identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
2B3F9391A9EA53B885FC5DB0C916B293.text	2B3F9391A9EA53B885FC5DB0C916B293.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glossoloma jostii J. L. Clark 2026	<div><p>Glossoloma jostii J. L. Clark sp. nov.</p><p>Fig. 1</p><p>Diagnosis.</p><p>Differs from all other congeners by its scandent subwoody habit (vs. the more common terrestrial unbranched shrub habit), brittle leaf blades (vs. chartaceous in most other congeners), apically constricted corolla (vs. non-ampliate or broad as in G. subglabrum), uniformly red corollas (vs. uniformly yellow, or yellow with red lobes), and valvate calyx lobes (vs. conduplicate calyx lobes where each lobe is appressed to an adjacent lobe and folded lengthwise in G. puroana and G. recaldeorum).</p><p>Type.</p><p>Ecuador • Tungurahua: Cantón Baños, parroquia Río Verde, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.30317&amp;materialsCitation.latitude=-1.4407501" title="Search Plazi for locations around (long -78.30317/lat -1.4407501)">Bosque Protector Cerro Candelaria (Fundación EcoMinga), upper Pastaza watershed, high ridgeline above canyon, trail between Cerro Candelaria permanent plot # 1 (2200 m) and high camp (2600 m)</a>, 1°26'26.7"S, 78°18'11.4"W, 2200–2664 m, 9 March 2016, J. L. Clark, J. A. Mayr &amp; D. A. Neill 14803 (holotype: QCA!; isotypes: E!, ECUAMZ!, G!, MO!, NY!, SEL! [barcode SEL 089267], US!) .</p><p>Description.</p><p>Elongate scandent nomadic climbers. Stems elongate and subwoody, terete in cross section, 3–6 mm in diameter, sparsely to densely pilose. Leaves opposite, equal in a pair; petioles 1–3 cm long, green, terete in cross-section, densely pilose; blade brittle (especially when dry), oblong, 5–10 × 1.5–3 cm, coriaceous, adaxially and abaxially green, apex acute to acuminate, base acute, margin serrate, 4–6 pairs of secondary veins, hispid abaxially and adaxially. Inflorescences reduced to a single axillary flower. Flowers resupinate, with elongate pendent pedicels, 1.6–2.2 cm long. Calyx lobes uniformly dark red, pilose on the outside and glabrous on the inside, lobes 5, nearly free, fused at the base for 2–4 mm, valvate to separate, and clasping corolla tube, lower lobe relatively smaller and narrow, ca. 1.3 × 0.4 cm, the other four lobes broadly ovate, apex acute, margins serrate, ca. 1.5 × 1.3 cm. Corolla tube ampliate on upper surface and apically constricted, appearing perpendicular relative to calyx, 2.7–3.8 cm long, gibbous at base, appearing laterally compressed, 2.0– 3.5 mm wide, outside uniformly pilose, inside mostly glabrous with minute glandular trichomes apically, throat elliptic in cross section, lobes 5, subequal, margins entire to serrulate, lobes reflexed, 8–11 × 9–12 mm, uniformly red. Androecium of 4 didynamous stamens, included, filaments broad and flat, ca. 3.0 cm long, adnate to the corolla tube base for 4 mm, white, glabrous; anthers oblong, coherent by the lateral walls, dehiscing longitudinally, 4.2–6.0 × 0.7–2.0 mm. Gynoecium with a single bilobed dorsal gland; ovary superior, 4.0–5.0 × 4.0–5.0 mm, cone-shaped, puberulent; style stout, included, 3.0 cm long; stigma stomatomorphic. Fruit not observed.</p><p>Phenology.</p><p>Collected with flowers in March.</p><p>Etymology.</p><p>The specific epithet honors Lou Jost, botanist, conservationist, self-taught mathematical biologist, and co-founder of the EcoMinga Foundation. His efforts have been instrumental in protecting montane forests throughout Ecuador, particularly in the Pastaza Valley on the eastern slopes of the Andes. Jost has contributed extensively to the discovery and description of plant species, especially orchids, and his vision and leadership have been pivotal in conserving biodiversity hotspots in the region. This epithet commemorates his dedication to science, conservation, and the protection of remnant Andean forests, including the locality of the three new species described here.</p><p>Distribution.</p><p>Glossoloma jostii is currently only known from the Cerro Candelaria reserve near Río Verde in Bosque Protector Cerro Candelaria, a private reserve managed by Fundación EcoMinga on the eastern slopes of the Ecuadorian Andes.</p><p>Comments.</p><p>The combination of uniformly red corollas with a nomadic scandent habit is rare in Glossoloma . An undescribed western Andean species also shares the scandent habit and uniformly red corollas of G. jostii, but differs by its green calyx lobes (vs. red in G. jostii) and a uniformly bright yellow throat (vs. the uniformly red inner and outer surfaces in G. jostii; Fig. 1 B). Some other species of Glossoloma also have uniformly red corollas, but they are readily distinguished from G. jostii by their unbranched, terrestrial subshrub habit and elongate tubular corollas without an apical constriction (vs. ampliated corollas that are apically constricted in G. jostii; Fig. 1 A). Examples of unbranched terrestrial subshrub species with red corolla include G. baguense (L. E. Skog) J. L. Clark, G. subglabrum J. L. Clark, G. oblongicalyx (J. L. Clark &amp; L. E. Skog) J. L. Clark, and G. panamense (C. V. Morton) J. L. Clark. One example of a terrestrial species of Glossoloma characterized with uniformly red corollas and apically constricted corolla tubes is Glossoloma ichthyoderma (Hanst.) J. L. Clark. Glossoloma jostii can be distinguished from G. ichthyoderma by its scandent, branched shoots (Fig. 1 D). In contrast, G. ichthyoderma consistently has erect, unbranched shoots, with a terrestrial or epiphytic habit. Vegetatively, G. jostii is differentiated by brittle leaves with a hispid indumentum (Fig. 1 C &amp; Table 1). The leaves in G. puroana are chartaceous (not brittle) with a sericeous indumentum on the leaf (vs. hispid indumentum in G. jostii; Table 1).</p></div>	https://treatment.plazi.org/id/2B3F9391A9EA53B885FC5DB0C916B293	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Clark, John L.	Clark, John L. (2026): Commemorating a conservation trifecta: Three new species of Glossoloma (Gesneriaceae) honoring a donor, a family of forest stewards, and a conservation visionary. PhytoKeys 271: 173-185, DOI: 10.3897/phytokeys.271.181141
5F0725EEE1E05852AA62CA3F6FBDCD91.text	5F0725EEE1E05852AA62CA3F6FBDCD91.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glossoloma puroana J. L. Clark 2026	<div><p>Glossoloma puroana J. L. Clark sp. nov.</p><p>Fig. 2</p><p>Diagnosis.</p><p>Differs from all other congeners by the presence of a scandent subwoody habit (vs. terrestrial unbranched shrub), chartaceous leaf blades (vs. brittle leaf blades in G. jostii), presence of inflorescence bracts (vs. absence of inflorescence bracts in G. jostii), nearly tubular corolla that lacks an apical constriction (vs. ampliate and apically constricted corolla in G. jostii), a corolla tube with prominent longitudinal ridges (vs. smooth corolla in G. jostii), and cucullate calyx lobes (vs. valvate calyx lobes in G. jostii).</p><p>Type.</p><p>Ecuador • Tungurahua: Cantón Baños, parroquia Río Verde, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.30317&amp;materialsCitation.latitude=-1.4407501" title="Search Plazi for locations around (long -78.30317/lat -1.4407501)">Bosque Protector Cerro Candelaria (Fundación EcoMinga), upper Pastaza watershed, high ridgeline above canyon, trail between Cerro Candelaria permanent plot # 1 (2200 m) and high camp (2600 m)</a>, 1°26'26.7"S, 78°18'11.4"W, 2200–2664 m, 9 March 2016, J. L. Clark, J. A. Mayr &amp; D. A. Neill 14812 (holotype: QCA!; isotypes: ECUAMZ!, G!, MO!, NY!, SEL! [barcode SEL 089280], US!) .</p><p>Description.</p><p>Elongate scandent nomadic climbers. Stems elongate and quadrangular in cross section, 3–6 mm in diameter, sparsely pilose. Leaves opposite, equal in a pair; petiole 3–5.5 cm long, green, terete in cross-section, sparsely pilose; blade oblong, 15–20 × 4.5–7.0 cm, chartaceous, adaxially dark green, abaxially light green, apex acute to acuminate, base rounded, margin serrate, 5–7 pairs of secondary veins, abaxially and adaxially sericeous. Inflorescences reduced to 1 to 3 axillary flowers. Flowers resupinate, with elongate pendent pedicels, 2.8–3.7 cm long. Calyx uniformly red, sparsely pilose on the outside and glabrous on the inside, lobes 5, fused at the base for 2–4 mm, conduplicate with each lobe appressed to an adjacent lobe and folded lengthwise with the margin curved inward, clasping corolla tube, lower lobe relatively smaller and narrow, ca. 1.8 × 0.8 cm, the other four lobes broadly oblong, apex rounded, margins serrate, ca. 2.0 × 1.4 cm. Corolla tube ampliate on upper surface and slightly constricted apically, appearing perpendicular relative to calyx, 3.0–4.0 cm long, gibbous at base, with prominent longitudinal ridges, appearing laterally compressed, 2.5–4.0 mm wide, outside uniformly pilose, inside mostly glabrous with minute glandular trichomes apically, throat elliptic in cross section, lobes 5, subequal, margins entire to serrulate, lobes reflexed, 8–11 × 9–12 mm, tube yellow, basally suffused with red. Androecium of 4 didynamous stamens, included, filaments broad and flat, ca. 3.8 cm long, adnate to the corolla tube base for 4 mm, white, glabrous; anthers oblong, coherent by the lateral walls, dehiscing longitudinally, 4.3–6.2 × 0.8–2.2 mm. Gynoecium with a single bilobed dorsal gland; ovary superior, 4.2–5.2 × 4.2–5.2 mm, cone-shaped, pilose; style stout, included, 3.5 cm long; stigma stomatomorphic. Fruit not observed.</p><p>Phenology.</p><p>Collected with flowers in March.</p><p>Etymology.</p><p>The specific epithet honors Puro Coffee (UK), founded by Andy Orchard, whose support through the World Land Trust was instrumental in establishing Fundación EcoMinga’s Cerro Candelaria Reserve that safeguards all three species described here. In addition, Puro Coffee, through continued donations to the World Land Trust, supports the salary of a park guard at the reserve.</p><p>Distribution.</p><p>Glossoloma puroana is currently only known from the Cerro Candelaria reserve near Río Verde in Bosque Protector Cerro Candelaria, a private reserve managed by Fundación EcoMinga on the eastern slopes of the Ecuadorian Andes.</p><p>Comments.</p><p>Glossoloma puroana is characterized by elongate tubular corollas with prominent longitudinal ridges (Fig. 2 A). The corolla is yellow, suffused with red at the base externally, and marked internally with prominent red spots (Fig. 2 C). The calyx lobes are red and folded in a conduplicate manner (i. e., each lobe appressed to an adjacent lobe and folded lengthwise with the margin curved inward – Fig. 2 A &amp; D). Yellow corollas are common in Glossoloma, but this is the only species in the genus known to have both a scandent habit and corollas yellow suffused with red (Fig. 2). In addition, while most species of Glossoloma have terete stems, G. puroana is distinctive in having subquadrangular stems (Fig. 2 B &amp; D). The leaf blades in G. puroana are chartaceous with a sericeous indumentum (Table 1). In contrast, the leaves in G. jostii are stiff, brittle (especially when dry) and have a hispid indumentum (Table 1). Another diagnostic character of G. puroana is the presence of multiple flowers in the leaf axils (vs. single axillary flowers in G. jostii and G. recaldeorum) and presence of inflorescence bracts (vs. absence of inflorescence bracts in G. jostii and G. recaldeorum).</p></div>	https://treatment.plazi.org/id/5F0725EEE1E05852AA62CA3F6FBDCD91	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Clark, John L.	Clark, John L. (2026): Commemorating a conservation trifecta: Three new species of Glossoloma (Gesneriaceae) honoring a donor, a family of forest stewards, and a conservation visionary. PhytoKeys 271: 173-185, DOI: 10.3897/phytokeys.271.181141
FC3B7CDBEFD756A1A861107725B59350.text	FC3B7CDBEFD756A1A861107725B59350.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glossoloma recaldeorum J. L. Clark 2026	<div><p>Glossoloma recaldeorum J. L. Clark sp. nov.</p><p>Fig. 3</p><p>Diagnosis.</p><p>Differs from all other congeners by the presence of a scandent subwoody habit (vs. terrestrial unbranched shrub), small ovate leaves (vs. oblong leaves in G. jostii and G. puroana), a corolla tube that is rounded at the throat (vs. laterally compressed corolla tubes in G. jostii and G. puroana), and single axillary flowers (vs. multiple flowers per leaf axil in G. jostii and G. puroana).</p><p>Type.</p><p>Ecuador • Tungurahua: Canton Baños, parroquia Río Verde, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.30317&amp;materialsCitation.latitude=-1.4407501" title="Search Plazi for locations around (long -78.30317/lat -1.4407501)">Bosque Protector Cerro Candelaria (Fundación EcoMinga), upper Pastaza watershed, high ridgeline above canyon, trail between Cerro Candelaria permanent plot # 1 (2200 m) and high camp (2600 m)</a>, 1°26'26.7"S, 78°18'11.4"W, 2200–2664 m, 9 March 2016, J. L. Clark, J. A. Mayr &amp; D. A. Neill 14818 (holotype: QCA!; isotypes: CAS!, E!, ECUAMZ!, G!, MO!, NY! SEL!, US!) .</p><p>Description.</p><p>Elongate scandent nomadic climbers. Stems elongate and terete in cross section, 2–4 mm in diameter, sparsely pilose. Leaves opposite, equal in a pair; petiole 4–10 mm long, red, terete in cross-section, sparsely pilose; blade ovate, 3–4 × 1.5–2.0 cm, coriaceous, adaxially dark green, abaxially light green, apex acute, base rounded, margin sparsely serrate, 3–5 pairs of secondary veins, abaxially and adaxially pilose. Inflorescences reduced to a single axillary flower. Flowers resupinate, with elongate pendent to horizontal 2.5–3.3 cm long pedicels. Calyx mostly light green with reddish base, sparsely pilose on the outside and glabrous on the inside, lobes 5, fused at the base for 2–4 mm, conduplicate with each lobe appressed to an adjacent lobe and folded lengthwise with the margin curved inward, clasping corolla tube, lower lobe relatively smaller and narrow, ca. 1.0 × 0.4 cm, the other four lobes broadly oblong, apex acute, margins serrate, ca. 1.3 × 0.8 cm. Corolla tube ampliate on upper surface and slightly constricted apically, appearing perpendicular relative to calyx, 2.5–3.2 cm long, gibbous at base, 2.5–4.0 mm wide, outside uniformly pilose, inside mostly glabrous with minute glandular trichomes apically, throat rounded in cross section, lobes 5, subequal, margins entire to serrulate, lobes reflexed, 8–11 × 9–12 mm, uniformly yellow. Androecium of 4 didynamous stamens, included, filaments broad and flat, ca. 3.1 cm long, adnate to the corolla tube base for 3 mm, white, glabrous; anthers oblong, coherent by the lateral walls, dehiscing longitudinally, 3.9–5.7 × 0.6–1.8 mm. Gynoecium with a single bilobed dorsal gland; ovary superior, 3.9–4.9 × 3.0–4.0 mm, cone-shaped, puberulent; style stout, included, 3.0 cm long; stigma stomatomorphic. Fruit not observed.</p><p>Phenology.</p><p>Collected with flowers in March.</p><p>Etymology.</p><p>The specific epithet honors the Recalde family of El Placer, the nearest community to the location of the species described here. Over the years, the brothers Luis, Fausto, Jesús, and Abdón Recalde, and later their children Darwin, Santiago, and Diana Recalde, have served as park guards protecting Cerro Candelaria Reserve, and have been instrumental in promoting a positive attitude in communities towards promoting forest conservation. Their keen passion for nature and their extraordinary observations have led to the discovery of many new species, especially frogs.</p><p>Distribution.</p><p>Glossoloma recaldeorum is currently only known from the Cerro Candelaria reserve near Río Verde in Bosque Protector Cerro Candelaria, a private reserve managed by Fundación EcoMinga on the eastern slopes of the Ecuadorian Andes.</p><p>Comments.</p><p>Glossoloma recaldeorum is distinguished by its small ovate leaves (Fig. 3 C), the smallest currently known in the genus. The petioles are typically red, in contrast to the green petioles of congeners. The corollas are yellow, as in G. puroana, but differ in being more rounded at the throat (Fig. 3 B) rather than laterally compressed (Fig. 2 C). In addition, the presence of solitary axillary flowers (Fig. 3 D) is unique within the genus, where most species bear multiple flowers per leaf axil (e. g., Fig. 2 D).</p></div>	https://treatment.plazi.org/id/FC3B7CDBEFD756A1A861107725B59350	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Clark, John L.	Clark, John L. (2026): Commemorating a conservation trifecta: Three new species of Glossoloma (Gesneriaceae) honoring a donor, a family of forest stewards, and a conservation visionary. PhytoKeys 271: 173-185, DOI: 10.3897/phytokeys.271.181141
