identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
BF0E9F26C63059398E7E4B1637104CD9.text	BF0E9F26C63059398E7E4B1637104CD9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thaiphusa mongkol Tan & Naiyanetr & Yeo 2026	<div><p>Thaiphusa mongkol sp. nov.</p><p>Figs 7, 8, 9, 10</p><p>Type material.</p><p>Holotype. Thailand • male (38.3 × 29.3 mm); Kanchanaburi Province, Thong Pha Phum District, Ban Pu Tae; Mongkol Wongkalasin leg.; 12 Aug. 1999; ZRC 2024.0331 . Paratypes. Thailand • 2 males (larger male 36.5 × 28.5 mm); same collection data as for holotype; ZRC 2024.0332 • 8 males (42.3 × 31.9, 41.7 × 32.0, 39.3 × 30.2, 37.4 × 28.9, 35.8 × 27.5, 35.5 × 28.0, 33.7 × 26.2, 33.7 × 26.2 mm); 3 females (40.8 × 31.9, 37.2 × 29.4, 34.2 × 26.6 mm); Kanchanaburi Province, Thong Pha Phum District, swamp forest, Weeyawat Jaitrong leg.; 30 May 2019; THNHM-IV-20172 • 1 male (36.8 × 28.5 mm); Kanchanaburi Province, Thong Pha Phum District; Sunchai Makchai leg.; 6 Nov. 2018; THNHM-IV-20173 .</p><p>Diagnosis.</p><p>Carapace (Fig. 7 A, B) transversely ovate, distinctly wider than long, high; dorsal surface (Fig. 7 C) strongly convex, inflated transversely, longitudinally, glabrous, smooth. Epigastric, postorbital cristae (Fig. 7 A, B) smooth, confluent; external orbital tooth outer margin concave, longer than inner margin; epibranchial tooth low, blunt, confluent with postorbital cristae; anterolateral margin rounded, subcristate; epistome posterior margin (Fig. 7 D) median tooth relatively broadly triangular, outer parts gently concave. Third maxilliped (Fig. 9 A) exopod flagellum half of merus width. Ambulatory dactyli (Fig. 7 A) elongated, slender. Male pleon (Fig. 8 B, C) narrowly triangular; pleonal somite 6 transversely rectangular, much wider than long, lateral margins sinuous. G 1 terminal article (Fig. 9 B – E) strongly bent outwards, relatively long, ~ 0.5 × length of subterminal article, tip tapered, with well-produced, broad dorsal flap for most part of terminal article, with broad gently convex apex at proximal third; subterminal article gradually narrowing from broad proximal part into slender, neck-like distal part, without cleft on outer margin, with distal part ~ 0.3 × length of proximal part.</p><p>Etymology.</p><p>The species is named after the collector of the holotype, Mr. Mongkol Wongkalasin. Used as noun in apposition.</p><p>Remarks.</p><p>Thaiphusa mongkol sp. nov., most closely resembles T. chantaburiensis in the G 1 terminal article being distinctly bent outwards at the median part as opposed to the G 1 terminal article being gently sinuous or almost straight in other Thaiphusa species. Thaiphusa mongkol sp. nov., however, can still be distinguished from T. chantaburiensis by the G 1 terminal article that is more strongly bent outwards almost to an angle of 90 ° (Fig. 9 B – E), as compared to the G 1 terminal article that is less strongly bent outwards, to only ~ 70 ° (cf. Chuensri 1973: fig. 6; Ng and Naiyanetr 1993: figs 18, 19). In addition, the proximal part of the G 1 terminal article of T. mongkol sp. nov., is without a lobed-shape bulge, and the dorsal flap is long, broad, with a gently convex apex in the proximal third (Fig. 9 D, E). This contrasts with that of T. chantaburiensis, which possesses a lobed-shaped bulge at the proximal part and a dorsal flap with a bluntly angular apex in the proximal third (cf. Chuensri 1973: fig. 6; Ng and Naiyanetr 1993: figs 18, 19). Moreover, Thaiphusa mongkol sp. nov., is currently known only from Thong Pha Phum district in Kanchanaburi Province, western Thailand, which is more than 400 km away from the locality of T. chantaburiensis, in Chanthaburi Province, eastern Thailand.</p><p>In life, Thaiphusa mongkol sp. nov., ranges in colouration, with a carapace that sports a uniformly bright yellow-orange colour (Fig. 10). This colouration is immediately different from both T. sirikit and T. reginamimus sp. nov., in the same area, which has a more distinctly patterned carapace, mainly ivory / cream in colour with a dark central patch (Fig. 6).</p><p>Distribution.</p><p>Thong Pha Phum district, Kanchanaburi province, western Thailand.</p></div>	https://treatment.plazi.org/id/BF0E9F26C63059398E7E4B1637104CD9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tan, Zhi Wan;Naiyanetr, Phaibul;Yeo, Darren C. J.	Tan, Zhi Wan, Naiyanetr, Phaibul, Yeo, Darren C. J. (2026): Six new species of terrestrial crabs (Crustacea, Decapoda, Potamidae) from Thailand. ZooKeys 1284: 49-86, DOI: 10.3897/zookeys.1284.182481
B31A9CD6AC4D5F6CB4FF1D7B94084C12.text	B31A9CD6AC4D5F6CB4FF1D7B94084C12.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thaiphusa reginamimus Tan & Naiyanetr & Yeo 2026	<div><p>Thaiphusa reginamimus sp. nov.</p><p>Figs 1, 2, 3, 5 A, C, 6</p><p>Type material.</p><p>Holotype. Thailand • male (42.0 × 29.5 mm); aquarium material purchased in Singapore by S. H. Tan on 24 Sep. 1996, originally collected in Kanchanaburi Province, Maenam Noi, near Ban Huai; Kittipong Jarutanin leg.; 1996; ZRC 2002.0109 . Paratypes. Thailand • 3 males (41.3 × 28.9, 41.0 × 29.2, 41.0 × 28.8 mm); 5 females (46.4 × 32.3–40.9 × 28.3 mm); same collection data as for holotype; ZRC 2002.0110 .</p><p>Non-type specimens.</p><p>Thailand • 1 male (38.0 × 26.6 mm); 1 female (45.8 × 31.3 mm); aquarium material purchased in Singapore by S. H. Tan on Oct. 1996; ZRC 1997.656 • 1 male (37.4 × 26.4 mm); Kanchanaburi Province, Thong Pha Phum District; Sunchai Makchai leg.; 2013; THNHM-IV-13089 • 1 female (39.0 × 28.0 mm); same collection data as for previous; THNHM-IV-13090 • 1 female (38.7 × 27.9 mm); same collection data as for previous; THNHM-IV-13091 • 1 female (34.1 × 24.2 mm); same collection data as for previous; THNHM-IV-13092 .</p><p>Diagnosis.</p><p>Carapace (Fig. 1 A, B) transversely ovate, distinctly wider than long, high; dorsal surface (Fig. 1 C) strongly convex longitudinally, distinctly inflated transversely, glabrous, smooth. Epigastric, postorbital cristae (Fig. 1 A, B) smooth, confluent; external orbital tooth outer margin distinctly longer than inner margin; epibranchial tooth low, blunt, confluent with postorbital cristae; anterolateral margin rounded, indistinct; epistome posterior margin (Fig. 1 D) median tooth relatively broadly triangular, outer parts gently concave. Third maxilliped (Fig. 3 A) exopod flagellum subequal to half of merus width. Ambulatory dactyli (Fig. 1 A) elongated, slender. Male thoracic sternite 3 / 4 groove demarcating suture prominent (Fig. 2 B – D). Male pleon (Fig. 2 B, C) narrowly triangular; pleonal somite 6 transversely rectangular, much wider than long, lateral margins convex. G 1 terminal article (Fig. 3 B – E) sinuous, relatively long, 0.52 × length of subterminal article, tip truncate, with low, broad dorsal flap, with broad, gently convex apex; subterminal article gradually narrowing from broad proximal part into slender, neck-like distal part, without forming shelf on outer margin, with distal part 0.49 × length of proximal part.</p><p>Etymology.</p><p>The species name is derived from the combination of Regina and mimus, queen and mimic in Latin, respectively, due to the similarity in morphology and colouration to Thaiphusa sirikit, which was named in honour of the late Queen and Queen Mother of Thailand. Name used as a noun in apposition.</p><p>Remarks.</p><p>Thaiphusa reginamimus sp. nov., is closest in external morphology to T. sirikit, which also occurs in Sai Yok and Thong Pha Phum districts, Kanchanaburi Province. In addition, its striking purple and white dorsal carapace colours also closely resemble those of T. sirikit . Thaiphusa reginamimus sp. nov., however, is distinguishable from T. sirikit by the following morphological characters: male thoracic sternite 3 / 4 groove demarcating suture prominent (Fig. 2 A, C) (vs male thoracic sternite 3 / 4 groove demarcating suture not distinct, Fig. 4 E; cf. Ng and Naiyanetr 1993: fig. 22 C); G 1 terminal article proportionally shorter, ~ 0.5 × length of subterminal article (Figs 3 B, 3 C, 5 A, 5 C) (vs G 1 terminal article proportionally longer, ~ 0.6 × length of subterminal article, Fig. 5 B, D; cf. Ng and Naiyanetr 1993: fig. 56 B, C); G 1 terminal article dorsal flap relatively higher and more strongly convex, appearing relatively shorter longitudinally (Figs 3 D, 3 E, 5 A, 5 C) (vs G 1 terminal article dorsal flap lower and more gently convex, appearing relatively longer longitudinally, Fig. 5 B, D; cf. Ng and Naiyanetr 1993: fig. 56 D, E); and G 1 subterminal article with narrowed, neck-like distal part subequal to or shorter than expanded proximal part (Figs 3 B, 3 C, 5 A, 5 C) (vs G 1 subterminal article with narrowed, neck-like distal part longer than expanded proximal part, Fig. 5 B, D; cf. Ng and Naiyanetr 1993: fig. 56 B – E). In addition, the live colouration of T. reginamimus sp. nov., varies and differs slightly from T. sirikit in that the carapace can exhibit a comparatively smaller, less defined dark patch, with the ambulatory legs that are paler in colouration, ranging from light orange to pale yellow (Fig. 6 A – D) (vs a carapace that has very strongly defined black and ivory coloured regions, with brightly coloured, ambulatory legs, usually red, sometimes orange in colour; Fig. 6 E; cf. Naiyanetr 1992: pl. 1).</p><p>Distribution.</p><p>Sai Yok and Thong Pha Phum districts, Kanchanaburi province, western Thailand.</p><p>Comparative material.</p><p>Thaiphusa sirikit (Naiyanetr, 1992) paratype. Thailand • male (46.7 × 32.2 mm); Kanchanaburi Province, Sai Yok District, Ban Nam Chon; P. Naiyanetr leg.; Nov. 1987; ZRC 1991.1880.</p></div>	https://treatment.plazi.org/id/B31A9CD6AC4D5F6CB4FF1D7B94084C12	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tan, Zhi Wan;Naiyanetr, Phaibul;Yeo, Darren C. J.	Tan, Zhi Wan, Naiyanetr, Phaibul, Yeo, Darren C. J. (2026): Six new species of terrestrial crabs (Crustacea, Decapoda, Potamidae) from Thailand. ZooKeys 1284: 49-86, DOI: 10.3897/zookeys.1284.182481
49134F33CB29554A93E9D712007E6298.text	49134F33CB29554A93E9D712007E6298.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thaipotamon nandidarbhai Tan & Naiyanetr & Yeo 2026	<div><p>Thaipotamon nandidarbhai sp. nov.</p><p>Figs 11, 12, 13</p><p>Thaipotamon nandidarbhi Naiyanetr, in Nandidarbha and Nandidarbha 2001 [nomen nudum].</p><p>Type material.</p><p>Holotype. Thailand • male (38.7 × 29.6 mm); Phrae Province, Sung Men District; Sermsak Nandidarbha leg.; Oct. 1997; ZRC 2024.0333 .</p><p>Diagnosis.</p><p>Carapace (Fig. 11 A, B) transversely sub-ovate, wider than long, CW / CL ratio 1.30, high; dorsal surface (Fig. 11 C) convex longitudinally, weakly convex transversely. Epigastric cristae (Fig. 11 A, B) distinct, smooth, not cristate, separated by broad, median Y-shaped furrow; epigastric cristae just anterior of postorbital cristae, separated by short weak, furrow, very weakly sloping posterolaterally; postorbital cristae distinct, margin entire, smooth, appearing strongly concave from dorsal view, outer edge with few small granules, confluent with anterolateral margins. External orbital tooth (Fig. 11 A, B) distinct, broadly triangular, outer margin straight, equal to length of inner margin, demarcated from rest of anterolateral margin by V-shaped cleft; epibranchial tooth produced, visible from dorsal view, not sharp. Anterolateral margins (Fig. 11 A, B) convex, subcristate, smooth. Frontal, orbital, anterolateral, branchial, mesogastric, urogastric, cardiac, intestinal regions smooth; suborbital region (Fig. 11 C) smooth with few small granules; pterygostomial regions granulose at lateral parts; subhepatic, sub-branchial regions granulose. Male pleonal somite 6 lateral margins gently sinuous in the holotype (Fig. 12 B, C). G 1 (Fig. 13 B – E) slender, sinuous; terminal article 0.50 × length of subterminal article, strongly curving outwards, with high and broad dorsal flap on proximal half, apex medial in position, with distal margin abruptly tapered, almost perpendicularly, flap 0.62 × length of terminal article (from ventral view), slopes gradually towards tip from dorsal view; terminal, subterminal articles separated by weak dilation. G 2 (Fig. 13 F) long, with distal article 0.75 × length of basal article.</p><p>Etymology.</p><p>This species is named after its collector and discoverer, Associate Professor Sermsak Nandidarbha, formerly of Chiang Mai Rajabhat University.</p><p>Remarks.</p><p>The species had been referred to several times under the name; “ Thaipotamon nandidarbhi ” in local Thai social media platforms and forums. In certain instances, the name was used in conjunction with the authority “ Naiyanetr, 1998 ” (gotoknow. org 2006). The usage of “ Thaipotamon nandidarbhi Naiyanetr, 1998 ” could very likely be attributed to several local Thai newspaper outlets, which reported on the discovery of a new freshwater crab and included the name “ Thaipotamon nandidarbhi ” and with Professor Phaibul Naiyanetr being mentioned as the authority, identifying it as a new species (Daily News 1998; ThaiNews 1998; Thairath 1998). While brief descriptions of the new species were provided in the news articles, no comparisons were made to any other species (see news articles attached in and translated from Nandidarbha and Nandidarbha 2001: 123–129). Provisions of Article 13.1. 1 of the ICZN (1999) Code requires that any designation of new species after 1930; “ be accompanied by a description or definition that states in words characters that are purported to differentiate the taxon ”. As such the name “ Thaipotamon nandidarbhi Naiyanetr, 1998 ” is not available for nomenclatural purposes.</p><p>There is a scientific paper by Nandidarbha and Nandidarbha (2001), published by the National Science and Technology Development Agency (NSTDA) of Thailand, that also contains “ Thaipotamon nandidarbhi ”. The article was mainly written in Thai and available as hard copies at the time of publishing. It was only recently digitised, with open access. In the introduction to this report, it was mentioned that there exists a crab species from Soong Men (Sung Men) District of Phrae Province, locally called “ Kam ”, meaning dark or deep, with brief descriptions and comparisons made to the common rice field crabs in the area with regards to the size and colouration of the carapace. Within appendix A of the report, there contains a letter from Professor Phaibul Naiyanetr regarding the taxonomy of the crab from Soong Men as a new species, and that the crab will be called “ Thaipotamon nandidarbi Naiyanetr ” (Nandidarbha and Nandidarbha 2001: 119). Thereafter in appendix B, the article again states that; “ This is a new species of freshwater crab found in Thailand, one that has never been reported or entered into the global taxonomy of freshwater crabs ” and that “ This discovery is the result of research by Associate Professor Sermsak Nandidarbha, a lecturer in the Department of Biology, Faculty of Science and Technology, Chiang Mai Rajabhat University ”. According to the report, a sample of the crab was sent to a crab expert in Thailand, Professor Phaibul Naiyanetr of the Department of Biology, Faculty of Science, Chulalongkorn University, for joint examination with experts from the Netherlands. The initial examination revealed it to be a new species of freshwater crab. Professor Phaibul Naiyanetr then sent the sample to the Netherlands, with detailed documentation for publication in Crustaceana, the International Journal of Crustacean Research. The species was given the name “ Thaipotamon nandidarbhi Naiyanetr. ” (translated from Nandidarbha and Nandidarbha, 2001: 121). The name “ Thaipotamon nandidarbhi ”, however, has never been formally published as intended; the paper supposedly sent to Crustaceana reported in Nandidarbha and Nandidarbha (2001) has never been published, nor was the name published in any other journal. While the name “ Thaipotamon nandidarbhi Naiyanetr, in Nandidarbha and Nandidarbha (2001) ” was used in conjunction with some form of description and comparisons, no types were actually designated and the repository of the specimens is unknown. As a result, Article 16.4. 1 of the ICZN (1999) Code, which requires a clear designation of a holotype or syntypes for the nominal taxon; and Article 16.4. 2, which requires an explicit statement of intent that they will be (or are) deposited in a collection, with the name and location of that collection clearly indicated, are both not fulfilled. The name “ Thaipotamon nandidarbhi Naiyanetr, in Nandidarbha and Nandidarbha, 2001 ” is thus also not available for nomenclature purposes. Although two spellings are used for the species in the report: Thaipotamon nandidarbi and Thaipotamon nandidarbhi, the spellings are likely a typographical error and here, we follow the report author’s actual name: Sermsak Nandidarbha. Therefore, the name is validated here, as Thaipotamon nandidarbhai sp. nov., with the necessary diagnosis and the designation of the holotype.</p><p>The distinctly hooked G 1 terminal article of Thaipotamon nandidarbhai sp. nov., shows similar degree of curving to that in the G 1 terminal articles of T. chulabhorn, T. varoonphornae, and T. wangsaphung sp. nov. (see later), with the distal end being perpendicular or almost perpendicular to the longitudinal axis (Figs 13 B – E, 24 B – E; cf. Naiyanetr 1993: fig. 1 D; Ng and Naiyanetr, 1993: fig. 54 B – F). This distinguishes the new species from the rest of its congeners, in which the G 1 terminal article is much less strongly curved outwards, with the tip terminating at an angle not perpendicular or almost perpendicular to the longitudinal axis (~ 30–70 °). Thaipotamon nandidarbhai sp. nov., is externally separated from T. chulabhorn, T. varoonphornae, and T. wangsaphung sp. nov., by the following differences: carapace appearing less strongly inflated (Fig. 11 C) (vs carapace appearing more strongly inflated, Fig. 21 C; cf. Naiyanetr 1993: fig. 1 A; Ng and Naiyanetr 1993: fig. 20 A); granulose sub-orbital, pterygostomial, sub-branchial regions (Fig. 11 C) (vs regions smooth or with very few, very faint granules, Fig. 21 C; cf. Naiyanetr 1993: fig. 1 A; Ng and Naiyanetr 1993: fig. 20 B); and male pleon somite 6 lateral margins gently sinuous (Fig. 12 C) (vs male pleon somite 6 lateral margins distinctly convex in T. varoonphornae and T. wangsaphung sp. nov., Fig. 22 C; cf. Ng and Naiyanetr 1993: fig. 20 C). It should be noted, however, that the differences in the male pleon somite 6 lateral margins do vary slightly across among individuals. Nevertheless, Thaipotamon nandidarbhai sp. nov., can be further separated from T. varoonphornae by these additional differences: G 1 terminal article relatively longer, 0.50 × length of subterminal article (Fig. 13 B – E) (vs G 1 terminal article relatively shorter, 0.46 × length of subterminal article; cf. Ng and Naiyanetr 1993: fig. 54 B – F); dorsal flap relatively broader, 0.62 × length of terminal article (Fig. 13 B – E) (vs dorsal flap relatively narrower, 0.57 × length of terminal article; cf. Ng and Naiyanetr 1993: 54 B – F); and dorsal flap sloping gradually towards tip of terminal article (Fig. 13 B – E) (vs dorsal flap not reaching tip of terminal article; cf. Ng and Naiyanetr 1993: 54 B – F). Additionally, Thaipotamon nandidarbhai sp. nov., can be separated from T. chulabhorn and T. wangsaphung sp. nov., by the following differences: G 1 terminal article relatively shorter, 0.50 × length of subterminal article (Fig. 13 B – E) (vs relatively longer, 0.53 × length of subterminal article in T. wangsaphung sp. nov., and reaching 0.60 × length in T. chulabhorn, Fig. 23 B – E; cf. Naiyanetr 1993: fig. 1 D); and G 1 terminal article dorsal flap distal margin more abruptly tapered, flap appearing symmetrical and semi-circular (Fig. 13 B – E) (vs dorsal flap distal margin gradually tapered, appearing less symmetrical, Fig. 23 B – E; cf. Naiyanetr 1993: fig. 1 D).</p><p>Distribution.</p><p>Phrae Province, northeastern Thailand.</p></div>	https://treatment.plazi.org/id/49134F33CB29554A93E9D712007E6298	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tan, Zhi Wan;Naiyanetr, Phaibul;Yeo, Darren C. J.	Tan, Zhi Wan, Naiyanetr, Phaibul, Yeo, Darren C. J. (2026): Six new species of terrestrial crabs (Crustacea, Decapoda, Potamidae) from Thailand. ZooKeys 1284: 49-86, DOI: 10.3897/zookeys.1284.182481
365E261700445FA69DB4FD760170BDD7.text	365E261700445FA69DB4FD760170BDD7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thaipotamon songkhwae Tan & Naiyanetr & Yeo 2026	<div><p>Thaipotamon songkhwae sp. nov.</p><p>Figs 14, 15, 16</p><p>Thaipotamon phitsanulok Kumjan &amp; Chamchoi, 2018 [nomen nudum].</p><p>Type material.</p><p>Holotype. Thailand • male (58.0 × 42.0 mm); Phitsanulok Province, Wat Bot District, Ban Yao; Chalong Nuichim leg.; Oct. 1997; ZRC 2024.0334 . Paratype. Thailand • 1 male (49.6 × 36.5 mm); same collection data as for holotype; ZRC 2024.0335 .</p><p>Diagnosis.</p><p>Carapace (Fig. 14 A, B) transversely sub-ovate, wider than long, CW / CL ratio 1.35–1.38, high; dorsal surface (Fig. 14 C) convex longitudinally, appearing concave transversely due to inflated branchial regions. Epigastric cristae distinct, smooth, not cristate, very weakly separated by broad, median furrow, appearing almost straight; epigastric cristae just anterior of postorbital cristae, very weakly separated by furrow; postorbital cristae distinct, margin entire, smooth, appearing concave from dorsal view, outer edge smooth, confluent with anterolateral margins. External orbital tooth (Fig. 14 A, B) distinct, broadly triangular, outer margin convex, equal to length of inner margin, demarcated from rest of anterolateral margin by broad cleft; epibranchial tooth very weakly produced, not clearly visible from dorsal view. Anterolateral margins (Fig. 14 A, B) convex, subcristate, smooth. Frontal, orbital, anterolateral, branchial, mesogastric, urogastric, cardiac, intestinal, suborbital, pterygostomial regions (Fig. 14 B, C) smooth; subhepatic, sub-branchial (Fig. 14 C) regions weakly covered with striae. Male pleonal somite 6 lateral margins convex (Fig. 15 C). G 1 (Fig. 16 B – E) broad, gently sinuous; subterminal article broad with cleft on outer margin at distal end; terminal article 0.58 × length of subterminal article, very gently curving outwards, with high, broad dorsal flap on proximal half, apex slightly skewed towards proximal portion, with relatively gradually tapered distal margin, flap 0.60 × length of terminal article (from ventral view), slopes gradually towards tip from dorsal view; terminal article separated from subterminal article by weak dilation, weakly produced collar. G 2 (Fig. 16 F) long, with distal article 0.63 × length of basal article.</p><p>Etymology.</p><p>The species is named after its local name, “ Pu Song Khwae ”, “ Pu ” meaning crab and “ Song Khwae ” being the local name for the type locality, Phitsanulok Province, in local Thai dialect. The name is used as a noun in apposition.</p><p>Remarks.</p><p>The new species, Thaipotamon songkhwae has been referred to frequently in various social media / forum posts, as well as in an ecotourism management paper, published in Thai (Kumjan and Chamchoi 2018), as “ Thaipotamon phitsanulok ”. The two names are most definitely referring to the same species, since all the material and accounts are from the same locality in Wat Bot District, Phitsanulok province. “ Thaipotamon phitsanulok Kumjan &amp; Chamchoi, 2018 ”, however, is not an available name. Provisions of Article 13.1. 1 of the ICZN (1999) Code require that any designation of new species; “ be accompanied by a description or definition that states in words characters that are purported to differentiate the taxon ”, and Article 16.1 of the ICZN (1999) Code requires the names to “ be explicitly indicated as intentionally new ”. Since no diagnosis or comparisons were provided nor was the name explicitly stated as new, the conditions for both articles were not met. Furthermore, “ Thaipotamon phitsanulok Kumjan &amp; Chamchoi, 2018 ” also did not fulfil Articles 16.4. 1 and 16.4. 2 of the ICZN (1999) Code, which respectively require a clear designation of a holotype or syntypes for the nominal taxon; and an explicit statement of intent that they will be (or are) deposited in a collection, with the name and location of that collection indicated. As the conditions for all articles listed above are not fulfilled, the name “ Thaipotamon phitsanulok ” is not available for nomenclatural purposes. The Thaipotamon species from Wat Bot District, Phitsanulok Province is validated herein and described as Thaipotamon songkhwae sp. nov.</p><p>Thaipotamon songkhwae sp. nov., together with T. suvankorni sp. nov. (see below), differs from all other congeners by the G 1 terminal article dorsal flap having a relatively more gradually tapered distal margin as compared to the distal margin of G 1 terminal article dorsal flap of the congeners, which is more abruptly tapered and the dorsal flap apex being skewed towards the proximal portion rather than being at the median portion (Figs 13 B – E, 16 B – E, 19 B – E, 23 B – E; cf. Ng and Naiyanetr 1993: figs 51–54). Other than the different shape of G 1 terminal article dorsal flap, T. songkhwae sp. nov., can be further differentiated from T. siamense, which has a similarly gently curving G 1 terminal article in the following differences: G 1 terminal article relatively longer, 0.58 × length of subterminal article (Fig. 13 B – E) (vs G 1 terminal article relatively shorter, 0.52 × length of subterminal article; cf. Ng and Naiyanetr 1993: figs 18 A, 19 A); and dorsal flap relatively broader, 0.60 × length of terminal article (Fig. 13 B – E) (vs dorsal flap relatively narrower, 0.58 × length of terminal article; cf. Ng and Naiyanetr 1993: figs 18 A, 19 A).</p><p>Thaipotamon songkhwae sp. nov., most closely resembles T. suvankorni sp. nov., in terms of G 1 morphology. However, Thaipotamon songkhwae sp. nov., can be distinguished from T. suvankorni sp. nov., by the following differences: carapace relatively more transverse, ~ 1.35–1.38 × broader than long (Fig. 14 A, B) (vs carapace relatively less transverse, ~ 1.20–1.30 × broader than long, Fig. 17 A, B); external orbital tooth relatively more broadly triangular (Fig. 14 B) (vs external orbital tooth relatively more acutely triangular, Figs 17 B, 20 A); cervical grooves and grooves between epibranchial and postorbital cristae indistinct (Fig. 14 A, B) (vs cervical grooves and grooves between epibranchial and postorbital cristae shallow but distinct, Figs 17 B, 20 A); G 1 subterminal article with cleft on outer margin at distal end (Fig. 16 B) (vs G 1 subterminal article without cleft on outer margin, Fig. 19 B); G 1 terminal article relatively longer, 0.58 × length of subterminal article (Fig. 16 B – E) (vs G 1 terminal article relatively shorter, 0.54 × length of subterminal article, Fig. 19 B – E); G 1 terminal article relatively less curved outwards (Fig. 16 B – E) (vs G 1 terminal article relatively more strongly curved outwards; Fig. 19 B – E); G 1 terminal article tapers to narrow, pointed end (Fig. 16 B – E) (vs G 1 terminal article tapers to broader, rounded end, Fig. 19 B – E); G 1 terminal article dorsal flap relatively broader, 0.60 × length of terminal article (Fig. 16 D) (vs G 1 terminal article dorsal flap relatively narrower, 0.57 × length of terminal article, Fig. 19 D); and G 1 terminal article dorsal flap extends all the way to tip of terminal article (Fig. 16 D) (vs G 1 terminal article dorsal flap not reaching tip of terminal article, Fig. 19 D).</p><p>Distribution.</p><p>Phitsanulok Province, northern Thailand.</p></div>	https://treatment.plazi.org/id/365E261700445FA69DB4FD760170BDD7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tan, Zhi Wan;Naiyanetr, Phaibul;Yeo, Darren C. J.	Tan, Zhi Wan, Naiyanetr, Phaibul, Yeo, Darren C. J. (2026): Six new species of terrestrial crabs (Crustacea, Decapoda, Potamidae) from Thailand. ZooKeys 1284: 49-86, DOI: 10.3897/zookeys.1284.182481
CD3017566AC4581898583D1BA0FB9A5D.text	CD3017566AC4581898583D1BA0FB9A5D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thaipotamon suvankorni Tan & Naiyanetr & Yeo 2026	<div><p>Thaipotamon suvankorni sp. nov.</p><p>Figs 17, 18, 19, 20</p><p>Type material.</p><p>Holotype. Thailand • male (45.5 × 35.0 mm); Chachoengsao Province, Tha Ta Kiep Wildlife Sanctuary; Ubonwan Booncharm leg.; 26 May 1993; ZRC 2024.0336 . Paratypes. Thailand • 1 male (38.7 × 29.9 mm); 2 females (41.3 × 32.3, 36.4 × 30.4 mm); same collection data as for holotype; ZRC 2024.0337 .</p><p>Diagnosis.</p><p>Carapace (Figs 17 A, 17 B, 20 A) transversely sub-ovate, wider than long, CW / CL ratio 1.20–1.30, high; dorsal surface (Fig. 17 C) convex transversely and longitudinally. Epigastric cristae (Fig. 17 C) distinct, smooth, not cristate, separated by broad, median Y-shaped furrow, gently sloping posterolaterally; epigastric cristae just anterior of postorbital cristae, separated by short weak, furrow; postorbital cristae distinct, margin entire, smooth, appearing weakly concave from dorsal view, outer edge smooth, confluent with anterolateral margins. External orbital tooth (Fig. 17 B) distinct, narrowly triangular, outer margin straight, equal to length of inner margin, demarcated from rest of anterolateral margin by V-shaped cleft; epibranchial tooth (Fig. 17 B) small, visible from dorsal view as nodule. Anterolateral margins convex, not cristate, smooth. Frontal, orbital, anterolateral, branchial, mesogastric, urogastric, cardiac, intestinal, suborbital, pterygostomial regions smooth; subhepatic, sub-branchial regions (Fig. 17 C) covered with weak striae. Male pleonal somite 6 lateral margins convex (Fig. 18 C). G 1 (Fig. 19 B – E) slender, sinuous; terminal article 0.54 × length of subterminal article, gently curving outwards, with high and broad dorsal flap on proximal half, apex slightly skewed towards proximal portion, with relatively gradually tapered distal margin, flap 0.57 × length of terminal article (from ventral view), not reaching tip from dorsal view; terminal article not distinctly separated from subterminal article by dilation. G 2 (Fig. 19 F) long, with distal article 0.65 × length of basal article. Female vulvae (Fig. 20 C) large, oval, recessed, opening directed mesially, located at mesial end of sternite 6, approximtely half the length of sternite 6, with short, calcified eave and soft membranous flap-like protrusion.</p><p>Etymology.</p><p>This species is named in honour of Mr. Phairot Suvankorn, former director of the Department of Royal Forestry, Ministry of Agriculture, Thailand.</p><p>Remarks.</p><p>The general G 1 structure of Thaipotamon suvankorni sp. nov., resembles that of T. songkhwae sp. nov. They can nevertheless be separated by other G 1 characters, as well as some carapace characters (see earlier remarks for Thaipotamon songkhwae sp. nov.). Thaipotamon suvankorni sp. nov., is also similar to T. siamense in the relatively weakly curved G 1 terminal article; however, T. suvankorni sp. nov., still clearly differs from T. siamense by the following differences: G 1 terminal article relatively longer, 0.54 × length of subterminal article (Fig. 19 B – E) (vs G 1 terminal article relatively shorter, 0.52 × length of subterminal article; cf. Ng and Naiyanetr 1993: figs 18 A, 19 A); and G 1 terminal article dorsal flap apex appears slightly skewed towards proximal portion, with relatively gradually tapered distal margin (Fig. 19 D) (vs G 1 terminal article dorsal flap apex medial, with abruptly tapered distal margin; cf. Ng and Naiyanetr 1993: fig. 19 A). Since female specimens are available for this species, vulvae (Fig. 20 C) characters, which are potentially diagnostic, have also been included in the Diagnosis for future taxonomic reference.</p><p>Distribution.</p><p>Chachoengsao Province, eastern Thailand.</p></div>	https://treatment.plazi.org/id/CD3017566AC4581898583D1BA0FB9A5D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tan, Zhi Wan;Naiyanetr, Phaibul;Yeo, Darren C. J.	Tan, Zhi Wan, Naiyanetr, Phaibul, Yeo, Darren C. J. (2026): Six new species of terrestrial crabs (Crustacea, Decapoda, Potamidae) from Thailand. ZooKeys 1284: 49-86, DOI: 10.3897/zookeys.1284.182481
7413DBF938DF5888885FE85507407428.text	7413DBF938DF5888885FE85507407428.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thaipotamon wangsaphung Tan & Naiyanetr & Yeo 2026	<div><p>Thaipotamon wangsaphung sp. nov.</p><p>Figs 21, 22, 23, 24</p><p>Type material.</p><p>Holotype. Thailand • male (40.7 × 28.7 mm); Loei Province, Wang Saphung District, Ban Na Luang, Phu Luang; P. Naiyanetr leg.; 14 Oct. 1998; ZRC 2024.0338 . Paratypes. Thailand • 1 male (36.6 × 27.0 mm); 1 female (33.0 × 24.9 mm); same collection data as for holotype; ZRC 2024.0339 .</p><p>Diagnosis.</p><p>Carapace (Figs 21 A, 21 B, 24 A) transversely sub-ovate, wider than long, CW / CL ratio 1.33–1.37, high; dorsal surface (Fig. 21 C) convex longitudinally and transversely. Epigastric cristae distinct, smooth, not cristate, separated by broad, median Y-shaped furrow, sloping posterolaterally strongly; epigastric cristae anterior of postorbital cristae, barely separated by furrow; postorbital cristae distinct, margin entire, smooth, appearing concave from dorsal view, outer edge smooth, confluent with anterolateral margins. External orbital tooth (Fig. 21 B) distinct, narrowly triangular, outer margin straight, subequal to length of inner margin, demarcated from rest of anterolateral margin by V-shaped cleft; epibranchial tooth (Fig. 21 B) weakly produced, visible from dorsal view as nodule. Anterolateral margins convex, smooth. Frontal, orbital, anterolateral, branchial, mesogastric, urogastric, cardiac, intestinal, suborbital, pterygostomial regions smooth; subhepatic, sub-branchial regions (Fig. 21 C) weakly granulose. Male pleonal somite 6 lateral margins convex (Fig. 22 C). G 1 (Fig. 23 B – E) slender, sinuous; terminal article 0.53 × length of subterminal article, strongly curving outwards, with high and broad dorsal flap on proximal half, apex medial in position, with proximal margin abruptly tapered, almost perpendicularly, with relatively gradually tapered distal margin, flap 0.57 × length of terminal article (from ventral view), reaching tip from dorsal view; terminal article not distinctly separated from subterminal article by dilation. G 2 (Fig. 23 F) long, with distal article 0.61 × length of basal article. Female vulvae (Fig. 24 C) large, semi-circular, recessed, opening directed mesially, located at mesial end of sternite 6, more than half length of sternite 6, with calcified eave and large, soft membranous flap-like protrusion, obscuring opening.</p><p>Etymology.</p><p>The species name is named after the locality it was collected from, Wang Saphung District in Loei Province, northeastern Thailand. The name is used as a noun in apposition.</p><p>Remarks.</p><p>Thaipotamon wangsaphung sp. nov., most closely resembles T. holthuisi Naiyanetr &amp; Yeo, 2010, in its G 1 terminal article, which features a high, broad dorsal flap on proximal half, with apex in median position, distal portion bent outwards (Fig. 23 B – E; cf. Naiyanetr and Yeo 2010: fig. 2). Thaipotamon wangsaphung sp. nov., can nevertheless be distinguished from T. holthuisi by the following characters: epigastric cristae separated from postorbital cristae by shallow grooves (Figs 21 B, 24 A) (vs epigastric cristae appearing to be confluent with postorbital cristae; cf. Naiyanetr and Yeo 2010: fig. 1 A); epigastric cristae sloping posterolaterally in dorsal view (Figs 21 B, 24 A) (vs epigastric cristae parallel to frontal margin in dorsal view; cf. Naiyanetr and Yeo 2010: fig. 1 A); postorbital cristae concave in dorsal view (Figs 21 B, 24 A) (vs postorbital cristae subparallel to frontal margin in dorsal view; cf. Naiyanetr and Yeo 2010: fig. 1 A); G 1 terminal article relatively longer, 0.53 × length of subterminal article (Fig. 23 B – E) (vs G 1 terminal article relatively shorter, 0.51 × length of subterminal article; cf. Naiyanetr and Yeo 2010: fig. 2); and G 1 terminal article dorsal flap distal margin gently tapering with a long, broad, gently convex apex (Fig. 23 D) (vs G 1 terminal article dorsal flap distal margin abruptly tapered, almost perpendicularly; cf. Naiyanetr and Yeo 2010: fig. 2 C). Since a female specimen is available for this species, vulvae (Fig. 24 C) characters, which are potentially diagnostic, have also been included in the Diagnosis for future taxonomic reference.</p><p>Distribution.</p><p>Loei Province, northeastern Thailand.</p></div>	https://treatment.plazi.org/id/7413DBF938DF5888885FE85507407428	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tan, Zhi Wan;Naiyanetr, Phaibul;Yeo, Darren C. J.	Tan, Zhi Wan, Naiyanetr, Phaibul, Yeo, Darren C. J. (2026): Six new species of terrestrial crabs (Crustacea, Decapoda, Potamidae) from Thailand. ZooKeys 1284: 49-86, DOI: 10.3897/zookeys.1284.182481
