded, apical lobes; acute laminae on terminal margins of telson and its cleft.</p><p>Distribution and habitat</p><p>(Fig. 7 E). Type locality is Caribbean, Curaçao, Reef Sint Marie, 12°12.73'N, 69°05.10'W, 0–0.5 m depth (Wittmann and Wirtz 2019). This species previously known only from six localities off the island of Curaçao plus one from Bonaire. The present records from caves in Bequia (Station 6) and Curaçao (Station 17) extend the known range to 12– 13°N, 61– 69°W, depth 0–28 m. The mysids are known from sublittoral to littoral euhaline waters where they form swarms closely hovering above the sea floor over diverse substrates, also found inside small caves (Wittmann and Wirtz 2019). The present first record from a large diveable cave, namely the Full Moon Cave in Bequia (Station 6), is based on only two specimens among a large swarm of M. columbiae . The great dominance of non-cave records classifies M. antillarum as trogloxene.</p></div>	https://treatment.plazi.org/id/7BFD586AB7CB5152A5C16937F51E0A02	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
2F13270764265053B1505210BBD8DA0F.text	2F13270764265053B1505210BBD8DA0F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mysidium (Orientomysidium) columbiae (Zimmer 1915)	<div><p>Mysidium (Orientomysidium) columbiae (Zimmer, 1915)</p><p>Fig. 9 L, M</p><p>Diamysis columbiae Zimmer, 1915 a: 172–174, figs 23–29 (outdated senior synonym); Zimmer 1915 b: 215 (in synonymy).</p><p>Mysidia columbiae: Zimmer 1915 b: 215, fig. 18 (recombination with generic junior homonym).</p><p>Mysidium columbiae: W. M. Tattersall 1951: 222, 223 (revised combination); Brattegard 1969: 86 –88, fig. 27 B (in part, record, description); Price and Heard 2004: 154, fig. 3 I (distribution); Miyashita and Calliari 2014: 9, Table 1 (distribution).</p><p>Mysidium colombiae: Price and Heard 2009: 938 (misspelling, distribution).</p><p>Mysidium (Orientomysidium) columbiae: Wittmann in Wittmann &amp; Wirtz, 2019: 27–30, tables 2, 3, figs 9,13E – J (subgeneric combination, description, in key); Wittmann and Ariani 2019: suppl. (fluorite statoliths, biogeography).</p><p>nec Mysidium columbiae: Brattegard 1969: fig. 27 E (male pleopod 4 not matching).</p><p>Material examined.</p><p>Lesser Antilles — Martinique • 20 ♀♀ ad., 9 ♂♂ ad., 2 ♀♀ subad. (MNHN -IU-2025-2830) from Zeb Cave (Station 2) — SW-Bequia • 4 ♂♂ ad., 3 ♂♂ subad., 8 ♀♀ subad., 15 imm. (ZMH -K-066770) from Full Moon Cave (Station 6) — Guadeloupe • 3 ♀♀ ad. (BL = 7.6–8.1 mm), 2 ♂♂ ad. (BL = 7.0– 7.5 mm) from Barracuda Cave (Station 11) • 5 ♀♀ ad., 5 ♂♂ ad., 2 ♂♂ subad., 3 imm. (NHMW -ZOO-CR-31418) from Cathedral Cave (Station 10) . Leeward Antilles — Curaçao • 1 ♀ subad. (BL = 3.5 mm) from entrance of small cave at Sun Reef (Station 17) .</p><p>DNA data.</p><p>At the COI locus, 15 individuals were amplified (545 and 658 bp fragments), three from Full Moon Cave (Station 6), Bequia (one single haplotype), two from Zeb Cave (Station 2) in Martinique (2 haplotypes) and ten (4 haplotypes) from Cathedral Cave (Station 10), Guadeloupe (GenBank PV 990313 – PV 990327). All substitutions are synonymous. Aligned 453 bp together with other two Mysidium species from this study (Fig. 8), the Martinique and Guadeloupe individuals are differentiated but grouped, with only ≈ 1 % divergence between the two islands (geographic distance 230 km). The divergence between these two and the Bequia sequences, however, is as high as ≈ 11 % for a minimum geographic distance (Bequia-Martinique) of 170 km. This indicates a much stronger barrier to M. columbiae ’ s gene flow south of Martinique than north of this island.</p><p>Short diagnosis.</p><p>Mysidium with cornea diameter 1.5–2.0 × as long as terminal segment of antennular trunk; only males with anterior margin of antennular trunk dorsally with rounded, shield-like, mediodistal extension; antennal scale extending far beyond antennular trunk (again more than in Fig. 9 A); thoracic endopod 3 with 3 - segmented carpopropodus; male pleopod 4 with 3 - segmented exopod, its sympod with strong endite at 2 / 5 sympod length from basis; telson subrectangular with distinctly concave lateral margins and with distinct apical cleft separating two broadly rounded, apical lobes; blunt laminae on terminal margins of telson and its cleft.</p><p>Distribution and habitat</p><p>(Fig. 7 F). Type locality is Caribbean coast of Colombia, Cartagena, ca 10°22.2'N, 75°31.2'W (Zimmer 1915 a; coordinates estimated by Wittmann and Wirtz 2019). Widely distributed in coastal marine waters of the western Atlantic: Bahamas, Caribbean, southern Gulf of Mexico and Brazil, 24°N to 8°S (Price and Heard 2004; Miyashita and Calliari 2014; Wittmann and Wirtz 2019; present data). The present samples from small up to diveable caves in Bequia (Station 6), Martinique (Station 2), Guadeloupe (Stations 10, 11) and Curaçao (Station 17) are within this range. Wittmann and Wirtz (2019) observed swarms hovering around and between corals during daytime and dispersing over the sea floor during the night in 3–26 m depth. Some of the present records were made during daytime from sometimes big swarms at the entrance of caves or at mid-distance from the entrance in the semi-dark zone. Nonetheless, this species is considered trogloxene due to the great dominance of non-cave records in the extensive literature.</p></div>	https://treatment.plazi.org/id/2F13270764265053B1505210BBD8DA0F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
A888E979599C55CE833B3E8D1224C2B1.text	A888E979599C55CE833B3E8D1224C2B1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mysidium Dana 1852	<div><p>Genus Mysidium Dana, 1852</p><p>Figs 7, 8, 9</p><p>Mysidia Dana, 1850: 130 (new genus, diagnosis; junior homonym of the hemipteran genus Mysidia Westwood, 1840); Zimmer 1915 b: 211, 215 (invalid restoration of the homonym Mysidia Dana, 1850).</p><p>Macromysis: Dana 1852: 638, 653 (diagnosis; incorrect assignment to Macromysis White, 1847, in turn a replacement name of the homonym Themisto Goodsir, 1842, and simultaneously a junior synonym of Praunus Leach, 1814); Meland et al. 2015: Table 1 (classification history).</p><p>Mysidium Dana, 1852: 638, 653 (replacement name Mysidium proposed in footnotes for occupied Mysidia); W. M. Tattersall 1951: 222 (Mysidium acknowledged); Wittmann and Wirtz 2019: 3 –6 (revision, diagnosis).</p><p>Diagnosis.</p><p>Shortened after Wittmann and Wirtz (2019): Anisomysini with well-developed, normal eyes; antennular trunk normal, without accessory flagellum; antennal scale setose all around, with small terminal segment; thoracopods 3–8 normal, endopods with 2 - or 3 - segmented carpopropodus; pleopods 1–5 of females and pleopods 1–3, 5 of males reduced to vestigial, setose, unsegmented rods; male pleopod 4 with distinct sympod, unsegmented endopod and 3 - or 4 - segmented, slender exopod; terminal two segments of exopod each bearing a large modified seta; uropods normal, setose all around, without spines; telson longer than broad, with entire or incised terminal margin, with spines and laminae on distal half; terminal margin with continuous series of densely-set spines and laminae.</p><p>Species inventory.</p><p>Eleven species are acknowledged at world-wide scale; bold italics highlight seven species recorded from Caribbean caves: Mysidium (Mysidium) gracile (Dana, 1852) (type species), M. (M.) integrum W. M. Tattersall, 1951, M. (M.) cubanense Băcescu &amp; Ortiz, 1984, M. (M.) rubroculatum Băcescu &amp; Ortiz, 1984, M. (M.) rickettsi Harrison &amp; Bowman, 1987, M. (M.) triangulare Wittmann in Wittmann &amp; Wirtz, 2019, M. (M.) winfieldi Hendrickx, Hernández-Payán &amp; Gómez-Gutierrez, 2023, M. (Occimysidium) pumae Ortiz, Hendrickx &amp; Winfield, 2017, M. (Orientomysidium) columbiae (Zimmer, 1915), M. (O.) antillarum Wittmann in Wittmann &amp; Wirtz, 2019, and M. iliffei Băcescu, 1991 . See also below key to species from Caribbean caves.</p><p>DNA data</p><p>(Fig. 8). Sequences of mitochondrial COI were obtained for three species of Mysidium . No other COI sequences are available in GenBank of any Mysidium or closely related genera.</p></div>	https://treatment.plazi.org/id/A888E979599C55CE833B3E8D1224C2B1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
05A7CA6673345F4DA5434498D1142644.text	05A7CA6673345F4DA5434498D1142644.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mysidium iliffei Bacescu 1991	<div><p>Mysidium iliffei Băcescu, 1991</p><p>Fig. 9 I</p><p>Mysidium iliffei Băcescu, 1991: 3–5 (original description); Ortiz et al. 2017 a: Table 1 (diagnostic characters); Wittmann and Wirtz 2019: 37, 38, Table 2 (revised definition, type locality, distribution, in key).</p><p>Mysidium illifei: Băcescu, 1991: species name misspelled in legend of fig. 1.</p><p>Note.</p><p>The types of M. iliffei are missing; there is no entry in the inventory booklet of the MINGA, and no other material is available. This poorly known species was not assigned at the subgeneric level by Wittmann and Wirtz (2019).</p><p>Short diagnosis.</p><p>Based on adult females and subadult males. Mysidium with cornea diameter 1.5 × as long as terminal segment of antennular trunk; antennal scale not extending beyond antennular trunk; thoracic endopod 3 with 3 - segmented carpopropodus; telson rectangular with approx. straight lateral margins and with distinct apical cleft separating two broadly rounded, apical lobes; acute laminae on terminal margins of telson and its cleft.</p><p>Distribution</p><p>(Fig. 7 G). Băcescu (1991) first described M. iliffei from material collected with M. cubanense by T. M. Iliffe in Jamaica. According to the sampling protocol by T. M. Iliffe (pers. comm. 09 March 2018; excerpt published by Wittmann and Wirtz 2019) the materials of M. iliffei were sampled from mysid swarms in the entrance of Joseph’s Caves, touristic sea cave in Jamaica, 18°16.08'N, 78°22.09'W, depth 1–3 m. The species is provisionally classified as a troglophile.</p></div>	https://treatment.plazi.org/id/05A7CA6673345F4DA5434498D1142644	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
D677D726DD765AEBAC208A3683DA34B2.text	D677D726DD765AEBAC208A3683DA34B2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Palaumysinae Wittmann 2013	<div><p>Subfamily Palaumysinae Wittmann, 2013</p><p>Mancomysini Băcescu &amp; Iliffe, 1986 b: 32 (diagnosis, no type genus); Ortiz et al. 2012: 988 (in key); Meland et al. 2015: 18, table 3 (nomen nudum, synonymy).</p><p>Mancomysinae: Meland and Willassen 2007: 1099, app. A (molecular phylogeny, revised rank); Petrescu and Wittmann 2009: 58 (type collection); Chevaldonné et al. 2015: fig. 5 (molecular systematics).</p><p>Palaumysinae Wittmann, 2013: 391 (replacement name); Wittmann et al. 2014: 332, 340, figs 4, 5, 21 (diagnosis, morphology, taxonomy); Ortiz and Lalana 2018: 67 (in list, Cuba); Kou et al. 2025: 5 (molecular phylogeny).</p><p>Diagnosis.</p><p>Updated from Wittmann et al. (2014). Mysidae with well-developed eyes, cornea calotte-shaped; antennular trunk robust, no male lobe; mesial antennular flagellum rudimentary, with 2–5 segments depending on species; antennal scale vestigial, unsegmented; labrum rostrally rounded, no distinct rostral process; thoracic endopods 3–8 with carpus separated from propodus by a transverse articulation, unless (in most species) carpopropodus undivided; two pairs of well-developed oostegites; penes of moderate size; all pleopods reduced to uniramous, unsegmented rods in both sexes, pleopod 4 of adult males with large, modified apical seta; both rami of the uropods undivided, endopod with statocyst as far as known composed of fluorite; telson short (sub) triangular, without or with small medio-apical indentation, its lateral margins bare except for each ending in a spine or tooth-like projection in paramedian position, no additional spines, no setae.</p><p>Type genus.</p><p>Palaumysis Băcescu &amp; Iliffe, 1986 .</p><p>Genus inventory.</p><p>Gironomysis Ortiz, García-Debrás &amp; Pérez, 1997 (1 species) and Palaumysis (5).</p></div>	https://treatment.plazi.org/id/D677D726DD765AEBAC208A3683DA34B2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
AB007AB1B0E25D0F90F6ECFC82D437FD.text	AB007AB1B0E25D0F90F6ECFC82D437FD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Palaumysis antillensis Wittmann & Chevaldonné 2026	<div><p>Palaumysis antillensis sp. nov.</p><p>Figs 12 H – J, 21 –28</p><p>Type material.</p><p>Lesser Antilles — SW-Bequia • holotype, ♂ ad. (BL = 1.43 mm; NHMW -ZOO-CR-31412) from Full Moon Cave (Station 4) • allotype, egg-bearing ♀ (BL = 1.60 mm; MNHN -IU-2025-2833), sampling data as for holotype • paratypes, 9 ♀♀ ad., 1 ♀ subad., 8 ♂♂ ad., 11 imm., 1 juv. (MNHN -IU-2025-2834), sampling data as for holotype • paratypes, 6 ♀♀ ad., 11 ♂♂ ad., 13 imm. (NHMW -ZOO-CR-31425), sampling data as for holotype • paratypes, 10 ♂♂ ad. (BL = 1.27–1.67 mm), 10 ♀♀ ad. (BL = 1.60–1.87 mm) (MINGA MYS 457), sampling data as for holotype • paratypes, 9 ♀♀ ad., 8 ♂♂ ad., 13 imm. (ZMH -K-066759), sampling data as for holotype — Guadeloupe • paratypes, 8 ♀♀ ad., 10 ♂♂ ad., 12 imm. (MNHN -IU-2025-2835) from Cathedral Cave (Station 13) • paratypes, 9 ♀♀ ad., 2 ♀ subad., 5 ♂♂ ad., 9 imm., 5 juv. (NHMW -ZOO-CR-31413), sampling data as for preceding • paratypes, 16 ♀♀ ad., 1 ♀ subad., 7 ♂♂ ad., 6 imm. (ZMH -K-066760), sampling data as for preceding .</p><p>Other material.</p><p>Cuba • 1 ♀ ad. (BL = 1.87 mm) from the anchialine El Brinco Cave (Station 23) .</p><p>Diagnosis.</p><p>Palaumysis with mesial antennular flagellum 3 - to 5 - segmented, apical segment 1 / 5–1 / 6 flagellum length; length of antennal scale 3–4 × maximum width, scale terminally with one long and one or two short setae, the long seta 0.6–1.0 × scale length, lateral margin of scale with none or one short seta; post-cervical lateral margin of carapace with projecting lobes; pleopod 1 with only moderately-sized setae in both sexes, no large plumose seta; apex of male pleopod 4 with one large and one short seta, the former reaching to or beyond pleopod 5 and bearing bilateral series of small cils from submedian to subapical portions, lateral margin of pleopod 4 with 0–3 short setae.</p><p>Description.</p><p>Adult ♀♀ with BL in the range of 1.53–1.92 mm (n = 119), ♂♂ with BL of 1.05–1.67 mm (n = 129) in the material from the Lesser Antilles. Cephalothorax comprises 34–43 % BL, pleon (without telson) 46–56 %, telson 5–8 %, carapace (without rostrum) 22–33 %, and rostrum 3–6 %. Clypeus transversely ellipsoidal, wider than long, with short, triangular mid-rostral protrusion (Figs 22, 24 D). Thoracic sternites without mid-ventral processes in both sexes, no spines, no setae.</p><p>Carapace (Figs 21 B, 21 C, 22, 23 A, 23 E). Rostrum as long as 0.3–0.6 × terminal segment of antennular trunk in dorsal view. The cervical sulcus forms a deep transverse incision (Fig. 23 A). The length of the post-cervical part of the carapace is 0.9–1.2 × the cervical part measured without rostrum. The posterior part with comparatively large, projecting lateral lobes (Fig. 23 E). Posterior margin slightly concave (Fig. 23 E) to slightly convex (Fig. 21 B), leaving ultimate four or five thoracomeres mid-dorsally exposed (in adults vs 1–4 in juveniles and immatures).</p><p>Eyes (Figs 21 E, 21 F, 22). Eye length 0.12–0.22 mm (n = 20), this is 8–15 % BL. Cornea calotte-shaped in dorsal view, diameter of the calotte 0.16–0.39 mm, length 0.07–0.17 mm, this is 0.6–1.3 × length of eyestalk (cornea not included). Cornea circular (Fig. 21 D) to weakly oval (Fig. 21 A) in lateral view, length 0.9–1.4 × maximum width. A small number of specimens in the present material appear light-adapted based on the very few pigments in the crystalline zone (Fig. 21 E) of the cornea; most of the other specimens appear dark-adapted by displaying more pigment between the crystalline cones (Fig. 21 F) upon external inspection (for light adaptation by slow pigment migration see Land 2004). OB completely internal, located at half-length of mesial margin of eyestalk, ellipsoidal to oviform with length 26–33 µm and width 15–22 µm (n = 8; Fig. 22).</p><p>Antennulae (Figs 22, 24 A, B). Antennular trunk non-dimorphic, 3 - segmented, with few setae, no spines, no teeth. Trunk extends 22–50 % its length beyond eyes in normal orientation (Fig. 21 B). Measured along dorsal midline, the basal segment is 30–43 % of trunk length, median segment 10–18 %, and terminal segment 47–56 %. Basal segment distolaterally with only one plumose seta, no apophysis. Median segment shows oblique segmental borders with both flanking segments in the sagittal plane, no apophysis, no setae. Terminal segment 1.6–2.1 × as long as wide; distolateral edge with a large plumose seta. Only this segment mid-dorsally with a subterminal setose apophysis bearing three or four small barbed setae (Fig. 24 A). The rudimentary mesial antennular flagellum 3 - to 5 - segmented; if 5 - segmented with second segment shortest and fourth segment longest, otherwise (Fig. 24 A) with first segment shortest and second segment longest. Apical segment 1 / 5–1 / 6 flagellum length, its apex with long seta accompanied by 2–4 short setae. The lateral antennular flagellum normal, its basal segment with 3–5 long, apically blunt aesthetascs.</p><p>Antennae (Fig. 24 C). Antennal sympod 2 - segmented, simple, no spines, no setae. Peduncle 3 - segmented, basal segment contributes 26–34 % to total length of peduncle, median segment 36–43 %, and terminal segment 28–33 %. Antennal scale vestigial, unsegmented, length is 3.1–4.7 × maximum width. Terminal margin truncate, with large bare seta subterminally to terminally accompanied by a small one; lateral margin with none or one small seta. Scale not reaching to distal margin of median segment of peduncle.</p><p>Mouthparts (Figs 23 A, 24 E – K, 25 A). Labrum (Fig. 24 E) rostrally well rounded; caudally weakly rugged and with small, stiff bristles; fields of setae on caudal and oral faces. Mandibles (Fig. 24 F – H). Proximal segment of palp 10–13 % palp length, no setae. Length of median segment 3.1–4.1 × its maximum width and 55–65 % palp length. Distal half of mesial margin of median segment with none or one sparsely barbed whip seta, lateral margin along distal 3 / 4 with four setae of that type, transverse terminal margin without seta (Fig. 24 F). Terminal segment well setose, 26–31 % palp length; its mesial margin with long smooth seta (longer than the segment). Pars molaris with moderately developed grinding surface. Left and right mandibles differ in armature with teeth and spines (Fig. 24 H vs Fig. 24 G). Pars incisiva of left mandible with four or five teeth, digitus mobilis with 2–4 teeth, and pars centralis with two or three spines bearing stiff bristles. Pars incisiva of right mandible with three or four teeth, digitus mobilis with three or four teeth, and pars centralis with complex of 7–10 bare teeth densely set on a large common basis. Labium (Fig. 24 I) normal, comprising two hairy lobes with stiff bristles on part of mesial face, no spines.</p><p>Foregut (Fig. 23 A – D). Gross structure as described by Wittmann and Abed-Navandi (2019) for Heteromysis domusmaris Wittmann &amp; Abed-Navandi, 2019 . Lateralia anteriorly with apically pronged spines (Fig. 23 C), each with small denticles along distal 1 / 2–2 / 3; more caudally with group of four or five unilaterally weakly serrated spines (Fig. 23 D). Dorsolateral infoldings with group formed by two or three long spines bearing unequally strong teeth along distal 1 / 2–4 / 5 of shaft (Fig. 23 B). Four of eight dissected foreguts were (almost) empty; the remaining were filled to ≈ 10–30 % with mostly unidentified organic material (detritus) and mineral particles.</p><p>Maxillula (Fig. 24 J, K) normal; distal segment terminally with 10–12 bare spines; subterminally with three setae bearing long barbs on their distal half; no pores detected. Endite of maxillula terminally with four or five strong, distally spiny setae proximally accompanied by one barbed seta; oral margin with two distally spiny setae, the most proximal one modified as in Fig. 24 K and pointing to the mouth field.</p><p>Maxilla (Fig. 25 A) normal; endopod with bare lateral margin, not counting a single seta subterminally on the distal segment; this segment 1.6–1.8 × as long as maximum width; proximal segment mesially with three basally barbed whip setae with thick handles. Lateral margin of exopod with five or six plumose whip setae, terminal margin with two plumose whip setae longer than the lateral setae.</p><p>Thoracopods (general, Figs 25 B – K, 26 A – E). Both sexes with flagellum of exopods 1–8 with 6 or 7, 7, 7, 6 or 7, 7, 7, 7, and 6 or 7 segments, respectively. Basal plates expanded, 1.6–2.4 × as long as wide, distolateral corner well rounded (Fig. 25 B). Thoracopod 1 with leaf-like, smooth epipod. Total length of endopods increases in series of endopods 1–7, while endopod 8 is shorter than endopods 5–7. Endopods 3–8 with unsegmented carpopropodus, with minute triangular scales (Fig. 26 E) on rostral face near and on lateral margin of ischium and merus (Figs 25 G, 26 D). Endopods 3–4 each with three paradactylar whip setae, endopods 5–8 with two or three such setae. Flagellum in most of these setae furnished with minute cils, while handle with normal-sized cils (barbs). Cils of the handle spine-like, each accompanied by a tiny cil (Fig. 25 I) in some paradactylar setae only in endopod 3. Dactylus 1 (Fig. 25 C) with strong, weakly bent nail; distal fourth of the inner (concave) margin microserrated by minute stiff cils except for a short, most apical bare portion in this nail (Fig. 25 D). Dactylus 2 (Fig. 25 F) with strong nail bent near apex; distal third of this nail with outer (convex) margin microserrated by minute stiff cils which are longer than those of nail 1. Dactyli 3–8 (Fig. 25 H, K) with long, slender, needle-shaped, weakly bent nail, distal 1 / 2–2 / 3 of the outer (convex) margin of this nail microserrated by minute teeth. Length increases from nail 1 to nail 7 (series of nails 1–3, 7 in Fig. 25 C, F, H, K), nail 8 shorter than nail 7. Endopod 3 is 1 / 4 BL (Fig. 25 G). When stretched anteriorly, endopod 8 reaches to the eyes, when stretched posteriorly to pleopod 5 or shortly beyond.</p><p>Maxillipeds (Fig. 25 B – F). Tip of coxa of first maxilliped (thoracic endopod 1, Fig. 25 B – D) with one short whip seta with barbed handle. Basis with large, prominent endite that is densely setose on and near its mesial margin. Ischium and merus without endites. Basis of second maxilliped (thoracic endopod 2, Fig. 25 E, F) with mesially projecting endite bearing one large, sparsely barbed seta. Combined praeischium plus ischium are 0.6–0.8 × length of merus, carpopropodus plus dactylus 0.7–0.8 × merus. Dactylus large, with comparatively meager brush formed by few basally barbed whip setae and even fewer bare setae.</p><p>Marsupium (Fig. 26 A – C). Oostegite 1 (Fig. 26 A) arising from thoracopod 7 represents a simple unfolded plate; lower margin of rostral half with series of whip setae with barbed handles; caudal half furnished with only few small whip setae. Oostegite 2 (Fig. 26 B) from thoracopod 8 with inward (mesially) folded caudal portion forming the caudal wall of the marsupial chamber; lower margin with series of barbed whip setae (Fig. 26 C) decreasing in length caudally. No setae on outer face of marsupium.</p><p>Penes (Fig. 26 D) simple, 0.5–0.6 × length of ischium 8. Terminally well rounded, only one barbed seta on distal third.</p><p>Pleon and pleopods (Figs 26 F – M, 27 A, 27 D). Pleomeres 1–5 showing 0.4, 0.3–0.4, 0.3–0.4, 0.3–0.4 and 0.4–0.5 times the length of pleomere 6, respectively. Paramedian pairs of muscle structures, visible as near-pentagonal images (arrows in Fig. 27 D), located in pleomeres 2–4 between neuronal cord and intestine. With some variation, length (without setae) increases in order of pleopods 2–5, 1 in females (Fig. 26 F – J) and in order of pleopods 2, 3, 1, (5 ≤ 4) in males (Figs 26 K – M, 27 A). Length of the respective pleopods 1–3, 5 approx. the same in both sexes, while pleopod 4 is mostly larger in males than in females. All pleopods of both sexes with 2–5 barbed setae (plus other setae). Apex of pleopods 1–3, 5 in both sexes and of pleopod 4 in most females with a short spine-like, bare seta. Apex of male pleopod 4 with none or one short seta in addition to the large modified seta, the latter reaching to or beyond pleopod 5 (Fig. 27 A); lateral margin of pleopod 4 with 0–3 short setae. A large seta at apex of pleopod 4 also found in many immatures and in three of 25 adult females inspected in this respect. Strong size differences of the apical seta between right and left pleopods 4 were also recorded. The pleopods alone clearly do not yield sufficient data for firm distinction between sexes in this species.</p><p>Tail fan (Fig. 27 B, C). Scutellum paracaudale short, triangular, tip narrowly rounded; not (Fig. 27 C) or only marginally, laterally covering part of telson. The exopod of uropods measures 0.9–1.1 (mostly 0.9) × length of endopod. In part due to inserting more caudally, the endopod extends by 0.04–0.2 × its length beyond exopod and 0.6–0.8 × beyond telson; exopod 0.5–0.7 × its length beyond telson. Exopod 5.3–6.8 × longer than wide; proximal 2 / 3 of lateral margin bare, distal third with 3–4 (mostly 3) setae increasing in size distally (Fig. 27 B). Diameter of the spherical fluorite statoliths is 15–28 μm (n = 20). Telson length 0.9–1.2 × its maximum width or 0.4–0.6 × length of pleomere 6. Telson length 2.1–4.2 × maximum thickness at anal bulge (Fig. 27 C), rendering the telson clearly wedge-shaped in lateral view. The apical spines are 4–8 % telson length.</p><p>Notes on the Cuban material.</p><p>The only specimen, an adult female, available from Cuba was completely dissected. Mesial antennular flagellum 4 - segmented, apical segment 1 / 6 flagellum length; length of antennal scale 3.3 × maximum width, terminal margin of scale with one long and one short seta, the long seta 0.6 × scale length, lateral margin of scale with one additional short seta; post-cervical lateral margin of carapace with projecting lobe; pleopod 1 with three setae, the largest seta 0.9 × pleopod length. The length of the longest seta at the terminal margin of the antennal scale is at the lower diagnostic limit of P. antillensis sp. nov., remaining features are well within limits. Numbers of segments and size of the apical segment of the mesial antennular flagellum exclude P. bahamensis .</p><p>DNA data.</p><p>Only two DNA sequences are available for Palaumysis in GenBank, at the 18 S nuclear locus: one for P. bahamensis (AM 422517) collected at Long Island, Bahamas and one for P. simonae (AM 422516) collected at Palau, Micronesia, both originating from Meland and Willassen (2007). The two Lesser Antilles populations of P. antillensis sp. nov. sampled here (Bequia and Guadeloupe) are geographically ≈ 400 km apart and ≈ 2000 km from the Cuban occurrence reported here. The Antillean populations are also at least 1500 km away from the known populations of P. bahamensis . The Bequia and Guadeloupe 18 S sequences (GenBank PV 990185 – PV 990200) obtained (from 16 individuals) are all identical with the total sequence length of 813 bp. Unfortunately, no DNA data were available for the more distant Cuban population.</p><p>When aligning the Antilles 18 S sequences with published ones, they are 0.37 % divergent from P. bahamensis . As reflected in divergence between P. bahamensis and P. simonae, which is 1.24 %, 18 S must be considered a conserved gene in Palaumysis as in most other mysids (Lejeusne and Chevaldonné 2006; Rastorgueff et al. 2014). The divergence data corresponds well with the minor morphological differences noted here between the W-Atlantic sister species P. bahamensis and P. antillensis sp. nov. compared with the differences between this W-Atlantic pair and the W-Pacific P. simonae .</p><p>Derivatio nominis.</p><p>The species name of P. antillensis sp. nov. is a Latinized adjective with a feminine ending, referring to the occurrence in islands of the Lesser Antilles.</p><p>Distribution and habitat</p><p>(Figs 10 I, 28). Type locality is the submarine Full Moon Cave (Station 4) in Bequia. The species was also recorded from the submarine Cathedral Cave (Station 13) in Guadeloupe and the anchialine El Brinco Cave (Station 23) in Cuba, total range 13– 22°N, 61– 81°W, 13–15 m depth. During the day the mysids formed dense swarms (Fig. 28) in dark recesses inside large caves. The species is classified as a troglophile or a stygophile.</p><p>Remarks.</p><p>The new species differs geographically and, as far as known, also genetically from the above described closest congener P. bahamensis by 3 - to 5 - segmented (vs 2 - segmented) mesial antennular flagellum with apical segment measuring 1 / 5–1 / 6 (vs 4 / 5) flagellum length, antennal scale with its longest seta longer (vs shorter) than 1 / 2 scale length, and apex of male pleopod 4 with one large and one short seta, the former bearing bilateral series of small cils from submedian to subapical portions vs apex with large smooth seta only.</p><p>The new species differs from the W-Pacific P. simonae (described above) by large seta accompanied by one or two short setae at the apex of the antennal scale (vs total of only one or two minute setae), apex of pleopod 1 with only moderately-sized setae in both sexes, no large seta (vs large plumose seta present in male) and length of antennal scale 3–4 × (vs 4–5 ×) maximum width.</p><p>The NW-Pacific P. philippinensis and P. pilifera share a (sub) rectangular posteroventral edge of the carapace and tip of antennal scale with only one seta vs well-rounded edge of carapace and tip of antennal scale with two or three setae in the new species.</p></div>	https://treatment.plazi.org/id/AB007AB1B0E25D0F90F6ECFC82D437FD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
ECE6D534F5615A1D8B269AB1CCA788B6.text	ECE6D534F5615A1D8B269AB1CCA788B6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Palaumysis Bacescu & Iliffe. A - D. Habitus 1986	<div><p>Genus Palaumysis Băcescu &amp; Iliffe, 1986</p><p>Figs 12, 13, 14, 15, 16, 17, 18, 19, 20, 21, 22, 23, 24, 25, 26, 27, 28</p><p>Palaumysis Băcescu &amp; Iliffe, 1986 b: 31 (diagnosis); Hanamura and Kase 2002: 254, 255 (revision, revised definition).</p><p>Diagnosis.</p><p>Modified after Hanamura and Kase (2002) for inclusion of features of P. bahamensis Băcescu &amp; Iliffe, 1986, and P. antillensis sp. nov. Palaumysinae with long, well-developed lateral flagellum of antennula with 3–5 apically blunt aesthetascs on basal segment in both sexes; antennal scale with a few setae, no spines, no teeth; carapace short, leaving posterior four or five thoracomeres mid-dorsally exposed (in adults); thoracic endopods 3–8 with unsegmented carpopropodus; pleopod 4 with long apical seta in adult males (rarely in adult females); endopod of uropods setose all around, longer than exopod, no spines; lateral margin of exopod with only few setae on distal half.</p><p>Type species.</p><p>Palaumysis simonae Băcescu &amp; Iliffe, 1986, by monotypy.</p><p>Species inventory.</p><p>Five species now included at a world-wide scale; bold italics highlight two species recorded from Caribbean caves: Palaumysis pilifera Hanamura &amp; Kase, 2003, P. philippinensis Hanamura &amp; Kase, 2002, P. antillensis sp. nov., P. bahamensis Pesce &amp; Iliffe, 2002, and P. simonae Băcescu &amp; Iliffe, 1986 . See also below global key to the genera and species of Palaumysinae .</p><p>Differentiation between Palaumysis species.</p><p>The now five known species of Palaumysis are well distinguished by a range of minor but significant characters. Three morpho-geographic groups may be distinguished: (1) the tropical W-Pacific P. simonae with the antennal scale bearing only one or two minute setae at the apex vs a large seta accompanied by 0–2 small setae at the apex in all remaining species; (2) NW-Pacific species, namely P. philippinensis and P. pilifera with a (sub) rectangular posteroventral edge of the carapace vs well-rounded edge in groups 1 and 3; (3) subtropical NW-Atlantic species, namely P. bahamensis and P. antillensis sp. nov., characterized by combination of a large seta accompanied by 0–2 small setae at the apex of the antennal scale as in (2) and a rounded posteroventral carapace edge as in (1).</p><p>Remarks.</p><p>Băcescu and Iliffe (1986 b) noted upon original description of P. simonae “ ... ± pentagonal zones on the ventral side of each pleonite ... ” and listed these “ zones ” among the main diagnostic features for the definition of the tribe Mancomysini Băcescu &amp; Iliffe, 1986, today considered a nomen nudum due to the missing type genus and, consequently, replaced by Palaumysinae Wittmann, 2013 . As shown above, near-pentagonal images are formed by pairs of paramedian muscle structures inside pleomeres, also found in P. antillensis sp. nov. (Fig. 27 D) and Bermudamysis speluncola (Fig. 36 E). Hanamura and Kase (2002) made additional corrections upon redescription based on types of P. simonae . Aesthetascs drawn by Băcescu and Iliffe (1986 b: fig. 2 A) as inserting on a prominence of the distal margin of the antennular trunk actually insert on the lateral antennular flagellum as usual in Mysidae (Fig. 38 A, B). Upon original description of P. bahamensis, Pesce and Iliffe (2002: 273, fig. 13) followed Băcescu and Iliffe (1986 b) by displacing the aesthetascs to the same position. Both author groups interpreted the respective mid-dorsal prominence as a rudiment of the (ventral) male lobe without discussing that this prominence is positioned dorsally and present in both sexes. In contrast, the prominence is here interpreted as the mid-dorsal lobe in subterminal position on the antennular trunk (Figs 24 A, 38 A); it bears no such aesthetascs but a few small barbed setae as in most species of Mysidae; moreover, this lobe bears teeth or spines in addition to setae in many genera other than Palaumysis . Upon original description of the closely related Gironomysis lalanai, Ortiz et al. (1997: fig. 12) drew the aesthetascs in the correct position but nonetheless followed the erroneous interpretation as remnants of the male lobe by Băcescu and Iliffe (1986 b).</p></div>	https://treatment.plazi.org/id/ECE6D534F5615A1D8B269AB1CCA788B6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
03D01721BF835DD8A916427B1B0BBD44.text	03D01721BF835DD8A916427B1B0BBD44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Palaumysis bahamensis Pesce & Iliffe 2002	<div><p>Palaumysis bahamensis Pesce &amp; Iliffe, 2002</p><p>Figs 12 K – M, 17 –20</p><p>Palaumysis bahamensis Pesce &amp; Iliffe, 2002: 273–275, figs 7–13 (original description); Meland and Willassen 2007: fig. 3 (phylogeny, rRNA sequencing); Chevaldonné et al. 2015: fig. 5 (molecular systematics).</p><p>Material examined.</p><p>Bahamas • 1 ♀ ad. (BL = 1.42 mm; NHMW -ZOO-CR-31426) on slides, from submarine cave ( Station 21); this station is ≈ 260 km SE of the type locality .</p><p>Types, type locality, and distribution</p><p>(Fig. 10 J). The types are missing since the 2009 l’Aquila earthquake, when they were probably destroyed (Diana Galassi pers. comm. 2025). Type locality by primary monotypy is Bahamas, South Andros Island, Grassy Cay, submarine cave Atlantis Blue Hole, 23°50.12'N, 77°27.76'W, depth 60–70 m (Pesce and Iliffe 2002; coordinates estimated). Efforts to obtain materials from the type locality repeatedly failed. This mysid is still known only from marine caves in the Bahamas. The present record from a submarine cave at Long Island (Bahamas, Station 21) extends the known range to 23– 24°N, 75– 77°W, depth 45– 70 m.</p><p>Revised diagnosis.</p><p>Essentially based on adult females and on a few characteristics of the only male specimen known from the description by Pesce and Iliffe (2002): Palaumysis with mesial antennular flagellum 2 - segmented, apical segment 4 / 5 flagellum length; length of antennal scale 3 × maximum width, scale terminally with one long and one short seta, the long seta &lt;1 / 2 scale length, lateral margin of scale with one short seta; post-cervical lateral margins of carapace with projecting lobes; apex of male pleopod 4 with large smooth seta reaching to pleopod 5, no additional seta.</p><p>Supplementary description.</p><p>Based on single adult female (BL = 1.42 mm) from Long Island, Bahamas. Pesce and Iliffe (2002) reported BL values of 1.9–2.5 mm for five specimens without indication of measurement methods and differentiation between sexes. Pigment not retracted from crystalline bodies of the eye.</p><p>Cephalothorax (Figs 17 A – C, 18). Basal segment of the 2 - segmented mesial antennular flagellum without setae; terminal segment with three short whip setae along mesial margin; apex with one long and three shorter, widely distributed whip setae (Fig. 17 B). Carapace (Figs 17 A, 18 D) short, leaving ultimate four thoracomeres mid-dorsally exposed; mid-terminal margin marked by arrow in Fig. 17 A. Mouthparts as in Fig. 18 E – K, foregut as in Fig. 19.</p><p>Thoracopods (Fig. 20 A – I). Flagellum of exopods 1–8 with 6, 6 or 7, 7, 6, 6, 6, 7, and 7 segments, respectively. Endopods 3–8 with unsegmented carpopropodus (Fig. 20 E, G). Dactyli 1, 2 with normal-sized, weakly bent strong nail (Fig. 20 B, D); concave margin of nail 1 weakly microserrated on distal third by stiff cils (Fig. 20 B), nail 2 with longer stiff cils along distal half (Fig. 20 D). Nails 3–8 long, needle-shaped, weakly bent; the needles microserrated by minute teeth along median to subapical portions (Fig. 20 F, H). Thoracic endopods 3–8 with minute triangular scales on ischium and merus as in P. antillensis sp. nov. (Fig. 26 D, E).</p><p>Marsupium (Figs 17 A, 20 G, 20 I). Oostegites 1, 2 from thoracopods 7, 8 representing simple unfolded plates. Lower margin (to the left in Fig. 20 G) with series of comparatively few whip setae furnished with a few barbs at handle and minute denticles (spines) along flagellum (Fig. 20 I); no other setae modifications.</p><p>Pleon. As in P. simonae (Fig. 16 A, B), each pleurite 6 of P. bahamensis bears an apically blunt to acute, triangular scutellum paracaudale. In dorsal view the scutella mark the caudolateral edges of the pleon (Fig. 17 D).</p><p>Pleopods (Fig. 20 J – M). Length (without setae) decreasing from pleopods 1 to 3 and increasing from 3 to 4, pleopod 5 subequal to pleopod 4. Pleopods 1–5 with 2–4 small barbed setae on distal half and one smooth seta at or near apex. The smooth seta strongly spine-like as in Fig. 20 J – M. The smooth seta of pleopod 4 approx. as long as in pleopods 2, 3.</p><p>Tail fan (Fig. 17 D). The endopod of uropods extends 0.1–0.2 × its length beyond exopod and 0.7–0.8 × beyond telson; exopod 0.5–0.7 × its length beyond telson. Exopod 7.0 × longer than wide; proximal 2 / 3 of lateral margin bare, distal third with three setae increasing in length distally. Diameter of the spherical fluorite statoliths is 18 μm (n = 2). Telson 1 / 2 as long as pleomere 6, its length 1.1 × maximum width.</p></div>	https://treatment.plazi.org/id/03D01721BF835DD8A916427B1B0BBD44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
92F49FA0ECA159539974F0B7D96FC3AD.text	92F49FA0ECA159539974F0B7D96FC3AD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Palaumysis simonae Bacescu & Iliffe. Modified 1986	<div><p>Palaumysis simonae Băcescu &amp; Iliffe, 1986</p><p>Figs 12 N – P, 13 –16</p><p>Palaumysis simonae Băcescu &amp; Iliffe, 1986 b: 31–34, fig. 2 (original description); Hanamura and Kase 2002: 255–257, figs 1, 2 (revised description); Hanamura and Kase 2003: 151, 152, Table 1, figs 4 a, b, 5 (supplementary description); Hanamura and De Grave 2004: 66, 67 (records); Meland and Willassen 2007: fig. 3 (phylogeny, rRNA sequencing); Petrescu and Wittmann 2009: 58 (type repository, etymology); Wittmann 2013: Table 1, fig. 4 A (male genital apparatus); Wittmann et al. 2014: figs 5, 42 (female habitus, molecular phylogeny); Chevaldonné et al. 2015: fig. 5 (molecular systematics); Wittmann and Ariani 2019: suppl. (statolith composition, biogeography); Wittmann 2023: table SS 1 (eye structure); Kou et al. 2025: figs 8–10 (molecular phylogeny).</p><p>Note.</p><p>This non-Caribbean species treated here for differentiation from P. bahamensis and P. antillensis sp. nov.</p><p>Types examined.</p><p>Palau Archipelago • holotype, ♀ ad. (BL = 1.20 mm; MINGA 49449) from Chandelier Cave (Station 36: type locality) • paratypes, 2 ♀♀ ad. (BL = 1.10–1.35 mm; MINGA 49448), sampling data as for holotype .</p><p>Non-types examined.</p><p>Palau Archipelago • 1 ♀ ad., 4 ♂♂ ad., 4 imm. (MNHN -IU-2025-2836) from unnamed cave near Soft Coral Arch (Station 37) • 4 ♀♀ ad., 2 ♂♂ ad., 2 imm. (NHMW -ZOO-CR-31419), sampling data as for preceding • 3 ♀♀ ad., 3 ♂♂ ad., 1 ♀ subad., 2 imm. (ZMH -K-066764), sampling data as for preceding • 1 ♀ ad., 1 ♂ ad., 2 imm. (MNHN -IU-2025-2837) from Ishura Cave (Station 38) • 2 ♀♀ ad., 1 ♂ ad., 1 imm. (NHMW -ZOO-CR-31427), sampling data as for preceding • 1 ♂ ad., 3 imm. (ZMH -K-066765), sampling data as for preceding .</p><p>Distribution.</p><p>This species so far known only from submarine caves in the Palau archipelago.</p><p>Revised diagnosis.</p><p>Palaumysis with mesial antennular flagellum 2 - to 4 - segmented, basal segment shortest, apical segment with 4 / 5 flagellum length in 2 - segmented flagella vs 1 / 5 in 3 - to 4 - segmented flagella; length of antennal scale 4–5 × maximum width, apex with one or two (mostly 2) minute setae, lateral margins bare; post-cervical lateral margins of carapace without or with small, projecting lobe; apex of pleopod 1 with large plumose (barbed) seta only in males caudally overreaching pleopod 2; apex of male pleopod 4 with one large and one short seta, the former reaching at most to apex of pleopod 5 and bearing several minute cils near 2 / 3 length, lateral margin of pleopod 4 with one or two short setae; pleopod 4 mostly normal in adult females, but male-like with large apical seta in a few specimens with fully developed brood pouch.</p><p>Notes on holotype</p><p>(Fig. 13 B). Small brush of aesthetascs on basal segment of mesial antennular flagellum. Cornea spherical with diameter measuring 10 % BL in lateral view. Pigment not retracted from cornea. Paramedian pairs of muscle structures, visible as one or two near-pentagonal images (as in Fig. 16 D) per segment, located in pleomeres 2–5 between neuronal cord and intestine.</p><p>Supplementary description.</p><p>♀♀ ad. with BL range 1.42–1.77 mm, TL of 1.52–1.93 mm (n = 11) and ♂♂ ad. with BL range 0.97–1.62 mm, TL of 1.25–1.88 mm (n = 12) in the present material. Hanamura and Kase (2002) obtained BL data within the present ranges. Băcescu and Iliffe (1986 b) reported TL of 1.6–2 mm without differentiation between sexes.</p><p>Cephalothorax (Fig. 13). Basal segment of the 2 - to 4 - segmented mesial antennular flagellum without setae; second to penultimate segments (if more than two segments) each with one or two short whip setae at disto-mesial edge; apex with one long plus 3–5 short whip setae (Fig. 14 A). Carapace short (Fig. 13 A), leaving four or five ultimate thoracic segments dorsally exposed. Pigment not or only weakly retracted from crystalline bodies of the eye. Labrum rostrally convex, broadly rounded.</p><p>Thoracopods. Flagellum of exopods 1–8 with 6, 7, 7, 7, 7, 7, 7, and 6 or 7 segments, respectively. Endopods 3–8 with unsegmented carpopropodus. Dactylus 1 with strong nail, length as in Fig. 25 C though less bent; distal half of inner (concave) margin of this nail microserrated by minute stiff cils; nails 2–8 as given below for P. antillensis sp. nov. (Fig. 25 F, H, K). Thoracic endopods 3–8 with minute triangular scales on ischium and merus as in P. antillensis sp. nov. (Fig. 26 D, E).</p><p>Pleon (Figs 15, 16 A, B, D). Paramedian pairs of muscle structures as in holotype. Sternites 4, 5 of both sexes with small, obliquely transverse, ventral elevation between pleopods and lateral margin (small arrows in Fig. 15 A, B, D). These elevations individually varying from widely blunt (sternite 4 in Fig. 15 B) to ending in or bearing a spine-like, caudally facing process (sternite 5 in Fig. 15 D). Sternites 1–3 without such processes in both sexes. Pleurite 6 bilaterally shortly in front of caudal margin with apically blunt to acute, triangular scutellum paracaudale in lateral position, laterally (Fig. 16 A) only marginally covering part of the telson. Both scutella marking the distolateral edges of the pleon in ventral (arrow in Fig. 16 B) as well as dorsal views.</p><p>Pleopods of adult females (Fig. 15 A). Length (without setae) decreasing from pleopods 1–3 and increasing from 3–5. Pleopods 1–5 with 2–4 small barbed setae on distal half and one smooth seta at or near apex, the latter varying individually in length and from seta-like to strongly spine-like (as in Fig. 15 E) in all pleopods. The smooth seta of pleopod 4 in two of 11 available adult females attaining almost the length of the apical seta typical of males.</p><p>Pleopods of adult males (Fig. 15 B – E). Length decreasing from pleopods 1–3 and increasing from 3 to 4. Pleopod 4 longest, pleopod 1 second longest, pleopod 5 subequal to pleopod 1. Distal half of pleopod 1 bilaterally with total of four or five small barbed setae; apex with one large plumose seta (Fig. 15 C), the latter approx. as long or longer (Fig. 15 B) than the large apical seta of pleopod 4. Pleopods 2, 3, 5 each with 2–4 small barbed setae on distal half and one smooth seta at or near apex, the latter varying individually in length and from seta-like to strongly spine-like (Fig. 15 E) only in these pleopods. Pleopod 4 with one large seta bearing several minute cils near 2 / 3 length and with an additional small barbed seta at apex; distal half of lateral margin with two or three barbed setae, mesial margin bare (Fig. 15 D).</p><p>Tail fan (Fig. 16 C). The endopod of uropods extends 0.08–0.13 × its length beyond exopod and 0.6–0.7 × beyond telson; exopod 0.5–0.7 × its length beyond telson. Statoliths composed of fluorite, diameter 20–28 µm (n = 23 including 15 measurements by Wittmann and Ariani 2019). Exopod 5.2–6.6 × longer than wide; proximal 2 / 3 of lateral margin bare, distal third with three setae increasing in size distally (Fig. 16 C). Telson length 0.8–1.2 × maximum width or 0.4–0.6 × length of pleomere 6. The apical spines are 8–13 % telson length.</p></div>	https://treatment.plazi.org/id/92F49FA0ECA159539974F0B7D96FC3AD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
41220AEDE5CF5B3A9CB88A893D01DC76.text	41220AEDE5CF5B3A9CB88A893D01DC76.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parvimysis Brattegard 1969	<div><p>Genus Parvimysis Brattegard, 1969</p><p>Fig. 6</p><p>Parvimysis Brattegard, 1969: 74, table 5 (diagnosis); Bowman 1977: 37 (downgrading to subgenus level, in key); Henderson and Bamber 1983: 143 (retaining genus level); Nouvel et al. 1999: 79 (in list of genera); Wittmann 2018: 542, 543 (diagnosis, in key, vaterite statoliths); Price 2020: 974, 979, 980 (in key to genera, key to species); Wittmann 2020: 3, 4 (revision, species inventory, Caribbean, West Atlantic, key to species).</p><p>Antromysis (Parvimysis): Bowman 1977: 37; Bowman 1981 a: Table 1 (in list); Modlin 1987 a: 110 (species record); Camp et al. 1998: 126 (in checklist).</p><p>Antromysis: Müller 1993: 209, 210 (listed Parvimysis in synonymy); Vázquez-Bader 2000: 48 (attributed the type species Parvimysis bahamensis Brattegard, 1969, to Antromysis).</p><p>Short diagnosis.</p><p>Diamysini with eyes well-developed, cornea well pigmented; antennular trunk terminally with the usual two flagella in both sexes, no accessory flagellum; antennal scale setose all around, with small terminal segment; thoracic endopods 1–8 with unsegmented carpus and propodus; female pleopods 1–5 and male pleopods 1–3, 5 representing simple setose unsegmented rods; male pleopod 4 biramous, endopod forming an unsegmented bilobate plate, exopod 2 - to 3 - segmented, apical segment with a single strong seta, in certain species accompanied by a small seta, remaining segments without seta; uropods without spine; telson small, subtriangular to trapezoid, terminally truncate to distinctly emarginated; lateral margins with spines, latero-terminal edges each with one larger spine; terminal margin bare or armed with diverse cuticle structures.</p><p>Species inventory.</p><p>Fourteen species are acknowledged; bold italics highlight two species recorded from Caribbean caves: Parvimysis bahamensis Brattegard, 1969 (type species), P. almyra Brattegard, 1977, P. pisciscibus Henderson &amp; Bamber, 1983, P. amazonica Wittmann, 2018, P. fittkaui Wittmann, 2018, P. fluviatilis Wittmann, 2018, P. lacustris Wittmann, 2018, P. macrops Wittmann, 2018, P. tridens Wittmann, 2018, P. brattegardi Wittmann, 2020, P. laminata Wittmann, 2020, P. nuda Wittmann, 2020, P. ornata Wittmann, 2020, and P. pricei Wittmann, 2020 . See also key below to species from Caribbean caves.</p></div>	https://treatment.plazi.org/id/41220AEDE5CF5B3A9CB88A893D01DC76	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
C065A6E0DDEE5B1B9B6AFA4F13B2904E.text	C065A6E0DDEE5B1B9B6AFA4F13B2904E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parvimysis brattegardi Wittmann 2020	<div><p>Parvimysis brattegardi Wittmann, 2020</p><p>Fig. 6 A, B</p><p>Parvimysis brattegardi Wittmann, 2020: 14–18, tables 1, 2, figs 6, 7 (original description, Curaçao); Wittmann 2023: table SS 1 (sensory organs, trogloxene status).</p><p>Material examined.</p><p>Leeward Antilles — Curaçao • 22 ♀♀ ad. (BL = 2.5–3.5 mm), 44 ♂♂ ad. (BL = 1.8–2.5 mm), 21 subad., 10 imm., 6 juv. from swarms in entrance of two small caves ( Station 17) .</p><p>Short diagnosis.</p><p>Parvimysis with subtriangular, apically rounded rostrum; antennal scale with basal segment not reaching to end of antennular trunk; eyes moderately sized, maximum cornea diameter 1.5–1.7 × length of terminal segment of antennular trunk in dorsal view; exopod of male pleopod 4 three-segmented, ending in large modified seta plus a minute lobe bearing a minute seta; telson trapezoid to subtriangular with weakly concave terminal margin lined by 4–12 laminae that are 0.2–0.5 × length of latero-apical spines.</p><p>Distribution</p><p>(Fig. 3 G). In sublittoral marine waters of Curaçao, 12°N, 69°W, depth 1–32 m, swarms on reef flat and slope, among algae, in rock recesses and small caves (Wittmann 2020, 2023).</p></div>	https://treatment.plazi.org/id/C065A6E0DDEE5B1B9B6AFA4F13B2904E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
0B7A62550BF75FD99762F18D41145071.text	0B7A62550BF75FD99762F18D41145071.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parvimysis laminata Wittmann. Slightly 2020	<div><p>Parvimysis laminata Wittmann, 2020</p><p>Fig. 6 C, D</p><p>Parvimysis sp. B: Wittmann and Ariani 2019: fig. 1 E, F, suppl. (vaterite statoliths, biogeography).</p><p>Parvimysis laminata Wittmann, 2020: 10–14, tables 1, 2, figs 3–5 (original description, Curaçao); Wittmann 2023: table SS 1 (sensory organs, trogloxene status).</p><p>Material examined.</p><p>Lesser Antilles — SW-Bequia • 4 ♀♀ ad. (ZMH -K-066766) from Full Moon Cave (Station 5) • 3 ♀♀ ad., 1 ♂ ad. (NHMW -ZOO-CR-31414), sampling data as for preceding • 2 ♀♀ ad., 2 ♂♂ ad. (MNHN -IU-2025-2839), sampling data as for preceding — Guadeloupe • 9 ♀♀ ad. (BL = 2.6–2.9 mm), 13 ♂♂ ad. (BL = 1.7–2.4 mm) (MINGA MYS 458) from Cathedral Cave (Station 9) • 3 ♀♀ ad. (BL = 2.4–2.8 mm), 3 ♂♂ ad. (BL = 1.9–2.3 mm) (MNHN -IU-2025-2840) from Barracuda Cave (Station 11) . Leeward Antilles — Curaçao • 10 ♀♀ ad. (BL = 2.7–3.0 mm), 10 ♂♂ ad. (BL = 2.1–2.5 mm) (ZMBN 135401) inside marine cave at Playa Lagun (Station 19 — type locality) • 20 ♀♀ ad., 14 ♂♂ ad. (ZMH -K-066774) from entrance of small caves at Sun Reef (Station 17) • 15 ♂♂ ad., 17 ♀♀ ad., 2 imm. (MNHN -IU-2025-2838), sampling data as for preceding .</p><p>Short diagnosis.</p><p>Parvimysis with short, broadly rounded rostrum; antennal scale with basal segment extending beyond antennular trunk; eyes very large, maximum cornea diameter 1.8–2.4 × length of terminal segment of antennular trunk in dorsal view; exopod of male pleopod 4 three-segmented, ending in large modified seta plus a minute lobe bearing a minute seta; telson trapezoid, terminally truncate or slightly concave, its terminal margin lined by 9–16 laminae that are 0.4–0.8 × length of latero-apical spines.</p><p>Distribution and habitat</p><p>(Fig. 3 H). In euhaline coastal waters of the Leeward Antilles (Curaçao and Bonaire: Stations 15, 17, 19) and the Lesser Antilles (Bequia and Guadeloupe: Stations 5, 9, 11), total range 12– 16°N, 61– 69°W, 1–32 m depth. Recorded from the open sea floor where it hovers in swarms and loose aggregations a few cm above the substrate, also found over and inside rock recesses (Wittmann 2020). Present records from the semi-dark zone of diveable caves (Stations 5, 9, 11). The mysids appeared colorless in situ. In the Barracuda Cave (Station 11) they were found close to the reddish Bermudamysis caribbaea sp. nov. Classified as trogloxene by Wittmann (2023), but inclusion of the new records points rather to a troglophile status.</p></div>	https://treatment.plazi.org/id/0B7A62550BF75FD99762F18D41145071	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
C0912470918E59B1AAC847036B8F2B28.text	C0912470918E59B1AAC847036B8F2B28.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Platyops Bacescu & Iliffe. A - D. Right 1986	<div><p>Genus Platyops Băcescu &amp; Iliffe, 1986</p><p>Figs 29 - 32</p><p>Platyops Băcescu &amp; Iliffe, 1986 a: 100 (diagnosis); Pesce et al. 1994: 117 (in list of subterranean fauna); Fukuoka 2006: fig. 1 (taxonomy, geographic distribution); San Vicente and Monniot 2014: 334, 341 (taxonomy, in key); Wittmann et al. 2014: 341 (taxonomy, transfer to Mysidetini); Daneliya 2021: 45 (revision, transfer to Heteromysini).</p><p>Type species.</p><p>Platyops sterreri Băcescu &amp; Iliffe, 1986, by monotypy. Syntypes here examined and lectotype defined.</p><p>Revised diagnosis.</p><p>Mysidetini with large, subquadrangular to subcircular, flattened eyestalks, no eye papilla; cornea or its rudiments implanted laterally; antennular trunk with normal setae (including, if any, whip setae), no flagellate spines, no backward-directed modified setae; antennal scale, pleopods and uropods setose, no spines, no teeth; thoracic endopod 3 non-prehensile, with 2 - to 3 - segmented carpopropodus; endopods 4–8 normal, praeischium without distal projection, ischium and merus without flagellate spines; marsupial chamber formed by oostegites from sympods 7–8; penes tube-like, &lt;1 / 5 BL; structure of pleopods non-dimorphic, reduced to unsegmented setose plates with inconspicuous rudiments of pseudobranchial lobes; uropods normal, both rami undivided; telson trapezoid to triangular with bare proximal half, only distal ≤ 1 / 2 of lateral margins with spines, its transverse terminal margin not incised or with an only minute incision, no laminae, no setae.</p><p>Species inventory.</p><p>Four species are acknowledged and given in below key to species; bold italics highlight two species recorded from Caribbean caves: Platyops sterreri Băcescu &amp; Iliffe, 1986, P. simplex (Hanamura &amp; Kase, 2001), P. stenoura (Hanamura &amp; Kase, 2004) and P. kumejimensis (Shimomura &amp; Fujita, 2020), comb. nov. The latter species recombined from Heteromysoides kumejimensis Shimomura &amp; Fujita, 2020, due to conforming with the revised diagnosis of the genus Platyops above, unlike Heteromysis and Deltamysis . This recombination is necessary since Daneliya (2021) synonymized the genus Heteromysoides with Heteromysis and divided it between Heteromysis, Platyops, and Deltamysis Bowman &amp; Orsi, 1992 .</p><p>Diagnosis and affiliation of the genus Platyops .</p><p>Daneliya (2021) stated upon revision of the genus Platyops “ The most characteristic feature of this genus is the eye structure, particularly the presence of a strong anterior spine, directed forward ”. As shown below, there is no such spine in the types of the type species P. sterreri . Actually, a spine is feigned only in lateral view (Figs 30 A, 31 A), but clearly no spine-like process is visible in dorsal view (Fig. 31 C). The laterally wide, transverse, anterior margin of the distally wedge-shaped eyestalk was correctly interpreted already upon original description by Băcescu and Iliffe (1986 a) as “ ... strongly flattened chisel-like … ”. The here-confirmed absence of a spine clearly contradicts the revision by Daneliya (2021). For the revised diagnosis of the genus, see above.</p><p>By including Heteromysoides dennisi Bowman, 1985, whose eyestalks bear a spine (Fig. 43 A) and which has a prehensile endopod 3 (Fig. 43 B), Daneliya (2021) amplified the diagnosis of Platyops in favor of transferring this genus from the tribe Mysidetini to the Heteromysini . The present examination of the type specimens of the type species P. sterreri at the MINGA confirms a normal non-prehensile structure of this endopod and the absence of an eyestalk spine (Figs 30, 31). Consequently, the genus Platyops is here retransferred to the Mysidetini, where it had been previously placed by Wittmann et al. (2014), mainly based on the non-prehensile structure of thoracic endopod 3 (Figs 30 A, 30 B, 31 B).</p><p>Affiliation of Heteromysoides kumejimensis Shimomura &amp; Fujita, 2020 .</p><p>Daneliya (2021) synonymized the genus Heteromysoides Băcescu, 1968, with the genus Heteromysis S. I. Smith, 1873, based on morphology of the type species Heteromysoides spongicola Băcescu, 1968 . He did not mention the publication of Shimomura and Fujita (2020), so Heteromysoides kumejimensis may have escaped the revision by Daneliya (2021) merely by temporal overlap. This species fits the above definition of the genus Platyops by the antennular trunk bearing only normal setae, non-prehensile thoracic endopod 3 with 3 - segmented carpopropodus, non-dimorphic pleopods reduced to setose plates and by a trapezoid, terminally slightly convex telson with bare basal half (Fig. 29 G). Structure of thoracic endopod 3 and telson exclude affiliation with Heteromysis . Rather than perpetuation of a ‘ ghost’ combination, we propose recombination as Platyops kumejimensis (Shimomura &amp; Fujita, 2020) .</p></div>	https://treatment.plazi.org/id/C0912470918E59B1AAC847036B8F2B28	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
FD303C23C5835C8FB0AF582C16CFD1F6.text	FD303C23C5835C8FB0AF582C16CFD1F6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Platyops stenoura (Hanamura & Kase 2004)	<div><p>Platyops stenoura (Hanamura &amp; Kase, 2004)</p><p>Fig. 29 J, K</p><p>Heteromysis stenoura Hanamura &amp; Kase, 2004: 2148–2151, figs 2, 3 (original description, Grand Cayman Island).</p><p>Platyops stenoura: Daneliya 2021: 45, 46 (revised assignment, new combination).</p><p>Short diagnosis.</p><p>Platyops with eyestalks forming thick plates with sharp spiniform mid-anterior projection, comparatively small cornea implanted laterally, cornea composed of numerous densely-set ommatidia; antennal scale undivided; thoracic sympod 3 subequal to sympod 2; endopod 3 less stout than endopod 2, endopod 3 non-prehensile, with 3 - segmented carpopropodus; penes tube-like, ≈ 1 / 6 BL; endopod of uropods approx. as long as exopod; telson trapezoid, length without spines 1.8 × maximum width near basis, terminal width 1 / 4 maximum width; telson with bare proximal half, distal half with four or five spines on each lateral margin; transverse terminal margin with pair of large distolateral spines flanking a paramedian pair of spines with 1 / 2–2 / 3 length of the distolateral spines.</p><p>Distribution</p><p>(Fig. 10 K). Known only from the submarine cave ‘ Mouse Trap’ (type locality) ≈ 400 m off the western coast of Grand Cayman Island, 19°19.25'N, 81°23.38'W, depth 9–11 m (as Heteromysis stenoura in Hanamura and Kase 2004).</p></div>	https://treatment.plazi.org/id/FD303C23C5835C8FB0AF582C16CFD1F6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
4FC238B257DD53B98DFA9679C0867D86.text	4FC238B257DD53B98DFA9679C0867D86.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Platyops sterreri Bacescu & Iliffe 1986	<div><p>Platyops sterreri Băcescu &amp; Iliffe, 1986</p><p>Figs 29 A – D, 30 –32</p><p>Platyops sterreri Băcescu &amp; Iliffe, 1986 a: 100–102, fig. 2 A – L (original description); Pesce et al. 1994: 117 (in list of stygophile species); Wittmann and Ariani 2019: suppl. (statolith composition); Daneliya 2021: 45, 46 (taxonomy); Iliffe and Calderón-Gutiérrez 2021: Table 1 (stygobiont species, Bermuda); Wittmann 2023: table SS 1 (non-visual eyestalk organs).</p><p>Type material examined.</p><p>Bermuda • Here designated lectotype, ♀ ad. (BL = 1.9 mm, TL = 2.4 mm, MINGA MYS 448, Fig. 30 A) from Castle Grotto (Station 28) • paralectotype, 1 damaged ♀ ad. (BL = 2.1 mm, MINGA MYS 449), sampling data as for lectotype .</p><p>Other material.</p><p>Bermuda • 1 ♀ ad. (also with BL 2.1 mm, MINGA MYS 451) from Walsingham Cave (Station 29) — Lesser Antilles Guadeloupe • 1 ♂ ad. (BL = 2.0 mm) on slides (NHMW -ZOO-CR-31428) from Cathedral Cave (Station 14) .</p><p>Identification of the types.</p><p>Previously, there was no entry in the inventory of the MINGA, and the species is therefore not listed by Petrescu and Wittmann (2009). Vials with handwritten labels by Mihai Băcescu were recently identified by Iorgu Petrescu (pers. comm. 2025) as containing the syntypes of P. sterreri .</p><p>An additional vial with one female specimen (MINGA MYS 451) is labeled “ Sus 1 ♀ j ..... st. 6 Platyops mirandus ”, whereby “ st. 6 " refers to Walsingham Cave in Băcescu and Iliffe (1986 a). Platyops mirandus is an unpublished nomen nudum, and this material therefore cannot considered part of the type series. Nonetheless, this specimen is here determined as P. sterreri .</p><p>Revised diagnosis.</p><p>Platyops with eyestalks forming thick plates with distal 1 / 5–1 / 6 obliquely truncate in the transversal plane, resulting in dorsal face extending more anteriorly than the ventral face, no eye papilla, no spine, no teeth; antennal scale with small apical segment; thoracic sympod 3 subequal to sympod 2; endopod 3 less stout than endopod 2, endopod 3 not prehensile, with 2 - to 3 - segmented carpopropodus; penes tube-like, only 1 / 20 BL; endopod of uropods approx. as long as exopod; telson trapezoid, length without spines 1.3–1.5 × maximum width near basis, terminal width 1 / 4 maximum width; telson with proximal 2 / 3 bare, distal 1 / 3 of each lateral margin with two or three spines, truncate terminal margin with pair of large distolateral spines flanking a paramedian pair of shorter spines, minute median notch present.</p><p>Supplementary notes on females.</p><p>Lectotype from Castle Grotto (Station 28): rostrum measures 3 % BL, carapace without rostrum 30 %, cephalothorax 38 %, pleon without telson 49 %, and telson 10 %. Eyestalks form thick plates with distal 1 / 5 obliquely truncate in the transversal plane. Eye 2.0 × longer than wide, structure as described below for males. Cornea fully functional, lens-shaped in dorsal view (as in Fig. 31 C, D), diameter 0.10 mm, length (thickness) 0.06 mm; this is 1 / 2 length of eyestalk (cornea not included). Cornea dorsally flattened and even flatter ventrally as seen in lateral view, length 1.6 × maximum width. Pigment partly retracted from cornea. Thoracic endopod 3 non-prehensile, thicker than endopods 5, 7. Carpopropodus 3, 5, 7 with 3, 3, 4 segments, respectively, slenderness increasing caudally. Pleopod 5 measures 1 / 16 BL, reaching 1 / 2 length of pleomere 6.</p><p>Non-type from Walsingham Cave (Station 29): carpopropodus of thoracic endopods 3–8 with 3, 6, 4, 4, 4 and 4 segments, respectively. Endopod 4 more slender than the remaining endopods.</p><p>First description of the male.</p><p>Based on non-type from Guadeloupe (Station 14). Cephalothorax comprises 36 % BL, pleon (without telson) 44 %, telson 17 %, carapace (without rostrum) 31 %, and rostrum 3 %. Clypeus pentagonal, wider than long (lower dashed line in Fig. 32 A). Thoracic sternites without mid-ventral processes, no spines, no setae.</p><p>Carapace. Rostrum broadly rounded, as long as 0.4 × the terminal segment of antennular trunk in dorsal view. The cervical sulcus forms a deep transverse incision (Fig. 31 A). The post-cervical part of carapace as long as 1.2 × cervical part measured along midline without rostrum. Caudolateral margin of carapace broadly rounded; posterior margin slightly concave, leaving the ultimate thoracomere mid-dorsally exposed.</p><p>Eyes (Fig. 31 A, C, D). Eye 11 % BL, subquadrangular with transverse, weakly convex, anterior margin in dorsal view (Fig. 31 C, D). Eye forming a 1.8 × longer than wide, proximally rectangular trapezoid with obliquely truncate distal margin; this renders dorsal margin longer than ventral margin in lateral view (Fig. 31 A). Cornea fully functional, lens-shaped in dorsal view (Fig. 31 C, D), diameter quite variable, 0.6–0.9 × eyestalk length (cornea not included), cornea length (thickness) ≈ 1 / 3 length of eyestalk. Cornea oval in lateral view, length 1.9 × maximum width (ventral margin covered by left antennular trunk in Fig. 31 A).</p><p>Antennula (Fig. 32 A). Antennular trunk 3 - segmented, well setose, no spines, no teeth. Trunk extends 0.5–0.6 × its length beyond eyes in normal orientation. Measured along dorsal midline, the basal segment is 42 % trunk length, median segment 20 %, and terminal segment 38 %. Lateral margin of basal segment at half-length with two minute barbed setae; subterminally with a rectangular, distally setose lobe anteriorly approaching but not extending beyond rostral margin of the median segment. On dorsal face near mesial margin distally with two setose apophyses. Dorsal face of median segment disto-mesially with one setose apophysis. Terminal segment with the usual mid-dorsal lobe bearing a few barbed setae, no spines, no teeth. Lateral margin bare, disto-mesial edge with diverse barbed and smooth setae. Appendix masculina well setose despite its small size, only 8 % length of terminal segment. Flagella damaged.</p><p>Antenna (Fig. 32 B). Sympod 2 - segmented, no setae. Distolateral edge extended into a tooth-like projection. Sympod with two triangular acutely pointed teeth subterminally below antennal scale; one digitiform lobe above basis of peduncle (dashed line in Fig. 32 B). Peduncle 3 - segmented, basal segment contributes 17 % to total length of peduncle, median segment 39 %, and terminal segment 44 %. Antennal scale with small terminal segment, total length of scale 5 × maximum width. Scale extends 1 / 4 its length beyond antennal peduncle and 0.2–0.3 × its length beyond antennular trunk.</p><p>Mouthparts. Labrum rostrally rounded. Proximal segment of left mandibular palp 14 % palp length, no setae. Length of median segment 2.7 × its maximum width and 64 % palp length. Central half of mesial margin of median segment with three basally barbed whip setae, central half of lateral margin with four setae of that type, lateral margin with an additional subterminal seta, transverse terminal margin with two setae on mesial corner. Terminal segment well setose, 22 % palp length. Pars molaris of left mandible with well-developed grinding surface. Pars incisiva and digitus mobilis each with ≈ 3 teeth, pars centralis with four spines bearing stiff bristles. Right mandible, labium and both maxillulae damaged. Distal 2 / 3 of endopod of maxilla all around well setose; its distal segment 1.7–1.8 × as long as maximum width; proximal segment mesially with four basally barbed whip setae with thick handle. Lateral margin of exopod with 10 or 11 plumose whip setae, terminal margin with two such setae.</p><p>Foregut. Gross structure as in Fig. 23. Lateralia anteriorly each with apically pronged spines, with bare shaft; more caudally with group of four or five unilaterally weakly serrated spines as in Fig. 23 D. Dorsolateral infoldings each with pair of long spines unilaterally bearing series of small teeth increasing in size distally along distal 4 / 5 of shaft.</p><p>Thoracopods (Figs 30 B, 31 B). Width decreases from thoracic sympod 1 to 5 and then increases from 5 to 8. Flagellum of exopods 3–7 with 8, 9, 9, 9, 8 segments, respectively, remaining exopods damaged or broken. Basal plates expanded, 2–3 × as long as wide, distolateral corner well rounded. Total length of endopods increases in series of endopods 1–3, 6, 8, remaining endopods broken. Thoracopod 1 with leaf-like, smooth epipod. Tip of coxa of endopod 1 with one short whip seta with barbed handle. Basis with large, prominent endite that is densely setose on mesial and terminal margins. Ischium and merus with short endites. Nail 1 thick, slightly bent, approx. as long as dactylus. Basis of endopod 2 with weakly developed mesial endite bearing two sparsely barbed setae. Combined praeischium plus ischium are 0.8–0.9 × length of merus, carpopropodus plus dactylus 1.1–1.2 × merus. Dactylus large, densely setose, mainly by whip setae with handle basally, bilaterally bearing stiff barbs. Nail smooth, approx. as long as dactylus; nail 2 thinner than nail 1. Endopod 3 non-prehensile, its carpopropodus, if 2 - segmented (Figs 30 B, 31 B), with proximal segment longer than combined distal segment and dactylus; terminal margin of distal segment with only one paradactylar whip seta (plus other setae) bearing barbed handle and smooth flagellum; nail 3 smooth, weakly bent, approx. as long as distal segment of carpopropodus. Endopod 6 slender; carpopropodus 3 - segmented with proximal segment shorter than combined median and distal segments; terminal margin of distal segment with two smooth, paradactylar whip setae (plus other setae); nail as in endopod 3. Penes as in Fig. 30 C.</p><p>Pleon and pleopods (Figs 31 A, 32 C – E). Pleomeres 1–5 are 0.8, 0.6, 0.6, 0.7 and 0.8 times the length of pleomere 6, respectively. Length of pleopods 1–5 increasing distally. Pleopod 5 (Fig. 32 E) measures 1 / 12 BL and 0.9 × length of pleomere 5, thus reaching to 2 / 3 caudal extension of pleomere 6. Pleopod 5 not fully visualized in Fig. 31 A. Pleopods 1–4 (Fig. 32 C, D) with barbed (plumose) setae only on distal half; pleopod 5 with setae on distal 2 / 3.</p><p>Tail fan (Fig. 32 F, G). Scutellum paracaudale short, sinusoid, laterally covering part of the telson basis. Both rami of uropods setose all around. Exopod approx. as long as endopod, each caudally reaching 0.4 × their length beyond telson. Exopod 7 × longer than wide. Diameter of the spherical fluorite statoliths 34–38 μm. Telson 17 % BL, equaling 1.1 × length of pleomere 6. Telson length 1.5 × maximum width; distolateral spines 22 % telson length.</p><p>DNA data.</p><p>Only one 622 bp mitochondrial COI sequence was obtained from a single individual from Cathedral Cave (GenBank PV 990329).</p><p>Distribution and habitat</p><p>(Fig. 10 L). Type locality not stated originally. Here fixed as Bermuda, submarine cave Castle Grotto (Station 28). The species was first described by Băcescu and Iliffe (1986 a) from two submarine caves 4 km apart in Bermuda: Castle Grotto and Green Bay Cave. The present record from Cathedral Cave (Station 14) in Guadeloupe is the first from outside Bermuda, namely 1770 km away. Total distribution range 16– 32°N, 62– 65°W, 0.5–18 m depth. From submarine caves of Bermuda (Băcescu and Iliffe 1986 a) and Lesser Antilles; at Bermuda also found over the open sea floor during the night (Wittmann and Ariani 2019; Wittmann 2023). At Guadeloupe, the species was not directly observed in situ, but captured together with large numbers of Palaumysis antillensis sp. nov. with which it shares the dark zone habitat of Cathedral Cave (Station 14). Platyops sterreri is considered a stygophile (Pesce et al. 1994) or a troglophile (Wittmann 2023).</p></div>	https://treatment.plazi.org/id/4FC238B257DD53B98DFA9679C0867D86	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wittmann, Karl J.;Chevaldonné, Pierre	Wittmann, Karl J., Chevaldonné, Pierre (2026): Biodiversity, complete inventory, taxonomy and keys to the species of Mysidae (Crustacea, Mysida) from marine caves of the Caribbean with revision of three genera and description of three new species. ZooKeys 1283: 93-204, DOI: 10.3897/zookeys.1283.169111
