taxonID	type	description	language	source
65816340F7325716A4F5A61E28DD3442.taxon	type_taxon	Type species. Houcosa zhaoi sp. nov.	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
65816340F7325716A4F5A61E28DD3442.taxon	etymology	Etymology. The generic name is a compound noun derived from the ancient Chinese mythical creatures ‘ hou’ (犼) and ‘ - cosa’, a common ending for Lycosidae genera. The gender is feminine.	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
65816340F7325716A4F5A61E28DD3442.taxon	diagnosis	Diagnosis. Houcosa gen. nov. is similar to Loongcosa gen. nov. in having a modified palp tibia, 2 pairs of spines on the male palp tibia, a biforked basoembolic apophysis (BEA, Figs 2, 12, 14), absent median apophysis and conductor, an extended posterior septum of the epigynes (Figs 3 C, D, 13 C, D, 15 C, D), but differs by the tibia as long as cymbium (Fig. 2 A – C) (vs 1 / 3 length, Figs 12 A – C, 14 A – C), the long and membranous terminal apophysis located on the anterior side of the embolus (TA, Fig. 2 F) (vs short and sclerotized, located on the posterior side of the embolus; Figs 12 F, 14 F), the thin embolus (Fig. 2 F) (vs broad, Figs 12 B, D, F, 14 B, D, F), the small and semicircular tegular apophysis (TeA, Figs 2 B – E) (vs absent), the triangular retrolateral tegular apophysis (RTeA, Fig. 2 A – E) (vs strongly sclerotized and grooved, Figs 12 B – E, 14 B – E), the small and semicircular septum (Se, Fig. 3 C) (vs large and almost tongue-shaped, Figs 13 C, 15 C), the S-shaped stalk of the spermatheca (SS, Fig. 3 D) (vs arc-shaped, Figs 14 D, 15 D), large and spherical head of the spermatheca (HS, Fig. 3 D) (vs small and spherical or elongate, Figs 14 D, 15 D).	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
65816340F7325716A4F5A61E28DD3442.taxon	description	Description. Carapace brown, pear-shaped. Eyes region black. Cervical groove and radial furrows indistinct (Fig. 3 A, B). Chelicerae brown, with 3 teeth on both margins. Labium brown, with dark base, longer than wide. Endites brown, longer than wide. Sternum brown and scutellate, with sparse black setae. Legs brown, with black pigmentation. Leg formula: 4132 or 4123. Opisthosoma oval, dorsum black brown, with lanceolate cardiac mark in anterior half, venter brown. Palp (Fig. 2): Tibia modified, rounded ventro-proximally and dorso-distally with 2 pairs of spines: pair of dorsal and pair of prolateral spines. Cymbium with short tip, cymbial claws absent. Subtegulum (St) located postero-prolaterally. Sperm duct waved. Tegulum with distal triangular apophysis (RTeA). Tegular apophysis (TeA) small, semicircular. Conductor and median apophysis lacking. Embolic division complex, palea lacking, with large and heavily sclerotized basoembolic apophysis with 2 horned-like extensions (BEA), terminal apophysis (TA) membranous. Embolus (E) originated on dorsal part of embolic division, whip-shaped. Epigyne (Fig. 3 C, D): Epigynal plate slightly wider than long, fovea absent, septum (Se) small with semicircular posterior part. Spermathecae S-shaped with long stalks (SS) and suboval heads (HS).	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
65816340F7325716A4F5A61E28DD3442.taxon	distribution	Distribution. China (Guangxi) (Fig. 16).	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
B6EB9187054E5B6F85477B51F3DF5493.taxon	description	Common name. 赵氏犼蛛 Figs 2, 3, 16	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
B6EB9187054E5B6F85477B51F3DF5493.taxon	etymology	Etymology. The specific epithet is taken from the family name of Mr Wenqi Zhao (Nanchang, China), who collected this species.	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
B6EB9187054E5B6F85477B51F3DF5493.taxon	diagnosis	Diagnosis. See genus diagnosis.	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
B6EB9187054E5B6F85477B51F3DF5493.taxon	description	Description. Male (holotype, Fig. 3 A). Total length 6.85. Prosoma 3.21 long, 2.43 wide; opisthosoma 3.51 long, 2.43 wide. Carapace greyish brown, lacking distinct pattern, with lighter median band in posterior part. Eye sizes and interdistances: AME 0.12, ALE 0.12, PME 0.37, PLE 0.32; AME – AME 0.07, AME – ALE 0.07, PME – PME 0.30, PME – PLE 0.35. Clypeus height 0.22. Legs yellow brown, with black pigmentation. Leg measurements: I 8.69 (2.24, 3.10, 2.23, 1.12); II 8.40 (2.23, 2.89, 2.09, 1.19); III 8.63 (2.23, 2.69, 2.56, 1.15); IV 11.87 (2.84, 3.74, 3.78, 1.51). Opisthosoma oval. Dorsum greyish brown, with yellow brown markings. Venter yellow brown. Palp (Fig. 2). Tibia with 2 pairs of prolateral spines. Embolus whip-shaped. Basoembolic apophysis biforked. No conductor. Terminal apophysis long and membranous. Tegular apophysis small and semicircular. Retrolateral tegular apophysis triangular. Female (paratype SWUC - T-LY- 19 - 02, Fig. 3 B). Total length 8.15. Prosoma 3.62 long, 2.79 wide; opisthosoma 4.35 long, 2.96 wide. Carapace with distinct light median band, wide in cephalic part as wide as PME – PME, and thin in posterior part. Eye sizes and interdistances: AME 0.10, ALE 0.13, PME 0.38, PLE 0.34; AME – AME 0.13, AME – ALE 0.10, PME – PME 0.32, PME – PLE 0.38. Clypeus 0.15 high. Leg measurements: I 8.70 (2.54, 3.13, 2.08, 0.95); II 8.54 (2.43, 3.12, 1.90, 1.09); III 8.18 (2.26, 2.55, 2.25, 1.12); IV 12.59 (3.08, 4.10, 3.92, 1.49). Epigyne (Fig. 3 C, D). Septum slightly longer than wide with semicircular posterior part. Copulatory openings located between septum and lateral walls. Stalk of spermatheca S-shaped, head of spermatheca suboval. Fertilization ducts crescent-shaped.	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
B6EB9187054E5B6F85477B51F3DF5493.taxon	distribution	Distribution. Currently known only from the type locality, China (Guangxi, Fig. 15).	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
B35E26550BCB5C80A3A1D2C3DDEA4665.taxon	description	Figs 4, 6 A, B, 16	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
B35E26550BCB5C80A3A1D2C3DDEA4665.taxon	diagnosis	Diagnosis. This species is similar to K. hikosanensis (Fig. 5) but differs from the latter by the thin and short retrolateral tegular apophysis (RTeA, Fig. 4 A, B) (vs thick and long), and the semicircular septum (Se, Fig. 4 C) (vs almost triangular).	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
B35E26550BCB5C80A3A1D2C3DDEA4665.taxon	description	Description. See Tanaka (1985, 2009). Habitus, male palps and epigynes as in Figs 4, 5 A, B.	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
B35E26550BCB5C80A3A1D2C3DDEA4665.taxon	distribution	Distribution. Japan (Fig. 16).	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
59F9C141E34C5BBEB23C28DB64EA8A55.taxon	description	Figs 5, 6 C, D, 16	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
59F9C141E34C5BBEB23C28DB64EA8A55.taxon	diagnosis	Diagnosis. See the diagnosis of K. fujiii (Tanaka, 1985).	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
59F9C141E34C5BBEB23C28DB64EA8A55.taxon	description	Description. See Tanaka (1985). Habitus, male palps and epigynes as in Fig. 5.	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
59F9C141E34C5BBEB23C28DB64EA8A55.taxon	distribution	Distribution. Japan (Fig. 16).	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
E6B0BDE9C085523DA72344D3BFF26E6D.taxon	description	Common name. 宁波鲲蛛 Figs 1 A – C, 8, 9, 16	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
E6B0BDE9C085523DA72344D3BFF26E6D.taxon	diagnosis	Diagnosis. This species is similar to K. zhui in having hook-like posterior arm of the retrolateral tegular apophysis and a spherical head of the spermatheca (Figs 8 B – E, 9 D, 10 B – E, 11 D), but differs from the latter by the thick tegular apophysis (TeA, Fig. 8 A – E) (vs thin, Fig. 10 A – E), the almost triangle-like terminal apophysis (TA, Fig. 8 F) (vs fingernail-like, Fig. 10 F), the digital-like anterior arm of retrolateral tegular apophysis in retrolateral view (RTeA, Fig. 8 B – E) (vs triangular-like, Fig. 10 B – E), the semicircular copulatory openings (CO, Fig. 9 C) (vs slit-like, Fig. 11 C), and the C-shaped stalk of the spermathecal (SS, Fig. 9 D) (vs S-shaped, Fig. 11 D).	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
E6B0BDE9C085523DA72344D3BFF26E6D.taxon	description	Description. Male (Fig. 9 A). Total length 8.62. Carapace 4.44 long, 3.31 wide; opisthosoma 3.99 long, 2.62 wide. Carapace greyish brown with distinct light median band. Eye sizes and interdistances: AME 0.13, ALE 0.13, PME 0.35, PLE 0.30; AME – AME 0.10, AME – ALE 0.08, PME – PME 0.30, PME – PLE 0.40. Clypeus 0.24 high. Leg measurements: I 11.11 (3.09, 4.08, 2.60, 1.34); II 10.59 (2.88, 3.87, 2.47, 1.37); III 10.88 (3.05, 3.63, 2.91, 1.29); IV 15.24 (3.89, 4.99, 4.60, 1.76). Palp (Fig. 8). Tegular apophysis strong, finger-shaped. Terminal apophysis strong and triangle-like, under the embolus. Embolus needle-shaped, with abrupt tip. Retrolateral tegular apophysis with two arms (digital-like anterior arm and hook-like posterior arm) and with a groove on the dorsal surface (maybe functioning as conductor). Female (Fig. 9 B). Total length 9.20. Carapace 3.96 long, 3.05 wide; opisthosoma 5.21 long, 3.51 wide. Carapace brown with light stripe behind cephalic part and wide band anteriorly, light band with pair of dark dots and longitudinal stripe. Eye sizes and interdistances: AME 0.15, ALE 0.15, PME 0.35, PLE 0.29; AME – AME 0.12, AME – ALE 0.08, PME – PME 0.30, PME – PLE 0.37. Clypeus 0.33 high. Leg measurements: I 9.90 (2.87, 3.60, 2.28, 1.15); II 7.76 (2.24, 2.71, 1.80, 1.01); III 9.89 (2.68, 3.28, 2.42, 1.51); IV 12.88 (3.49, 4.30, 3.87, 1.22). Epigyne (Figs 9 C, D). Septum under epigynal plate. Copulatory openings semicircular, located on shoulders of septum. Stalk of spermatheca C-shaped, head of spermatheca spherical. Fertilization ducts crescent-shaped.	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
E6B0BDE9C085523DA72344D3BFF26E6D.taxon	distribution	Distribution. China (Anhui, Zhejiang) (Fig. 16).	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
0EAE2F16AE6F5D45A24C479875FD32C2.taxon	type_taxon	Type species. Arctosa ningboensis Yin, Bao & Zhang, 1996.	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
0EAE2F16AE6F5D45A24C479875FD32C2.taxon	etymology	Etymology. The generic name is a compound noun derived from the ancient Chinese mythical creatures ‘ kun’ (鲲) and ‘ - cosa’, a common ending for Lycosidae genera. The gender is feminine.	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
0EAE2F16AE6F5D45A24C479875FD32C2.taxon	diagnosis	Diagnosis. Kuncosa gen. nov. is similar to Loongcosa gen. nov. in having 2 pairs of spines in male palp tibia, a developed retrolateral tegular apophysis (Figs 4 A, B, 5 A, B, 8 A – E, 10 A – E, 12 A – E, 14 A – E), a broad septum (Figs 4 C, D, 5 C, D, 9 C, D, 11 C, D, 13 C, D, 15 C, D), but differs from the later by tibia as half-length of cymbium (Figs 4 A, B, 5 A, B, 8 A – C, 10 A – C) (vs 1 / 3 length, figs 12 A – C, 14 A – C), the thin embolus lacking pars pendula (Figs 8 F, 10 F) (vs broad with pars pendula, Figs 12 B, D, F, 14 B, D, F), the comma-shaped basoembolic apophysis (BEA, Figs 8 F, 10 F) (vs bifurcated, Figs 12 A – F, 14 A – F), the strong and triangular terminal apophysis (TA, Figs 8 F, 10 F) (vs thin, and finger-shaped, Figs 12 F, 14 F), the tegular apophysis present (TeA, Figs 4 A, B, 5 A, B, 8 A – E, 10 A – E) (vs absent), the bifurcated retrolateral tegular apophysis (RTeA, Figs 4 A, B, 5 A, B, 8 A – E, 10 A – E) (vs strong sclerotized and grooved, Figs 12 B – E, G, 14 B – E, G), truncated posterior margin of septum (Figs 4 C, D, 5 C, D, 7, 9 C, D, 11 C, D) (vs with a tongue like apophysis, Figs 13 C, D, 15 C, D).	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
0EAE2F16AE6F5D45A24C479875FD32C2.taxon	description	Description. Carapace brown with light median band. Eyes region black. Fovea vertical. Cervical groove and radial furrows indistinct. Chelicerae elongate, brown, with 3 teeth on both margins. Labium yellow brown, with dark base. Endites yellow brown. Sternum brown and scutellate, with sparse black setae. Legs brown, with black pigmentation. Leg formula: 4132. Opisthosoma oval. Dorsum dark brown, with lanceolate cardiac mark in anterior half, and black irregular markings in posterior half. Venter brown. Palp (Figs 4 A, B, 5 A, B, 8, 10): Tibia with 2 pairs of spines, and rounded disto-dorsal edge. Cymbium asymmetric, with short tip and lacking claws. Sperm duct weavy. Tegulum with long tegular apophysis (TeA) and short bifurcate retrolateral apophysis (RTeA). Conductor and median apophysis lacking. Embolic division complex, palea absent, terminal apophysis (TA) triangle- or fingernail-like, Basoembolic apophysis (BEA) strongly sclerotized Embolus needle-shaped, originated from proventral side, lacking pars pendula and with abrupt tip. Epigyne (Figs 4 C, D, 5 C, D, 7, 9 C, D, 11 C, D): Epigynal plate wider than long, fovea indistinct, covered by broad septum and oval. Stalk of spermatheca (SS) C- or S-shaped, head of spermatheca (HS) suboval.	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
0EAE2F16AE6F5D45A24C479875FD32C2.taxon	distribution	Distribution. China (Anhui, Chongqing, Guizhou, Hubei, Hunan, Jiangxi, Jiangsu, Zhejiang), southern part of Japan and Korea. (Fig. 16)	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
7AEB3522A4E9563FBF4E62E237429439.taxon	description	Common name. 朱氏鲲蛛 Figs 1 D, E, 10, 11, 16	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
7AEB3522A4E9563FBF4E62E237429439.taxon	diagnosis	Diagnosis. See the diagnosis of K. ningboensis (Yin, Bao & Zhang, 1996).	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
7AEB3522A4E9563FBF4E62E237429439.taxon	description	Description. Male (Fig. 11 A). Total length 7.40. Prosoma 4.13 long, 3.17 wide; opisthosoma 3.68 long, 2.38 wide. Carapace greyish brown. Eye sizes and interdistances: AME 0.14, ALE 0.14, PME 0.31, PLE 0.30; AME – AME 0.10, AME – ALE 0.07, PME – PME 0.28, PME – PLE 0.35. Clypeus height 0.22. Leg measurements: I 10.81 (3.01, 4.10, 2.53, 1.17); II 10.27 (2.90, 3.77, 2.42, 1.18); III 10.50 (2.88, 3.57, 2.85, 1.20); IV 14.14 (3.63, 4.85, 4.22, 1.44). Palp (Fig. 10). Tegular apophysis strong, finger-shaped. Terminal apophysis strong and fingernail-like, under the embolus. Embolus needle-shaped, with abrupt tip. Retrolateral tegular apophysis with two arms (triangular-like anterior arm and hook-like posterior arm) and with a groove on the dorsal surface (maybe functioning as conductor). Female (Fig. 11 B). Total length 9.40. Prosoma 4.47 long, 3.31 wide; opisthosoma 5.43 long, 3.83 wide. Eye sizes and interdistances: AME 0.17, ALE 0.15, PME 0.37, PLE 0.33; AME – AME 0.13, AME – ALE 0.09, PME – PME 0.31, PME – PLE 0.41. Clypeus height 0.28. Leg measurements: I 10.46 (2.95, 3.94, 2.43, 1.14); II 9.77 (2.89, 3.45, 2.32, 1.11); III 10.26 (2.97, 3.49, 2.64, 1.16) IV 14.07 (3.77, 4.51, 4.19, 1.60). Epigyne (Figs 11 C, D). Septum under epigynal plate, T-shaped. Copulatory openings slit-like, located on the shoulders of septum. Stalk of spermatheca S-shaped, head of spermatheca spherical. Fertilization ducts crescent-shaped.	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
7AEB3522A4E9563FBF4E62E237429439.taxon	distribution	Distribution. China (Chongqing, Guizhou, Hubei, Hunan, Jiangxi, Jiangsu).	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
D6F77E0E949F5739BE130590766AA948.taxon	description	Common name. 齿盾龙蛛 Figs 1 F, G, 12, 13, 16	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
D6F77E0E949F5739BE130590766AA948.taxon	diagnosis	Diagnosis. This species differs from L. wuyiensis by the short and C-shaped basoembolic apophysis with 2 arms in ventral view (BEA, Fig. 12 B, D, F) (vs long, hook-shaped with 3 arms, Fig. 14 B, D – F), epigyne with an anterior hood (AH, Fig. 13 C) (vs with 2 hoods, Fig. 15 C), and septum lacking stalk (Fig. 13 C) (vs having a stalk, Fig. 15 C).	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
D6F77E0E949F5739BE130590766AA948.taxon	description	Description. Male (Fig. 13 A). Total length 6.38. Carapace 3.48 long, 2.77 wide; opisthosoma 2.91 long, 1.99 wide. Carapace greyish brown with distinct light median band and submarginal stripes, median band wide in cephalic part, with pair of dark dots, thoracic part of median band thin. Eye sizes and interdistances: AME 0.12, ALE 0.13, PME 0.41, PLE 0.33; AME – AME 0.13, AME – ALE 0.07, PME – PME 0.31, PME – PLE 0.32. Clypeus height 0.21. Leg measurements: I 8.41 (2.34, 3.01, 2.11, 0.95); II 8.05 (2.25, 2.72, 2.10, 0.98); III 8.28 (2.26, 2.65, 2.33, 1.04); IV 11.64 (2.79, 3.71, 3.81, 1.33). Palp (Fig. 12): Basoembolic apophysis with 2 arms, anterior (aa) and posterior (pa), almost as long as tegulum. Embolus broad, short and arc-shaped. Terminal apophysis short, sclerotized and located posterior to embolus. Retrolateral tegular apophysis sclerotized, twice as long as wide, with a groove on the dorsal surface. Female (Fig. 13 B). Total length 7.25. Carapace 3.42 long, 2.75 wide; opisthosoma 3.91 long, 2.99 wide. Colouration as in male. Eye sizes and interdistances: AME 0.13, ALE 0.13, PME 0.41, PLE 0.35; AME – AME 0.12, AME – ALE 0.09, PME – PME 0.33, PME – PLE 0.36. Clypeus height 0.15. Leg measurements: I 8.92 (2.54, 3.25, 2.12, 1.01); II 8.44 (2.54, 3.11, 1.87, 0.92); III 8.86 (2.68, 2.60, 2.36, 1.22); IV 12.18 (2.90, 4.01, 3.74, 1.53). Epigyne (Fig. 13 C, D). Epigynal plate almost as long as wide, anterior hood (AH) unpaired. Septum originated beneath hood. Copulatory openings semicircular. Stalk of spermatheca C-shaped, head of spermatheca spherical, ca 1.7 times wider than stalk.	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
D6F77E0E949F5739BE130590766AA948.taxon	distribution	Distribution. China (Hunan, Guizhou, Guangxi) (Fig. 16).	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
8DD212E2E6685E0FAA3F298989EAA91E.taxon	type_taxon	Type species. Pardosa dentitegulum Yin, Peng, Xie, Bao & Wang, 1997.	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
8DD212E2E6685E0FAA3F298989EAA91E.taxon	etymology	Etymology. The generic name is a compound noun derived from the Chinese totem ‘ loong’ (龙) and ‘ - cosa’, a common ending in Lycosidae genera. The gender is feminine.	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
8DD212E2E6685E0FAA3F298989EAA91E.taxon	diagnosis	Diagnosis. Loongcosa gen. nov. is similar to Houcosa gen. nov. (Figs 2, 3) and Kuncosa gen. nov. (Figs 4 – 11) in having 2 pairs of spines on the male palpal tibia, a strong basoembolic apophysis (BEA), Figs 2, 12, 14), an extended posterior septum of the epigyne (Figs 3 C, D, 13 C, D, 15 C, D), but differs from the latter two genera by the short tibia, 1 / 3 the length of the cymbium (Figs 12 A – C, 14 A – C) (vs as long as in Houcosa and as half-length in Kuncosa), tegular apophysis absent (TeA, Figs 12 A – E, 14 A – E) (vs present in both Houcosa and Kuncosa), terminal apophysis short, sclerotized, spine-like and located on posteriorly to embolus (TA, Figs 12 F, 14 F) (vs long, membranous, needlelike and located anteriorly to embolus in Houcosa and strong, triangular in Kuncosa), wide and heavily sclerotized embolus (Em, Figs 12 B, D, F, 14 B, D, F) (vs thin in both Houcosa and Kuncosa), the strong sclerotized and grooved retrolateral tegular apophysis (RTeA, Figs 12 B – E, G, 14 B – E, G) (vs triangular in Houcosa and bifurcated in Kuncosa), large and almost tongue-shaped septum with a kind of scape (Sc, Figs 13 C, 15 C) (vs small, semicircular in Houcosa and truncated posterior margin in Kuncosa), the arc-shaped stalk of the spermatheca (SS, Figs 13 D, 15 D) (vs S-shaped in Houcosa), and a small head of the spermatheca (SH, Figs 13 D, 15 D) (vs large and suboval in Houcosa).	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
8DD212E2E6685E0FAA3F298989EAA91E.taxon	description	Description. Carapace brown with distinct light median band, median band wide in cephalic part, with pair of dark dots, thoracic part of median band thin (Figs 13 A, B, 15 A, B). Chelicerae brown, with 3 teeth on both margins. Labium brown, with dark base. Endites brown. Sternum brown, with sparse black setae. Legs brown, with black pigmentation. Leg formula: 4132. Opisthosoma oval, dorsum black brown, with yellow setae anteriorly, and 5 pairs of white markings in posterior half when live, venter brown. Palp (Figs 12, 14): Tibia ca 1.4 times longer than wide in lateral view, with 2 pairs of spines, disto-dorsal part and proximo-ventral part rounded. Lacking claws; Bulb ca 1.4 longer than wide; Tegular apophysis, conductor and median apophysis lacking. Terminal apophysis (TA) short, spine-like, sclerotized and located posterior to embolus. Basoembolic apophysis (BEA) large and heavily sclerotized, with 2 or 3 distinct arms. Embolus short, wide, heavily sclerotized, roundly bent, tip pointed. Retrolateral tegular apophysis strongly sclerotized and grooved. Epigyne (Figs 13 C, D, 15 C, D): Epigynal plate almost as long as wide; anterior part heavily sclerotized, fovea distinct anteriorly; anterior margin with one or pair of anterior hoods (AH), septum broad either with stalk or without, posteriorly with a kind of scape (Sc). Stalk of spermatheca C-shaped, head of spermatheca spherical or irregular.	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
8DD212E2E6685E0FAA3F298989EAA91E.taxon	distribution	Distribution. China (Anhui, Fujian, Hunan, Guizhou, Guangxi, Shaanxi, Zhejiang) (Fig. 16).	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
60FDB4D5DD285D2EA4FA3CDFCBF64195.taxon	description	Common name. 武夷龙蛛 Figs 1 H, I, 14, 15, 16	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
60FDB4D5DD285D2EA4FA3CDFCBF64195.taxon	diagnosis	Diagnosis. See the diagnosis of L. dentitegulum (Yin, Peng, Xie, Bao & Wang, 1997).	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
60FDB4D5DD285D2EA4FA3CDFCBF64195.taxon	description	Description. Male (Fig. 15 A). Total length 6.25. Carapace 3.29 long, 2.67 wide; opisthosoma 2.93 long, 1.79 wide. Carapace greyish brown, with median band, submarginal stripe absent. Opisthosoma lacking distinct cardiac mark. Eye sizes and interdistances: AME 0.11, ALE 0.11, PME 0.36, PLE 0.34; AME – AME 0.11, AME – ALE 0.08, PME – PME 0.31, PME – PLE 0.31. Clypeus 0.18 high. Leg measurements: I 8.11 (2.17, 2.90, 2.00, 1.04); II 7.80 (2.15, 2.68, 1.97, 1.00); III 7.96 (2.15, 2.56, 2.20, 1.05); IV 11.33 (2.87, 3.64, 3.48, 1.34). Palp (Fig. 14). Basoembolic apophysis with three arms: dorsal (da), posterior (pa), and anterior (aa). Embolus (E) very broad, terminal part almost straight, ca 2 times longer than wide, tip abrupt. Terminal apophysis short, sclerotized, and located on the posterior side of the embolus. Retrolateral tegular apophysis sclerotized, twice as long as wide, with a groove on the dorsal surface. Female (Fig. 15 B). Total length 9.09. Carapace 3.59 long, 2.80 wide; opisthosoma 5.25 long, 3.82 wide. Colouration as in male. Eye sizes and interdistances: AME 0.11, ALE 0.11, PME 0.44, PLE 0.35; AME – AME 0.13, AME – ALE 0.09, PME – PME 0.36, PME – PLE 0.35. Clypeus height 0.20. Leg measurements: I 8.34 (2.41, 3.07, 1.87, 0.99); II 7.99 (2.32, 2.83, 1.88, 0.96); III 8.04 (2.25, 2.55, 2.21, 1.03); IV 11.42 (2.83, 3.73, 3.51, 1.35). Epigyne (Fig. 15 C, D). Epigynal plate ca 1.3 times wider than long, fovea distinct, septum with distinct stalk. Stalk of spermatheca C-shaped, head of spermatheca elongate.	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
60FDB4D5DD285D2EA4FA3CDFCBF64195.taxon	distribution	Distribution. China (Anhui, Fujian, Hunan, Guizhou, Shaanxi, Zhejiang) (Fig. 16).	en	Wang, Lu-Yu, Marusik, Yuri M., Zhang, Zhi-Sheng (2025): Three new genera of wolf spiders (Araneae, Lycosidae) living on the forest floor in East Asia. ZooKeys 1239: 123-145, DOI: 10.3897/zookeys.1239.152834
