taxonID	type	description	language	source
73128785FFB7FFE32E87FF38DEFAC978.taxon	type_taxon	Type species: — Calea myrtifolia (Candolle 1836: 670) Baker (1884: 260).	en	Bueno, Vinicius R., Reis-Silva, Genilson A., Nakajima, Jimi N., Gostel, Morgan R., Heiden, Gustavo (2025): A new narrower circumscription and a nomenclatural synopsis for Calea subg. Meyeria (Asteraceae: Neurolaeneae). Phytotaxa 683 (3): 273-280, DOI: 10.11646/phytotaxa.683.3.8, URL: https://doi.org/10.11646/phytotaxa.683.3.8
73128785FFB7FFE32E87FF38DEFAC978.taxon	description	So far, Calea sect. Meyeria included nineteen species (Bueno & Heiden 2022 a, 2022 b). Based on the phylogenetic reconstruction presented in Bueno (2023), the section is redefined with a narrower morphological circumscription proposed here to include shrubs with a cymose capitulescence, bearing capitula with four or more foliaceous outer phyllaries and having the cypsela longer than the pappus. Under this new circumscription, the subgenus comprises nine taxa with eight species (C. funkiana, C. marginata, C. myrtifolia, C. parvifolia, C. phyllolepis, C. pruskiana, C. subintegerrima, C. triantha) and one nothospecies (C. × parviantha). A monophyletic C. subg. Meyeria requires the exclusion of the following 11 species: Calea arachnoidea, C. chodatii Hassler (1915: 174), C. grandiflora, C. heteropappa Pruski (1988: 346 – 347), C. ilienii Malme (1933: 94 – 95), C. kristiniae Pruski (1984: 98 – 102), C. monocephala Dusén (1910: 32), C. nervosa G. M. Barroso (1975: 109), C. quadrifolia Pruski & Urbatsch (1988: 341), C. semirii (1998: 698), and C. sessilifolia. These species were moved into another subgenera and sections according to Bueno (2023). Pruski (2023) resurrected Meyeria and segregated it from Calea based on morphological characters. Based on Bueno (2023), we disagree with the recognition of Meyeria as distinct of Calea and synonymize the species that Pruski (2023) first combined to Meyeria, which are part of Calea. subg. Meyeria.	en	Bueno, Vinicius R., Reis-Silva, Genilson A., Nakajima, Jimi N., Gostel, Morgan R., Heiden, Gustavo (2025): A new narrower circumscription and a nomenclatural synopsis for Calea subg. Meyeria (Asteraceae: Neurolaeneae). Phytotaxa 683 (3): 273-280, DOI: 10.11646/phytotaxa.683.3.8, URL: https://doi.org/10.11646/phytotaxa.683.3.8
73128785FFB7FFE32E87FCBEDDF3CE71.taxon	materials_examined	Holotype: — BRAZIL. Minas Gerais: Santana do Riacho, Serra do Cipó, 28 February 1991, S. Mayo, M. C. E. Amaral, I. Cordeiro & J. R. Pirani CFSC 7026 (SPF [SPF 18618]!; isotypes MG [MG 144560], digital image!; SP [SP 031897]!; UEC [UEC 124309]!).	en	Bueno, Vinicius R., Reis-Silva, Genilson A., Nakajima, Jimi N., Gostel, Morgan R., Heiden, Gustavo (2025): A new narrower circumscription and a nomenclatural synopsis for Calea subg. Meyeria (Asteraceae: Neurolaeneae). Phytotaxa 683 (3): 273-280, DOI: 10.11646/phytotaxa.683.3.8, URL: https://doi.org/10.11646/phytotaxa.683.3.8
73128785FFB7FFE32E87FCBEDDF3CE71.taxon	discussion	Note: — The isotype from the MG herbarium was found after the publication of this species and therefore is added here.	en	Bueno, Vinicius R., Reis-Silva, Genilson A., Nakajima, Jimi N., Gostel, Morgan R., Heiden, Gustavo (2025): A new narrower circumscription and a nomenclatural synopsis for Calea subg. Meyeria (Asteraceae: Neurolaeneae). Phytotaxa 683 (3): 273-280, DOI: 10.11646/phytotaxa.683.3.8, URL: https://doi.org/10.11646/phytotaxa.683.3.8
73128785FFB7FFE32E87FBA8DE72C257.taxon	materials_examined	Holotype: — BRAZIL. São Paulo / Paraná, 1828 – 1829, F. Sellow s. n. (P [P 02140741]!; isotype G [G 00456488, fragment], digital image!).	en	Bueno, Vinicius R., Reis-Silva, Genilson A., Nakajima, Jimi N., Gostel, Morgan R., Heiden, Gustavo (2025): A new narrower circumscription and a nomenclatural synopsis for Calea subg. Meyeria (Asteraceae: Neurolaeneae). Phytotaxa 683 (3): 273-280, DOI: 10.11646/phytotaxa.683.3.8, URL: https://doi.org/10.11646/phytotaxa.683.3.8
73128785FFB7FFE32E87FBA8DE72C257.taxon	discussion	Note: — Calea marginata was proposed by Blake (1937) based on the type specimens of Meyeria longifolia Candolle. He proposed the new name because the specific epithet was already preoccupied by C. longifolia Gardner (1848: 418). The superfluous name C. barrosoana was proposed by Robinson (1979) as a new name for M. longifolia Candolle, since he was unaware of Blake’s previous publication. In the protologue, the type specimen collector of C. marginata is not indicated. The specimen label of P 02140741 has only an indication of C. Gaudichaud-Beaupré ’ s name, which could suggest that he was the collector of the specimen. However, most of the materials assigned as collections done by him should be interpreted as belonging to his personal herbarium, as some of them were, for example, originally collected by F. Sellow (Moraes 2020). Furthermore, it is known that C. Gaudichaud-Beaupré has never collected in the habitat of this species in the “ campos gerais ” of the Paraná and São Paulo states border (Moraes 2020). So, it is assumed that the type specimen corresponds to a collection by F. Sellow that belonged to Gaudichaud-Beaupré personal herbarium. Furthermore, the specimen could not have been collected in 1833, as Sellow had already passed away at that given date. Pruski (2023) indicates that P 02140741 would be a specimen collected in 1833, however this year refers to the year when Gaudichaud - Beaupré incorporated this Sellow collection into his personal herbarium (Moraes 2023). According to Herter (1933), Sellow traveled between 1828 and 1829 through the regions where this species naturally occurs from Curitiba (Paraná state) to Itapeva (São Paulo state) through the “ Caminho das Tropas ”, an ancient cattle trading route in Brazil linking South and Southeastern Brazil from the 17 th to the 19 th century (Souza 2014). Considering the timespan when Sellow may have collected the specimen, this area referred in the label as São Paulo today is part of the state of Paraná, which was established in 1853 following the emancipation of land that was formerly part of the state of São Paulo. The available data does not allow mapping the precise locality nor can the collection year be ascertained. To reflect this uncertainty, we provide a range of possible collection years and the collection locality as the border area between these two states.	en	Bueno, Vinicius R., Reis-Silva, Genilson A., Nakajima, Jimi N., Gostel, Morgan R., Heiden, Gustavo (2025): A new narrower circumscription and a nomenclatural synopsis for Calea subg. Meyeria (Asteraceae: Neurolaeneae). Phytotaxa 683 (3): 273-280, DOI: 10.11646/phytotaxa.683.3.8, URL: https://doi.org/10.11646/phytotaxa.683.3.8
73128785FFB6FFE22E87FF51D908CE1A.taxon	materials_examined	Lectotype (designated here): — BRAZIL. Minas Gerais: Diamantina, 1833, A. C. Vauthier 320 (GDC [00456458], digital image!; isolectotype G [00456458], digital image!; G [00222839], digital image!; GH [00589157], digital image!; GH [00589158], digital image!; P [02140745]!; P [02140746]!; P [02140747]!; W [0043963], digital image!).	en	Bueno, Vinicius R., Reis-Silva, Genilson A., Nakajima, Jimi N., Gostel, Morgan R., Heiden, Gustavo (2025): A new narrower circumscription and a nomenclatural synopsis for Calea subg. Meyeria (Asteraceae: Neurolaeneae). Phytotaxa 683 (3): 273-280, DOI: 10.11646/phytotaxa.683.3.8, URL: https://doi.org/10.11646/phytotaxa.683.3.8
73128785FFB6FFE22E87FF51D908CE1A.taxon	materials_examined	Lectotype (designated by Pruski 2023): — BRAZIL. Minas Gerais: Catas Altas, Serra do Caraça, L. Riedel 1427 (LE [not found]; isolectotypes G [00412260], digital image!; K [K 000323152]!; P [P 02469708]!; P [P 02469709]!; P [P 02469710]!). Remaining syntypes: — BRAZIL. Minas Gerais: Antônio Carlos, Sítio João Ayres, A. Glaziou 11054 (K [K 01103654]!; P [P 02469714]!). BRAZIL. Minas Gerais: São João Del-Rei, 1830, F. Sellow 1339 (K [K 001103655]. BRAZIL. Minas Gerais: Congonhas, K. F. P. Martius [not seen]. Minas Gerais: São João Del-Rei, K. F. P. Martius [not seen]. Minas Gerais: Ouro Preto, Cachoeira do Campo, 1840, P. Claussen s. n. (K [K 000323384]!; K [000323152]!; K [K 02469709]!; P [P 02469715]; P [P 02469718]!).	en	Bueno, Vinicius R., Reis-Silva, Genilson A., Nakajima, Jimi N., Gostel, Morgan R., Heiden, Gustavo (2025): A new narrower circumscription and a nomenclatural synopsis for Calea subg. Meyeria (Asteraceae: Neurolaeneae). Phytotaxa 683 (3): 273-280, DOI: 10.11646/phytotaxa.683.3.8, URL: https://doi.org/10.11646/phytotaxa.683.3.8
73128785FFB6FFE22E87FF51D908CE1A.taxon	discussion	Note: — Pruski (2023) indicated that C. myrtifolia has a holotype, but the Vauthier 320 collection corresponds to several syntypes: G 00456458, G 00222839, GDC 00456458, GH 00589157, GH 00589158, P 02140745, P 02140746, P 02140747, W 0043963. Therefore, it is necessary to choose a lectotype from the available syntypes. The specimen chosen is from the G-DC herbarium where Candolle was based. The protologue information indicates the type specimen of C. myrtifolia as the collection A. C. Vauthier 370; however, the number 370 is a typo, since we noticed that the correct handwritten number is 320. Pruski (2023) has synonymized and lectotypified C. myrtifolia var. paucidentata. We found additional isolectotypes not cited by Pruski (2023): P 02469708, P 02469709, P 02469710. Pruski (2023) indicated the materials from L. Riedel as a lectotype and cited those from P. Claussen as remaining syntypes. The following collections are also synonyms: A. Glaziou 11054, F. Sellow 1339 and the two of K. F. P. Martius already mentioned.	en	Bueno, Vinicius R., Reis-Silva, Genilson A., Nakajima, Jimi N., Gostel, Morgan R., Heiden, Gustavo (2025): A new narrower circumscription and a nomenclatural synopsis for Calea subg. Meyeria (Asteraceae: Neurolaeneae). Phytotaxa 683 (3): 273-280, DOI: 10.11646/phytotaxa.683.3.8, URL: https://doi.org/10.11646/phytotaxa.683.3.8
73128785FFB6FFE22E87FB41DE44CCD9.taxon	materials_examined	Lectotype (designated here): — BRAZIL. São Paulo-Paraná: 1828 – 1829, F. Sellow s. n. (P [P 02140753]!; isolectotype B, destroyed, photo in F [F 244778], digital image!; G [G 00456484], digital image!).	en	Bueno, Vinicius R., Reis-Silva, Genilson A., Nakajima, Jimi N., Gostel, Morgan R., Heiden, Gustavo (2025): A new narrower circumscription and a nomenclatural synopsis for Calea subg. Meyeria (Asteraceae: Neurolaeneae). Phytotaxa 683 (3): 273-280, DOI: 10.11646/phytotaxa.683.3.8, URL: https://doi.org/10.11646/phytotaxa.683.3.8
73128785FFB6FFE22E87FB41DE44CCD9.taxon	discussion	Note: — The syntypes of C. parvifolia were found in P and G herbaria. The specimen deposited in B was lost in a fire during World War II (Grimé & Plowman 1986), but a photograph of it is available at F. The type specimen of this species was interpreted as collected by C. Gaudichaud-Beaupré, whereas it was in fact collected by F. Sellow (Moraes 2020) and later incorporated in his personal herbarium. The precise locality of this collection was also difficult to identify. The available data does not allow us to map the precise locality nor can the collection year be ascertained. To reflect this uncertainty, we present the range of possible years and the collection locality as the border between the states of Paraná and São Paulo, as discussed for C. marginata. Pruski (2023) considered P 02140753 as a holotype, but the existence of F 244778 and G 00456484 implies that these materials were syntypes. Thus, we designate the specimen deposited in P as the lectotype, since the duplicate deposited in G is incomplete. The specimen Dusén 2809 deposited in F [F 0 BN 015383] displays the name Calea paranensis, which was never published (nom. in sched.). This specimen is actually C. parvifolia.	en	Bueno, Vinicius R., Reis-Silva, Genilson A., Nakajima, Jimi N., Gostel, Morgan R., Heiden, Gustavo (2025): A new narrower circumscription and a nomenclatural synopsis for Calea subg. Meyeria (Asteraceae: Neurolaeneae). Phytotaxa 683 (3): 273-280, DOI: 10.11646/phytotaxa.683.3.8, URL: https://doi.org/10.11646/phytotaxa.683.3.8
73128785FFB6FFE52E87F901D82BCB0F.taxon	materials_examined	Lectotype (designated by Pruski 2023): — BRAZIL. Rio Grande do Sul: F. Sellow s. n. (K [K 000323167]!).	en	Bueno, Vinicius R., Reis-Silva, Genilson A., Nakajima, Jimi N., Gostel, Morgan R., Heiden, Gustavo (2025): A new narrower circumscription and a nomenclatural synopsis for Calea subg. Meyeria (Asteraceae: Neurolaeneae). Phytotaxa 683 (3): 273-280, DOI: 10.11646/phytotaxa.683.3.8, URL: https://doi.org/10.11646/phytotaxa.683.3.8
73128785FFB6FFE52E87F901D82BCB0F.taxon	discussion	Note: — The types of Calea phyllolepis were not found in K. A record of a possible type specimen was found based on one photo from B in the Berlin Negatives collection at F. However, as this photograph was taken from a specimen deposited in B that was destroyed, it is not accessible anymore (Grimé & Plowman 1986). Unfortunately, it was not possible to find any remaining specimen collected by F. Sellow numbered as 1041, 2534 or 3140 in the K herbarium, neither in other herbaria consulted. According to Herter (1945), the numbers that Baker (1884) indicated as types do not refer to specimens collected in São Paulo state. These numbers from Sellow are from a period when he was traveling by boat between Rio Pardo and Porto Alegre, in the state of Rio Grande do Sul, Brazil. Calea phyllolepis occurs naturally and is well represented by several collections along this area, in accordance with the species description, numbering and information provided by Herter (1945). Additionally, this species does not occur in São Paulo and Paraná states (Reis-Silva et al. 2023). Therefore, it is assumed that the type specimens were in fact collected in the state of Rio Grande do Sul.	en	Bueno, Vinicius R., Reis-Silva, Genilson A., Nakajima, Jimi N., Gostel, Morgan R., Heiden, Gustavo (2025): A new narrower circumscription and a nomenclatural synopsis for Calea subg. Meyeria (Asteraceae: Neurolaeneae). Phytotaxa 683 (3): 273-280, DOI: 10.11646/phytotaxa.683.3.8, URL: https://doi.org/10.11646/phytotaxa.683.3.8
73128785FFB1FFE52E87F8ACD8ABCD87.taxon	materials_examined	Holotype: — BRAZIL. Paraná, Jaguariaíva, campo próximo à ponte do Rio das Mortes, PR- 151, 16 December 1991, A. C. Cervi et al. 3546 (MBM [MBM 161398]!; isotypes: CGMS [44616], digital image!; CPAP [10996], digital image!; UPCB [33501], digital image!).	en	Bueno, Vinicius R., Reis-Silva, Genilson A., Nakajima, Jimi N., Gostel, Morgan R., Heiden, Gustavo (2025): A new narrower circumscription and a nomenclatural synopsis for Calea subg. Meyeria (Asteraceae: Neurolaeneae). Phytotaxa 683 (3): 273-280, DOI: 10.11646/phytotaxa.683.3.8, URL: https://doi.org/10.11646/phytotaxa.683.3.8
73128785FFB1FFE52E87FEB4DEECCBBC.taxon	materials_examined	Holotype: — BRAZIL. São Paulo: Campos do Jordão, Trilha do rio Sapucaí, Parque Estadual de Campos do Jordão, 4 February 1988, M. J. Robim & J. P. M. Carvalho 530 (UEC [UEC 124311]!; isotype SPSF [SPSF 11784]!).	en	Bueno, Vinicius R., Reis-Silva, Genilson A., Nakajima, Jimi N., Gostel, Morgan R., Heiden, Gustavo (2025): A new narrower circumscription and a nomenclatural synopsis for Calea subg. Meyeria (Asteraceae: Neurolaeneae). Phytotaxa 683 (3): 273-280, DOI: 10.11646/phytotaxa.683.3.8, URL: https://doi.org/10.11646/phytotaxa.683.3.8
73128785FFB1FFE52E87FDE0DEC1C9B3.taxon	materials_examined	Lectotype (designated by Bueno & Heiden 2022): — BRAZIL. Paraná: Jaguariaíva, 5 February 1910, P. H. K. Dusén 9187 (GH [GH 00589132], digital image!; isolectotypes G [G 00222849], digital image!; S [S-R- 875], digital image!).	en	Bueno, Vinicius R., Reis-Silva, Genilson A., Nakajima, Jimi N., Gostel, Morgan R., Heiden, Gustavo (2025): A new narrower circumscription and a nomenclatural synopsis for Calea subg. Meyeria (Asteraceae: Neurolaeneae). Phytotaxa 683 (3): 273-280, DOI: 10.11646/phytotaxa.683.3.8, URL: https://doi.org/10.11646/phytotaxa.683.3.8
73128785FFB1FFE52E87FDE0DEC1C9B3.taxon	discussion	Bueno & Heiden (2022 b) proposed the new status and combination of Calea subintegerrima with a description, mapped occurrence, and morphological comparisons. While designating the lectotype, they cited an incorrect collection number (9189) but the correct barcode number (GH 00589132), so that the lectotypification was unambiguous and therefore effective. Pruski (2023) rectified this citation error by indicating that the correct collection number is P. H. K. Dusén 9187. Pruski (2023) claimed that this combination is invalid because the “ basionym literature reference ” was “ incorrect ”. However, the reference was complete and correct, except for the lacking journal series number (ser. 3), which does not make the citation ambiguous because the publication year indicates the series.	en	Bueno, Vinicius R., Reis-Silva, Genilson A., Nakajima, Jimi N., Gostel, Morgan R., Heiden, Gustavo (2025): A new narrower circumscription and a nomenclatural synopsis for Calea subg. Meyeria (Asteraceae: Neurolaeneae). Phytotaxa 683 (3): 273-280, DOI: 10.11646/phytotaxa.683.3.8, URL: https://doi.org/10.11646/phytotaxa.683.3.8
73128785FFB1FFE52E87FBE8DC38CD77.taxon	materials_examined	Lectotype (designated by Pruski 2005: 2027): — BRAZIL. São Paulo: Cunha, J. Velloso s. n. (t. 120, Vell Flumin. [Icones] 8, digital image!).	en	Bueno, Vinicius R., Reis-Silva, Genilson A., Nakajima, Jimi N., Gostel, Morgan R., Heiden, Gustavo (2025): A new narrower circumscription and a nomenclatural synopsis for Calea subg. Meyeria (Asteraceae: Neurolaeneae). Phytotaxa 683 (3): 273-280, DOI: 10.11646/phytotaxa.683.3.8, URL: https://doi.org/10.11646/phytotaxa.683.3.8
73128785FFB1FFE52E87FBE8DC38CD77.taxon	materials_examined	Lectotype (designated by Pruski 2005: 2027): — BRAZIL. São Paulo: between Mogi das Cruzes and São Paulo, Nov 1833, P. Lund 866 (G-DC [G 00456737], digital image!; isolectotypes C [C 10007025], digital image!; C [C 10007027], digital image!; S [S-R- 3508], digital image!).	en	Bueno, Vinicius R., Reis-Silva, Genilson A., Nakajima, Jimi N., Gostel, Morgan R., Heiden, Gustavo (2025): A new narrower circumscription and a nomenclatural synopsis for Calea subg. Meyeria (Asteraceae: Neurolaeneae). Phytotaxa 683 (3): 273-280, DOI: 10.11646/phytotaxa.683.3.8, URL: https://doi.org/10.11646/phytotaxa.683.3.8
73128785FFB1FFE52E87FBE8DC38CD77.taxon	discussion	Note: — Calea triantha is the only scientific name listed in this study for which its type is not based on a plant specimen. The whereabouts of the original specimens studied by Vellozo is unknown, but the plates published earlier (Vellozo 1831) are accepted as types for the new taxa described by him in the Flora Fluminensis. According to Pruski (2005), this plate undoubtedly represents this species and we agree with his findings. The monumental work of Vellozo (1829 [1825], 1831, 1881) in Flora Fluminensis describes the flora of the state of Rio de Janeiro, which occupied an area whose state borders has changed since the time of publication. According to Pastore et al. (2021), the specimen on which the illustration is based upon was collected in Cunha, São Paulo, where it still can be found. This species does not occur within the current borders of the state of Rio de Janeiro (Reis-Silva et al. 2023). The protologue of M. hispida Candolle (1836) and the study by Pruski (2005) do not indicate the collection site of the type specimen, but we found this information in Baker (1884). Interestingly, this synonym was collected between Mogi das Cruzes and São Paulo in São Paulo state, which are nearby Cunha that currently lies within São Paulo state boundaries.	en	Bueno, Vinicius R., Reis-Silva, Genilson A., Nakajima, Jimi N., Gostel, Morgan R., Heiden, Gustavo (2025): A new narrower circumscription and a nomenclatural synopsis for Calea subg. Meyeria (Asteraceae: Neurolaeneae). Phytotaxa 683 (3): 273-280, DOI: 10.11646/phytotaxa.683.3.8, URL: https://doi.org/10.11646/phytotaxa.683.3.8
