taxonID	type	description	language	source
76788795FFA06448FF3D4B9DCB34FA8D.taxon	description	urn: lsid: zoobank. org: act: 4 CE 72345 - 51 C 8 - 44 A 4 - 8 BFB-D 1 E 14705 CD 0 C (Figs 2 A – B; 3 A – D)	en	Gogshelidze, Mariam, Novák, János (2026): Two new cavernicolous Ephippiochthonius species from Caucasian caves in Georgia (Arachnida: Pseudoscorpiones). Zootaxa 5785 (1): 164-172, DOI: 10.11646/zootaxa.5785.1.7, URL: https://doi.org/10.11646/zootaxa.5785.1.7
76788795FFA06448FF3D4B9DCB34FA8D.taxon	materials_examined	Type locality. Caucasus, Georgia, Imereti region, Kharagauli, Zemo Imereti Plateau karst massif, Kozmani cave. (42 ° 06 ’ 03.2 ” N, 43 ° 17 ’ 18.6 ” E; 586 m a. s. l.).	en	Gogshelidze, Mariam, Novák, János (2026): Two new cavernicolous Ephippiochthonius species from Caucasian caves in Georgia (Arachnida: Pseudoscorpiones). Zootaxa 5785 (1): 164-172, DOI: 10.11646/zootaxa.5785.1.7, URL: https://doi.org/10.11646/zootaxa.5785.1.7
76788795FFA06448FF3D4B9DCB34FA8D.taxon	distribution	Distribution. Georgia: Kozmani cave.	en	Gogshelidze, Mariam, Novák, János (2026): Two new cavernicolous Ephippiochthonius species from Caucasian caves in Georgia (Arachnida: Pseudoscorpiones). Zootaxa 5785 (1): 164-172, DOI: 10.11646/zootaxa.5785.1.7, URL: https://doi.org/10.11646/zootaxa.5785.1.7
76788795FFA06448FF3D4B9DCB34FA8D.taxon	etymology	Etymology. The species is named after the village name Boriti, where Kozmani Cave is located.	en	Gogshelidze, Mariam, Novák, János (2026): Two new cavernicolous Ephippiochthonius species from Caucasian caves in Georgia (Arachnida: Pseudoscorpiones). Zootaxa 5785 (1): 164-172, DOI: 10.11646/zootaxa.5785.1.7, URL: https://doi.org/10.11646/zootaxa.5785.1.7
76788795FFA06448FF3D4B9DCB34FA8D.taxon	diagnosis	Diagnosis (♂). A small-sized hypogean Ephippiochthonius species. Chelicera almost the same size as carapace, movable cheliceral finger without isolated apical tooth (di) and prominent spinneret, lyrifissure ldb present; without eyes or eye-spots, anterior margin of carapace with four macrosetae; chelal hand depressed at level of ib / isb, with a distinct hump distal of ib / isb and moderate slope between trichobothria ib / isb and eb; fixed finger with 16 – 19 pointed teeth; movable finger with 15 – 17 teeth, teeth of proximal third decreases in size; pedipalpal femur (♂) 7.9 times longer than broad, length (♂) 0.52 mm; chela (♂) 5.9 times longer than deep, length (♂) 0.74 mm; ratio between movable chelal finger / chelal hand (♂) 1.5 – 1.7; all chelal lyrifissure patterns present in their standard number.	en	Gogshelidze, Mariam, Novák, János (2026): Two new cavernicolous Ephippiochthonius species from Caucasian caves in Georgia (Arachnida: Pseudoscorpiones). Zootaxa 5785 (1): 164-172, DOI: 10.11646/zootaxa.5785.1.7, URL: https://doi.org/10.11646/zootaxa.5785.1.7
76788795FFA06448FF3D4B9DCB34FA8D.taxon	materials_examined	Type material examined. Holotype: ♂, ISUIZ-PSEUD- 044, Georgia, Imereti region, Zemo Imereti plateau, Kozmani cave (42 ° 06 ’ 03.2 ” N 43 ° 17 ’ 18.6 ” E; 586 m a. s. l.), 27. XI. 2021, E. Maghradze leg. Paratype: 1 ♂, CaBOl- 1036046 same locality as holotype, 14. IX. 2021, E. Maghradze leg.	en	Gogshelidze, Mariam, Novák, János (2026): Two new cavernicolous Ephippiochthonius species from Caucasian caves in Georgia (Arachnida: Pseudoscorpiones). Zootaxa 5785 (1): 164-172, DOI: 10.11646/zootaxa.5785.1.7, URL: https://doi.org/10.11646/zootaxa.5785.1.7
76788795FFA06448FF3D4B9DCB34FA8D.taxon	description	Description. A small sized hypogean species. Integument depigmented; very weakly hipsid granulation on lateral surfaces of carapace, on cheliceral hand and bases of chelal fingers. Carapace slightly longer than broad and constricted posteriorly; medial part of anterior margin weakly prominent, without epistome, and strongly dentate; without eyes or eyespots. Chaetotaxy: 4 preocular microsetae, formula mm 4 mm: 6: 4: 2: 2; lyrifissures: 2 pairs at anterior margin, 1 pair at the posterior margin (Fig. 3 A). Chelicera slightly shorter than carapace, with 5 setae and 2 lateratl microsetae on hand; hand with one ventral and five dorsal lyriffisures, ldb present. Fixed finger with seven teeth, decreasing in size proximally. Movable finger without an isolated subapical tooth (di), with six teeth decreasing in size proximally; spinneret prominent; seta gl 0.85 mm from base of movable finger (Fig. 3 B). Rallum with 6 serrated blades. Serrula exterior with 15 blades, serrula interior 13 blades. Chaetotaxy of tergites 4: 4: 4: 4: 4 – 6: 4 – 6: 6: 4 – 6: 4 – 6: 4 – 6, tergite IX with two long and two normal sized seta. Chaetotaxy of sternites 12: 8: 8: 4: 4: 4: 4: 4: 4, genital opening with 7 – 7 setae on each side and 4 + 4 internal glandular setae (Fig. 3 C). Pedipalpal coxa with four setae and two microsetae, distal marginal seta of disk broken, only base visible; coxa I 4 + 4; coxa II 4 + 4 and 9 + 9 spines; coxa III 6 + 6 seta and 6 + 6 spine; coxa IV 6 + 6 seta; intercoxal tubercle bisetose. Pedipalp with femoral chaetotaxy 5: 6: 2: 6: 1. Chela with hand depressed at level of ib / isb with a distinct hump distad of ib / isb and moderate slope between trichobothria ib / isb and eb; dorsa – lantiaxial surface of the chelal hand between the hump and trichobothria eb / esb flattened, limited by distinct edge; ventral interocondylar bow-like protuberance (ip) present; width equal to depth; chaetotaxy 4: 5: 3, setae ph 1, ph 2, and ph 4 present; seta ph 3 absent, setae ih 1, ih 3 and ih 4 approximately level with trichobothria ib / isb; distal end of hand and bases of chelal fingers with sclerotized condylar complex. Fixed finger with 19 pointed teeth, the distal one small, of these the 6 proximal teeth decreasing in size, most with dental canals (Fig. 3 D); tip of finger with an accessory tooth (td) on antiaxial face; tip of fixed chelal finger of male with a deep hollow on paraxial face and subdistal protuberance (sp); one pair of long antiaxial sensory setae (as) at the base (only base visible), both distad of lyrifissure fb, distance between them 0.03 mm; 3 teeth at the level of est / it occupying 0.1 mm distance between successive apices 0.033 mm. Movable finger with 17 teeth, distal two thirds of finger with pointed teeth, the distal tooth very tiny on a low protuberance; proximal third with six – seven tiny teeth reaching halfway without dental canals; coupled sensilla pc slightly proximal to b. Distance between b – sb 0.09; sb – st 0.18; st – t 0.02. Measurements (in mm) and ratios. Male holotype. Body 1.13. Carapace 0.36 / 0.33 (1.1 ×). Chelicera 0.32 / 0.13 (2.4 ×), movable finger 0.15; ratio of chelicera / carapace 0.86 x. Pedipalp: femur 0.52 / 0.06 (7.9 ×), patella 0.18 / 0.09 (1.91 ×), chela 0.74 / 0.12 (5.9 ×), hand 0.29 (2.3 ×), movable finger 0.44; ratio of movable finger / hand 1.5 ×, femur / movable finger 1.1 ×, femur / carapace 1.4 ×, chela / carapace 2.0 ×, chela / femur 1.4 ×.	en	Gogshelidze, Mariam, Novák, János (2026): Two new cavernicolous Ephippiochthonius species from Caucasian caves in Georgia (Arachnida: Pseudoscorpiones). Zootaxa 5785 (1): 164-172, DOI: 10.11646/zootaxa.5785.1.7, URL: https://doi.org/10.11646/zootaxa.5785.1.7
76788795FFA06448FF3D4B9DCB34FA8D.taxon	discussion	Remarks. The new species is assigned to the genus Ephippiochthonius based on the following characters: chela lagyniform (ephippiochthonian), seta ph 3 in the proximal portion of chelal hand absent, medial protuberance (ip) present between chelal condyles and also presence of subapical protuberance (sp) and subdistal hollow. The newly described species from Kozmani Cave differs from E. kovali in having its carapace slightly narrowed in posterior part (almost square in E. kovali), and in having no epistome (weakly expressed epistome in E. kovali). Ephippiochthonius boritiensis sp. nov. differs from E. vargovitshi and E. dbari in having no isolated subapical tooth (di) (present in E. vargovitshi and E. dbari) and by its completely absent eyes (both pairs reduced cuticular area in E. vargovitshi, first pair reduced to cuticular area and second pair absent in E. dbari and E. kovali) (Turbanov & Kolesnikov 2023).	en	Gogshelidze, Mariam, Novák, János (2026): Two new cavernicolous Ephippiochthonius species from Caucasian caves in Georgia (Arachnida: Pseudoscorpiones). Zootaxa 5785 (1): 164-172, DOI: 10.11646/zootaxa.5785.1.7, URL: https://doi.org/10.11646/zootaxa.5785.1.7
76788795FFA0644BFF3D4AEDCFC0FED5.taxon	type_taxon	Type species. Scorpio tetrachelatus Preyssler, 1790	en	Gogshelidze, Mariam, Novák, János (2026): Two new cavernicolous Ephippiochthonius species from Caucasian caves in Georgia (Arachnida: Pseudoscorpiones). Zootaxa 5785 (1): 164-172, DOI: 10.11646/zootaxa.5785.1.7, URL: https://doi.org/10.11646/zootaxa.5785.1.7
76788795FFA56449FF3D4B18C9EAFB40.taxon	description	urn: lsid: zoobank. org: act: 6664 FB 95 - BC 00 - 461 E-B 3 FC-A 41 F 75 D 4 ABE 6 (Figs 2 C – D; 4 A – C)	en	Gogshelidze, Mariam, Novák, János (2026): Two new cavernicolous Ephippiochthonius species from Caucasian caves in Georgia (Arachnida: Pseudoscorpiones). Zootaxa 5785 (1): 164-172, DOI: 10.11646/zootaxa.5785.1.7, URL: https://doi.org/10.11646/zootaxa.5785.1.7
76788795FFA56449FF3D4B18C9EAFB40.taxon	materials_examined	Type locality. Georgia, Caucasus, Imereti Region, Chiatura, Zemo Imereti Plateau. (42 ° 19 ’ 02.2 ” N 43 ° 20 ’ 20.0 ” E; 717 m a. s. l.).	en	Gogshelidze, Mariam, Novák, János (2026): Two new cavernicolous Ephippiochthonius species from Caucasian caves in Georgia (Arachnida: Pseudoscorpiones). Zootaxa 5785 (1): 164-172, DOI: 10.11646/zootaxa.5785.1.7, URL: https://doi.org/10.11646/zootaxa.5785.1.7
76788795FFA56449FF3D4B18C9EAFB40.taxon	distribution	Distribution. Georgia: Tsintskila Cave.	en	Gogshelidze, Mariam, Novák, János (2026): Two new cavernicolous Ephippiochthonius species from Caucasian caves in Georgia (Arachnida: Pseudoscorpiones). Zootaxa 5785 (1): 164-172, DOI: 10.11646/zootaxa.5785.1.7, URL: https://doi.org/10.11646/zootaxa.5785.1.7
76788795FFA56449FF3D4B18C9EAFB40.taxon	etymology	Etymology. The specific name is derived from the Greek τραπέζιον (trapezion, meaning “ little table ” or, “ trapezoid ”), in reference to the trapezoid-shaped carapace.	en	Gogshelidze, Mariam, Novák, János (2026): Two new cavernicolous Ephippiochthonius species from Caucasian caves in Georgia (Arachnida: Pseudoscorpiones). Zootaxa 5785 (1): 164-172, DOI: 10.11646/zootaxa.5785.1.7, URL: https://doi.org/10.11646/zootaxa.5785.1.7
76788795FFA56449FF3D4B18C9EAFB40.taxon	diagnosis	Diagnosis. A medium-sized hypogean Ephippiochthonius species. Chelicera smaller than carapace, movable cheliceral finger with isolated apical tooth (di) and prominent spinneret, lyrifissure ldb present; without eyes or eyespots, anterior margin of carapace with two microsetae; chelal hand depressed at level of ib / isb, with a distinct hump distal of ib / isb and moderate slope between trichobothria ib / isb and eb; fixed finger with 20 pointed teeth; movable finger with eight teeth, teeth of proximal third decreases in size; pedipalpal femur (♀) 6.6 times longer than broad, length (♀) 0.69 mm; chela (♀) 2.19 times longer than deep, length (♀) 0.40 mm; ratio between movable chelal finger / chelal hand (♀) 3.3; all chelal lyrifissure patterns present in their standard number.	en	Gogshelidze, Mariam, Novák, János (2026): Two new cavernicolous Ephippiochthonius species from Caucasian caves in Georgia (Arachnida: Pseudoscorpiones). Zootaxa 5785 (1): 164-172, DOI: 10.11646/zootaxa.5785.1.7, URL: https://doi.org/10.11646/zootaxa.5785.1.7
76788795FFA56449FF3D4B18C9EAFB40.taxon	materials_examined	Type material examined. Holotype: ♀ ISUIZ-PSEUD- 122, Georgia, Imereti region, Zemo Imereti plateau, Tsintskila cave (42 ° 19 ’ 02.2 ” N, 43 ° 20 ’ 20.0 ” E; 717 m a. s. l.), 20. VIII. 2022, leg. Cave Invertebrate Research Group (Institute of Zoology, Ilia State University).	en	Gogshelidze, Mariam, Novák, János (2026): Two new cavernicolous Ephippiochthonius species from Caucasian caves in Georgia (Arachnida: Pseudoscorpiones). Zootaxa 5785 (1): 164-172, DOI: 10.11646/zootaxa.5785.1.7, URL: https://doi.org/10.11646/zootaxa.5785.1.7
76788795FFA56449FF3D4B18C9EAFB40.taxon	description	Description. A medium-sized hypogean species. Integument depigmented; weakly hipsid granulation on anterior surfaces of carapace, on cheliceral hand and bases of chelal fingers. Carapace as long as broad and distinctly constricted posteriorly; medial part of anterior margin weakly prominent, without epistome, and strongly dentate; without eyes or eyespots. Chaetotaxy: 4 preocular microsetae, formula mm 4 mm: 6: 4: 2: 2; lyrifissures: two pairs at anterior margin, one pair at the posterior margin (Fig. 4 A). Chelicera slightly shorter than carapace, with five setae and two lateral microsetae on hand; hand with 1 ventral and 5 dorsal lyriffisures, ldb present. Fixed finger with 14 teeth, decreasing in size proximally. Movable finger without an isolated subapical tooth (di), with 6 teeth decreasing in size proximally; spinneret prominent; seta gl 0.11 mm from base of movable finger (Fig. 4 B). Rallum with 11 serrated blades. Serrula exterior with 14 blades, serrula interior 12 blades. Chaetotaxy of tergites 4: 4: 4: 4: 6: 6: 6: 6: 6: 6, chaetotaxy of sternites 9: 8:?: 8: 8: 6: 6: 6: 6: 6, 1 + 1 spiracles. Pedipalpal coxa with 3 setae and 3 microsetae, distal marginal seta of disk 0.07 mm long; coxa I 3 + 3; coxa II 4 + 4 and 12 + 10 spines; coxa III 5 + 5 seta and 4 + 4 spine; coxa IV 6 + 6 seta; intercoxal tubercle bisetose. Pedipalp with femoral chaetotaxy 3: 6: 3: 5: 1. Chela with hand depressed at level of ib / isb with a distinct hump distad of ib / isb and moderate slope between trichobothria ib / isb and eb; dorsa – lantiaxial surface of the chelal hand between the hump and trichobothria eb / esb flattened, limited by distinct edge; ventral interocondylar bow-like protuberance (ip) present; width equal to depth; chaetotaxy 4: 5: 3, seta ph ₃ lacking, setae ih ₁ ih ₃ and ih ₄ approximately level with trichobothria ib / isb; distal end of hand and bases of chelal fingers with sclerotized condylar complex. Fixed finger with 20 pointed teeth, the distal one small, of these the 6 proximal teeth decreasing in size, most with dental canals (Fig. 4 C); tip of finger with an accessory tooth (td) on antiaxial face; tip of fixed chelal finger of male with a deep hollow on paraxial face and subdistal protuberance (sp); one pair of long antiaxial sensory setae (as) at the base (0.02 mm long), both distad of lyrifissure fb, distance between them 0.01 mm; 4 teeth at the level of est / it occupying 0.1 mm distance between successive apices 0.028 – 0.033 mm. Movable finger with seven teeth, almost the same size. Coupled sensilla pc slightly proximal to b. Distance between ib / isb and esb approximately equal to that between ib / isb and the base of the hand; distance between st and sb 1.8 times longer than that between sb and b. Measurements (in mm) and ratios. Female holotype. Body 1.17. Carapace 0.46 / 0.45 (1.0 ×). Chelicera 0.39 / 0.18 (2.1 ×), movable finger 0.20; ratio chelicera / carapace 0.78. Pedipalp: femur 0.69 / 0.10 (6.6 ×), patella 0.27 / 0.11 (2.4 ×), chela 0.40 / 0.06 (5.9 ×), hand 0.29 (6.6 ×), movable finger 0.59 x; ratio of movable finger / hand 2.3 x, femur / movable finger 1.1 x, femur / carapace 1.5 x, chela / carapace 0.8 x, chela / femur 0.5 x.	en	Gogshelidze, Mariam, Novák, János (2026): Two new cavernicolous Ephippiochthonius species from Caucasian caves in Georgia (Arachnida: Pseudoscorpiones). Zootaxa 5785 (1): 164-172, DOI: 10.11646/zootaxa.5785.1.7, URL: https://doi.org/10.11646/zootaxa.5785.1.7
76788795FFA56449FF3D4B18C9EAFB40.taxon	discussion	Remarks. The newly described species from Tsintskila Cave can be destinguished from E. boratiensis sp. nov. based on its movable cheliceral finger having an isolated apical tooth (di) (di absent in E. boritiensis sp. nov.), and having 20 pointed teeth on fixed chelal finger (16 – 19 teeth in E. boritiensis). Ephippiochthonius trapezoides sp. nov. can be diferrentiated from E. kovali, E. vargovitshi and E. dbari based on its compelately absent eyes (both pairs reduced to smooth cuticular area with tapetum in E. vargovitshi, first pair reduced to smooth cuticular area with tapetum and second pair entirely absent in E. dbari and E. kovali).	en	Gogshelidze, Mariam, Novák, János (2026): Two new cavernicolous Ephippiochthonius species from Caucasian caves in Georgia (Arachnida: Pseudoscorpiones). Zootaxa 5785 (1): 164-172, DOI: 10.11646/zootaxa.5785.1.7, URL: https://doi.org/10.11646/zootaxa.5785.1.7
