taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
6711721DFFF74D1EFF63FC51FB70FF53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18019030/files/figure.png	https://doi.org/10.5281/zenodo.18019030	FIGURES 1–4. Onychargia atrocyana in nature. (1), a mature ♂ (Thong Nuoc Nature Reserve, Ea Hleo, Dak Lak Prov., 16.x.2023); (2), an intermediate immature ♂ (Thong Nuoc Nature Reserve, Ea Hleo, Dak Lak Prov., 16.x.2023); (3), an intermediate immature ♂ (U Minh Thuong National Park, An Giang Prov., 21.ix.2024); (4), a pair (Bao Loc, Lam Dong Prov., 11.iv.2024). Photographed by Q.T. Phan.	FIGURES 1–4. Onychargia atrocyana in nature. (1), a mature ♂ (Thong Nuoc Nature Reserve, Ea Hleo, Dak Lak Prov., 16.x.2023); (2), an intermediate immature ♂ (Thong Nuoc Nature Reserve, Ea Hleo, Dak Lak Prov., 16.x.2023); (3), an intermediate immature ♂ (U Minh Thuong National Park, An Giang Prov., 21.ix.2024); (4), a pair (Bao Loc, Lam Dong Prov., 11.iv.2024). Photographed by Q.T. Phan.	2025-12-17	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan		Zenodo	biologists	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan			
6711721DFFF74D1EFF63FC51FB70FF53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18019085/files/figure.png	https://doi.org/10.5281/zenodo.18019085	FIGURES 31–39. Comparing of anal appendages of Onychargia spp. [31–33] O. atrocyana (Bao Loc, Lam Dong Prov., 2.viii.2015, Q.T. Phan leg.), [34–36] O. priydak (Thuy Tung, Ea H’leo, Dak Lak Prov., 16.x.2023, Q.T. Phan leg.) and [37–39] O. hungeri (holotype male). (31, 34, 37), appendages in lateral view; (32, 35, 38), appendages in dorsal view (paraprocts not shown); (33, 36, 39), appendages (including valvules) in ventral view (cerci not shown).	FIGURES 31–39. Comparing of anal appendages of Onychargia spp. [31–33] O. atrocyana (Bao Loc, Lam Dong Prov., 2.viii.2015, Q.T. Phan leg.), [34–36] O. priydak (Thuy Tung, Ea H’leo, Dak Lak Prov., 16.x.2023, Q.T. Phan leg.) and [37–39] O. hungeri (holotype male). (31, 34, 37), appendages in lateral view; (32, 35, 38), appendages in dorsal view (paraprocts not shown); (33, 36, 39), appendages (including valvules) in ventral view (cerci not shown).	2025-12-17	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan		Zenodo	biologists	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan			
6711721DFFF74D1EFF63FC51FB70FF53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18019087/files/figure.png	https://doi.org/10.5281/zenodo.18019087	FIGURES 40–45. Comparing of male’s genital ligula [40, 42, 44] and female’s posterior pronotal lobe [41, 43, 45] of Onychargia spp. (40–41), O. atrocyana (Bao Loc, Lam Dong Prov., 2.viii.2015, Q.T. Phan leg.); (42–43), O. priydak (Thuy Tung, Ea H’leo, Dak Lak Prov., 16.x.2023, Q.T. Phan leg.); (44–45), O. hungeri (paratype female).	FIGURES 40–45. Comparing of male’s genital ligula [40, 42, 44] and female’s posterior pronotal lobe [41, 43, 45] of Onychargia spp. (40–41), O. atrocyana (Bao Loc, Lam Dong Prov., 2.viii.2015, Q.T. Phan leg.); (42–43), O. priydak (Thuy Tung, Ea H’leo, Dak Lak Prov., 16.x.2023, Q.T. Phan leg.); (44–45), O. hungeri (paratype female).	2025-12-17	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan		Zenodo	biologists	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan			
6711721DFFF74D1EFF63FC51FB70FF53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18019092/files/figure.png	https://doi.org/10.5281/zenodo.18019092	FIGURE 51. Distribution map of Onychargia spp. in Vietnam.	FIGURE 51. Distribution map of Onychargia spp. in Vietnam.	2025-12-17	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan		Zenodo	biologists	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan			
6711721DFFF44D1EFF63FB88FAA4F8BB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18019038/files/figure.png	https://doi.org/10.5281/zenodo.18019038	FIGURES 5–6. Onychargia priydak in nature.(5), a mature ♂ (Thong Nuoc Nature Reserve, Ea Hleo, Dak Lak Prov., 16.x.2023); (6), a pair (Bao Loc, Lam Dong Prov., 11.iv.2024). Photographed by Q.T. Phan.	FIGURES 5–6. Onychargia priydak in nature.(5), a mature ♂ (Thong Nuoc Nature Reserve, Ea Hleo, Dak Lak Prov., 16.x.2023); (6), a pair (Bao Loc, Lam Dong Prov., 11.iv.2024). Photographed by Q.T. Phan.	2025-12-17	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan		Zenodo	biologists	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan			
6711721DFFF44D1EFF63FB88FAA4F8BB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18019085/files/figure.png	https://doi.org/10.5281/zenodo.18019085	FIGURES 31–39. Comparing of anal appendages of Onychargia spp. [31–33] O. atrocyana (Bao Loc, Lam Dong Prov., 2.viii.2015, Q.T. Phan leg.), [34–36] O. priydak (Thuy Tung, Ea H’leo, Dak Lak Prov., 16.x.2023, Q.T. Phan leg.) and [37–39] O. hungeri (holotype male). (31, 34, 37), appendages in lateral view; (32, 35, 38), appendages in dorsal view (paraprocts not shown); (33, 36, 39), appendages (including valvules) in ventral view (cerci not shown).	FIGURES 31–39. Comparing of anal appendages of Onychargia spp. [31–33] O. atrocyana (Bao Loc, Lam Dong Prov., 2.viii.2015, Q.T. Phan leg.), [34–36] O. priydak (Thuy Tung, Ea H’leo, Dak Lak Prov., 16.x.2023, Q.T. Phan leg.) and [37–39] O. hungeri (holotype male). (31, 34, 37), appendages in lateral view; (32, 35, 38), appendages in dorsal view (paraprocts not shown); (33, 36, 39), appendages (including valvules) in ventral view (cerci not shown).	2025-12-17	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan		Zenodo	biologists	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan			
6711721DFFF44D1EFF63FB88FAA4F8BB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18019087/files/figure.png	https://doi.org/10.5281/zenodo.18019087	FIGURES 40–45. Comparing of male’s genital ligula [40, 42, 44] and female’s posterior pronotal lobe [41, 43, 45] of Onychargia spp. (40–41), O. atrocyana (Bao Loc, Lam Dong Prov., 2.viii.2015, Q.T. Phan leg.); (42–43), O. priydak (Thuy Tung, Ea H’leo, Dak Lak Prov., 16.x.2023, Q.T. Phan leg.); (44–45), O. hungeri (paratype female).	FIGURES 40–45. Comparing of male’s genital ligula [40, 42, 44] and female’s posterior pronotal lobe [41, 43, 45] of Onychargia spp. (40–41), O. atrocyana (Bao Loc, Lam Dong Prov., 2.viii.2015, Q.T. Phan leg.); (42–43), O. priydak (Thuy Tung, Ea H’leo, Dak Lak Prov., 16.x.2023, Q.T. Phan leg.); (44–45), O. hungeri (paratype female).	2025-12-17	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan		Zenodo	biologists	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan			
6711721DFFF44D1EFF63FB88FAA4F8BB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18019030/files/figure.png	https://doi.org/10.5281/zenodo.18019030	FIGURES 1–4. Onychargia atrocyana in nature. (1), a mature ♂ (Thong Nuoc Nature Reserve, Ea Hleo, Dak Lak Prov., 16.x.2023); (2), an intermediate immature ♂ (Thong Nuoc Nature Reserve, Ea Hleo, Dak Lak Prov., 16.x.2023); (3), an intermediate immature ♂ (U Minh Thuong National Park, An Giang Prov., 21.ix.2024); (4), a pair (Bao Loc, Lam Dong Prov., 11.iv.2024). Photographed by Q.T. Phan.	FIGURES 1–4. Onychargia atrocyana in nature. (1), a mature ♂ (Thong Nuoc Nature Reserve, Ea Hleo, Dak Lak Prov., 16.x.2023); (2), an intermediate immature ♂ (Thong Nuoc Nature Reserve, Ea Hleo, Dak Lak Prov., 16.x.2023); (3), an intermediate immature ♂ (U Minh Thuong National Park, An Giang Prov., 21.ix.2024); (4), a pair (Bao Loc, Lam Dong Prov., 11.iv.2024). Photographed by Q.T. Phan.	2025-12-17	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan		Zenodo	biologists	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan			
6711721DFFF44D1EFF63FB88FAA4F8BB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18019092/files/figure.png	https://doi.org/10.5281/zenodo.18019092	FIGURE 51. Distribution map of Onychargia spp. in Vietnam.	FIGURE 51. Distribution map of Onychargia spp. in Vietnam.	2025-12-17	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan		Zenodo	biologists	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan			
6711721DFFF54D10FF63F9DEFD99FF7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18019053/files/figure.png	https://doi.org/10.5281/zenodo.18019053	FIGURES 7–8. Habitus of (7), holotype ♂ and (8), paratype ♀ of Onychargia hungeri sp. nov.	FIGURES 7–8. Habitus of (7), holotype ♂ and (8), paratype ♀ of Onychargia hungeri sp. nov.	2025-12-17	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan		Zenodo	biologists	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan			
6711721DFFF54D10FF63F9DEFD99FF7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18019064/files/figure.png	https://doi.org/10.5281/zenodo.18019064	FIGURES 9–14. Structures of the holotype ♂ [9–11] and paratype ♀ [12–14] of Onychargia hungeri sp. nov. (9, 12), head in frontal view; (10, 13), head and thorax in lateral view; (11, 14), thorax in dorsal view.	FIGURES 9–14. Structures of the holotype ♂ [9–11] and paratype ♀ [12–14] of Onychargia hungeri sp. nov. (9, 12), head in frontal view; (10, 13), head and thorax in lateral view; (11, 14), thorax in dorsal view.	2025-12-17	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan		Zenodo	biologists	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan			
6711721DFFF54D10FF63F9DEFD99FF7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18019075/files/figure.png	https://doi.org/10.5281/zenodo.18019075	FIGURES 15–22. Appendages and genital ligula of Onychargia hungeri sp. nov. [15–20] holotype ♂ and [21–22] paratype ♀. (15), appendages in lateral view; (16), appendages in dorsolateral view; (17), appendages in dorsal view; (18), appendages in ventral view; (19), genital ligula in lateral view; (20), genital ligula in ventral view; (21), abdominal tip in dorsal view; (22), abdominal tip in lateral view.	FIGURES 15–22. Appendages and genital ligula of Onychargia hungeri sp. nov. [15–20] holotype ♂ and [21–22] paratype ♀. (15), appendages in lateral view; (16), appendages in dorsolateral view; (17), appendages in dorsal view; (18), appendages in ventral view; (19), genital ligula in lateral view; (20), genital ligula in ventral view; (21), abdominal tip in dorsal view; (22), abdominal tip in lateral view.	2025-12-17	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan		Zenodo	biologists	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan			
6711721DFFF54D10FF63F9DEFD99FF7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18019077/files/figure.png	https://doi.org/10.5281/zenodo.18019077	FIGURES 23–27. Onychargia hungeri sp. nov. in the type location. (23), an early immature ♂ (16.iv.2025); (24), an intermediate immature ♂ (16.iv.2025); (25), a mature ♂ (16.iv.2025); (26), a mature ♀ (25.vii.2025); (27), a mating pair (23.iv.2025). Photographed by M.K. Tran.	FIGURES 23–27. Onychargia hungeri sp. nov. in the type location. (23), an early immature ♂ (16.iv.2025); (24), an intermediate immature ♂ (16.iv.2025); (25), a mature ♂ (16.iv.2025); (26), a mature ♀ (25.vii.2025); (27), a mating pair (23.iv.2025). Photographed by M.K. Tran.	2025-12-17	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan		Zenodo	biologists	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan			
6711721DFFF54D10FF63F9DEFD99FF7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18019081/files/figure.png	https://doi.org/10.5281/zenodo.18019081	FIGURES 28–30. Location and type locality of Onychargia hungeri sp. nov. (28), location of the type location in Vietnam; (29), a female ovipositing in tandem with a male; (30), the first author taking photo in the habitat of the new species.	FIGURES 28–30. Location and type locality of Onychargia hungeri sp. nov. (28), location of the type location in Vietnam; (29), a female ovipositing in tandem with a male; (30), the first author taking photo in the habitat of the new species.	2025-12-17	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan		Zenodo	biologists	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan			
6711721DFFF54D10FF63F9DEFD99FF7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18019085/files/figure.png	https://doi.org/10.5281/zenodo.18019085	FIGURES 31–39. Comparing of anal appendages of Onychargia spp. [31–33] O. atrocyana (Bao Loc, Lam Dong Prov., 2.viii.2015, Q.T. Phan leg.), [34–36] O. priydak (Thuy Tung, Ea H’leo, Dak Lak Prov., 16.x.2023, Q.T. Phan leg.) and [37–39] O. hungeri (holotype male). (31, 34, 37), appendages in lateral view; (32, 35, 38), appendages in dorsal view (paraprocts not shown); (33, 36, 39), appendages (including valvules) in ventral view (cerci not shown).	FIGURES 31–39. Comparing of anal appendages of Onychargia spp. [31–33] O. atrocyana (Bao Loc, Lam Dong Prov., 2.viii.2015, Q.T. Phan leg.), [34–36] O. priydak (Thuy Tung, Ea H’leo, Dak Lak Prov., 16.x.2023, Q.T. Phan leg.) and [37–39] O. hungeri (holotype male). (31, 34, 37), appendages in lateral view; (32, 35, 38), appendages in dorsal view (paraprocts not shown); (33, 36, 39), appendages (including valvules) in ventral view (cerci not shown).	2025-12-17	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan		Zenodo	biologists	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan			
6711721DFFF54D10FF63F9DEFD99FF7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18019090/files/figure.png	https://doi.org/10.5281/zenodo.18019090	FIGURES 46–50. Photos of Onychargia cf. hungeri outside Vietnam. (46), in Sha Lo Tung, Hong Kong, by Dr. Ying Kin Ken So; (47), in Yilan, Taiwan, by ku6777 (Source: https://www.inaturalist.org/observations/298352858); (48), in Sanzhaozhen, Macao, by kenkwan0208 (Source: https://www.inaturalist.org/observations/295608385); (49), in Shuen Wan, Shenzen, by treefong (Source: https://www.inaturalist.org/observations/231366494); (50), in Xiuying District, Haikou, Hainan, by zicky (Source: https://www.inaturalist.org/observations/292090559).	FIGURES 46–50. Photos of Onychargia cf. hungeri outside Vietnam. (46), in Sha Lo Tung, Hong Kong, by Dr. Ying Kin Ken So; (47), in Yilan, Taiwan, by ku6777 (Source: https://www.inaturalist.org/observations/298352858); (48), in Sanzhaozhen, Macao, by kenkwan0208 (Source: https://www.inaturalist.org/observations/295608385); (49), in Shuen Wan, Shenzen, by treefong (Source: https://www.inaturalist.org/observations/231366494); (50), in Xiuying District, Haikou, Hainan, by zicky (Source: https://www.inaturalist.org/observations/292090559).	2025-12-17	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan		Zenodo	biologists	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan			
6711721DFFF54D10FF63F9DEFD99FF7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18019087/files/figure.png	https://doi.org/10.5281/zenodo.18019087	FIGURES 40–45. Comparing of male’s genital ligula [40, 42, 44] and female’s posterior pronotal lobe [41, 43, 45] of Onychargia spp. (40–41), O. atrocyana (Bao Loc, Lam Dong Prov., 2.viii.2015, Q.T. Phan leg.); (42–43), O. priydak (Thuy Tung, Ea H’leo, Dak Lak Prov., 16.x.2023, Q.T. Phan leg.); (44–45), O. hungeri (paratype female).	FIGURES 40–45. Comparing of male’s genital ligula [40, 42, 44] and female’s posterior pronotal lobe [41, 43, 45] of Onychargia spp. (40–41), O. atrocyana (Bao Loc, Lam Dong Prov., 2.viii.2015, Q.T. Phan leg.); (42–43), O. priydak (Thuy Tung, Ea H’leo, Dak Lak Prov., 16.x.2023, Q.T. Phan leg.); (44–45), O. hungeri (paratype female).	2025-12-17	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan		Zenodo	biologists	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan			
6711721DFFF54D10FF63F9DEFD99FF7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18019030/files/figure.png	https://doi.org/10.5281/zenodo.18019030	FIGURES 1–4. Onychargia atrocyana in nature. (1), a mature ♂ (Thong Nuoc Nature Reserve, Ea Hleo, Dak Lak Prov., 16.x.2023); (2), an intermediate immature ♂ (Thong Nuoc Nature Reserve, Ea Hleo, Dak Lak Prov., 16.x.2023); (3), an intermediate immature ♂ (U Minh Thuong National Park, An Giang Prov., 21.ix.2024); (4), a pair (Bao Loc, Lam Dong Prov., 11.iv.2024). Photographed by Q.T. Phan.	FIGURES 1–4. Onychargia atrocyana in nature. (1), a mature ♂ (Thong Nuoc Nature Reserve, Ea Hleo, Dak Lak Prov., 16.x.2023); (2), an intermediate immature ♂ (Thong Nuoc Nature Reserve, Ea Hleo, Dak Lak Prov., 16.x.2023); (3), an intermediate immature ♂ (U Minh Thuong National Park, An Giang Prov., 21.ix.2024); (4), a pair (Bao Loc, Lam Dong Prov., 11.iv.2024). Photographed by Q.T. Phan.	2025-12-17	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan		Zenodo	biologists	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan			
6711721DFFF54D10FF63F9DEFD99FF7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18019038/files/figure.png	https://doi.org/10.5281/zenodo.18019038	FIGURES 5–6. Onychargia priydak in nature.(5), a mature ♂ (Thong Nuoc Nature Reserve, Ea Hleo, Dak Lak Prov., 16.x.2023); (6), a pair (Bao Loc, Lam Dong Prov., 11.iv.2024). Photographed by Q.T. Phan.	FIGURES 5–6. Onychargia priydak in nature.(5), a mature ♂ (Thong Nuoc Nature Reserve, Ea Hleo, Dak Lak Prov., 16.x.2023); (6), a pair (Bao Loc, Lam Dong Prov., 11.iv.2024). Photographed by Q.T. Phan.	2025-12-17	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan		Zenodo	biologists	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan			
6711721DFFF54D10FF63F9DEFD99FF7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18019092/files/figure.png	https://doi.org/10.5281/zenodo.18019092	FIGURE 51. Distribution map of Onychargia spp. in Vietnam.	FIGURE 51. Distribution map of Onychargia spp. in Vietnam.	2025-12-17	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan		Zenodo	biologists	Tran, Minh Khang;Huong, Le Thi Thu;Phan, Quoc Toan			
