taxonID	type	description	language	source
695B87E7F1143446C7FAF9C808F8F91E.taxon	type_taxon	TYPE SPECIES. — Cerithium leve Quoy & Gaimard, 1834 (non Perry, 1811) by subsequent designation (Crosse 1888)	en	Merle, Didier, Pacaud, Jean-Michel, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H. (2026): New campanilids (Mollusca: Gastropoda) from the Ranikot Group (upper Paleocene / early Eocene, Sindh, Pakistan). Comptes Rendus Palevol 25 (1): 1-15, DOI: 10.5852/cr-palevol2026v25a1, URL: https://doi.org/10.5852/cr-palevol2026v25a1
695B87E7F1143448C7C3F8E9080BFBF3.taxon	description	(Figs 2; 3) urn: lsid: zoobank. org: act: D 5 A 4 D 99 C- 2 D 50 - 462 B-B 1 F 6 - E 5 F 9 AE 4 DCD 4 D	en	Merle, Didier, Pacaud, Jean-Michel, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H. (2026): New campanilids (Mollusca: Gastropoda) from the Ranikot Group (upper Paleocene / early Eocene, Sindh, Pakistan). Comptes Rendus Palevol 25 (1): 1-15, DOI: 10.5852/cr-palevol2026v25a1, URL: https://doi.org/10.5852/cr-palevol2026v25a1
695B87E7F1143448C7C3F8E9080BFBF3.taxon	materials_examined	TYPE MATERIAL. — Holotype. Pakistan • Lower Indus Basin, Sindh Province, Jamshoro district; Lakhra Dome, Old Indus section (stn 1), uppermost Lakhra Formation; early Eocene (Ypresian); CPAG. RAN. I. 93 (cast MNHN. F. A 93720), H: 112 mm (apex and early whorls broken, 10 whorls), D: 46.8 mm; Fig. 3 A, B. Paratype. Pakistan • 1 spm; same data as for the holotype; CPAG. RAN. I. 88 (cast MNHN. F. A 93714), H: 56.1 mm (shell incomplete, 9 whorls), D: 27.1 mm; Figs 2 D, 3 D • 1 spm; idem; CPAG. RAN. I. 89 (cast MNHN. F. A 93715), H: 34.4 mm (shell incomplete, 7 whorls), D: 14.8 mm; Figs 3 G • 1 spm; idem; CPAG. RAN. I. 90 (cast MNHN. F. A 93716), H: 19.3 mm (shell incomplete, 7 whorls), D: 11.4 mm; Figs 2 A, 3 F • 1 spm; idem; CPAG. RAN. I. 91 (cast MNHN. F. A 93718), H: 23.3 mm (shell incomplete, 6 whorls), D: 12.1 mm; Figs 2 B, 3 E • 1 spm; idem; CPAG. RAN. I. 92 (cast MNHN. F. A 93719), H: 33.7 mm (shell incomplete, 9 whorls), 11.8 mm; Figs 2 C, 3 H • 1 spm; idem; CPAG. RAN. I. 94 (cast MNHN. F. A 93722), H: 78.6 mm (shell incomplete, 10 whorls), D: 33 mm; Fig. 3 C. OTHER EXAMINED MATERIAL. — Pakistan • 5 spms; same data as for the holotype; MNHN. F. A 93713 • 6 spms; idem; MNHN. F. A 93717 • 8 spms; idem; MNHN. F. A 93721.	en	Merle, Didier, Pacaud, Jean-Michel, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H. (2026): New campanilids (Mollusca: Gastropoda) from the Ranikot Group (upper Paleocene / early Eocene, Sindh, Pakistan). Comptes Rendus Palevol 25 (1): 1-15, DOI: 10.5852/cr-palevol2026v25a1, URL: https://doi.org/10.5852/cr-palevol2026v25a1
695B87E7F1143448C7C3F8E9080BFBF3.taxon	etymology	ETYMOLOGY. — Dedicated to Grégoire Métais (CNRS), main contributor to the 2011, 2012 and 2014 Franco-Pakistani fieldtrips to Sindh.	en	Merle, Didier, Pacaud, Jean-Michel, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H. (2026): New campanilids (Mollusca: Gastropoda) from the Ranikot Group (upper Paleocene / early Eocene, Sindh, Pakistan). Comptes Rendus Palevol 25 (1): 1-15, DOI: 10.5852/cr-palevol2026v25a1, URL: https://doi.org/10.5852/cr-palevol2026v25a1
695B87E7F1143448C7C3F8E9080BFBF3.taxon	biology_ecology	TYPE HORIZON. — Diversified mollusc bed of the uppermost Lakhra Formation (top of the Old Indus section, stn 1), early Eocene (Ypresian).	en	Merle, Didier, Pacaud, Jean-Michel, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H. (2026): New campanilids (Mollusca: Gastropoda) from the Ranikot Group (upper Paleocene / early Eocene, Sindh, Pakistan). Comptes Rendus Palevol 25 (1): 1-15, DOI: 10.5852/cr-palevol2026v25a1, URL: https://doi.org/10.5852/cr-palevol2026v25a1
695B87E7F1143448C7C3F8E9080BFBF3.taxon	distribution	TYPE LOCALITY. — Pakistan, Sindh Province, Jamshoro district; Lakhra Dome, Old Indus section (stn 1). DISTRIBUTION. — Pakistan, Sindh, Lakhra Formation at Old Indus section (stn 1); Leilan and Jhirak (Cossmann & Pissarro 1909), early Eocene (Ypresian).	en	Merle, Didier, Pacaud, Jean-Michel, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H. (2026): New campanilids (Mollusca: Gastropoda) from the Ranikot Group (upper Paleocene / early Eocene, Sindh, Pakistan). Comptes Rendus Palevol 25 (1): 1-15, DOI: 10.5852/cr-palevol2026v25a1, URL: https://doi.org/10.5852/cr-palevol2026v25a1
695B87E7F1143448C7C3F8E9080BFBF3.taxon	description	DESCRIPTION Medium sized, elongate, conical shell of more than ten convex teleoconch whorls (based on the holotype); apical angle 30 °. Antepenultimate and penultimate whorls slightly more inflated that earlier spire whorls. Last whorl nearest 39.4 % total height (holotype). Whorls separated by narrowly impressed suture. Protoconch and first teleoconch whorls unknown. Spiral sculpture (composite description). Specimen MNHN. F. A 93716 (H: 19.3 mm, D: 11.4 mm, seven whorls, Fig. 2 A) on whorls 1 to 5, P 1 to P 3 present; P 1 close-set to adapical suture and more prominent than P 2 and P 3; P 1 - P 2 fine but well-marked; P 3 close-set to abapical suture. On whorl no. 6 (see Fig. 2 A), appearance of s 2 and appearance of weakly nodular varical ribs on P 1. On whorl no. 7, P 1 - P 2 merging – Specimen MNHN. F. A 93718 (H: 23.3 mm, D: 12.1 mm, six whorls, Fig. 2 B) on whorl nos. 1 - 2, P 1 to P 3. On whorl no. 3, P 1 and P 2 merging. On whorls nos. 4 - 6 (see Fig. 2 B), P 1 - P 2 fused and nodular varical ribs devoid of traces of spiral cord becoming more prominent on whorl no. 6 – Specimen MNHN. F. A 93719 (H: 33.7 mm, D: 11.8 mm, six whorls, Fig. 2 C), on whorl no. 2, appearance of two tertiary cords (t 1 + t 2) between P 1 and P 2. On whorl no. 4, P 1, t 1, t 2 fusing. On whorl no. 5, P 2 fusing with P 1, t 1 + t 2 already fused. Appearance of s 2. On whorl no. 6, disappearance of cord except P 3 – SpecimenMNHN. F. A 93714 (H: 56.1 mm, D: 27.1 mm, nine whorls, Fig. 2 D), on whorl no. 1, P 1 - P 3. On whorl no. 2, P 1 and P 2 fusing. On whorl no. 3 to no. 5, P 1 and P 2 fused and P 3 close set to the abapical suture. On whorls no. 6 to no. 9, no spiral sculpture – Holotype MNHN. F. A 93714 (H: 112 mm, D: 46.8 mm, ten whorls, Fig. 3 A, B) no cord even on early whorls. Axial sculpture appearing approximately after disappearance of spiral cords and consisting of 15 moderately thick varicose ribs on penultimate and last whorls, and between 12 - 16 ribs on preceding whorls. First ribs more varicose on adapical part of whorls. Aperture narrow 54.8 % of diameter and 48.8 % of length of last whorl. Anal canal indistinct. Columella weakly concave, with two folds. Siphonal canal short.	en	Merle, Didier, Pacaud, Jean-Michel, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H. (2026): New campanilids (Mollusca: Gastropoda) from the Ranikot Group (upper Paleocene / early Eocene, Sindh, Pakistan). Comptes Rendus Palevol 25 (1): 1-15, DOI: 10.5852/cr-palevol2026v25a1, URL: https://doi.org/10.5852/cr-palevol2026v25a1
695B87E7F1143448C7C3F8E9080BFBF3.taxon	diagnosis	COMPARISONS Cossmann & Pissarro (1909) identified similar specimens from the Lakhra Formation as? Cerithium (Campanile) subsemicostatum d’Archiac & Haime, 1854. Indeed, they are reminiscent of that species, which is also a Campanile and displays nodular varical ribs (see d’Archiac & Haime 1854: pl. 29, fig. 5). However, Campanile subsemicostatum differs in its more numerous ribs (8 - 9 on the ventral side, instead 6 - 7 in C. metaisi Merle & Pacaud, n. sp.), in its opistocyrt ribs and in its wider apical angle. Moreover, d’Archiac & Haime (1854: 300) noticed that the matrix containing Campanile subsemicostatum is associated with Nummulites garansensis (accepted as Nummulites garansiana Joly & Leymerie, 1848), a Rupelian Large Benthic Foraminifera (LBF) and for this reason, Cossmann & Pissarro (1909: B 53) placed a query after their identification. Considering their morphological differences and also their distant stratigraphic age, there is no doubt that C. subsemicostatum (d’Archiac & Haime, 1854) and C. metaisi Merle & Pacaud, n. sp. belong to two distinct species. Campanile metaisi Merle & Pacaud, n. sp. is reminiscent to Campanile cornucopiae (J. Sowerby, 1818) from the middle Eocene of the Paris Basin, Normandy and Hampshire in its medium size and in its nodular varices. However, Campanile cornucopiae differs by its early whorls that are more ornamented (see Wrigley 1940, figs 1, 4 and Pacaud et al. 2014: pl. 8).	en	Merle, Didier, Pacaud, Jean-Michel, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H. (2026): New campanilids (Mollusca: Gastropoda) from the Ranikot Group (upper Paleocene / early Eocene, Sindh, Pakistan). Comptes Rendus Palevol 25 (1): 1-15, DOI: 10.5852/cr-palevol2026v25a1, URL: https://doi.org/10.5852/cr-palevol2026v25a1
695B87E7F11A3449C73BF8E90F4DFD4D.taxon	description	urn: lsid: zoobank. org: act: 0 DA 1619 A-ACDE- 4170 - BD 12 - B 7 F 2 AAF 647 FA	en	Merle, Didier, Pacaud, Jean-Michel, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H. (2026): New campanilids (Mollusca: Gastropoda) from the Ranikot Group (upper Paleocene / early Eocene, Sindh, Pakistan). Comptes Rendus Palevol 25 (1): 1-15, DOI: 10.5852/cr-palevol2026v25a1, URL: https://doi.org/10.5852/cr-palevol2026v25a1
695B87E7F11A3449C73BF8E90F4DFD4D.taxon	type_taxon	TYPE SPECIES. — Campanistylus lakhraensis Merle & Pacaud, n. gen., n. sp.	en	Merle, Didier, Pacaud, Jean-Michel, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H. (2026): New campanilids (Mollusca: Gastropoda) from the Ranikot Group (upper Paleocene / early Eocene, Sindh, Pakistan). Comptes Rendus Palevol 25 (1): 1-15, DOI: 10.5852/cr-palevol2026v25a1, URL: https://doi.org/10.5852/cr-palevol2026v25a1
695B87E7F11A3449C73BF8E90F4DFD4D.taxon	etymology	ETYMOLOGY. — From Campanile and stylus (from Greek name στύΛΟς meaning column reflecting the straight-sided shape of the early teleconch whorls). Gender, male.	en	Merle, Didier, Pacaud, Jean-Michel, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H. (2026): New campanilids (Mollusca: Gastropoda) from the Ranikot Group (upper Paleocene / early Eocene, Sindh, Pakistan). Comptes Rendus Palevol 25 (1): 1-15, DOI: 10.5852/cr-palevol2026v25a1, URL: https://doi.org/10.5852/cr-palevol2026v25a1
695B87E7F11A3449C73BF8E90F4DFD4D.taxon	diagnosis	DIAGNOSIS. — Medium sized shell; early spire very slender turriculate, resembling a column, followed by whorls gradually increasing in growth rate resulting in broader, more convex whorls. Overall aspect of shell, turriculate, coleoconid. Whorls separated by narrowly impressed suture. Delicate spiral sculpture present on around 25 early teleoconch whorls, disappearing on later whorls. Series of cords including P 1, P 2, s 2, P 3. Surface of cords usually smooth, sometimes slightly granulose. Aperture narrow, ovoid, compressed adapically. Anal canal short, narrow. Columella weakly concave, one fold. Siphonal canal moderately short, bent and dorsally recurved. Outer lip prosocyrt, not thickened. Remains of periostracum imprints. Residual colour pattern consisting of red stripes, approximately parallel, with inclination varying from slightly sinuous to sigmoidal.	en	Merle, Didier, Pacaud, Jean-Michel, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H. (2026): New campanilids (Mollusca: Gastropoda) from the Ranikot Group (upper Paleocene / early Eocene, Sindh, Pakistan). Comptes Rendus Palevol 25 (1): 1-15, DOI: 10.5852/cr-palevol2026v25a1, URL: https://doi.org/10.5852/cr-palevol2026v25a1
695B87E7F11B3449C4C1FD2E08A7F830.taxon	description	(Figs 5; 6) urn: lsid: zoobank. org: act: 501 A 660 A-F 529 - 48 F 8 - 8 D 9 D-D 49 A 3571 A 5 F 5	en	Merle, Didier, Pacaud, Jean-Michel, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H. (2026): New campanilids (Mollusca: Gastropoda) from the Ranikot Group (upper Paleocene / early Eocene, Sindh, Pakistan). Comptes Rendus Palevol 25 (1): 1-15, DOI: 10.5852/cr-palevol2026v25a1, URL: https://doi.org/10.5852/cr-palevol2026v25a1
695B87E7F11B3449C4C1FD2E08A7F830.taxon	materials_examined	TYPE MATERIAL. — Holotype. Pakistan • Lower Indus Basin, Sindh Province, Jamshoro district; Lakhra Dome, East of Lakhra village section (stn 2), lower Lakhra Formation; early Eocene (Ypresian); CPAG. RAN. I. 95 (cast MNHN. F. A 93729); H: 136.8 mm (c. 32 whorls), D: 40.1 mm; Fig. 6 A, B. Paratypes. Pakistan • 1 spm; same data as the holotype; CPAG. RAN. I. 102 (cast MNHN. F. A 93726), H: 31.9 mm (shell incomplete, 6 whorls), D: 17.9 mm; Fig. 5 D • 1 spm; idem; CPAG. RAN. I. 103 (cast MNHN. F. A 93725), H: 57.1 mm (shell incomplete, 8 whorls), D: 27.1 mm; Fig. 5 C • 1 spm; idem; CPAG. RAN. I. 104 (cast MNHN. F. A 93727), H: 29.2 mm (shell incomplete, 7 whorls), D: 14 mm; Fig. 5 B • 1 spm; idem; CPAG. RAN. I. 105 (cast MNHN. F. A 93728), H: 58.6 mm (shell incomplete, 8 whorls), D: 29.7 mm; Fig. 5 A • 1 spm; same data as for the holotype; CPAG. RAN. I. 96 (cast MNHN. F. A 98131), H: 159.4 mm (c. 27 whorls), D: 43.3 mm; Fig. 6 C • 1 spm; idem; CPAG. RAN. I. 97 (cast MNHN. F. A 98129), H: 114.9 mm (shell incomplete, 6 whorls), D: 43.8 mm • 1 spm; idem; CPAG. RAN. I. 98 (cast MNHN. F. A 93712), H: 91.4 mm (shell incomplete, 3 whorls), D: 46.1 mm; Fig. 6 H, I • 1 spm; idem; CPAG. RAN. I. 99 (cast MNHN. F. A 98132), H: 115.4 mm (shell incomplete, 7 whorls), D: 44.1 mm; Fig. 6 F, G • 1 spm; idem; CPAG. RAN. I. 100 (cast MNHN. F. A 93711), H: 98.2 mm (c. 23 whorls), D: 28.8 mm; Fig. 6 D, E • 1 spm; idem; CPAG. RAN. I. 101 (cast MNHN. F. A 93724), H: 17.8 mm (shell incomplete, 3 whorls), D: 12.3 mm. OTHER EXAMINED MATERIAL. — Pakistan • 1 spm; same data as the holotype; MNHN. F. A 93696 • 1 spm; idem; MNHN. F. A 93697 • 1 spm; idem; MNHN. F. A 93698 • 2 spms; idem; MNHN. F. A 93699 • 1 spm; idem; MNHN. F. A 93700 • 2 spms; idem; MNHN. F. A 93701 • 3 spms; idem; MNHN. F. A 93702 • 1 spm; idem; MNHN. F. A 93704 • 5 spms; idem; MNHN. F. A 93705 • 1 spm; idem; MNHN. F. A 93706 • 3 spms; idem; MNHN. F. A 93707 • 2 spms; idem; MNHN. F. A 93708 • 1 spm; idem; MNHN. F. A 93709 • 2 spms; idem; MNHN. F. A 93710.	en	Merle, Didier, Pacaud, Jean-Michel, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H. (2026): New campanilids (Mollusca: Gastropoda) from the Ranikot Group (upper Paleocene / early Eocene, Sindh, Pakistan). Comptes Rendus Palevol 25 (1): 1-15, DOI: 10.5852/cr-palevol2026v25a1, URL: https://doi.org/10.5852/cr-palevol2026v25a1
695B87E7F11B3449C4C1FD2E08A7F830.taxon	etymology	ETYMOLOGY. — From the type formation: Lakhra Formation.	en	Merle, Didier, Pacaud, Jean-Michel, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H. (2026): New campanilids (Mollusca: Gastropoda) from the Ranikot Group (upper Paleocene / early Eocene, Sindh, Pakistan). Comptes Rendus Palevol 25 (1): 1-15, DOI: 10.5852/cr-palevol2026v25a1, URL: https://doi.org/10.5852/cr-palevol2026v25a1
695B87E7F11B3449C4C1FD2E08A7F830.taxon	biology_ecology	TYPE HORIZON. — Diversified molluscs bed, base of the East-Lakhra village section (stn 2), lower Lakhra Formation, early Eocene (Ypresian).	en	Merle, Didier, Pacaud, Jean-Michel, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H. (2026): New campanilids (Mollusca: Gastropoda) from the Ranikot Group (upper Paleocene / early Eocene, Sindh, Pakistan). Comptes Rendus Palevol 25 (1): 1-15, DOI: 10.5852/cr-palevol2026v25a1, URL: https://doi.org/10.5852/cr-palevol2026v25a1
695B87E7F11B3449C4C1FD2E08A7F830.taxon	distribution	TYPE LOCALITY. — Pakistan, Sindh Province, Jamshoro district; Lakhra Dome, East-Lakhra village section (stn 2). DISTRIBUTION. — Lakhra Formation, early Eocene (Ypresian).	en	Merle, Didier, Pacaud, Jean-Michel, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H. (2026): New campanilids (Mollusca: Gastropoda) from the Ranikot Group (upper Paleocene / early Eocene, Sindh, Pakistan). Comptes Rendus Palevol 25 (1): 1-15, DOI: 10.5852/cr-palevol2026v25a1, URL: https://doi.org/10.5852/cr-palevol2026v25a1
695B87E7F11B3449C4C1FD2E08A7F830.taxon	description	DESCRIPTION Medium sized shell (H = 136 mm) of more than 33 teleoconch whorls starting with 24 turriculate whorls resembling a column, followed by eight whorls becoming increasingly convex; overall aspect of shell, turriculate, coleoconid. Antepenultimate and penultimate whorls slightly more inflated that earlier spire whorls. Apical angle 18 - 19 °. Last whorl 29.3 % total height (holotype). Whorls separated by narrowly impressed suture. Protoconch and first teleoconch whorls unknown. Delicate spiral sculpture present on early teleoconch whorls, disappearing on last whorls – Detailed composite description of the spiral sculpture: holotype MNHN. F. A 93729 (Fig. 6 A, B, H: 136.8 mm), primary cords P 1 - P 3 present until 20 th whorl, P 1 close-set to the adapical suture more prominent than P 2 and P 3, P 2 less prominent. On whorl no. 21, appearance of s 2 between P 2 and P 3, P 2 less prominent than the other primary cords. From whorl no. 22 to no. 24 (Fig. 6 B), s 2 becoming larger and subequal to P 2, s 2 and P 2 less prominent that the other cords; small granules visible on cords. On whorls nos. 25 - 26 (Fig. 6 B), P 1, P 2 and s 2 starting to fade gradually, P 3 still prominent. On whorl no. 27 (Fig. 6 B), loss of spiral sculpture – Paratype MNHN. F. A 93728 (Fig. 5 A, H: 58.6 mm) on whorls no. 1 to no. 4, P 1 - P 3 and s 2 present; P 1 prominent, close-set to adapical suture; P 3 close-set to abapical suture; s 2 close-set to P 2 and less prominent. On whorls nos. 5 - 6, P 1, P 2 and s 2 starting to fade gradually, P 3 still prominent. On whorl no. 7, P 1 tending to fuse with P 2. On whorl 8, loss of the spiral sculpture – Paratype MNHN. F. A 93727 (Fig. 5 B, H: 29.2 mm), on whorls no. 1 to no. 3, P 1 - P 3 and s 2 present; P 1 prominent, close-set to adapical suture; P 3 close-set to abapical suture; s 2 close-set to P 2 and less prominent. On whorl nos. 4 - 5, P 1, P 2 and s 2 starting to fade, P 3 still prominent. On whorl no. 6, loss of the spiral sculpture – Paratype MNHN. F. A 93725 (Fig. 5 C, H: 57.1 mm), on whorls no. 1 to no. 3, P 1 - P 3 and s 2 present; P 1 prominent, close-set to adapical suture; P 3 prominent close-set to abapical suture; P 2 and s 2 weaker; small granules on P 2, s 2 and P 3. On whorl no. 4, P 1, P 2 to tending to fuse and s 2 starting to fade gradually, P 3 still prominent. On whorl no. 5, only P 3. On whorl no. 6, loss of the spiral sculpture – Paratype MNHN. F. A 93726 (Fig. 5 D, H: 31.9 mm), on whorls nos. 1 - 2, P 1 - P 3 and s 2 present; P 1 close-set to adapical suture; P 3 close-set to abapical suture; s 2 close to P 2 and less prominent. On whorl no. 3, P 1 and P 2 tending to fuse. On whorl no. 4, only P 3. On whorl no. 5, total loss of the spiral sculpture – No axial sculpture, except some growing striae. Aperture narrow, ovoid, compressed adapically, of 88.7 % of diameter and 84.33 % of length of last whorl. Anal canal short and narrow. Columella weakly concave with one fold; columellar lip slightly erect; parietal lip adherent. Siphonal canal moderately short, bent, turning left and dorsally curved. Outer lip prosocyrt, not thickened. Remains of periostracum imprints (paratype MNHN. F. A 93711). Residual colour pattern consisting of red stripes, approximately parallel, with inclination varying from slightly sinuous to sigmoidal (paratype MNHN. F. A 98132, Fig. 6 F, G).	en	Merle, Didier, Pacaud, Jean-Michel, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H. (2026): New campanilids (Mollusca: Gastropoda) from the Ranikot Group (upper Paleocene / early Eocene, Sindh, Pakistan). Comptes Rendus Palevol 25 (1): 1-15, DOI: 10.5852/cr-palevol2026v25a1, URL: https://doi.org/10.5852/cr-palevol2026v25a1
