taxonID	type	description	language	source
4AF2797C5ED45B8689B23066B3088AD2.taxon	description	Figs 4 A – H, 10 B, 11 D – F, 12 A – C; Suppl. material 3 Chinese vernacular name: 黃紋亞力姬蜂	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
4AF2797C5ED45B8689B23066B3088AD2.taxon	diagnosis	Diagnosis. This species can be distinguished from congeners by the combination of the following characters: smaller ocelli (OD = 0.15 – 0.19 mm; OOL / OD = 1.2 – 1.6); fore wing areolet trapezoid with stalk, receiving 2 m-cu at distal corner (Fig. 10 B); fore wing 1 cu-a almost vertical, opposite or slightly distad to M & RS (Fig. 10 B); lateromedian longitudinal carinae of propodeum present posteriorly with the area petiolaris opened (Fig. 3 E); T 1 2.7 – 3.5 × its posterior width; S 9 with subtriangular median area weakly sclerotized (Fig. 11 D); mesosoma and metasoma generally black, with latero-anterior corners of mesoscutum, ventral subtriangular marking of mesopleuron (enclosing black area), and median markings on tergites behind T 2 yellow (Figs 4 A, D – F, 12 A – C). This new species is similar to the Asian species A. flaviventris Li & Sun, 2022, A. albimaculatus Sheng, Sun & Li, 2020, and A. zangicus Sheng, Sun & Li, 2020 which share black bodies and pale markings on the latero-anterior corners of mesoscutum. It can be distinguished from A. flaviventris by the following characters: lateromedian longitudinal carinae of propodeum absent on the anterior and median portions (Fig. 4 E) (present and combined as single carina medially in A. flaviventris); metasomal tergites and ventral portion of mesopleuron with yellow markings (Figs 4 A, F, 12 B) (without in A. flaviventris). It can also be distinguished from A. albimaculatus by the following characters: face and mandibles yellow (Figs 4 C, 12 A) (face white and mandibles entirely black in A. albimaculatus); fore and mid legs each with yellow coxae and reddish-brown other part (Fig. 4 A) (generally black with white tibiae in A. albimaculatus); latero-anterior corners of mesoscutum, ventral portion of mesopleuron, and metasomal tergites with yellow markings (Figs 4 A, D, F, 12 B, C) (marking on mesoscutal corners white, other markings absent in A. albimaculatus); antenna without white band (Fig. 4 A) (with white band in A. albimaculatus); and lateromedian longitudinal carinae of propodeum absent on the anterior and median portions (Fig. 4 E) (present in A. albimaculatus). Lastly, it can be distinguished from A. zangicus by the following characters: areolet present (Fig. 10 B) (absent in A. zangicus); face and clypeus yellow (Figs 4 C, 12 A) (face black and clypeus yellow in A. zangicus); truncate ventral margin of clypeus (Figs 4 C, 12 A) (concave in A. zangicus); metasomal tergites black with yellow markings (Fig. 4 F) (generally reddish-brown in A. zangicus).	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
4AF2797C5ED45B8689B23066B3088AD2.taxon	description	Description. The measurements were based on Taiwanese specimens (1 female and 3 males). Female (holotype). Head (Figs 4 A – C, 12 A): matt and granulate, HW / HL = 1.8; ocelli normal, with OD = 0.2 mm, POL / OD = 0.8, OOL / OD = 1.4, POL / OOL = 0.6; face matt and granulate, FW / FH = 1.6; clypeus polished and smooth with sparse punctures and subventral median elevation, rounded on ventral margin, CLW / CLH = 2.6; MSL / BMW = 0.5; mandible minutely coriaceous with minute punctures, teeth equal in length; flagellum with 45 segments; average ratio of basal five flagellomeres length 2.5: 1.4: 1.1: 1.2: 1.0. Mesosoma (Figs 4 A, D, E, 12 B, C): matt and granulate; pronotum with epomia weak, rugose at dorso-anterior corner; mesoscutum with MSSL / MSSW = 1.2, notauli short and indistinct, present near anterior margin; scutellum with SCL / SCW = 1.3, lateral carina absent; epicnemial carina weak, ~ 0.7 × height of mesopleuron; metapleuron with pleural carina and submetapleural carina strong and complete; juxtacoxal carina present posteriorly; propodeum with spiracle circular, maximum axis 1.0 × as minimum axis; anterior and posterior transverse carinae and lateral longitudinal carina absent; lateromedian longitudinal carina absent on anterior and median portions, present on posterior ~ 0.3 with area petiolaris opened anteriorly; average ratio of hind tarsomere length 4.0: 2.0: 1.5: 1.0: 1.0. Wings (Fig. 10 B): fore wing length 8.3 mm; areolet open and trapezoid with stalk 0.5 as long as 2 rs-m, receiving 2 m-cu at distal corner; RMI = 0.7; 1 cu-a almost vertical, opposite or slightly distad to M & RS, with BNI = 0.2. Hind wing length 6.0 mm; NI = 2.2; distal hamuli 6 – 7. Metasoma (Fig. 4 F – H): matt and granulate, except tergites behind T 2 weakly coriaceous; T 1 3.5 × as long as posterior width, 6.6 × as long as anterior width, 2.3 × as long as length of T 2; T 1 with latero-median carina absent, dorso-lateral carina and ventro-lateral carina strong and complete, spiracle at around middle of T 1, glymma distinct; T 2 0.8 × as long as posterior width, 1.3 × as long as anterior width, gastrocoeli shallow and indistinct, thyridia almost circular; ovipositor sheath 6.0 × as long as its maximum width in lateral view, as long as apical depth of metasoma. Color (Figs 4, 10 B): head, mesosoma, and metasoma generally black, and legs generally reddish-brown, except face, dorsal 1 / 2 of clypeus, malar space, mandibles, palpi, dorso-posterior corner of pronotum, latero-anterior corners of mesoscutum, dorso-anterior corner and ventral subtriangular marking (except enclosing area) of mesopleuron, tegula, scutellum, postscutellum, fore and mid coxae and trochanters, dorsal surface of hind coxa, hind second trochanter, median markings on tergites behind T 2 yellow; antenna, marking on ventral 1 / 2 of clypeus, tinged color of tergites behind T 2, ovipositor sheath reddish-brown; ventral side of hind coxa, hind trochanter, and hind femur black. Wings hyaline, veins blackish-brown, and pterostigma reddish-brown. Male. General structure and color similar to female, except notauli long and distinct, lateromedian longitudinal carinae of propodeum vestigial on median portion, and clypeus completely yellow, subtriangular marking with enclosing black area on ventral mesopleuron more distinct (Fig. 11 A – C). Male genitalia with gonostyle tapered and rounded apically, S 9 weakly concave on posterior margin, with subtriangular median area weakly sclerotized (Fig. 11 D – F). HW / HL = 1.8 – 1.9 (1.8 ± 0.06); OD = 0.15 – 0.19 (0.17 ± 0.02) mm, POL / OD = 0.5 – 0.9 (0.7 ± 0.2), OOL / OD = 1.2 – 1.6 (1.4 ± 0.18), POL / OOL = 0.4 – 0.6 (0.5 ± 0.09); FW / FH = 1.4 – 1.8 (1.6 ± 0.17), CLW / CLH = 3.0 – 4.0 (3.6 ± 0.54), MSL / BMW = 0.52 – 0.54 (0.52 ± 0.01); flagellum with 43 – 45 segments; average ratio of basal five flagellomeres length 2.3: 1.3: 1.2: 1.1: 1.0; MSSL / MSSW = 1.2 – 1.3 (1.3 ± 0.04); SCL / SCW = 1.1 – 1.2 (1.1 ± 0.07); maximum axis of propodeal spiracles 1.0 – 1.2 (1.1 ± 0.1) × as minimum axis; average ratio of hind tarsomere length 4.1: 2.2: 1.8: 1.0: 1.2; fore wing length 7.2 – 7.8 (7.5 ± 0.29) mm; areolet with stalk 0.4 – 0.6 (0.5 ± 0.1) × as long as 2 rs-m; RMI = 0.7 – 0.8 (0.8 ± 0.04); BNI = 0.1 – 0.2 (0.2 ± 0.03); hind wing length 5.2 – 5.6 (5.4 ± 0.23) mm; NI = 2.0 – 2.7 (2.4 ± 0.36); distal hamuli 6 – 7; T 1 2.7 – 3.1 (2.9 ± 0.16) × as long as posterior width, 6.1 – 6.6 (6.3 ± 0.29) × as long as anterior width, 1.3 – 1.6 (1.5 ± 0.11) × as long as length of T 2; T 2 1.2 – 1.4 (1.2 ± 0.1) × as long as posterior width, 1.6 – 1.9 (1.7 ± 0.13) × as long as anterior width.	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
4AF2797C5ED45B8689B23066B3088AD2.taxon	distribution	Distribution. Taiwan (Nantou).	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
4AF2797C5ED45B8689B23066B3088AD2.taxon	etymology	Etymology. The specific name flavomaculatus is derived from the Latin words flavo - (yellow) and maculatus (stained or spotted), referring to the yellow markings in the body color pattern of this new species. The name is an adjective.	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
DC23034A8D1952D2BEED552AEFA0D9C5.taxon	diagnosis	Diagnosis. According to Townes (1970), Gauld (1997), and Wharton (2015), this genus can be distinguished from other genera of Mesoleiini by the combination of the following characters: sculpture of head and mesosoma generally matt, granulate or minutely coriaceous; mandibular teeth usually equal in length (lower tooth longer in few Holarctic species and all known Neotropical species); notaulus usually distinct at least near anterior margin of mesoscutum; fore wing areolet usually present; fore wing 1 cu-a usually opposite to M & RS; T 1 long and slender, usually longer than 2.5 × its posterior width; T 1 with latero-median carinae absent, dorsal lateral carina distinct, and glymma always distinct; lateromedian longitudinal carinae of propodeum distinct posteriorly and forming an area petiolaris with median part of posterior transverse carinae. This genus is similar to Barytarbes, as some Alexeter species share the absence of anterior and median portions of the lateromedian longitudinal carinae on the propodeum and have mandibles with the lower tooth longer than the upper. However, Barytarbes typically features a shorter and broader T 1 and a propodeum with weaker lateromedian longitudinal carinae in the posterior part. Additionally, some Barytarbes species lack a glymma on the T 1, distinguishing them from Alexeter. Townes (1970) also mentioned that some Alexeter species are challenging to separate from genera such as Campodorus, Lagarotis, and Alcochera. Generally, Alexeter can be separated from Campodorus by having a clypeus with sharp, truncate or rounded ventral margin, flat in lateral view or with subventral transverse ridge or median elevation (clypeus strongly bulging medially and subapically with bilobed ventral margin in Campodorus), and areolet usually present (always absent in Campodorus). It can also be separated from Lagarotis in having mesopleuron granulate or minutely coriaceous (heavily sculptured, e. g., minutely rugose, in Lagarotis). Lastly, it can be separated from Alcochera by the mandible with teeth usually equal in length (lower tooth always longer than upper tooth in Alcochera), and notaulus distinct at least near anterior margin (weak in Alcochera).	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
FD6321A67A0C581BB8A6402A1ABEE64D.taxon	description	Figs 5 A – H, 10 C, 11 G – I; Suppl. material 3 Chinese vernacular name: 郁薇亞力姬蜂	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
FD6321A67A0C581BB8A6402A1ABEE64D.taxon	diagnosis	Diagnosis. This species can be distinguished from congeners by the combination of the following characters: fore wing length usually shorter than 10.0 mm (7.8 – 10.4 mm); ocelli large (OD = 0.18 – 0.25 mm; OOL / OD = 0.6 – 0.9); POL / OOL = 0.8 – 1.1; female with flagellum segments 46; fore wing areolet trapezoid with stalk, receiving 2 m-cu at distal corner (Fig. 10 C); fore wing 1 cu-a almost vertical, opposite or slightly distad to M & RS (Fig. 10 C); lateromedian longitudinal carinae of propodeum present posteriorly with the area petiolaris opened (Fig. 5 E); T 1 3.1 – 4.3 × its posterior width; head generally yellowish-white with frons and vertex brown to blackish-brown (Fig. 5 A, B); mesosoma, legs, and metasoma generally yellowish-brown, with mesoscutum having three distinct blackish-brown longitudinal stripes on the lobes (Fig. 5 A, D). This new species is most similar to A. clavator (Müller, 1776), A. monticola sp. nov., and A. mediolobus sp. nov. which share generally yellowish color in head, mesosoma, and metasoma, but can be distinguished from them by the POL / OOL ratio (0.8 – 1.1 vs 1.3 in Chinese A. clavator (Li and Sun 2022) and 1.1 – 1.5 in A. mediolobus sp. nov.), female OOL / OD ratio (0.7 – 0.9 vs 0.5 – 0.7 in A. mediolobus sp. nov. and A. monticola sp. nov.), length-to-posterior-width ratio of T 1 (3.1 – 4.3 vs 2.5 – 3.1 in A. mediolobus sp. nov.), female flagellomere counts (46 vs 40 – 43 in A. mediolobus sp. nov.), color patterns of mesoscutum (three stripes on each lobe vs absent in A. clavator and one on the median lobe in A. mediolobus sp. nov.), gena (yellowish-white vs yellowish-brown to brown in other species), vertex (brown vs yellowish-brown or orange in A. monticola sp. nov.), and fore and mid coxae (yellowish-white vs yellowish-brown in A. monticola sp. nov.). For other yellowish- or reddish-brown species A. nebulator (Thunberg, 1822), A. gracilentus (Holmgren, 1857), and A. luteifrons (Cresson, 1868), this new species can be easily distinguished from them by having a yellowish-white head (black in these species) and three blackish-brown stripes on the lobes of mesoscutum (stripes absent in these species). A comparative table of the measurements, ratios, and colors of this new species and other yellowish- or reddish-brown Alexeter species are provided in Suppl. material 3. Additionally, this new species may also be confused with two species of the genus Barytarbes, B. fulvus Sheng & Schönitzer, 2008 and B. nigrimaculatus Sheng & Sun, 2017, which share a generally yellowish-brown body color and the absence of lateromedian longitudinal carinae on the anterior and median portions of propodeum. However, this new species can be distinguished from both Barytarbes species by having mandibular teeth equal in length (Fig. 5 C) (lower tooth longer in both species), T 1 with distinct glymma (Fig. 5 G) (glymma absent in B. fulvus and indistinct in B. nigrimaculatus), T 1 3.1 – 4.3 × its posterior width (~ 1.9 × in B. nigrimaculatus, 4.0 × in B. fulvus), mesoscutum with three distinct blackish-brown longitudinal stripes (Fig. 5 D) (without stripes on mesoscutum in B. fulvus), and metasomal tergites without black markings (Fig. 5 A) (with black markings in B. nigrimaculatus).	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
FD6321A67A0C581BB8A6402A1ABEE64D.taxon	description	Description. The measurements were based on Taiwanese specimens (3 females and 7 males). Female. Head (Fig. 5 A – C): matt and minutely coriaceous, HW / HL = 1.6 – 1.9 (1.9, 1.8 ± 0.15); ocelli large, with OD = 0.20 – 0.25 (0.21, 0.22 ± 0.03) mm, POL / OD = 0.5 – 0.9 (0.9, 0.7 ± 0.19), OOL / OD = 0.7 – 0.9 (0.8, 0.8 ± 0.11), POL / OOL = 0.8 – 1.1 (1.1, 0.9 ± 0.18); face matt and minutely coriaceous, FW / FH = 1.37 – 1.44 (1.4, 1.41 ± 0.05); clypeus flat, polished and smooth with sparce punctures, truncate on ventral margin, CLW / CLH = 2.9 – 3.3 (3.3, 3.1 ± 0.28); MSL / BMW = 0.3 – 0.7 (0.3, 0.5 ± 0.21); mandible smooth with sparce punctures, teeth equal in length; flagellum with 46 (46) segments; average ratio of basal five flagellomeres length 2.5: 1.3: 1.1: 1.1: 1.0. Mesosoma (Fig. 5 A, D, E): matt and minutely coriaceous; pronotum with epomia weak; mesoscutum with MSSL / MSSW = 1.1 – 1.2 (1.1, 1.1 ± 0.07), notauli short and distinct near anterior margin; scutellum with SCL / SCW = 1.2 – 1.3 (1.2, 1.3 ± 0.01), lateral carina present anteriorly; epicnemial carina strong, ~ 0.6 × height of mesopleuron; metapleuron with pleural carina and submetapleural carina complete; juxtacoxal carina absent; propodeum with spiracle circular, maximum axis 1.0 – 1.1 (1.0, 1.04 ± 0.06) × as minimum axis; anterior transverse carinae absent; posterior transverse carina absent but vestigial medially; lateromedian longitudinal carinae absent on anterior and median portions, present on posterior ~ 0.3 – 0.4 with area petiolaris opened anteriorly; lateral longitudinal carinae vestigial posteriorly; average ratio of hind tarsomere length 4.1: 2.2: 1.7: 1.0: 1.2. Wings (Fig. 10 C): fore wing length 8.7 – 10.4 (8.94, 9.4 ± 0.91) mm; areolet open and trapezoid with stalk 0.4 – 0.5 (0.4, 0.4 ± 0.05) as long as 2 rs-m, receiving 2 m-cu at distal corner; RMI = 0.5 – 0.8 (0.8, 0.7 ± 0.12); 1 cu-a almost vertical, opposite or slightly distad to M & RS, with BNI = 0.1 – 0.2 (0.2, 0.2 ± 0.02). Hind wing length 6.8 – 7.9 (6.77, 7.2 ± 0.62) mm; NI = 1.7 – 3 (1.7, 2.2 ± 0.71); distal hamuli 5 – 6 (6 in left and 5 in right wing). Metasoma (Fig. 5 F – H): matt and minutely coriaceous; T 1 3.1 – 3.8 (3.1, 3.3 ± 0.43) × as long as posterior width, 6.2 – 8.2 (6.2, 7.0 ± 1.07) × as long as anterior width, 1.3 – 1.5 (1.3, 1.4 ± 0.09) × as long as length of T 2; T 1 with latero-median carina absent, dorso-lateral carina present basally, ventro-lateral carina complete, spiracle at around middle of T 1, glymma distinct; T 2 1.1 – 1.4 (1.2, 1.2 ± 0.13) × as long as posterior width, 1.9 – 2.3 (2.1, 2.1 ± 0.2) × as long as anterior width, gastrocoeli indistinct, thyridia deep and semi-circular; ovipositor sheath 3.3 – 5.6 (3.3, 4.6 ± 1.21) × as long as its maximum width in lateral view, shorter than apical depth of metasoma. Color (Figs 5, 10 C): head yellowish-white, except antenna yellowish-brown; vertex brown; frons and areas between lateral ocelli and eyes blackish-brown. Mesosoma, legs, and metasoma generally yellowish-brown, except pronotum, ventral and anterior area of mesopleuron, mesoscutum, scutellum, fore and mid coxae, and trochanters yellowish-white, three longitudinal stripes on lobes of mesoscutum and T 1 dark brown to blackish-brown. Wings hyaline, veins blackish-brown, and pterostigma pale yellowish-brown. Male. General structure and color similar to female. Male genitalia with gonostyle tapered and rounded apically, S 9 weakly concave on posterior margin, completely sclerotized (Fig. 11 G – I). HW / HL = 1.7 – 1.9 (1.8 ± 0.08); OD = 0.18 – 0.23 (0.21 ± 0.02) mm, POL / OD = 0.6 – 0.9 (0.7 ± 0.12), OOL / OD = 0.6 – 0.8 (0.7 ± 0.09), POL / OOL = 0.9 – 1.1 (1 ± 0.08); FW / FH = 1.3 – 1.7 (1.4 ± 0.13), CLW / CLH = 3.0 – 3.6 (3.3 ± 0.29), MSL / BMW = 0.3 – 0.4 (0.3 ± 0.05); flagellum with 42 – 47 segments; average ratio of basal five flagellomeres length 2.3: 1.3: 1.1: 1.1: 1.0; MSSL / MSSW = 1.1 – 1.3 (1.2 ± 0.09); SCL / SCW = 1.0 – 1.4 (1.2 ± 0.12); maximum axis of propodeal spiracles 1.0 – 1.2 (1.1 ± 0.09) × as minimum axis; average ratio of hind tarsomere length 3.8: 2.0: 1.6: 1.0: 1.1; fore wing length 7.8 – 9.7 (8.9 ± 0.63) mm; areolet with stalk 0.3 – 0.5 (0.4 ± 0.06) × as long as 2 rs-m; RMI = 0.6 – 0.8 (0.7 ± 0.09); BNI = 0.1 – 0.2 (0.2 ± 0.03); hind wing length 5.7 – 7.0 (6.4 ± 0.47) mm; NI = 1.8 – 3.0 (2.5 ± 0.43); distal hamuli 4 – 7; T 1 3.5 – 4.3 (3.9 ± 0.29) × as long as posterior width, 7.0 – 10.4 (8.2 ± 1.18) × as long as anterior width, 1.3 – 1.5 (1.4 ± 0.06) × as long as length of T 2; T 2 1.2 – 1.6 (1.4 ± 0.16) × as long as posterior width, 1.6 – 2.6 (2.2 ± 0.32) × as long as anterior width.	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
FD6321A67A0C581BB8A6402A1ABEE64D.taxon	distribution	Distribution. Taiwan (Hualien andTaichung).	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
FD6321A67A0C581BB8A6402A1ABEE64D.taxon	etymology	Etymology. The eponym of the specific name hsiaoae is the Chinese family name ‘ Hsiao’ combined with the feminine suffix ‘ - ae ’. It is dedicated to memory the deceased collector of the type series, Ms. Yu-Wei Hsiao. The name is a noun in the genitive case.	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
83E0F047930952A78AD333EE4828E7EA.taxon	description	Figs 6 A – H, 10 D, 11 J – L; Suppl. material 3 Chinese vernacular name: 中褐亞力姬蜂	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
83E0F047930952A78AD333EE4828E7EA.taxon	diagnosis	Diagnosis. This species can be distinguished from congeners by the combination of the following characters: fore wing length usually shorter than 10.0 mm (7.7 – 10.1 mm); ocelli large (OD = 0.20 – 0.27 mm; OOL / OD = 0.5 – 0.7); POL / OOL = 1.1 – 1.5; female with flagellum segments 40 – 43; fore wing areolet trapezoid with stalk, receiving 2 m-cu at distal corner (Fig. 10 D); fore wing 1 cu-a almost vertical and opposite to M & RS (Fig. 10 D); lateromedian longitudinal carinae of propodeum present posteriorly with the area petiolaris almost closed (Fig. 6 E); posterior transverse carina present (Fig. 6 E); T 1 2.5 – 3.1 × its posterior width; head, mesosoma, legs, and metasoma generally brown to yellowish-brown, with mesoscutum having single blackish-brown longitudinal stripe on the median lobe (Fig. 6 A, D). This new species is most similar to A. clavator, A. hsiaoae sp. nov., and A. monticola sp. nov. in body color but can be distinguished from them by the POL / OOL ratio (1.1 – 1.5 vs 0.8 – 1.1 in A. hsiaoae sp. nov. and 0.8 – 1.2 in A. monticola sp. nov.), female OOL / OD ratio (0.5 – 0.7 vs 0.7 – 0.9 in A. hsiaoae sp. nov.), length-to-posterior-width ratio of T 1 (2.5 – 3.1 vs 3.1 – 4.3 in A. hsiaoae sp. nov., 3.4 – 3.9 in A. monticola sp. nov., and ~ 3.3 in Chinese A. clavator), female flagellomere counts (40 – 43 vs 46 in A. hsiaoae sp. nov. and 46 – 52 in A. monticola sp. nov.), color patterns of mesoscutum (one stripe on the median lobe vs absent in A. clavator and three stripes on each lobe in A. hsiaoae sp. nov. and A. monticola sp. nov.), gena (brown vs yellowish-white in A. hsiaoae sp. nov. and sometimes orange in A. monticola sp. nov.), and fore and mid coxae (yellowish-brown vs yellowish-white in A. hsiaoae sp. nov.). This new species can also be distinguished from other yellowish- and reddish-brown species A. nebulator, A. gracilentus, and A. luteifrons by having yellowish-brown head (black in these species) and the blackish-brown stripe on the median lobe of mesoscutum (yellowish-brown in these species). A comparative table of the measurements, ratios, and colors of this new species and other yellowish- or reddish-brown Alexeter species are provided in Suppl. material 3.	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
83E0F047930952A78AD333EE4828E7EA.taxon	description	Description. The measurements are based on Taiwanese specimens (6 females and 1 male). Female. Head (Fig. 6 A – C): matt and minutely coriaceous, HW / HL = 1.7 – 2.1 (1.7, 1.9 ± 0.12); ocelli large, with OD = 0.20 – 0.27 (0.27, 0.25 ± 0.03) mm, POL / OD = 0.6 – 0.7 (0.6, 0.7 ± 0.05), OOL / OD = 0.5 – 0.7 (0.5, 0.5 ± 0.06), POL / OOL = 1.1 – 1.5 (1.1, 1.2 ± 0.13); face matt and minutely coriaceous, FW / FH = 1.4 – 1.6 (1.5, 1.5 ± 0.08); clypeus matt and evenly punctate with subventral median elevation, truncate or slightly concave on ventral margin, CLW / CLH = 3.0 – 3.5 (3.5, 3.2 ± 0.19); MSL / BMW = 0.3 – 0.4 (0.3, 0.3 ± 0.04); mandible densely punctate in dorsal surface, teeth equal in length; flagellum with 40 – 43 (43) segments; average ratio of basal five flagellomeres length 2.2: 1.2: 1.1: 1.1: 1.0. Mesosoma (Fig. 6 A, D, E): matt and granulate; pronotum with epomia absent, carinate at dorso-anterior corner; mesoscutum with MSSL / MSSW = 1.1 – 1.2 (1.1, 1.2 ± 0.06), notauli short, distinct near anterior margin; scutellum with SCL / SCW = 1.0 – 1.4 (1.4, 1.3 ± 0.15), lateral carina present at base; epicnemial carina strong, ~ 0.7 × height of mesopleuron; metapleuron with pleural carina and submetapleural carina complete; juxtacoxal carina present posteriorly; propodeum with spiracle suboval, maximum axis 1.0 – 1.2 (1.0, 1.1 ± 0.08) × as minimum axis; anterior transverse carina absent; posterior transverse carina present medially; lateromedian longitudinal carinae absent on anterior and median portions, present on posterior ~ 0.2 with area petiolaris almost closed; lateral longitudinal carinae weak; average ratio of hind tarsomere length 3.9: 2.1: 1.6: 1.0: 1.2. Wings (Fig. 10 D): fore wing length 7.7 – 10.1 (10.12, 9.4 ± 0.84) mm; areolet open and trapezoid with stalk 0.3 – 0.4 (0.3, 0.3 ± 0.05) as long as 2 rs-m, receiving 2 m-cu at distal corner; RMI = 0.6 – 0.7 (0.7, 0.6 ± 0.05); 1 cu-a almost vertical and opposite to M & RS, with BNI = 0.1 – 0.2 (0.1, 0.1 ± 0.04). Hind wing length 5.6 – 7.4 (7.39, 6.8 ± 0.6) mm; NI = 1.9 – 2.8 (2.4 ± 0.32); distal hamuli 6 – 8 (6). Metasoma (Fig. 6 F – H): matt and coriaceous; T 1 2.5 – 3.1 (2.8, 2.9 ± 0.22) × as long as posterior width, 5.8 – 8.0 (7.7, 7.0 ± 0.91) × as long as anterior width, 1.4 – 1.5 (1.5, 1.5 ± 0.03) × as long as length of T 2; T 1 with latero-median carina absent, dorso-lateral carina present on anterior 0.1 and posterior 0.2, ventro-lateral carina complete, spiracle at around middle of T 1, glymma distinct; T 2 0.9 – 1.3 (1.1, 1.2 ± 0.14) × as long as posterior width, 1.4 – 2.1 (1.8, 1.8 ± 0.22) × as long as anterior width, gastrocoeli indistinct, thyridia indistinct and circular; ovipositor sheath 2.7 – 5.5 (3.5, 3.5 ± 1.02) × as long as its maximum width in lateral view, shorter than apical depth of metasoma. Color (Fig. 6, 10 D): head, mesosoma, legs, and metasoma generally brown, except face, dorsal 0.7 of clypeus, malar space, gena, mandible base, and dorso-posterior corner of pronotum yellowish-white; palpi, pronotum, dorso-anterior corner and speculum of mesopleuron, fore and mid coxae, and all trochanters and tarsi pale yellowish-brown; frons, vertex, median lobe of mesoscutum in anterior 1 / 2, lateral sides of scutellum blackish-brown. Wings hyaline, veins blackish-brown, pterostigma pale yellowish-brown. Male. General structure and color similar to female. Male genitalia with gonostyle tapered and rounded apically, S 9 weakly concave on posterior margin, completely sclerotized (Fig. 11 J – L). HW / HL = 1.9; OD = 0.25 mm, POL / OD = 0.6, OOL / OD = 0.5, POL / OOL = 1.3; FW / FH = 1.5, CLW / CLH = 3.0, MSL / BMW = 0.4; flagellum segments broken; average ratio of basal five flagellomeres length 2.2: 1.3: 1.1: 1.2: 1.0; MSSL / MSSW = 1.1; SCL / SCW = 1.2; maximum axis of propodeal spiracles 1.1 × as minimum axis; average ratio of hind tarsomere length 3.7: 2.1: 1.6: 1.0: 1.3; fore wing length 8.9 mm; areolet with stalk 0.1 × as long as 2 rs-m; RMI = 0.3; BNI = 0.1; hind wing length 6.6 mm; NI = 1.9; distal hamuli 6; T 1 2.9 × as long as posterior width, 8.9 × as long as anterior width, 1.5 × as long as length of T 2; T 2 1.3 × as long as posterior width, 1.7 × as long as anterior width.	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
83E0F047930952A78AD333EE4828E7EA.taxon	distribution	Distribution. Taiwan (Hualien and Nantou).	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
83E0F047930952A78AD333EE4828E7EA.taxon	etymology	Etymology. The specific name mediolobus is derived from the Latin medio - (meaning medial) plus lobus (meaning lobe). It refers to the median lobe of mesoscutum in this species with a distinct blackish-brown marking. The name is an adjective.	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
2975436C34585226ABF2F6E7E24AD1C0.taxon	description	Figs 7 A – H, 10 E; Suppl. material 3 Chinese vernacular name: 山亞力姬蜂	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
2975436C34585226ABF2F6E7E24AD1C0.taxon	diagnosis	Diagnosis. This species can be distinguished from congeners by the combination of the following characters: fore wing length longer than 10.0 mm (10.7 – 12.0 mm); ocelli large (OD = 0.21 – 0.29 mm; OOL / OD = 0.5 – 0.8); POL / OOL = 0.8 – 1.2; female with flagellum segments 46 – 52; fore wing areolet triangular with stalk, receiving 2 m-cu at distal corner (Fig. 10 E); fore wing 1 cu-a vertical and opposite to M & RS (Fig. 10 E); lateromedian longitudinal carinae of propodeum present posteriorly and forming the area petiolaris opened (Fig. 7 E); posterior transverse carina absent (Fig. 7 E); metasomal T 1 3.4 – 3.9 × its posterior width; head, mesosoma, legs, and metasoma generally yellowish-brown to orange, with mesoscutum having three blackish-brown longitudinal stripes on the lobes (Fig. 7 A, D). This new species is most similar to A. clavator, A. hsiaoae sp. nov., and A. mediolobus sp. nov. in body color but can be distinguished by the POL / OOL ratio (0.8 – 1.2 vs 1.1 – 1.5 in A. mediolobus sp. nov.), female OOL / OD ratio (0.5 – 0.7 vs 0.7 – 0.9 in A. hsiaoae sp. nov.), length-to-posterior-width ratio of T 1 (3.4 – 3.9 vs 2.5 – 3.1 in A. mediolobus sp. nov., and ~ 3.3 in Chinese A. clavator), female flagellomere counts (46 – 52 vs 40 – 43 in A. mediolobus sp. nov.), color patterns of mesoscutum (three stripes on each lobe vs absent in A. clavator and one stripe on the median lobe in A. mediolobus sp. nov.), gena (orange or yellowish-brown vs yellowish-white in A. hsiaoae sp. nov.), and fore and mid coxae color (yellowish-brown vs yellowish-white in A. hsiaoae sp. nov.). This new species can also be distinguished from other yellowish- and reddish-brown species A. nebulator, A. gracilentus, and A. luteifrons by having yellowish-brown head (black in these species) and three brown stripes on the lobes of mesoscutum (stripes absent in these species). A comparative table of measurements, ratios, and colors of this new species and other yellowish- or reddish-brown Alexeter species are provided in Suppl. material 3.	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
2975436C34585226ABF2F6E7E24AD1C0.taxon	description	Description. The measurements are based on Taiwanese specimens (10 females). Female. Head (Fig. 7 A – C): matt and minutely coriaceous, HW / HL = 1.6 – 1.8 (1.8, 1.7 ± 0.08); ocelli large, with OD = 0.21 – 0.29 (0.24, 0.25 ± 0.03) mm, POL / OD = 0.6 – 0.8 (0.6, 0.7 ± 0.09), OOL / OD = 0.5 – 0.7 (0.7, 0.7 ± 0.09), POL / OOL = 0.8 – 1.2 (0.9, 1.0 ± 0.14); face matt and coriaceous, FW / FH = 1.3 – 1.6 (1.5, 1.5 ± 0.07); clypeus polished and smooth with sparce punctures and subventral median elevation, truncate on ventral margin, CLW / CLH = 3.0 – 3.8 (3.4, 3.4 ± 0.27); MSL / BMW = 0.4 – 0.5 (0.4, 0.4 ± 0.03); mandible smooth with sparce punctures, teeth equal in length; flagellum with 46 – 52 (49) segments; average ratio of basal five flagellomeres length 2.2: 1.3: 1.1: 1.0: 1.0. Mesosoma (Fig. 7 A, D, E): matt and granulate; pronotum with epomia absent, weakly and densely carinate at dorso-anterior corner; mesoscutum with MSSL / MSSW = 1.1 – 1.3 (1.3, 1.2 ± 0.06), notauli distinct on anterior ~ 0.3; scutellum with SCL / SCW = 0.9 – 1.4 (1.1, 1.2 ± 0.16), lateral carina absent; epicnemial carina strong, ~ 0.5 × height of mesopleuron; metapleuron with pleural carina and submetapleural carina complete; juxtacoxal carina absent; propodeum with spiracle circular or suboval, maximum axis 1.0 – 1.1 (1.1, 1.1 ± 0.05) × as minimum axis; anterior and posterior transverse carinae absent; lateromedian longitudinal carinae absent on anterior and median portions, present on posterior ~ 0.2 with area petiolaris opened anteriorly; lateral longitudinal carinae vestigial posteriorly; average ratio of hind tarsomere length 4.0: 2.0: 1.6: 1.0: 1.1. Wings (Fig. 10 E): fore wing length 10.7 – 12.0 (10.74, 11.2 ± 0.48) mm; areolet open and triangular with stalk 0.3 – 0.5 (0.4, 0.4 ± 0.05) as long as 2 rs-m, receiving 2 m-cu at distal corner; RMI = 0.6 – 1.0 (0.7, 0.7 ± 0.10); 1 cu-a vertical and almost opposite to M & RS, with BNI = 0.1 – 0.2 (0.1, 0.17 ± 0.03). Hind wing length 8.0 – 9.0 (8.01, 8.3 ± 0.37) mm; NI = 1.8 – 3.7 (2.1, 2.3 ± 0.61); distal hamuli 6 – 9 (6 in left and 7 in right wing). Metasoma (Fig. 7 F – H): matt and granulate; T 1 3.4 – 3.9 (3.9, 3.5 ± 0.16) × as long as posterior width, 6.7 – 8.2 (7.3, 7.5 ± 0.55) × as long as anterior width, 1.4 – 1.6 (1.4, 1.4 ± 0.06) × as long as length of T 2; T 1 with latero-median carina absent, dorso-lateral carina present on basal ~ 0.1 and apical 0.3, ventro-lateral carina complete, spiracle at around basal 0.4, glymma distinct; T 2 1.2 – 1.5 (1.5, 1.3 ± 0.08) × as long as posterior width, 1.9 – 2.4 (2.3, 2.1 ± 0.17) × as long as anterior width, gastrocoeli indistinct, thyridia long ellipse; ovipositor sheath 2.3 – 3.3 (2.8, 2.8 ± 0.26) × as long as its maximum width in lateral view, shorter than apical depth of metasoma. Color (Fig. 7, 8 E): head, mesosoma, legs, and metasoma generally yellowish-brown to orange, except face, clypeus, and malar space yellowish-white; frons (in most paratypes) and three longitudinal stripes on lobes of mesoscutum blackish-brown; metasoma in some paratypes brown. Wings hyaline, veins yellowish-brown, and pterostigma pale yellowish-brown. Male. Unknown.	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
2975436C34585226ABF2F6E7E24AD1C0.taxon	distribution	Distribution. Taiwan (Nantou).	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
2975436C34585226ABF2F6E7E24AD1C0.taxon	etymology	Etymology. The specific name monticola is derived from Latin, meaning existing or having a habitat in or on mountains. The name is an adjective.	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
1EE83A6EA8A058A192692EB8B22A063B.taxon	description	Figs 8 A – H, 10 F, 11 P, Q, 12 D – F; Suppl. material 3 Chinese vernacular name: 偽藏亞力姬蜂	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
1EE83A6EA8A058A192692EB8B22A063B.taxon	diagnosis	Diagnosis. This species can be distinguished from congeners by the combination of the following characters: fore wing length usually longer than 10.0 mm (9.6 – 12.2 mm); ocelli normal (OD = 0.19 – 0.25 mm; OOL / OD = 0.8 – 1.1); POL / OOL = 0.7 – 1.1; clypeus rounded on ventral margin, mandibles with lower tooth slightly longer than upper tooth (Figs 8 C, 12 D); fore wing areolet absent (Fig. 10 F); fore wing 1 cu-a inclivous and distad to M & RS (Fig. 10 F); lateromedian longitudinal carinae of propodeum complete, with the area petiolaris closed (Fig. 8 E); posterior transverse carina present (Fig. 8 E); male gonostyle broad (Fig. 11 Q); general body color reddish-brown or brown, with yellow markings on the latero-anterior corners of mesoscutum (Figs 8, 12 E, F). This new species is most similar to A. zangicus Sheng, Sun & Li, 2020 which shares the absence of fore wing areolet and the yellow markings on the latero-anterior corners of mesoscutum. It can be distinguished from A. zangicus by the following characters: body color generally brown to reddish-brown (Figs 8, 12 E, F) (black in A. zangicus); face yellowish-brown with median blackish-brown marking in female and yellow in male, clypeus entirely yellow (Figs 8 C, 12 D) (face black and clypeus yellow in A. zangicus); rounded ventral margin of clypeus (Figs 8 C, 12 D) (concave in A. zangicus). It is also similar to the red-type male of A. rufispeculus sp. nov. which shares the reddish-brown body color and mesoscutal yellow marking but differs in the following characters: clypeus rounded (Figs 8 C, 12 D) (truncate in A. rufispeculus); mandible with lower tooth slightly longer than upper tooth (Figs 8 C, 12 D) (equal in length in A. rufispeculus); areolet absent (Fig. 10 F) (present in A. rufispeculus); lateromedian longitudinal carinae of propodeum complete (Fig. 8 E) (absent on the anterior and median portions of propodeum in A. rufispeculus); T 1 2.3 – 3.0 × its posterior width (3.0 – 4.3 × in A. rufispeculus); and male gonostyle broad (Fig. 11 Q) (normal in A. rufispeculus).	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
1EE83A6EA8A058A192692EB8B22A063B.taxon	description	Description. The measurements are based on Taiwanese specimens (6 females and 1 male). Female. Head (Figs 8 A – C, 12 D): polished and minutely coriaceous, HW / HL = 1.6 – 1.8 (1.7, 1.7 ± 0.09); ocelli normal to large, with OD = 0.19 – 0.25 (0.19, 0.22 ± 0.02) mm, POL / OD = 0.6 – 1.0 (0.7, 0.8 ± 0.13), OOL / OD = 0.8 – 1.1 (1.1, 0.9 ± 0.12), POL / OOL = 0.7 – 1.1 (0.7, 0.9 ± 0.16); face polished and minutely coriaceous, FW / FH = 1.4 – 1.7 (1.7, 1.5 ± 0.1); clypeus flat, polished and smooth with sparse punctures, rounded on ventral margin, CLW / CLH = 2.8 – 3.3 (2.8, 3.1 ± 0.23); MSL / BMW = 0.3 – 0.5 (0.4, 0.4 ± 0.07); mandible smooth with sparce punctures, lower tooth slightly longer than upper tooth; flagellum with 42 – 48 (48) segments; average ratio of basal five flagellomeres length 2.5: 1.3: 1.2: 1.1: 1.0. Mesosoma (Figs 8 D, E, 12 E, F): matt and granulate with dense punctures, mesopleuron minutely coriaceous with weak rugae behind epicnemial carina; pronotum with epomia absent, rugose on dorso-anterior corner; mesoscutum with MSSL / MSSW = 1.2 – 1.3 (1.2, 1.2 ± 0.03), notauli long, reaching 1 / 2 of mesoscutum; scutellum with SCL / SCW = 1.0 – 1.4 (1.4, 1.2 ± 0.16), lateral carina absent; epicnemial carina weak, as long as height of mesopleuron; metapleuron with pleural carina and submetapleural carina complete; juxtacoxal carina long and jointing submetapleural carina anteriorly; propodeum with spiracle suboval, maximum axis 1.1 – 1.3 (1.1, 1.2 ± 0.11) × as minimum axis; anterior transverse carina absent; posterior transverse carina present medially at posterior ~ 0.4; lateromedian longitudinal carinae present on anterior and median portions, with area petiolaris closed; lateral longitudinal carinae well-developed; average ratio of hind tarsomere length 4.4: 2.2: 1.6: 1.0: 1.1. Wings (Fig. 10 F): fore wing length 10.1 – 12.2 (11.99, 11.4 ± 0.77) mm; areolet absent; RMI = 0.7 – 0.9 (0.9, 0.8 ± 0.11); 1 cu-a inclivous and distad to M & RS, with BNI = 0.2 – 0.3 (0.2, 0.3 ± 0.03). Hind wing length 7.4 – 8.8 (8.65, 8.4 ± 0.54) mm; NI = 2.3 – 4.3 (3.0, 3.1 ± 0.75); distal hamuli 7 – 8 (8). Metasoma (Fig. 8 F – H): polished and minutely coriaceous; T 1 2.3 – 2.8 (2.5, 2.7 ± 0.21) × as long as posterior width, 5.8 – 7.0 (6.2, 6.4 ± 0.45) × as long as anterior width, 1.5 – 1.7 (1.5, 1.6 ± 0.09) × as long as length of T 2; T 1 with latero-median carina absent, dorso-lateral carina weak, distinct anteriorly, ventro-lateral carina complete, spiracle at around middle of T 1, glymma distinct; T 2 0.9 – 1.2 (1.0, 1.0 ± 0.08) × as long as posterior width, 1.3 – 1.5 (1.4, 1.4 ± 0.07) × as long as anterior width, gastrocoeli indistinct, thyridia indistinct and semi-circular; ovipositor sheath 3.1 – 4.6 (4.6, 3.7 ± 0.59) × as long as its maximum width in lateral view, shorter than apical depth of metasoma. Color (Figs 8, 10 F): head, mesosoma, legs, and metasoma generally brown to reddish-brown, except frons, areas between lateral ocelli and eyes, median area of face, posterior area of mesoscutum, pronotum, dorsal portion of mesopleuron, ventro-apical metapleuron, glymma of T 1, and ovipositor sheath blackish-brown or tinged with blackish-brown; clypeus, mandibles, palpi, dorsal surface and dorso-posterior corner of pronotum, latero-anterior corners of mesoscutum, tegula, dorso-anterior corner of mesopleuron, scutellum, and mid and hind tarsi yellow. Wings hyaline, veins and pterostigma brown or blackish-brown. Male. General structure and color similar to female, except general color blackish-brown (Fig. 12 D – F); whole face, ventral markings of mesopleuron, fore and mid coxae, and anterior 1 / 2 of T 3 yellow; yellow markings on lateral anterior corners of mesoscutum more distinct and extend posteriorly through notauli (Fig. 12 E, F). Male genitalia with gonostyle broad, tapered and rounded apically (Fig. 11 P, Q). HW / HL = 1.9; OD = 0.25 mm, POL / OD = 0.5, OOL / OD = 0.7, POL / OOL = 0.7; FW / FH = 1.6, CLW / CLH = 2.5, MSL / BMW = 0.3; flagellum with 42 segments; average ratio of basal five flagellomeres length 2.6: 1.4: 1.2: 1.3: 1.0; MSSL / MSSW = 1.2; SCL / SCW = 1.1; maximum axis of propodeal spiracles 1.0 × as minimum axis; average ratio of hind tarsomere length 4.3: 2.1: 1.6: 1.0: 1.0; fore wing length 9.6 mm; RMI = 0.7; BNI = 0.3; hind wing length 6.7 mm; NI = 3.0; distal hamuli 7 – 8; T 1 3.0 × as long as posterior width, 6.2 × as long as anterior width, 1.7 × as long as length of T 2; T 2 1.0 × as long as posterior width, 1.5 × as long as anterior width.	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
1EE83A6EA8A058A192692EB8B22A063B.taxon	distribution	Distribution. Taiwan (Nantou).	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
1EE83A6EA8A058A192692EB8B22A063B.taxon	etymology	Etymology. The specific name pseudozangicus is derived from the name of the species most morphologically similar to this new species, A. zangicus Sheng, Sun & Li, 2020. The name is an adjective.	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
E67E4508455D594F816A1ECA66892E68.taxon	description	Figs 9 A – H, 10 G, 11 M – O, 12 G – K; Suppl. material 3 Chinese vernacular name: 鏡紅亞力姬蜂	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
E67E4508455D594F816A1ECA66892E68.taxon	diagnosis	Diagnosis. This species can be distinguished from congeners by the combination of the following characters: ocelli large (OD = 0.20 – 0.30 mm; OOL / OD = 0.6 – 0.9); POL / OOL = 0.6 – 1.1; clypeus truncate on ventral margin (Figs 9 C, 12 G); fore wing areolet triangular with stalk, receiving 2 m-cu at distal corner (Fig. 10 G); fore wing vertical, basad, opposite, or distad to M & RS (Fig. 10 G); lateromedian longitudinal carinae of propodeum present posteriorly with the area petiolaris opened (Fig. 10 E); posterior transverse carina present (Fig. 10 E); general body color variable from reddish-brown (red type) (Fig. 12 J, K) to blackish-brown (black type) (Figs 9, 12 G – I); speculum on the mesopleuron usually reddish-brown, rarely not (Figs 9 A, 12 I – K); males with yellow markings on the latero-anterior corners of mesoscutum (Fig. 12 H – J). The red-type male of this new species is similar to the female of A. pseudozangicus sp. nov. which shares the reddish-brown body color and yellow markings on the latero-anterior corners of the mesoscutum but differs in the following characters: clypeus truncate (Figs 9 C, 12 G) (rounded in A. pseudozangicus); mandible with both teeth equal in length (Figs 9 C, 12 G) (lower tooth slightly longer than upper tooth in A. pseudozangicus); areolet present (Fig. 10 G) (absent in A. pseudozangicus); lateromedian longitudinal carinae of propodeum absent on the anterior and median portions (Fig. 9 E) (complete in A. pseudozangicus); T 1 3.0 – 4.3 × its posterior width (2.3 – 3.0 × in A. pseudozangicus); and the width of male gonostyle normal (Fig. 11 O) (broad in A. pseudozangicus).	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
E67E4508455D594F816A1ECA66892E68.taxon	description	Description. The measurements were based on Taiwanese specimens (15 females and 8 males). Female. Head (Figs 9 A – C, 12 G): matt and granulate, HW / HL = 1.7 – 2.0 (1.9, 1.8 ± 0.11); ocelli large, with OD = 0.20 – 0.30 (0.26, 0.26 ± 0.03) mm, POL / OD = 0.4 – 0.8 (0.4, 0.5 ± 0.10), OOL / OD = 0.6 – 0.8 (0.7, 0.7 ± 0.06), POL / OOL = 0.6 – 1.1 (0.6, 0.8 ± 0.13); face matt and granulate, FW / FH = 1.2 – 1.5 (1.5, 1.4 ± 0.08); clypeus smooth with sparce punctures, truncate, and having transverse median ridge on ventral margin, CLW / CLH = 2.6 – 4.3 (3.7, 3.1 ± 0.43); MSL / BMW = 0.4 – 0.5 (0.4, 0.4 ± 0.04); mandible evenly punctate in dorsal surface, teeth equal in length or with lower tooth slightly longer than upper tooth; flagellum with 41 – 51 (49) segments; average ratio of basal five flagellomeres length 2.5: 1.3: 1.1: 1.1: 1.0. Mesosoma (Figs 9 A, D, E, 12 H – K): polished and granulate; pronotum with epomia absent, weakly rugose on dorso-anterior corner; mesoscutum with MSSL / MSSW = 1.1 – 1.3 (1.1, 1.2 ± 0.06), notauli short, distinct on anterior ~ 0.2; scutellum with SCL / SCW = 0.9 – 1.4 (1.4, 1.1 ± 0.15), lateral carina present anteriorly; epicnemial carina weak, ~ 1 / 2 as height of mesopleuron; metapleuron with pleural carina and submetapleural carina complete; juxtacoxal carina vestigial posteriorly; propodeum with spiracle circular to suboval, maximum axis 1.0 – 1.3 (1.0, 1.1 ± 0.09) × as minimum axis; anterior transverse carina absent; posterior transverse carina present medially; lateromedian longitudinal carinae absent on anterior and median portions, present on posterior ~ 0.25 with area petiolaris opened anteriorly; lateral longitudinal carinae absent; average ratio of hind tarsomere length 4.3: 2.2: 1.7: 1.0: 1.2. Wings (Fig. 10 G): fore wing length 8.4 – 10.9 (10.7, 10.0 ± 0.66) mm; areolet triangular with stalk 0.3 – 0.6 (0.4, 0.5 ± 0.09) as long as 2 rs-m, receiving 2 m-cu at distal corner; RMI = 0.6 – 0.9 (0.7, 0.7 ± 0.08); 1 cu-a vertical, basad, opposite, or distad to M & RS, with BNI = 0.1 – 0.3 (0.2, 0.2 ± 0.05). Hind wing length 6.2 – 8.2 (8.0, 7.5 ± 0.51) mm; NI = 1.6 – 3.0 (1.7, 2.2 ± 0.44); distal hamuli 6 – 8 (6). Metasoma (Fig. 9 F – H): polished and minutely coriaceous; T 1 3.0 – 4.2 (3.3, 3.2 ± 0.31) × as long as posterior width, 5.8 – 9.5 (6.7, 7.2 ± 1.06) × as long as anterior width, 1.3 – 1.7 (1.4, 1.4 ± 0.09) × as long as length of T 2; T 1 with latero-median carina absent, dorso-lateral carina and ventro-lateral carina complete and varied from weak to strong, spiracle at around middle of T 1, glymma distinct; T 2 1.1 – 1.5 (1.5, 1.3 ± 0.09) × as long as posterior width, 1.6 – 2.3 (2.0, 1.8 ± 0.17) × as long as anterior width, gastrocoeli shallow and indistinct, thyridia circular; ovipositor sheath 2.4 – 5.5 (5.5, 3.6 ± 0.88) × as long as its maximum width in lateral view, shorter than apical depth of metasoma. Color (Figs 9, 10 G, 12 K): variable, head, and legs generally reddish-brown, mesosoma and metasoma generally blackish-brown or brown in black-type specimens (including holotype) and reddish-brown in red-type specimens, except antenna, fore legs except tarsi, mid coxae, and trochanters yellowish-brown; face, clypeus (whole or in dorsal 1 / 2), mandibles, malar space, palpi, tegula, all tarsi, hind tibiae (whole or at least in basal 0.6) yellow or pale yellow; frons, ocellar area, ventral marking of clypeus and T 1 (in some specimens) blackish-brown; speculum and scutellum always reddish-brown; and metasomal tergites behind T 3 reddish-brown, sometimes tinged with red. Wings hyaline, veins blackish-brown or yellowish-brown, pterostigma pale yellowish-brown. Male. General structure and color similar to female, except latero-anterior margin of mesoscutum and ventral side of mesopleuron with pale yellow marking (Fig. 12 G – J). Male genitalia with gonostyle tapered and rounded apically, S 9 weakly concave on posterior margin, with subtriangular median area weakly sclerotized (Fig. 11 M – O). HW / HL = 1.6 – 2.0 (1.8 ± 0.18); OD = 0.24 – 0.28 (0.26 ± 0.01) mm, POL / OD = 0.5 – 0.7 (0.6 ± 0.08), OOL / OD = 0.6 – 0.9 (0.7 ± 0.08), POL / OOL = 0.6 – 1.0 (0.8 ± 0.14); FW / FH = 1.2 – 1.4 (1.3 ± 0.05), CLW / CLH = 2.7 – 3.2 (3.0 ± 0.2), MSL / BMW = 0.2 – 0.4 (0.3 ± 0.05); flagellum with 43 – 49 segments; average ratio of basal five flagellomeres length 2.3: 1.2: 1.1: 1.1: 1.0; MSSL / MSSW = 1.2 – 1.3 (1.2 ± 0.06); SCL / SCW = 1.0 – 1.2 (1.1 ± 0.06); maximum axis of propodeal spiracles 1.0 – 1.1 (1.0 ± 0.06) × as minimum axis; average ratio of hind tarsomere length 4.1: 2.2: 1.7: 1.0: 1.1; fore wing length 8.6 – 10.1 (9.5 ± 0.52) mm; RMI = 0.6 – 0.9 (0.7 ± 0.12); BNI = 0.1 – 0.2 (0.2 ± 0.04); hind wing length 6.5 – 7.5 (7 ± 0.41) mm; NI = 1.5 – 3.7 (2.4 ± 0.75); distal hamuli 6 – 8; T 1 2.9 – 4.1 (3.5 ± 0.34) × as long as posterior width, 5.8 – 8.0 (7.0 ± 0.73) × as long as anterior width, 1.3 – 1.6 (1.5 ± 0.08) × as long as length of T 2; T 2 1.3 – 1.6 (1.4 ± 0.13) × as long as posterior width, 1.8 – 2.4 (2.0 ± 0.2) × as long as anterior width.	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
E67E4508455D594F816A1ECA66892E68.taxon	distribution	Distribution. Taiwan (Yilan, Hualien, Nantou, Taichung, and Chiayi).	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
E67E4508455D594F816A1ECA66892E68.taxon	etymology	Etymology. The specific name rufispeculus is derived from the Latin words rufi - (meaning red) and speculus (meaning mirror, refer to speculum on the mesopleuron herein), referring to the reddish-brown speculum on the mesopleuron of this new species. The name is an adjective.	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
7D01260823945AB688ABEC2F69CB8984.taxon	description	Figs 3 A – H, 10 A, 11 A – C; Suppl. material 3 Chinese vernacular name: 朝鮮亞力姬蜂 / 朝亞力姬蜂	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
7D01260823945AB688ABEC2F69CB8984.taxon	diagnosis	Diagnosis. This species can be distinguished from congeners by the combination of the following characters: fore wing length usually longer than 10.0 mm; POL / OOL = 0.5 – 0.7; fore wing areolet triangular with stalk, receiving 2 m-cu at distal corner (Fig. 10 A); fore wing 1 cu-a slightly inclivous, distad or almost opposite to M & RS (Fig. 10 A); lateromedian longitudinal carinae of propodeum complete and distinct, almost parallel (Fig. 3 E); T 1 2.7 – 3.3 × its apical width; gonostyle truncate apically (Fig. 11 B); mesosoma black with scutellum reddish-brown; metasoma and legs reddish-brown except hind tarsi yellow (Fig. 3 A). This species is the only one in Alexeter with such large body size, black mesosoma, reddish-brown scutellum, and almost entirely reddish-brown legs and metasoma. The Asian species A. daisetsusanus Uchida, 1930, A. dorogawaensis Uchida, 1934, A. segmentarius (Fabricius, 1787), and A. fallax (Holmgren, 1857) share a general color pattern of black mesosoma and reddish-brown metasoma. However, the metasomas of these other species are black on at least the posterior tergites, whereas in A. shakojiensis, the posterior tergites of the metasoma are reddish-brown.	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
7D01260823945AB688ABEC2F69CB8984.taxon	distribution	Distribution. Korea, China, and Taiwan (new record: Hualien, Nantou, and Chiayi).	en	Chen, Hsuan-Pu, Huang, Chia-Lung, Shiao, Shiuh-Feng (2025): A taxonomic revision of the genus Alexeter Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae, Mesoleiini) from Taiwan, with descriptions of six new species. ZooKeys 1250: 315-358, DOI: 10.3897/zookeys.1250.156835
