identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
3EC5E69C56D6565899766C62F28AD62C.text	3EC5E69C56D6565899766C62F28AD62C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ceruchus ahualuo Huang, Liu & Wan 2026	<div><p>Ceruchus ahualuo Huang, Liu &amp; Wan sp. nov.</p><p>Figs 1, 2, 3, 5, 6</p><p>Chinese common name.</p><p>阿花洛角锹甲.</p><p>Type material.</p><p>Holotype: China • ♂; Yunnan Province: Weixi County, Mts. near Ahualuo River, elev. 2600 m, in decayed logs, 2024 - IV- 2, Baijun Li leg., deposited in AHU . Paratypes: China • 4 ♂, 2 ♀ Yunnan Province: Weixi County, Zhonglu Township, elev. 2750 m; in decayed logs, 2021 - III- 5, Yanchen Liu leg., (AHU) . China • 3 ♂ Yunnan Province: Weixi County, Mts. near Ahualuo River, elev. 2800–3000 m, in decayed logs, 2026 - V, Yujie Huang leg., (AHU) .</p><p>Description.</p><p>Male (Figs 1 A, 1 B, 1 D, 1 E, 2). Body length 13.5–17.0 mm. Body black, labial palps, maxillary palps and legs slightly paler. Pronotum wider than long, strongly convex, with relatively broad lateral margins. Elytra with 7 distinct longitudinal striae, humeral angles obtusely rounded. Scutellum semicircular. Legs slender; ventral surface of femora with sparse yellow setae; ventral apex of mesotibiae with a yellow setal brush.</p><p>Head with vertex bearing a distinct subconical depression anteromedially. Labrum triangular. Mandibles slender, longer than head, apex acute, inner margin clothed with yellow setae. Base of ventral margin with a triangular obtuse tooth directed vertically downward, smaller in minor individuals; a very small projection present at approximately apical two-thirds of mandibular length, more distinct in minor individuals. Dorsal margin near base with a large finger-like tooth (Fig. 2), curved upward and directed obliquely forward; one subquadrate obtuse denticle positioned immediately posterior to this large tooth toward base and directed horizontally.</p><p>Aedeagus. (Fig. 3 A – E). Parameres broad and plate-like, with bluntly rounded apices, curved dorsally. Median lobe moderately sclerotized and equal in length to parameres, broadened medially and tapering toward both base and apex. Basal piece stout, longer than parameres. Struts approximately as long as median lobe. Caudal margin of the basal piece of the aedeagus with a shallow median emargination.</p><p>Female. (Fig. 1 C, F). Body length 10.5–13.5 mm. Dark brown. Mandibles small, shorter than head. Dorsal margin of mandibular base forming a slightly upward-curved arched plate; apex forming an acute triangular denticle, base obtuse and quadrate, middle bearing a very indistinct small denticle, rendering arched plate shallowly trifurcate. Base of ventral margin without a tooth. Anteromedian portion of vertex with a very shallow depression. Pronotum subtrapezoidal, disc distinctly convex and ridge-like, flanked by a small convexity on each side of the elevation. Setae on inner margin of mandibles, mesofemora, and ventral apex of mesotibiae much less conspicuous and dense than in male. Other characters similar to those of male.</p><p>Female genitalia (Fig. 3 F). Hemisternites subtrapezoidal and plate-like, bearing small styli at apex. Accessory duct long, broad, and apically spherical. Spermatheca slightly sclerotized; spermathecal duct relatively thin and short, distinctly separated from and narrower than spermatheca; spermathecal gland slender.</p><p>Diagnosis.</p><p>Ceruchus ahualuo sp. nov. is similar to C. niger (Fig. 4 A, D), C. sinensis (Fig. 4 B, E), and C. yangi (Fig. 4 C, F) in body size and the density of setae on the mesofemora, but can be distinguished from them by the large finger-like tooth on the dorsal surface near the base of each mandible. This discrete mandibular character was not observed in other examined species of Ceruchus . In the male genitalia, the new species differs from C. yangi and C. niger by the caudal margin of the basal piece of the aedeagus with a shallow median emargination in ventral view (vs. caudal margin of the basal piece forming an M-shaped margin in ventral view; cf. Fig. 3 A and Huang et al. 2011: figs 134–139, 164–169). It also differs from C. sinensis by the median lobe swollen medially in lateral view (vs. median lobe not swollen medially in lateral view; cf. Fig. 3 E and Huang et al. 2011: figs 196–201).</p><p>Given the sympatric occurrence of C. ahualuo sp. nov. and C. niger, the COI data were used to further assess their separation. In the maximum-likelihood tree based on mitochondrial COI sequences, the new species was recovered as a distinct lineage and was clearly separated from C. niger (Fig. 5). The pairwise K 2 P distance between the two species was 0.1359, falling within the range of interspecific distances observed among the sampled Ceruchus species and much higher than the distance between the two C. minor sequences (0.0077; Table 2). These molecular results are consistent with the morphological differences described above and provide additional support for separating the new species from the sympatric C. niger .</p><p>Etymology.</p><p>The specific epithet ahualuo is a noun in apposition, derived from the name of the type locality of this species.</p><p>Distribution.</p><p>China: Yunnan.</p></div>	https://treatment.plazi.org/id/3EC5E69C56D6565899766C62F28AD62C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Huang, Yujie;Liu, Yanchen;Shen, Yagang;Feng, Xueyun;Wan, Xia	Huang, Yujie, Liu, Yanchen, Shen, Yagang, Feng, Xueyun, Wan, Xia (2026): Description of a new species of Ceruchus MacLeay (Coleoptera, Lucanidae), with taxonomic notes on the Chinese species of the genus. ZooKeys 1289: 353-365, DOI: 10.3897/zookeys.1289.190703
91B5AD32B5B45A03B41943EEED2B3DDA.text	91B5AD32B5B45A03B41943EEED2B3DDA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ceruchus MacLeay 1819	<div><p>Genus Ceruchus MacLeay, 1819</p><p>Ceruchus MacLeay, 1819: 115 . Type species: Lucanus chrysomelinus Hochenwarth, 1785 .</p><p>Platycerus Latreille, 1807: 133 . Preoccupied name: Platycerus Geoffroy, 1762 in Lucanidae .</p><p>Tarandus Megerle: Dejean, 1837: 194. Nomen nudum in Dejean’s Catalogue des coléoptères de la collection de M. le comte Dejean.</p><p>Material examined.</p><p>Ceruchus sinensis Nagel, 1933: Holotype. China • 1 ♂; Yunnan Province, near Mt Frances Gardner, west of Mekong, elev. 2438 m, 9. VII. 1922; Prof. J. W. Gregory leg.; MNHN . Non-type material. China • 1 ♂; Yunnan Province, Weixi County, Baijixun Township, 1. IV. 2021; Zhiwen Yu leg.; AHU .</p><p>Ceruchus minor Tanikado &amp; Okuda, 1994: Paratype. China • 1 ♀; Shaanxi Province, Qinling Mountains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.5&amp;materialsCitation.latitude=33.9167" title="Search Plazi for locations around (long 108.5/lat 33.9167)">Huxian</a>, elev. 1400–1600 m, 33.9167°N, 108.5000°E, 27. VI. 1993; Tabana leg.; MNHN . Non-type material. China • 1 ♂; Shaanxi Province, Qinling Mountains, Changan, elev. 1400–1600 m, 11–24. IV. 1993; M. Tabana leg.; MNHN . China • 3 ♂ 2 ♀; Shaanxi Province, Mei County, Honghegu, elev. 1600 m, III. 2025; Yanchen Liu leg.; AHU .</p><p>Ceruchus deuvei Boucher &amp; Král, 1997: Paratype. China • 1 ♀; Sichuan Province, Mts. Siguniang, Huangqiao-gou Valley, southwest of Lixian, elev. c. 2500 m, deciduous forest; MNHN . Non-type material. China • 1 ♂; Sichuan Province, Niuzhihe Fl., Pusiun, elev. 2200 m, 16. VI. 2000; Belousov leg.; MNHN .</p><p>Ceruchus niger Boucher &amp; Král, 1997: Holotype. China • 1 ♂; Yunnan Province, Zhongdian, elev. 3300 m, Abies forest, VII; Th. Deuve leg.; MNHN . Non-type material. China • 4 ♂ 3 ♀; Yunnan Province, Weixi County, Zhonglu Township, III. 2021; Yanchen Liu leg.; AHU . China • 2 ♂ 1 ♀; Yunnan, Shangri-La City, VI. 2018; collector unknown; AHU . China • 2 ♂; Yunnan Province, Weixi County, Zhonglu Township, III. 2026; Yujie Huang leg.; AHU .</p><p>Ceruchus yangi Huang, Imura &amp; Chen, 2011: Holotype. China • 1 ♂; Guizhou Province, Jiangkou County, Mt Fanjingshan, elev. c. 2000 m, 25. X. 2009; X. - D. Yang leg.; SHNU . Non-type material. China • 2 ♂; Chongqing, Nanchuan District, Mt Jinfoshan, elev. 1855 m, III. 2025; collector unknown; AHU . China • 2 ♂ 1 ♀; Guizhou Province, Leishan County, Mt Leigongshan, elev. 2000 m, IV. 2021; Yanchen Liu leg.; AHU .</p><p>Ceruchus yingqii Huang &amp; Chen, 2013: Holotype. China • 1 ♂; Sichuan Province, Xiaojin County, Rilong Township, Changping-gou Valley, elev. 3600 m, 6. VIII. 2011; H. Huang leg.; SHNU .</p><p>Ceruchus ganyanae Huang &amp; Chen, 2017: Holotype. China • 1 ♂; Sichuan Province, Muli County, near Kangwu Pass, elev. c. 3200 m, VI. 2011; Ganyan Yang leg.; SHNU .</p><p>Ceruchus motuoensis Huang, Chen, Tao &amp; Xiao, 2020: Holotype. China • 1 ♂; Xizang, Motuo County, 62 k, elev. 2754 m, 13. XI. 2019; R. - C. Tao &amp; C. - M. Xiao leg.; SHNU .</p><p>Ceruchus tabanai Okuda, 2008: Non-type material. China • 1 ♂ 1 ♀; Sichuan Province, Puge County Mt Luojishan, III. 2026; Yanchen Liu leg.; AHU .</p><p>Ceruchus reginae Boucher &amp; Král, 1997: Non-type material. China • 1 ♂; Yunnan Province, Fugong County, Shiyueliang Township, elev. 2900 m, 15. III. 2021; Yanchen Liu leg.; AHU .</p><p>Description.</p><p>Generic characters. Members of the genus Ceruchus are relatively small. Sexual dimorphism distinct. Body mostly reddish-brown or dark brown. Dorsum slightly convex, relatively smooth, and sparsely setose. Venter possesses more pronounced short setae. Compound eyes small and entire, lacking canthus. Antennae stout and not distinctly geniculate; scape relatively stout and curved, forming an angle greater than 90 degrees with pedicel. Pedicel longer than antennomeres 3–7; antennomeres 8–10 dilated, forming a 3 – segmented club. Both maxillary and labial palps considerably long; maxillary palps almost equal in length to antennae.</p><p>Male. Body length 9.0– 21.5 mm. Mandibles not longer than combined length of head and pronotum, slightly curved upwards, hollowed on inner-ventral side, and clothed with relatively long, dense setae; apex of mandibles acute; mandibles with a well-developed major tooth near middle or base, curved upward and directed subhorizontally or obliquely forward; one or two small, subquadrate, inwardly curved obtuse denticles present posterior to major tooth toward base; base of ventral mandibular margin usually with a triangular obtuse tooth directed almost vertically downward. Head relatively broad, with sparse large punctures; an oblique groove formed by a line of punctures present behind each eye; labrum small, usually triangular. Pronotum subrectangular, with distinct small punctures; anterior margin sinuate, posterior margin nearly straight, and lateral margins nearly parallel. Elytra with distinct longitudinal striae formed by punctures. Legs short; tibiae of all legs serrate and armed with sharp denticles; outer apical margin of protibia not bifurcate, bearing only a single acute tooth; inner apical margin with a well-developed spur curved downward; outer apical margins of mesotibia and metatibia without distinct bifurcation, inner apical margins with two asymmetrical outward-curving spurs.</p><p>Male genitalia. Internal sac not everted from middle to apex; instead, only a slender sac-like structure present within median lobe. Median lobe broad, forming a large plate with a rounded apex. Parameres plate-like and rounded. Basal piece stout, longer than parameres. Median struts variable in thickness and length.</p><p>Female. Body length 9.0– 15.5 mm. Mandibles short and small; dorsal margin of mandibular base forming a slightly upward-curved arched plate, apex forming an acute triangular denticle, base obtuse and quadrate, and middle bearing one small denticle, giving arched plate a trifurcate appearance; base of ventral margin without a tooth. Anteromedian portion of vertex with a shallow depression. Pronotum subtrapezoida, distinctly convex and ridge-like, flanked by a small convexity on each side. Setae on inner margin of mandibles, mesofemora, and ventral apex of mesotibiae much less conspicuous and dense than in male. Punctures on head, pronotum, and elytra deeper and denser than in male. Other characters similar to those of male.</p><p>Female genitalia. Hemisternites subtrapezoidal and plate-like, bearing small styli at apex. Accessory duct long, broad, and apically spherical. Spermatheca often slightly sclerotized; spermathecal duct relatively thin and short, distinctly separated from and narrower than spermatheca; spermathecal gland slender.</p><p>Distribution.</p><p>Holarctic.</p></div>	https://treatment.plazi.org/id/91B5AD32B5B45A03B41943EEED2B3DDA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Huang, Yujie;Liu, Yanchen;Shen, Yagang;Feng, Xueyun;Wan, Xia	Huang, Yujie, Liu, Yanchen, Shen, Yagang, Feng, Xueyun, Wan, Xia (2026): Description of a new species of Ceruchus MacLeay (Coleoptera, Lucanidae), with taxonomic notes on the Chinese species of the genus. ZooKeys 1289: 353-365, DOI: 10.3897/zookeys.1289.190703
