taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
B0061B8E72A95208849D496C52D445B9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1725115	https://doi.org/10.3897/zookeys.1286.194383.figure5	Figure 5. Phylogenetic relationships of the Pacific clade of the Pristimantis lacrimosus species group. Maximum likelihood tree obtained for the 16 S, 12 S, ND 1 and RAG 1 genes. Branch support (blue numbers) are shown as aLRT values (before the slash) and ultrafast bootstrap values (after the slash). For each individual, the species name is followed by the voucher number. In the new species, the voucher number is followed by the abbreviation of the province and the locality (all in Ecuador). Abbreviations: BO = Bolívar, EO = El Oro, ES = Esmeraldas, MA = Manabí, PI = Pichincha, SD = Santo Domingo. Photographs of the new species and similar species from the Ecuadorian Chocó are shown on the right.	Figure 5. Phylogenetic relationships of the Pacific clade of the Pristimantis lacrimosus species group. Maximum likelihood tree obtained for the 16 S, 12 S, ND 1 and RAG 1 genes. Branch support (blue numbers) are shown as aLRT values (before the slash) and ultrafast bootstrap values (after the slash). For each individual, the species name is followed by the voucher number. In the new species, the voucher number is followed by the abbreviation of the province and the locality (all in Ecuador). Abbreviations: BO = Bolívar, EO = El Oro, ES = Esmeraldas, MA = Manabí, PI = Pichincha, SD = Santo Domingo. Photographs of the new species and similar species from the Ecuadorian Chocó are shown on the right.	2026-07-24	Ron, Santiago R.;Paredes-Aguirre, Poulette;Paucar, Diego A.;López, Yaara;Apunte, Katherine;Ortega, Jhael A.		Zenodo	biologists	Ron, Santiago R.;Paredes-Aguirre, Poulette;Paucar, Diego A.;López, Yaara;Apunte, Katherine;Ortega, Jhael A.			
B0061B8E72A95208849D496C52D445B9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1725116	https://doi.org/10.3897/zookeys.1286.194383.figure6	Figure 6. Photographs showing color variation in living adult males of Pristimantis milpe sp. nov. A. QCAZ - A 77106, holotype, SVL = 24.77 mm; B. QCAZ - A 76585, paratype, SVL = 26.97 mm; C. QCAZ - A 76518, paratype, SVL = 26.06 mm; D. QCAZ - A 65561, SVL = 19.64 mm; E. QCAZ - A 67428, SVL = 26.47 mm. Left column: dorsolateral view; central column: dorsal view; right column: ventral view. F. QCAZ - A 79899; note significant color change within a period of less than 24 h. Locality data for all specimens is provided in the holotype and paratypes sections.	Figure 6. Photographs showing color variation in living adult males of Pristimantis milpe sp. nov. A. QCAZ - A 77106, holotype, SVL = 24.77 mm; B. QCAZ - A 76585, paratype, SVL = 26.97 mm; C. QCAZ - A 76518, paratype, SVL = 26.06 mm; D. QCAZ - A 65561, SVL = 19.64 mm; E. QCAZ - A 67428, SVL = 26.47 mm. Left column: dorsolateral view; central column: dorsal view; right column: ventral view. F. QCAZ - A 79899; note significant color change within a period of less than 24 h. Locality data for all specimens is provided in the holotype and paratypes sections.	2026-07-24	Ron, Santiago R.;Paredes-Aguirre, Poulette;Paucar, Diego A.;López, Yaara;Apunte, Katherine;Ortega, Jhael A.		Zenodo	biologists	Ron, Santiago R.;Paredes-Aguirre, Poulette;Paucar, Diego A.;López, Yaara;Apunte, Katherine;Ortega, Jhael A.			
B0061B8E72A95208849D496C52D445B9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1725117	https://doi.org/10.3897/zookeys.1286.194383.figure7	Figure 7. Photographs showing color variation in living adult / subadult females of Pristimantis milpe sp. nov. A. QCAZ - A 67424, paratype, adult female, SVL = 32.07 mm; B. QCAZ - A 45268, paratype, subadult female, SVL = 29.44 mm; C. QCAZ - A 74171, paratype, adult female, SVL = 34.21 mm; D. QCAZ - A 74172, paratype, adult female, SVL = 34.17 mm; E. QCAZ - A 51314, adult female, SVL = 33.13 mm. Left column: dorsolateral view; central column: dorsal view; right column: ventral view. Locality data for all specimens is provided in the paratypes section.	Figure 7. Photographs showing color variation in living adult / subadult females of Pristimantis milpe sp. nov. A. QCAZ - A 67424, paratype, adult female, SVL = 32.07 mm; B. QCAZ - A 45268, paratype, subadult female, SVL = 29.44 mm; C. QCAZ - A 74171, paratype, adult female, SVL = 34.21 mm; D. QCAZ - A 74172, paratype, adult female, SVL = 34.17 mm; E. QCAZ - A 51314, adult female, SVL = 33.13 mm. Left column: dorsolateral view; central column: dorsal view; right column: ventral view. Locality data for all specimens is provided in the paratypes section.	2026-07-24	Ron, Santiago R.;Paredes-Aguirre, Poulette;Paucar, Diego A.;López, Yaara;Apunte, Katherine;Ortega, Jhael A.		Zenodo	biologists	Ron, Santiago R.;Paredes-Aguirre, Poulette;Paucar, Diego A.;López, Yaara;Apunte, Katherine;Ortega, Jhael A.			
B0061B8E72A95208849D496C52D445B9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1725120	https://doi.org/10.3897/zookeys.1286.194383.figure10	Figure 10. Relationship between tympanum diameter and SVL for adult individuals of Pristimantis eremitus and P. milpe sp. nov. Note that most P. eremitus are above the regression line, indicating a larger tympanum relative to P. milpe sp. nov.	Figure 10. Relationship between tympanum diameter and SVL for adult individuals of Pristimantis eremitus and P. milpe sp. nov. Note that most P. eremitus are above the regression line, indicating a larger tympanum relative to P. milpe sp. nov.	2026-07-24	Ron, Santiago R.;Paredes-Aguirre, Poulette;Paucar, Diego A.;López, Yaara;Apunte, Katherine;Ortega, Jhael A.		Zenodo	biologists	Ron, Santiago R.;Paredes-Aguirre, Poulette;Paucar, Diego A.;López, Yaara;Apunte, Katherine;Ortega, Jhael A.			
B0061B8E72A95208849D496C52D445B9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1725122	https://doi.org/10.3897/zookeys.1286.194383.figure12	Figure 12. Color variation in preserved adult males of Pristimantis milpe sp. nov. in dorsal, ventral, and lateral view. A. QCAZ - A 77106, holotype, SVL = 24.8 mm; B. QCAZ - A 78622, SVL = 25.6 mm; C. QCAZ - A 29791, SVL = 25.3 mm; D. QCAZ - A 42346, SVL = 28.0 mm; E. QCAZ - A 29319, SVL = 25.9 mm; F. QCAZ - A 65562, paratype, SVL = 26.4 mm; G. QCAZ - A 77107, paratype, SVL = 25.0 mm; H. QCAZ - A 74169, SVL = 22.8 mm; I. QCAZ - A 80061, SVL = 28.6 mm; J. QCAZ - A 79958. SVL = 26.9 mm. Photos by Adrián Sunción-Gavilanez and Nayely Ramos-Miranda. Locality data for all specimens is provided in the holotype and paratype sections.	Figure 12. Color variation in preserved adult males of Pristimantis milpe sp. nov. in dorsal, ventral, and lateral view. A. QCAZ - A 77106, holotype, SVL = 24.8 mm; B. QCAZ - A 78622, SVL = 25.6 mm; C. QCAZ - A 29791, SVL = 25.3 mm; D. QCAZ - A 42346, SVL = 28.0 mm; E. QCAZ - A 29319, SVL = 25.9 mm; F. QCAZ - A 65562, paratype, SVL = 26.4 mm; G. QCAZ - A 77107, paratype, SVL = 25.0 mm; H. QCAZ - A 74169, SVL = 22.8 mm; I. QCAZ - A 80061, SVL = 28.6 mm; J. QCAZ - A 79958. SVL = 26.9 mm. Photos by Adrián Sunción-Gavilanez and Nayely Ramos-Miranda. Locality data for all specimens is provided in the holotype and paratype sections.	2026-07-24	Ron, Santiago R.;Paredes-Aguirre, Poulette;Paucar, Diego A.;López, Yaara;Apunte, Katherine;Ortega, Jhael A.		Zenodo	biologists	Ron, Santiago R.;Paredes-Aguirre, Poulette;Paucar, Diego A.;López, Yaara;Apunte, Katherine;Ortega, Jhael A.			
B0061B8E72A95208849D496C52D445B9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1725121	https://doi.org/10.3897/zookeys.1286.194383.figure11	Figure 11. Color variation in preserved adult / subadult females of Pristimantis milpe sp. nov. in dorsal, ventral, and lateral views. A. QCAZ - A 51314, paratype, adult, SVL = 33.1 mm; B. QCAZ - A 55069, subadult, SVL = 27.3 mm; C. QCAZ - A 67341, paratype, subadult female, SVL = 27.0 mm; D. QCAZ - A 67424, paratype, adult, SVL = 32.1 mm; E. QCAZ - A 58018, adult, SVL = 33.1 mm; F. QCAZ - A 32919, adult, SVL = 32.9 mm; G. QCAZ - A 78832, adult, SVL = 35.4 mm. Photos by Adrián Sunción-Gavilanez. Locality data for all specimens is provided in the paratypes section.	Figure 11. Color variation in preserved adult / subadult females of Pristimantis milpe sp. nov. in dorsal, ventral, and lateral views. A. QCAZ - A 51314, paratype, adult, SVL = 33.1 mm; B. QCAZ - A 55069, subadult, SVL = 27.3 mm; C. QCAZ - A 67341, paratype, subadult female, SVL = 27.0 mm; D. QCAZ - A 67424, paratype, adult, SVL = 32.1 mm; E. QCAZ - A 58018, adult, SVL = 33.1 mm; F. QCAZ - A 32919, adult, SVL = 32.9 mm; G. QCAZ - A 78832, adult, SVL = 35.4 mm. Photos by Adrián Sunción-Gavilanez. Locality data for all specimens is provided in the paratypes section.	2026-07-24	Ron, Santiago R.;Paredes-Aguirre, Poulette;Paucar, Diego A.;López, Yaara;Apunte, Katherine;Ortega, Jhael A.		Zenodo	biologists	Ron, Santiago R.;Paredes-Aguirre, Poulette;Paucar, Diego A.;López, Yaara;Apunte, Katherine;Ortega, Jhael A.			
B0061B8E72A95208849D496C52D445B9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1725123	https://doi.org/10.3897/zookeys.1286.194383.figure13	Figure 13. Advertisement call of Pristimantis milpe sp. nov. A. Oscillogram; B. Spectrogram; C. Power spectrum. Individual not collected.	Figure 13. Advertisement call of Pristimantis milpe sp. nov. A. Oscillogram; B. Spectrogram; C. Power spectrum. Individual not collected.	2026-07-24	Ron, Santiago R.;Paredes-Aguirre, Poulette;Paucar, Diego A.;López, Yaara;Apunte, Katherine;Ortega, Jhael A.		Zenodo	biologists	Ron, Santiago R.;Paredes-Aguirre, Poulette;Paucar, Diego A.;López, Yaara;Apunte, Katherine;Ortega, Jhael A.			
B0061B8E72A95208849D496C52D445B9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1725114	https://doi.org/10.3897/zookeys.1286.194383.figure4	Figure 4. Distribution of P. milpe sp. nov. and the type localities of P. latidiscus and P. subsigillatus. Localities of P. milpe sp. nov. are based on museum specimens. Blue diamonds are localities with specimens with confirmed identification (Museo de Zoología Pontificia Universidad Católica del Ecuador, QCAZ); red diamonds represent localities obtained from the GBIF database (identified as “ P. subsigillatus ”) from the Museum of Comparative Zoology Harvard University (MCZ), Museo Universidad Técnica Particular de Loja (MUTPL), Museo de Zoología Universidad San Francisco de Quito (ZSFQ), Museo de Zoología, Universidad Tecnológica Indoamérica (MZUTI), National Museum of Natural History (USNM), and University of Kansas Museum of Natural History (KU). GBIF localities require confirmation. The type localities of P. latidiscus and P. subsigillatus are also shown with a star (NHMUK).	Figure 4. Distribution of P. milpe sp. nov. and the type localities of P. latidiscus and P. subsigillatus. Localities of P. milpe sp. nov. are based on museum specimens. Blue diamonds are localities with specimens with confirmed identification (Museo de Zoología Pontificia Universidad Católica del Ecuador, QCAZ); red diamonds represent localities obtained from the GBIF database (identified as “ P. subsigillatus ”) from the Museum of Comparative Zoology Harvard University (MCZ), Museo Universidad Técnica Particular de Loja (MUTPL), Museo de Zoología Universidad San Francisco de Quito (ZSFQ), Museo de Zoología, Universidad Tecnológica Indoamérica (MZUTI), National Museum of Natural History (USNM), and University of Kansas Museum of Natural History (KU). GBIF localities require confirmation. The type localities of P. latidiscus and P. subsigillatus are also shown with a star (NHMUK).	2026-07-24	Ron, Santiago R.;Paredes-Aguirre, Poulette;Paucar, Diego A.;López, Yaara;Apunte, Katherine;Ortega, Jhael A.		Zenodo	biologists	Ron, Santiago R.;Paredes-Aguirre, Poulette;Paucar, Diego A.;López, Yaara;Apunte, Katherine;Ortega, Jhael A.			
