taxonID	type	description	language	source
73618790F357FFD6879FFDA82F7A7D60.taxon	vernacular_names	Common English name: Kirirom Bent-toed Gecko Common Khmer name:	en	Grismer, L. Lee, Neang, Thy, Samorn, Vireak, Song, Det, Stuart, Bryan L. (2026): A new species of the Cyrtodactylus intermedius group (Squamata: Gekkonidae) continues to underscore the high degree of site-specific endemism in the Cardamom Region, Cambodia. Zootaxa 5741 (3): 519-538, DOI: 10.11646/zootaxa.5741.3.5, URL: https://doi.org/10.11646/zootaxa.5741.3.5
73618790F357FFD6879FFDA82F7A7D60.taxon	description	Figure 5	en	Grismer, L. Lee, Neang, Thy, Samorn, Vireak, Song, Det, Stuart, Bryan L. (2026): A new species of the Cyrtodactylus intermedius group (Squamata: Gekkonidae) continues to underscore the high degree of site-specific endemism in the Cardamom Region, Cambodia. Zootaxa 5741 (3): 519-538, DOI: 10.11646/zootaxa.5741.3.5, URL: https://doi.org/10.11646/zootaxa.5741.3.5
73618790F357FFD6879FFDA82F7A7D60.taxon	materials_examined	Holotype. FMNH 263242 (field number HKV 64694), adult male, Cambodia, Kampong Speu Province, Phnom Sruoch District, Kirirom National Park, 11 ° 21 ' 30 " N 104 ° 02 ' 11 " E, 600 m asl, collected 6 June 2000 by Bryan L. Stuart, Dara An, and Vuthy Kong. Paratypes. FMNH 263239 – 263240 (two adult females), same data as holotype except near O Tasek Stream, 11 ° 18 ' 39 " N 104 ° 04 ' 42 " E, 500 – 600 m asl, collected 4 June 2000. CBC 03495 (adult female), same data as holotype except 11 º 19 ' 08 " N 104 º 06 ' 49 " E, 310 m asl, collected 20 October 2023 by Vireak Samorn and Det Song. Additional specimens. FMNH 263238 (unsexed juvenile), same data as paratypes FMNH 263239 – 263240. FMNH 263241 (unsexed juvenile), same data as holotype except near O Krang Snoul Stream, 11 ° 22 " 00 " N 104 ° 06 ' 28 " E, 300 – 400 m asl, collected 5 June 2000. These specimens were only used to describe the juvenile coloration and pattern.	en	Grismer, L. Lee, Neang, Thy, Samorn, Vireak, Song, Det, Stuart, Bryan L. (2026): A new species of the Cyrtodactylus intermedius group (Squamata: Gekkonidae) continues to underscore the high degree of site-specific endemism in the Cardamom Region, Cambodia. Zootaxa 5741 (3): 519-538, DOI: 10.11646/zootaxa.5741.3.5, URL: https://doi.org/10.11646/zootaxa.5741.3.5
73618790F357FFD6879FFDA82F7A7D60.taxon	diagnosis	Diagnosis (adults). Cyrtodactylus kiriromensis sp. nov. can be differentiated from all other species of the intermedius group by the combination of having 7 – 9 supralabials, 8 – 11 infralabials, 25 – 28 paravertebral tubercles, 17 – 19 longitudinal rows of dorsal tubercles, 38 or 39 ventral scales, seven expanded subdigital lamellae on the 4 th toe, 12 – 14 unmodified expanded subdigital lamellae on the 4 th toe, 19 – 21 total subdigital lamellae on the 4 th toe, five or six expanded subdigital lamellae on the 4 th finger, 11 – 13 unmodified expanded subdigital lamellae on the 4 th finger, 17 – 19 total subdigital lamellae on the 4 th finger, 14 – 29 enlarged femoral scales (both thighs combined), no femoral pores, seven or eight enlarged precloacal scales, seven or eight enlarged precloacal pores / pits, two or three enlarged post-precloacal scale rows, enlarged femoral and precloacal scales not continuous, body tubercles moderately keeled and not greatly reduced; caudal tubercles extend beyond base of tail, median subcaudals transversely expanded but not extending onto lateral caudal region, no digital pockets, dorsal color pattern not faded, no distinct dark-colored blotches on top of head, dark-colored body bands bearing lightened centers, no dark-colored markings on body between dark-colored body bands, dark-colored body bands approximately same width as light-colored interspaces and bordered by white tubercles, light-colored interspaces not reduced to thin bands, dark-colored caudal bands slightly wider than light-colored caudal bands, and light-colored caudal bands usually not bearing dark-colored markings.	en	Grismer, L. Lee, Neang, Thy, Samorn, Vireak, Song, Det, Stuart, Bryan L. (2026): A new species of the Cyrtodactylus intermedius group (Squamata: Gekkonidae) continues to underscore the high degree of site-specific endemism in the Cardamom Region, Cambodia. Zootaxa 5741 (3): 519-538, DOI: 10.11646/zootaxa.5741.3.5, URL: https://doi.org/10.11646/zootaxa.5741.3.5
73618790F357FFD6879FFDA82F7A7D60.taxon	description	Description of holotype. Adult male SVL 84.7 mm; head triangular in dorsal profile, distinct from neck, moderate in length, HL / SVL 0.29, slightly widened, HW / HL 0.66, moderately depressed, HD / HL 0.38; snout elongate, more than one-half head length, SN / HL 0.38; frontonasal region flat anteriorly, concave posteriorly concave at union with frontal region; frontal region deeply concave; rostrum grooved; anterior loreal region concave on both sides forming a narrow rostrum anteriorly in dorsal profile, canthus rounded, indistinctly inflated posteriorly; eye relatively large, rounded, protuberant, diameter 6.0 mm, slightly smaller than the distance from eye to ear, 6.2 mm, shorter than eye to nostril 7.0 mm; ear opening small, oriented forward, 2.1 mm in length; rostral rather broad, 3.8 mm in width, 2.1 mm in high, in contact with first supralabials laterally, supranasals and internasals dorsally, partially divided dorsally by a short linear furrow; supranasals two, right one somewhat rounded, left one hexagonal, separated by two internasals three total, one below intruding upper part of rostral; nostrils projecting outward, nasal valves visible posteriorly; external nares bordered by rostral anteriorly, three postnasals posteriorly, first supralabials ventrally, supranasals dorsally; supralabials eightR / nineL, subrectangular in shape, largest anteriorly, tapering posteriorly; infralabials 10 R / 10 L, large squarish anteriorly, rectangular posteriorly, tapering posteriorly, scales in interorbital region, top of head with small scales intermixed with small low tubercles, supraocular scales granular, varying in size; scales on loreal and frontonasal region slightly flattened, larger than those on top of head; a row of flat scales above supralabials larger than those on loreal; mental scale rounded anteriorly, pointed posteriorly, in contact with first infralabials laterally, postmentals posteriorly; two pairs of postmentals, first pair in broad contact, the second pair separated posteriorly by four gular scales; three rows of elongate, enlarged chinshields between infralabials and gular scales beginning from second postmental to the level of 8 th infralabial; gular scales, anterior throat scales, small, granular, subimbricate grading into larger imbricate pectoral and ventral scales. Body somewhat flattened in lateral profiles, AG / SVL 0.47; dorsal and ventral scales separated by distinct ventrolateral fold; scales on nape small, homogenous, granular, with large, conical, weekly keeled tubercles; scales on dorsum small, homogeneous, intermixed with large, raised, moderately keeled tubercles; 17 longitudinal rows of dorsal tubercles; 26 paravertebral tubercles; ventrolateral fold well-developed by a row of raised, enlarged discontinuous tubercles; scales on either side of ventrolateral fold, small same size as those on dorsum, grading to larger flat, imbricate, midventral scales; ventral scales flat, rounded, imbricate, larger than those on dorsum; enlarged poreless femoral scales 12 R / 12 L on each side, proximal femoral scales smaller than one-half size of distal femoral scales; seven enlarged pore-bearing precloacal scales; three rows of enlarged scales anterior to enlarged pore-bearing precloacal scales; enlarged femoral scales discontinuous with precloacal scales; three post precloacal scale rows; cloacal spurs 2 R / 2 L on based of tail. Limbs well-developed; forelimb relatively slender, FL 12.0 mm, approximately one third body length, FL / AG 0.30; scales on dorsal surface of forelimb relatively larger than those on dorsum, interspersed with low larger tubercles; scales on ventral surface of forelimb similar to those along ventrolateral folds, semi-granular; scales on mid-palmar region small, larger than those on base of ventral surface of first and fifth fingers; forelimb digits bearing strongly cured claws; expanded subdigital lamellae on fourth finger (6 R / 6 L), unmodified subdigital lamellae (11 R / 11 L), totaling 17 R / 17 L; hind limbs relatively long, TBL / AG 0.35; scales on dorsal surface of hind limbs small, granular intermixed with larger, more dense array of tubercles than those on forelimb; scales on anterior and ventral surface of thigh large, same size as ventral scales; hind limb with curved claws; expanded subdigital lamellae on fourth toes (7 R / 7 L); unmodified subdigital lamellae (13 R / 12 L), totaling 20 R / 19 L; scales on mid-plantar small, larger than those at its base. Tail thick anteriorly, TL 82.1 mm, regenerated posteriorly; TW 6.9 mm; ventral base of tail with hemipenial swellings; subcaudal scales transversely enlarged, undivided anteriorly, divided posteriorly, not extending to lateral surface of tail; subcaudal scales much larger than scales on lateral caudals; base of dorsal caudals small intermixed with large, 10 rows of highly raised, tubercles; dorsal caudal tubercles extending beyond base of tail with tubercles at its posterior margin. Color pattern in preservative (Fig. 5). Color of dorsal surface of rostrum, top of head, dorsum, flanks, limbs, tail light-brown; top of head lacking dark-brown blotches; posterior margin of nuchal loop rounded; anterior and posterior margins of nuchal loop border by white tubercles; four dark-brown body bands with lightened centers between nuchal loop and hind limb insertions and bordered by white tubercles; three dark-brown caudal bands; posterior section of tail unicolor tan, regenerated; isolated white tubercles on hind limbs; gular region, throat, venter, underside of limbs, and subcaudal region beige with faint dark-brown stippling. Color pattern in life of the paratype CBC 04395 is shown in Figure 5. Variation. The paratypes approach the holotype very closely in all aspects of coloration and pattern (Fig. 5). FMNH 263239 – 40 have complete tails. The posterior section of the tail in CBC 04395 is regenerated and somewhat mottled. Color pattern of the juveniles FMNH 263238 and 262241 is much bolder and contrasting. See Table 3 for variation in meristic among the type series.	en	Grismer, L. Lee, Neang, Thy, Samorn, Vireak, Song, Det, Stuart, Bryan L. (2026): A new species of the Cyrtodactylus intermedius group (Squamata: Gekkonidae) continues to underscore the high degree of site-specific endemism in the Cardamom Region, Cambodia. Zootaxa 5741 (3): 519-538, DOI: 10.11646/zootaxa.5741.3.5, URL: https://doi.org/10.11646/zootaxa.5741.3.5
73618790F357FFD6879FFDA82F7A7D60.taxon	distribution	Distribution. Cyrtodactylus kiriromensis sp. nov. is known only from the type locality at Kirirom National Park but is expected to range farther throughout the Sruoch Plateau.	en	Grismer, L. Lee, Neang, Thy, Samorn, Vireak, Song, Det, Stuart, Bryan L. (2026): A new species of the Cyrtodactylus intermedius group (Squamata: Gekkonidae) continues to underscore the high degree of site-specific endemism in the Cardamom Region, Cambodia. Zootaxa 5741 (3): 519-538, DOI: 10.11646/zootaxa.5741.3.5, URL: https://doi.org/10.11646/zootaxa.5741.3.5
73618790F357FFD6879FFDA82F7A7D60.taxon	etymology	Etymology. The specific epithet is a toponym of the type locality Kirirom National Park, Kampong Speu Province, Cambodia.	en	Grismer, L. Lee, Neang, Thy, Samorn, Vireak, Song, Det, Stuart, Bryan L. (2026): A new species of the Cyrtodactylus intermedius group (Squamata: Gekkonidae) continues to underscore the high degree of site-specific endemism in the Cardamom Region, Cambodia. Zootaxa 5741 (3): 519-538, DOI: 10.11646/zootaxa.5741.3.5, URL: https://doi.org/10.11646/zootaxa.5741.3.5
73618790F357FFD6879FFDA82F7A7D60.taxon	biology_ecology	Natural history. All specimens of the new species were found at night in disturbed or intact semi-evergreen forest, often near a stream, at 310 – 600 m asl. The holotype was found on a vertical tree trunk 5 m from a cascading stream in an open area at an old dam site within disturbed semi-evergreen forest. FMNH 263238 – 40 were found along a stream in semi-evergreen forest on a 1.2 m diameter boulder on the stream bank (FMNH 263238), on a 2 x 2 m rock face 1.5 m above the stream (FMNH 263239), and on a large rock face 2.5 m above the ground, 1.5 m from the stream (FMNH 263240). FMNH 263241 was found in disturbed semi-evergreen forest mixed with bamboo on a boulder approximately 100 m from a stream. CBC 03495 was found after a light rain in semi-evergreen forest on the base of a tree 0.3 m above the ground.	en	Grismer, L. Lee, Neang, Thy, Samorn, Vireak, Song, Det, Stuart, Bryan L. (2026): A new species of the Cyrtodactylus intermedius group (Squamata: Gekkonidae) continues to underscore the high degree of site-specific endemism in the Cardamom Region, Cambodia. Zootaxa 5741 (3): 519-538, DOI: 10.11646/zootaxa.5741.3.5, URL: https://doi.org/10.11646/zootaxa.5741.3.5
73618790F357FFD6879FFDA82F7A7D60.taxon	discussion	Comparisons. The phylogenetic relationships of Cyrtodactylus kiriromensis sp. nov. delimit it from all other species of intermedius group wherein it is part of a southern clade composed of C. bokorensis, C. borgattaorum, C. laangensis, C. phuquocensis, C. regicavernicolus and its sister species C. septimontium (Fig. 3) from which it differs by an uncorrected pairwise sequence divergence of 3.2 – 4.3 %. From the other species it differs by a range of 8.1 % from C. bokorensis to 8.9 % from C. laangensis. The ANOVA analyses recovered multiple statistically different mean morphometric and meristic values between C. kiriromensis sp. nov. and the other species of the southern clade (Table 2; Fig. 4). Cyrtodactylus kiriromensis sp. nov. differed from C. bokorensis, C. regicavernicolus, and C. laangensis in that the enlarged femoral scales were not separated, as opposed to being separated from the precloacal scales by smaller granular scales. It often differs further from C. bokorensis, C. laangensis, and C. septimontium in that proximal enlarged femoral scales are usually not less than one-half the size of the distal femoral scales. From C. regicavernicolus it differed by have a nearly unicolor top of head as opposed to the top of the head bearing dark markings. This characteristic was variable in C. bokorensis and C. laangensis. The distribution of the categorical characters are in Table 1 S. Table 4 provides summary statistics for the species discussed above. The more distantly related C. borgattaorum was not included in the statistical analyses due the low sample size of adults (n = 1) and C. phuquocensis was excluded due to notable potential differences in the counting of meristic characters.	en	Grismer, L. Lee, Neang, Thy, Samorn, Vireak, Song, Det, Stuart, Bryan L. (2026): A new species of the Cyrtodactylus intermedius group (Squamata: Gekkonidae) continues to underscore the high degree of site-specific endemism in the Cardamom Region, Cambodia. Zootaxa 5741 (3): 519-538, DOI: 10.11646/zootaxa.5741.3.5, URL: https://doi.org/10.11646/zootaxa.5741.3.5
