identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
695B87E7F1143446C7FAF9C808F8F91E.text	695B87E7F1143446C7FAF9C808F8F91E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Campanile P. Fischer 1884	<div><p>Genus Campanile P. Fischer, 1884</p><p>TYPE SPECIES. — Cerithium leve Quoy &amp; Gaimard, 1834 (non Perry, 1811) by subsequent designation (Crosse 1888)</p><p>(accepted as Cerithium symbolicum Iredale, 1917).</p></div>	https://treatment.plazi.org/id/695B87E7F1143446C7FAF9C808F8F91E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier;Pacaud, Jean-Michel;Lashari, Rafiq A.;Brohi, Imdad A.;Solangi, Sarfraz H.	Merle, Didier, Pacaud, Jean-Michel, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H. (2026): New campanilids (Mollusca: Gastropoda) from the Ranikot Group (upper Paleocene / early Eocene, Sindh, Pakistan). Comptes Rendus Palevol 25 (1): 1-15, DOI: 10.5852/cr-palevol2026v25a1, URL: https://doi.org/10.5852/cr-palevol2026v25a1
695B87E7F1143448C7C3F8E9080BFBF3.text	695B87E7F1143448C7C3F8E9080BFBF3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Campanile metaisi Merle & Pacaud 2026	<div><p>Campanile metaisi Merle &amp; Pacaud, n. sp.</p><p>(Figs 2; 3)</p><p>urn:lsid:zoobank.org:act: D5A4D99C-2D50-462B-B1F6-E5F9AE4DCD4D</p><p>? Cerithium (Campanile) subsemicostatum — Cossmann &amp; Pissarro 1909: 53-54, pl. 5, figs 9, 10 (non d’Archiac &amp; Haime, 1854).</p><p>TYPE MATERIAL. — Holotype. Pakistan • Lower Indus Basin, Sindh Province, Jamshoro district; Lakhra Dome, Old Indus section (stn 1), uppermost Lakhra Formation; early Eocene (Ypresian); CPAG. RAN.I.93 (cast MNHN. F.A93720), H: 112 mm (apex and early whorls broken, 10 whorls), D: 46.8 mm; Fig. 3A, B.</p><p>Paratype. Pakistan • 1 spm; same data as for the holotype; CPAG. RAN.I.88 (cast MNHN. F.A93714), H: 56.1 mm (shell incomplete, 9 whorls), D: 27.1 mm; Figs 2D, 3D • 1 spm; idem; CPAG. RAN.I.89 (cast MNHN. F.A93715), H: 34.4 mm (shell incomplete, 7 whorls), D: 14.8 mm; Figs 3G • 1 spm; idem; CPAG.RAN.I.90 (cast MNHN. F.A93716), H: 19.3 mm (shell incomplete, 7 whorls), D: 11.4 mm; Figs 2A, 3F • 1 spm; idem; CPAG.RAN.I.91 (cast MNHN. F.A93718), H: 23.3 mm (shell incomplete, 6 whorls), D: 12.1 mm; Figs 2B, 3E • 1 spm; idem; CPAG.RAN.I.92 (cast MNHN. F.A93719), H: 33.7 mm (shell incomplete, 9 whorls), 11.8 mm; Figs 2C, 3H • 1 spm; idem; CPAG.RAN.I.94 (cast MNHN. F.A93722), H: 78.6 mm (shell incomplete, 10 whorls), D: 33 mm; Fig. 3C .</p><p>OTHER EXAMINED MATERIAL. — Pakistan • 5 spms; same data as for the holotype; MNHN. F.A93713 • 6 spms; idem; MNHN. F.A93717 • 8 spms; idem; MNHN. F.A93721 .</p><p>ETYMOLOGY. — Dedicated to Grégoire Métais (CNRS), main contributor to the 2011, 2012 and 2014 Franco-Pakistani fieldtrips to Sindh.</p><p>TYPE HORIZON. — Diversified mollusc bed of the uppermost Lakhra Formation (top of the Old Indus section, stn 1), early Eocene (Ypresian).</p><p>TYPE LOCALITY. — Pakistan, Sindh Province, Jamshoro district; Lakhra Dome, Old Indus section (stn 1).</p><p>DISTRIBUTION. — Pakistan, Sindh, Lakhra Formation at Old Indus section (stn 1); Leilan and Jhirak (Cossmann &amp; Pissarro 1909), early Eocene (Ypresian).</p><p>DESCRIPTION</p><p>Medium sized, elongate, conical shell of more than ten convex teleoconch whorls (based on the holotype); apical angle 30°. Antepenultimate and penultimate whorls slightly more inflated that earlier spire whorls. Last whorl nearest 39.4% total height (holotype). Whorls separated by narrowly impressed suture. Protoconch and first teleoconch whorls unknown. Spiral sculpture (composite description). Specimen MNHN.F. A93716 (H: 19.3 mm, D: 11.4 mm, seven whorls, Fig. 2A) on whorls 1 to 5, P1 to P3 present; P1 close-set to adapical suture and more prominent than P2 and P3; P1-P2 fine but well-marked; P3 close-set to abapical suture. On whorl no. 6 (see Fig. 2A), appearance of s2 and appearance of weakly nodular varical ribs on P1. On whorl no. 7, P1-P2 merging – Specimen MNHN.F.A93718 (H: 23.3 mm, D: 12.1 mm, six whorls, Fig. 2B) on whorl nos. 1-2, P1 to P3. On whorl no. 3, P1 and P2 merging. On whorls nos. 4-6 (see Fig. 2B), P1-P2 fused and nodular varical ribs devoid of traces of spiral cord becoming more prominent on whorl no. 6 – Specimen MNHN.F.A93719 (H: 33.7 mm, D: 11.8 mm, six whorls, Fig. 2C), on whorl no. 2, appearance of two tertiary cords (t1+ t2) between P1 and P2. On whorl no. 4, P1, t1, t2 fusing. On whorl no. 5, P2 fusing with P1, t1 + t2 already fused. Appearance of s2. On whorl no. 6, disappearance of cord except P3 – SpecimenMNHN.F.A93714 (H: 56.1 mm, D: 27.1 mm, nine whorls, Fig. 2D), on whorl no. 1, P1-P3. On whorl no. 2, P1 and P2 fusing. On whorl no. 3 to no. 5, P1 and P2 fused and P3 close set to the abapical suture. On whorls no. 6 to no. 9, no spiral sculpture – Holotype MNHN.F.A93714 (H: 112 mm, D: 46.8 mm, ten whorls, Fig. 3A, B) no cord even on early whorls. Axial sculpture appearing approximately after disappearance of spiral cords and consisting of 15 moderately thick varicose ribs on penultimate and last whorls, and between 12-16 ribs on preceding whorls. First ribs more varicose on adapical part of whorls. Aperture narrow 54.8% of diameter and 48.8% of length of last whorl. Anal canal indistinct. Columella weakly concave, with two folds. Siphonal canal short.</p><p>COMPARISONS</p><p>Cossmann &amp; Pissarro (1909) identified similar specimens from the Lakhra Formation as? Cerithium (Campanile) subsemicostatum d’Archiac &amp; Haime, 1854 . Indeed, they are reminiscent of that species, which is also a Campanile and displays nodular varical ribs (see d’Archiac &amp; Haime 1854: pl. 29, fig. 5). However, Campanile subsemicostatum differs in its more numerous ribs (8-9 on the ventral side, instead 6-7 in C. metaisi Merle &amp; Pacaud, n. sp.), in its opistocyrt ribs and in its wider apical angle. Moreover, d’Archiac &amp; Haime (1854: 300) noticed that the matrix containing Campanile subsemicostatum is associated with Nummulites garansensis (accepted as Nummulites garansiana Joly &amp; Leymerie, 1848), a Rupelian Large Benthic Foraminifera (LBF) and for this reason, Cossmann &amp; Pissarro (1909: B53) placed a query after their identification. Considering their morphological differences and also their distant stratigraphic age, there is no doubt that C. subsemicostatum (d’Archiac &amp; Haime, 1854) and C. metaisi Merle &amp; Pacaud, n. sp. belong to two distinct species. Campanile metaisi Merle &amp; Pacaud, n. sp. is reminiscent to Campanile cornucopiae (J. Sowerby, 1818) from the middle Eocene of the Paris Basin, Normandy and Hampshire in its medium size and in its nodular varices. However, Campanile cornucopiae differs by its early whorls that are more ornamented (see Wrigley 1940, figs 1, 4 and Pacaud et al. 2014: pl. 8).</p></div>	https://treatment.plazi.org/id/695B87E7F1143448C7C3F8E9080BFBF3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier;Pacaud, Jean-Michel;Lashari, Rafiq A.;Brohi, Imdad A.;Solangi, Sarfraz H.	Merle, Didier, Pacaud, Jean-Michel, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H. (2026): New campanilids (Mollusca: Gastropoda) from the Ranikot Group (upper Paleocene / early Eocene, Sindh, Pakistan). Comptes Rendus Palevol 25 (1): 1-15, DOI: 10.5852/cr-palevol2026v25a1, URL: https://doi.org/10.5852/cr-palevol2026v25a1
695B87E7F11A3449C73BF8E90F4DFD4D.text	695B87E7F11A3449C73BF8E90F4DFD4D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Campanistylus Merle & Pacaud 2026	<div><p>Genus Campanistylus Merle &amp; Pacaud, n. gen.</p><p>urn:lsid:zoobank.org:act: 0DA1619A-ACDE-4170-BD12-B7F2AAF647FA</p><p>TYPE SPECIES. — Campanistylus lakhraensis Merle &amp; Pacaud, n. gen., n. sp.</p><p>SPECIES INCLUDED. — Type species.</p><p>ETYMOLOGY. — From Campanile and stylus (from Greek name στύΛΟς meaning column reflecting the straight-sided shape of the early teleconch whorls). Gender, male.</p><p>DIAGNOSIS. — Medium sized shell; early spire very slender turriculate, resembling a column, followed by whorls gradually increasing in growth rate resulting in broader, more convex whorls. Overall aspect of shell, turriculate, coleoconid. Whorls separated by narrowly impressed suture. Delicate spiral sculpture present on around 25 early teleoconch whorls, disappearing on later whorls. Series of cords including P1, P2, s2, P3. Surface of cords usually smooth, sometimes slightly granulose. Aperture narrow, ovoid, compressed adapically. Anal canal short, narrow. Columella weakly concave, one fold. Siphonal canal moderately short, bent and dorsally recurved. Outer lip prosocyrt, not thickened. Remains of periostracum imprints. Residual colour pattern consisting of red stripes, approximately parallel, with inclination varying from slightly sinuous to sigmoidal.</p></div>	https://treatment.plazi.org/id/695B87E7F11A3449C73BF8E90F4DFD4D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier;Pacaud, Jean-Michel;Lashari, Rafiq A.;Brohi, Imdad A.;Solangi, Sarfraz H.	Merle, Didier, Pacaud, Jean-Michel, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H. (2026): New campanilids (Mollusca: Gastropoda) from the Ranikot Group (upper Paleocene / early Eocene, Sindh, Pakistan). Comptes Rendus Palevol 25 (1): 1-15, DOI: 10.5852/cr-palevol2026v25a1, URL: https://doi.org/10.5852/cr-palevol2026v25a1
695B87E7F11B3449C4C1FD2E08A7F830.text	695B87E7F11B3449C4C1FD2E08A7F830.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Campanistylus lakhraensis Merle & Pacaud 2026	<div><p>Campanistylus lakhraensis Merle &amp; Pacaud, n. gen., n. sp.</p><p>(Figs 5; 6)</p><p>urn:lsid:zoobank.org:act: 501A660A-F529-48F8-8D9D-D49A3571A5F5</p><p>TYPE MATERIAL. — Holotype. Pakistan • Lower Indus Basin, Sindh Province, Jamshoro district; Lakhra Dome, East of Lakhra village section (stn 2), lower Lakhra Formation; early Eocene (Ypresian); CPAG. RAN.I.95 (cast MNHN. F.A93729); H: 136.8 mm (c. 32 whorls), D: 40.1 mm; Fig. 6A, B.</p><p>Paratypes. Pakistan • 1 spm; same data as the holotype; CPAG. RAN.I.102 (cast MNHN. F.A93726), H: 31.9 mm (shell incomplete, 6 whorls), D: 17.9 mm; Fig. 5D • 1 spm; idem; CPAG. RAN.I.103 (cast MNHN. F.A93725), H: 57.1 mm (shell incomplete, 8 whorls), D: 27.1 mm; Fig. 5C • 1 spm; idem; CPAG. RAN.I.104 (cast MNHN. F.A93727), H: 29.2 mm (shell incomplete, 7 whorls), D: 14 mm; Fig. 5B • 1 spm; idem; CPAG.RAN.I.105 (cast MNHN. F.A93728), H: 58.6 mm (shell incomplete, 8 whorls), D: 29.7 mm; Fig. 5A • 1 spm; same data as for the holotype; CPAG. RAN.I.96 (cast MNHN. F.A98131), H: 159.4 mm (c. 27 whorls), D: 43.3 mm; Fig. 6C • 1 spm; idem; CPAG.RAN.I.97 (cast MNHN. F.A98129), H: 114.9 mm (shell incomplete, 6 whorls), D: 43.8 mm • 1 spm; idem; CPAG.RAN.I.98 (cast MNHN. F.A93712), H: 91.4 mm (shell incomplete, 3 whorls), D: 46.1 mm; Fig. 6H, I • 1 spm; idem; CPAG.RAN.I.99 (cast MNHN. F.A98132), H: 115.4 mm (shell incomplete, 7 whorls), D: 44.1 mm; Fig. 6F, G • 1 spm; idem; CPAG.RAN.I.100 (cast MNHN. F.A93711), H: 98.2 mm (c. 23 whorls), D: 28.8 mm; Fig. 6D, E • 1 spm; idem; CPAG.RAN.I.101 (cast MNHN. F.A93724), H: 17.8 mm (shell incomplete, 3 whorls), D: 12.3 mm .</p><p>OTHER EXAMINED MATERIAL. — Pakistan • 1 spm; same data as the holotype; MNHN. F.A93696 • 1 spm; idem; MNHN. F.A93697 • 1 spm; idem; MNHN. F.A93698 • 2 spms; idem; MNHN. F.A93699 • 1 spm; idem; MNHN. F.A93700 • 2 spms; idem; MNHN. F.A93701 • 3 spms; idem; MNHN. F.A93702 • 1 spm; idem; MNHN. F.A93704 • 5 spms; idem; MNHN. F.A93705 • 1 spm; idem; MNHN. F.A93706 • 3 spms; idem; MNHN. F.A93707 • 2 spms; idem; MNHN. F.A93708 • 1 spm; idem; MNHN. F.A93709 • 2 spms; idem; MNHN. F.A93710 .</p><p>ETYMOLOGY. — From the type formation: Lakhra Formation.</p><p>TYPE HORIZON. — Diversified molluscs bed, base of the East-Lakhra village section (stn 2), lower Lakhra Formation, early Eocene (Ypresian).</p><p>TYPE LOCALITY. — Pakistan, Sindh Province, Jamshoro district; Lakhra Dome, East-Lakhra village section (stn 2).</p><p>DISTRIBUTION. — Lakhra Formation, early Eocene (Ypresian).</p><p>DESCRIPTION</p><p>Medium sized shell (H = 136 mm) of more than 33 teleoconch whorls starting with 24 turriculate whorls resembling a column, followed by eight whorls becoming increasingly convex; overall aspect of shell, turriculate, coleoconid. Antepenultimate and penultimate whorls slightly more inflated that earlier spire whorls. Apical angle 18-19°. Last whorl 29.3% total height (holotype). Whorls separated by narrowly impressed suture. Protoconch and first teleoconch whorls unknown. Delicate spiral sculpture present on early teleoconch whorls, disappearing on last whorls – Detailed composite description of the spiral sculpture: holotype MNHN.F.A93729 (Fig. 6A, B, H: 136.8 mm), primary cords P1-P3 present until 20th whorl, P1 close-set to the adapical suture more prominent than P2 and P3, P2 less prominent.On whorl no. 21, appearance of s2 between P2 and P3, P2 less prominent than the other primary cords. From whorl no. 22 to no. 24 (Fig. 6B), s2 becoming larger and subequal to P2, s2 and P2 less prominent that the other cords; small granules visible on cords. On whorls nos. 25-26 (Fig. 6B), P1, P2 and s2 starting to fade gradually, P3 still prominent. On whorl no. 27 (Fig. 6B), loss of spiral sculpture – Paratype MNHN.F.A93728 (Fig. 5A, H: 58.6 mm) on whorls no. 1 to no. 4, P1-P3 and s2 present; P1 prominent, close-set to adapical suture; P3 close-set to abapical suture; s2 close-set to P2 and less prominent. On whorls nos. 5-6, P1, P2 and s2 starting to fade gradually, P3 still prominent. On whorl no. 7, P1 tending to fuse with P2. On whorl 8, loss of the spiral sculpture – Paratype MNHN.F.A93727 (Fig. 5B, H: 29.2 mm), on whorls no. 1 to no. 3, P1-P3 and s2 present; P1 prominent, close-set to adapical suture; P3 close-set to abapical suture; s2 close-set to P2 and less prominent. On whorl nos. 4-5, P1, P2 and s2 starting to fade, P3 still prominent. On whorl no. 6, loss of the spiral sculpture – Paratype MNHN.F.A93725 (Fig. 5C, H: 57.1 mm), on whorls no. 1 to no. 3, P1-P3 and s2 present; P1 prominent, close-set to adapical suture; P3 prominent close-set to abapical suture; P2 and s2 weaker; small granules on P2, s2 and P3. On whorl no. 4, P1, P2 to tending to fuse and s2 starting to fade gradually, P3 still prominent. On whorl no. 5, only P3. On whorl no. 6, loss of the spiral sculpture – Paratype MNHN.F.A93726 (Fig. 5D, H: 31.9 mm), on whorls nos. 1-2, P1-P3 and s2 present; P1 close-set to adapical suture; P3 close-set to abapical suture; s2 close to P2 and less prominent. On whorl no. 3, P1 and P2 tending to fuse. On whorl no. 4, only P3. On whorl no. 5, total loss of the spiral sculpture – No axial sculpture, except some growing striae. Aperture narrow, ovoid, compressed adapically, of 88.7% of diameter and 84.33% of length of last whorl. Anal canal short and narrow. Columella weakly concave with one fold; columellar lip slightly erect; parietal lip adherent. Siphonal canal moderately short, bent, turning left and dorsally curved. Outer lip prosocyrt, not thickened. Remains of periostracum imprints (paratype MNHN.F. A93711). Residual colour pattern consisting of red stripes, approximately parallel, with inclination varying from slightly sinuous to sigmoidal (paratype MNHN.F. A98132, Fig. 6F, G).</p></div>	https://treatment.plazi.org/id/695B87E7F11B3449C4C1FD2E08A7F830	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier;Pacaud, Jean-Michel;Lashari, Rafiq A.;Brohi, Imdad A.;Solangi, Sarfraz H.	Merle, Didier, Pacaud, Jean-Michel, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H. (2026): New campanilids (Mollusca: Gastropoda) from the Ranikot Group (upper Paleocene / early Eocene, Sindh, Pakistan). Comptes Rendus Palevol 25 (1): 1-15, DOI: 10.5852/cr-palevol2026v25a1, URL: https://doi.org/10.5852/cr-palevol2026v25a1
