taxonID	type	description	language	source
6B5A7E5FCC56FFE3FF1E4C49FCDCF983.taxon	type_taxon	Type species. Sciara elegans Winnertz, 1867 — Monogr. Sciarinen: 27 – 28 (by original designation) [= Leptosciarella (Leptosciarella) scutellata (Staeger, 1840)].	en	Hebert, Florence, Muller, Valerie Ngo, Nel, Andre (2025): The oldest representative of sciarid genus Leptosciarella (Diptera) from the lowermost Eocene amber of Oise (France). Zootaxa 5739 (1): 42-52, DOI: 10.11646/zootaxa.5739.1.8, URL: https://doi.org/10.11646/zootaxa.5739.1.8
6B5A7E5FCC56FFE3FF1E4C49FCDCF983.taxon	discussion	Literature: Tuomikoski (1960: 17, 20) [as Trichosia (Leptosciarella); key and original description as subgenus]; Evenhuis (1994: 176) [as Trichosia (Leptosciarella); catalogue of fossil species]; Röschmann & Mohrig (1994: 127, 130) [as Trichosia (Leptosciarella); checklist of species in Baltic amber]; Mohrig & Röschmann (1994 84) [as Trichosia (Leptosciarella); revision of species in Baltic amber]; Röschmann & Mohrig (1995 a): 22, 23 [as Trichosia and Trichosia (Leptosciarella); revision of species in Saxonian amber]; Röschmann and Mohrig (1995 b: 162) [as Trichosia (Leptosciarella); additional records in Saxonian amber]; Menzel & Mohrig (1997: 4, 5) [as Leptosciarella; key and genus rank of Leptosciarella]; Mohrig and Menzel (1997: 44) [as Leptosciarella; key and revision of Palaearctic subgenera and species]; Menzel & Mohrig (2000: 88, 353) [as Leptosciarella; key, genus diagnosis and redescription]; Shin et al. (2013: 840, 842); Shin et al. (2019: 134); Vilkamaa et al. (2023: 13) [all as Leptosciarella; suprageneric classification of Leptosciarella].	en	Hebert, Florence, Muller, Valerie Ngo, Nel, Andre (2025): The oldest representative of sciarid genus Leptosciarella (Diptera) from the lowermost Eocene amber of Oise (France). Zootaxa 5739 (1): 42-52, DOI: 10.11646/zootaxa.5739.1.8, URL: https://doi.org/10.11646/zootaxa.5739.1.8
6B5A7E5FCC56FFE6FF1E4E89FE0AFE5B.taxon	description	urn: lsid: zoobank. org: act: A 73 B 561 A-F 94 F- 48 E 3 - 87 A 9 - 55176 E 63 B 3 A 2 Figures 1 – 7	en	Hebert, Florence, Muller, Valerie Ngo, Nel, Andre (2025): The oldest representative of sciarid genus Leptosciarella (Diptera) from the lowermost Eocene amber of Oise (France). Zootaxa 5739 (1): 42-52, DOI: 10.11646/zootaxa.5739.1.8, URL: https://doi.org/10.11646/zootaxa.5739.1.8
6B5A7E5FCC56FFE6FF1E4E89FE0AFE5B.taxon	materials_examined	Holotype. MNHN-F. A 97855 (male, well preserved), stored in the Palaeontological Collection, Muséum national d’Histoire naturelle, Paris.	en	Hebert, Florence, Muller, Valerie Ngo, Nel, Andre (2025): The oldest representative of sciarid genus Leptosciarella (Diptera) from the lowermost Eocene amber of Oise (France). Zootaxa 5739 (1): 42-52, DOI: 10.11646/zootaxa.5739.1.8, URL: https://doi.org/10.11646/zootaxa.5739.1.8
6B5A7E5FCC56FFE6FF1E4E89FE0AFE5B.taxon	etymology	Etymology. The species name ‘ menzeli’ is dedicated to Dr Frank Menzel, in recognition to his great help in producing this paper and his impressive work on the Sciaridae.	en	Hebert, Florence, Muller, Valerie Ngo, Nel, Andre (2025): The oldest representative of sciarid genus Leptosciarella (Diptera) from the lowermost Eocene amber of Oise (France). Zootaxa 5739 (1): 42-52, DOI: 10.11646/zootaxa.5739.1.8, URL: https://doi.org/10.11646/zootaxa.5739.1.8
6B5A7E5FCC56FFE6FF1E4E89FE0AFE5B.taxon	diagnosis	Diagnosis. Male characters only. Eyes dichoptic with a bridge; gonocoxite distinctly longer than wide; gonostylus shorter than gonocoxite, three times longer than wide; without prominent edge in middle of inner side; gonostylus tip straight, with a combination of curved, slender setae and very sharp longer setae, with a beak-shaped terminal tooth (megaseta sensu Hippa & Vilkamaa 1991: fig. 4) at inner angle; inner surface of stylus under tooth with awl-shaped setae.	en	Hebert, Florence, Muller, Valerie Ngo, Nel, Andre (2025): The oldest representative of sciarid genus Leptosciarella (Diptera) from the lowermost Eocene amber of Oise (France). Zootaxa 5739 (1): 42-52, DOI: 10.11646/zootaxa.5739.1.8, URL: https://doi.org/10.11646/zootaxa.5739.1.8
6B5A7E5FCC56FFE6FF1E4E89FE0AFE5B.taxon	materials_examined	Locality and horizon. The locality of amber origin (deposit of Oise) is located in the départment de l’Oise, région Hauts-de-France, France; Lowermost Eocene (Ypresian) (Nel & Brasero, 2010).	en	Hebert, Florence, Muller, Valerie Ngo, Nel, Andre (2025): The oldest representative of sciarid genus Leptosciarella (Diptera) from the lowermost Eocene amber of Oise (France). Zootaxa 5739 (1): 42-52, DOI: 10.11646/zootaxa.5739.1.8, URL: https://doi.org/10.11646/zootaxa.5739.1.8
6B5A7E5FCC56FFE6FF1E4E89FE0AFE5B.taxon	description	Description. Male. Head capsule roundish. Eye dichoptic, with closed eye bridge three facets wide. All three ocelli well developed. Antennae light brown, without any contrast between scape, pedicel and flagellum; pedicel roundish and slightly wider than high, not cylindrically elongated; 11 flagellomeres, first to seventh basal flagellomeres 5.3 times longer than wide, more distal ones 4.5 times longer than wide; necks of flagellomeres naked, wider than high, with an edge between basal portion and neck; surface of flagellomeres rough with deep pits, sensilla of two different lengths; setae on flagellomeres as long as flagellomere widths. Maxillary palp pale with three palpomeres, first and third palpomeres slender, second palpomere oval thickened; third palpomere very long, 1.6 times as long as the first and 2.7 times as long as the second palpomere; palpal sensilla on the first segment simple, without noticeable margin or depression; first palpomere with two bristles; second palpomere with two bristles, third with a longitudinal row of setae of unequal lengths and strong bristles at apex. Prefrons and clypeus slightly bulging. Thorax flatly arched, pale brown. Posterior pronotum apparently bare. Anterior pronotum with at least two setae. Mesonotum with setae in lateral, dorsocentral, and acrostichal rows with bare spaces between, but with very small setae in anterior part. Scutellum with four robust setae and few smaller and weaker setae. Anepimeron not expanded ventrally. Katepisternum triangular, as long as high, and without setae. Mesothoracic epimeron with its anterior margin ending at anterodorsal margin of episternum 3. Wing narrow, well developed, 1.6 mm long; length / width ratio 3.0. ‘ C’ (length of part of C distal to apex of R 4 + 5) = 0.75 x ‘ W’ (distance between apex of R 4 + 5 and wing apex). Wing membrane transparent and with numerous trichia, all posterior veins distinct; r-m and bm-m equal in length, without macrotrichia; Rs very short and perpendicular to R and r-m, situated in proximal wing half, about in 2 / 5 of total wing length; R 1 and R 4 + 5 with macrotrichia on dorsal side only, disposed in two rows except in one row near apex of R 4 + 5; R 1 approximately 0.5 times as long as R, ending in C well before branching point of M-fork; r-m as long as basal part of median vein; M 1 and M 2 diverging; numerous macrotrichia on M and CuA; anal area distinct, slightly wedge-shaped reduced. Halter brownish; with a short stalk; knob of halter distinctly longer than stalk. Legs yellowish brown, long and slender. Coxae with very few setae. Femora with dorsal parts covered with long setae and ventral parts almost bare. Fore tibia with anteroapical tibial organ and posteroapical spur, no conspicuously spinelike differentiated setae recognizable in the remaining setosity; tibial organ as a dark patch situated at the widest point about 3 / 4 times as wide as apex of fore tibia and weakly differentiated, only denser with stronger and darker setae (not depressed and without strongly curved margin). Pretarsus on the foreleg long, as long as the following four tarsal segments together. Apex of mid tibia with three robust setae. Apex of hind tibia with a posterodorsal row of eight setae and additional robust setae on ventral and posterior sides. Tibial spurs long and strong; both spurs on apex of mid and hind tibiae of equal lengths. Claws simple. Abdomen dark brown, tergites and sternites dense, strong, and long setose. Hypopygium about 1.6 times as high as wide at base, slightly paler than abdominal segments. Gonocoxites about as long as wide, inner side of gonocoxites with few short setae, outer side with long and robust setae; ventral, inner side of gonocoxites v-shaped and basally widely separated; base of gonocoxites ventrally without intercoxal bristle groups or lobe (s). Gonostylus shorter than gonocoxite, three times longer than wide, narrow, evenly curved, narrowed in apical third and with flattened gonostylar apex, apex straight; inner side of gonostylus closed, without hollows or lobes; gonostylar apex with a strong, beak-shaped tooth subapically on ventral side, above with coarse setae shorter than tooth; near tooth with two stronger megasetae at same level, below tooth with three to four hyaline, awl-like megasetae in upper third of gonostylar inner side. Tegmen wider than high, with flattened apex. Aedeagus short. Female unknown.	en	Hebert, Florence, Muller, Valerie Ngo, Nel, Andre (2025): The oldest representative of sciarid genus Leptosciarella (Diptera) from the lowermost Eocene amber of Oise (France). Zootaxa 5739 (1): 42-52, DOI: 10.11646/zootaxa.5739.1.8, URL: https://doi.org/10.11646/zootaxa.5739.1.8
