taxonID	type	description	language	source
622E6F09DD6BFFA8FCC61359F50278CB.taxon	description	(Figs. 1 – 5, Table 3) urn: lsid: zoobank. org: act: ECBED 1 B 6 - EC 60 - 440 D-A 826 - 89322 D 5 F 6674	en	Bianchi, Filipe Michels, Krein, Verônica, Rider, David, Grazia, Jocelia (2021): Finding Padaeus bovillus (Hemiptera: Pentatomidae): A Phylogenetic Placement and the Description of Its Sister Species. Zoological Studies 60 (11), No. 11: 1-13, DOI: 10.6620/ZS.2021.60-11, URL: http://dx.doi.org/10.5281/zenodo.12825774
622E6F09DD6BFFA8FCC61359F50278CB.taxon	materials_examined	Type Material: Holotype: ECUADOR: 1 male, Paramba, Ecuador / Coll. I. R. Sc. N. B, Equateur: Paramba, Ecuador (UFRG); 1 female, Paramba, Ecuador / Collection Rosenberg / NMNH.	en	Bianchi, Filipe Michels, Krein, Verônica, Rider, David, Grazia, Jocelia (2021): Finding Padaeus bovillus (Hemiptera: Pentatomidae): A Phylogenetic Placement and the Description of Its Sister Species. Zoological Studies 60 (11), No. 11: 1-13, DOI: 10.6620/ZS.2021.60-11, URL: http://dx.doi.org/10.5281/zenodo.12825774
622E6F09DD6BFFA8FCC61359F50278CB.taxon	etymology	Etymology: The name is allusive to the seclusion among the authors while describing this species (L. seclusus: shut up, separated, recluse, seclude). Most of the intellectual work was made during the pandemic of COVID- 19, with the authors respecting proper social distancing.	en	Bianchi, Filipe Michels, Krein, Verônica, Rider, David, Grazia, Jocelia (2021): Finding Padaeus bovillus (Hemiptera: Pentatomidae): A Phylogenetic Placement and the Description of Its Sister Species. Zoological Studies 60 (11), No. 11: 1-13, DOI: 10.6620/ZS.2021.60-11, URL: http://dx.doi.org/10.5281/zenodo.12825774
622E6F09DD6BFFA8FCC61359F50278CB.taxon	diagnosis	Diagnosis: Mitripus seclusus sp. n. may be distinguished from M. convergens and M. legionarius by each humeral angle developed laterally as a stout spine; from M. acutus by the clypeus being slightly longer than the mandibular plates, and the anterolateral margins of the pronotum entirely smooth; and from M. bovillus by the humeral angles not depressed anteriorly, and by the shape of the parameres (Fig. 3).	en	Bianchi, Filipe Michels, Krein, Verônica, Rider, David, Grazia, Jocelia (2021): Finding Padaeus bovillus (Hemiptera: Pentatomidae): A Phylogenetic Placement and the Description of Its Sister Species. Zoological Studies 60 (11), No. 11: 1-13, DOI: 10.6620/ZS.2021.60-11, URL: http://dx.doi.org/10.5281/zenodo.12825774
622E6F09DD6BFFA8FCC61359F50278CB.taxon	description	D e s c r i p t i o n: C o l o r a t i o n: D o r s a l s u r f a c e ochraceous with black punctures, giving an overall dark brown matte aspect; antennae ochraceous ventrally and basally, brownish dorsally. Scutellum darker basally than apically. Connexivum brownish, middle third ochraceous. Ventral surface yellowish ochraceous; head and abdomen impunctate, thorax with brownish-yellow punctures. Legs ochraceous with reddish-brown spots on femora and tibiae; each femur with brownish ring at apex; apex of each tarsomere brownish. Head: Clypeus slightly longer than mandibular plates, both rounded apically. Head tapering to apex, lateral margins of mandibular plates sinuous, concave near eyes. Ocelli red or yellow. Antennomere proportions: I <II <III (IV and V lacking). Anterior margins of bucculae truncate with sharp projections, posterior margins evanescent. Gena flat. Rostrum reaching anterior margins of metacoxae. Thorax: Anterolateral margin of pronotum smooth on anterior half, slightly sinuous; anterolateral angles developed as small yellowish-ochraceous spines, projected laterally. Each humeral angle produced laterally as a black stout spine. Pronotal cicatrices brownish, with an ochraceous spot posterior to each mesial angle. Basal angles of scutellum with small foveae. Apex of radial vein with a small ivory dot at endocorium. Membrane of hemelytra fumose, with subparallel veins. Evaporatoria each extending halfway from ostiole to metapleural lateral margin; surface impunctate, presenting gyrification near ostioles; lateral fold present; peritreme spout-like. Abdomen: Each posterolateral angle of connexiva developed as a tiny black spine. Posterolateral angles of urosternite VII slightly projected as a spine. Spiracles concolorous with abdominal disc. Male genitalia: Pygophore (Fig. 3 A – C): In dorsal view, trapezoidal; genital cup not well exposed; each posterolateral angle of pygophore projected as a small triangle, rounded apically; median projection of dorsal rim short. Dorsal rim interrupted by diagonal depressions flanking median projection of dorsal rim; lateral margins slightly sinuous, discontinuous near median projection of dorsal rim; superior process of dorsal rim exposed, rectangular; ventral rim concave with a triangular projection at middle. In posterior view, genial cup opening ellipsoid; dorsal rim concave, smooth; superior process of dorsal rim long, bladelike; transverse ridge concave medially, U-shaped; inferior layer of ventral rim medially straight, tumescent laterally. In ventral view, ventral rim concave with a triangular projection at middle; posterolateral angles projected posteriorly, slightly bifid. Tenth segment: posterior margin trapezoidal with setae; disc smooth; tubercles on basal third slightly developed. Parameres (Fig. 3 D – G): In dorsal and ventral view: each with basal apodeme smaller than crown; stem slightly smaller than crown; crown bending outward nearly at a 45 ° angle. Basal process of paramere narrow, rounded, with long setae; apical process of paramere enlarged basally, tapering towards truncate apex. In mesial and lateral views: apical process of paramere stout, triangular, with scale-like structures on its outer surface. Phallus (Fig. 3 H – J): Phallotheca tubular, slightly constricted basally; ventral basal processes of phallotheca quadrangular; dorsal processes of phallotheca hook-like, bent ventrally in about 45 °, short, not surpassing expanded conjunctiva; vesica process spout-like; conjunctiva lacking processes; ductus seminis distalis short, not exposed out of phallotheca. Female genitalia: In posteroventral view (Fig. 4 A), gonocoxites VIII with shallow punctures, concolorous with ochraceous disc; surface flat with a slightly depressed area near posterolateral margin; mesial margins straight, overlapping, with a brown spot apically; apex squared. Laterotergites VIII with black margins; apex spine-like. Gonocoxites IX trapezoid, about three times wider than long, anterior and lateral margins straight, posterior margin concave. Laterotergites IX rounded apically, mesial margins forming a right angle, lateral margins convex, slightly surpassing tergite VIII; segment X rectangular.	en	Bianchi, Filipe Michels, Krein, Verônica, Rider, David, Grazia, Jocelia (2021): Finding Padaeus bovillus (Hemiptera: Pentatomidae): A Phylogenetic Placement and the Description of Its Sister Species. Zoological Studies 60 (11), No. 11: 1-13, DOI: 10.6620/ZS.2021.60-11, URL: http://dx.doi.org/10.5281/zenodo.12825774
622E6F09DD6BFFA8FCC61359F50278CB.taxon	distribution	Distribution: Mitripus seclusus sp. n. is known only from Ecuador (Fig. 5).	en	Bianchi, Filipe Michels, Krein, Verônica, Rider, David, Grazia, Jocelia (2021): Finding Padaeus bovillus (Hemiptera: Pentatomidae): A Phylogenetic Placement and the Description of Its Sister Species. Zoological Studies 60 (11), No. 11: 1-13, DOI: 10.6620/ZS.2021.60-11, URL: http://dx.doi.org/10.5281/zenodo.12825774
622E6F09DD6FFFA5FC861178F3577E88.taxon	description	(Figs. 1 – 5, Table 3) urn: lsid: zoobank. org: act: 6910 E 7 CD-BB 6 F- 4222 - A 9 E 2 - F 2 E 155 BE 3 C 59	en	Bianchi, Filipe Michels, Krein, Verônica, Rider, David, Grazia, Jocelia (2021): Finding Padaeus bovillus (Hemiptera: Pentatomidae): A Phylogenetic Placement and the Description of Its Sister Species. Zoological Studies 60 (11), No. 11: 1-13, DOI: 10.6620/ZS.2021.60-11, URL: http://dx.doi.org/10.5281/zenodo.12825774
622E6F09DD6FFFA5FC861178F3577E88.taxon	materials_examined	Material examined: Type Material: Holotype female: COSTA RICA: Tuis, Cartago, Terralba 650 m., A. Pittier, BRIT. MUS. (NHMUK) TYPE HEM / 050. COSTA RICA: 1 m #, Costa Rica, Prov. Heredia, F. La Selva, 3 km S Pto. Viejo, 10 ° 26 ' N 84 ° 01 ' W / 25. III. 1987, H. A. Hespenheide / D. A. Rider Collection (DARC); 1 f #, Costa Rica, Pr. Heredia, Puerto Viejo, Finca La Selva / R J. Marquis, coll. No. I, 11. IX. 1986 / Piper aricianum (?) / NMNH; 1 f #, Collection Schild-Burgdorf, Costa Rica, San Carlos / Padaeus bovillus Distant / NMNH; 1 m # / Costa Rica: Cartago Prov., Mon. Nac. Guayabo, 22. XII. 1994 / M. J. Tauber, C. A. Tauber, P. J. Tauber Collectors / UC Berkeley EMEC 1240641. PANAMA: 1 m #, 1 f #, Panama: Bocas Del Toro Pr. 3 km. n. Continental Div. on Fortuna Hwy 925 m., 13. VII. 1996, A. R. Gillogly / EGER / Sibaria n. sp. Det. J. E. Eger, 1997 (UFRG).	en	Bianchi, Filipe Michels, Krein, Verônica, Rider, David, Grazia, Jocelia (2021): Finding Padaeus bovillus (Hemiptera: Pentatomidae): A Phylogenetic Placement and the Description of Its Sister Species. Zoological Studies 60 (11), No. 11: 1-13, DOI: 10.6620/ZS.2021.60-11, URL: http://dx.doi.org/10.5281/zenodo.12825774
622E6F09DD6FFFA5FC861178F3577E88.taxon	diagnosis	Diagnosis: Mitripus bovillus may be distinguished from M. convergens and M. legionarius by the humeral angles which are developed laterally as stout spines; from Mitripus acutus by the clypeus slightly longer than mandibular plates, anterolateral margins of pronotum entirely smooth; and from Mitripus seclusus by the humeral angles depressed anteriorly and the shape of parameres (Fig. 3).	en	Bianchi, Filipe Michels, Krein, Verônica, Rider, David, Grazia, Jocelia (2021): Finding Padaeus bovillus (Hemiptera: Pentatomidae): A Phylogenetic Placement and the Description of Its Sister Species. Zoological Studies 60 (11), No. 11: 1-13, DOI: 10.6620/ZS.2021.60-11, URL: http://dx.doi.org/10.5281/zenodo.12825774
622E6F09DD6FFFA5FC861178F3577E88.taxon	description	Redescription: Coloration: Dorsal surface ochraceous with black punctures, giving an overall dark brown matte aspect; head and thorax with ochraceous punctures; antennae ochraceous, antennomeres I – II with irregular dark brown spots, antennomeres III – IV dark on apical 3 / 4, and antennomere V dark brown on apical half. Scutellum dark brown on base. Connexivum blackish, middle third ochraceous. Ventral surface yellowish-ochraceous; abdomen impunctate. Legs ochraceous with brown spots on femora and tibiae; apex of each tarsomere brownish. Head: Clypeus slightly longer than mandibular plates, rounded apically. Head tapering to apex, lateral margins of mandibular plates sinuous, concave near eyes. Ocelli red or yellow. Antennomere proportions: I <II <III <IV = V. Anterior margins of bucculae truncated, each with sharp projection, posterior margins evanescent. Rostrum reaching metacoxae. Thorax: Anterolateral margins of pronotum smooth on anterior half, anterolateral angles developed as small yellowish-ochraceous spines, projected laterally. Each humeral angle produced laterally as a black, stout spine, somewhat depressed anteriorly. Pronotal cicatrices brownish, with ochraceous spot posterior to mesial angles. Basal angles of scutellum with small fovea. Apex of radial vein with a small ivory dot at endocorium. Hemelytral membrane fumose, with veins subparallel. Evaporatoria each extending halfway from ostiole to metapleural lateral margin, and present on posterior margin of mesopleuron; surface impunctate, with gyrification near ostiole; lateral fold present; peritreme spout-like. Abdomen: Posterolateral angles of connexiva developed as tiny black spines. Posterolateral angles of urosternite VII slightly projected as spines. Spiracles concolorous with abdominal disc. Male genitalia: Pygophore (Fig. 3 K – M): in dorsal view, pygophore trapezoidal; genital cup not well exposed; posterolateral angles of pygophore developed as rounded projections; median projection of dorsal rim short. Dorsal rim interrupted by diagonal depressions flanking median projection of dorsal rim, lateral margins slightly sinuous, discontinuous near median projection of dorsal rim; superior process of dorsal rim exposed. Transverse ridge slightly concave; ventral rim concave with triangular projection medially. In posterior view, genital cup opening ellipsoid; dorsal rim concave, smooth; superior process of dorsal rim truncate, bladelike.; transverse ridge concave medially, U-shaped; inferior layer of ventral rim tumescent laterally, medial triangular projection bent inward into genital cup. In ventral view, ventral rim biconcave, triangular projection medially; posterolateral angles produced, rounded. Tenth segment: Posterior margin trapezoidal with setae; disc smooth; tubercles on basal third slightly developed. Parameres (Fig. 3 N – Q): In dorsal and ventral views, each with basal apodeme smaller than crown; stem as long as crown; crown bending outward nearly at a 45 ° angle. Basal process of paramere not developed, with a few setae; apical process of paramere enlarged basally, tapering to truncate apex. In mesial and lateral views, apical process of paramere stout, triangular, with scale-like structures on its outer surface. Phallus (Fig. 3 R – T): phallotheca tubular, slightly constricted basally; ventral basal processes of phallotheca quadrangular; dorsal processes of phallotheca hook-like, bent ventrally in about a right angle, not surpassing expanded conjunctiva; vesica process spout-like; conjunctiva lacking processes; ductus seminis distalis short, not exposed out of phallotheca. Female genitalia: In posteroventral view (Fig. 4 B), gonocoxites VIII with shallow punctures concolorous with ochraceous disc; surface flat; mesial margins concave, exposing a scletorized area of gonapophyses VIII, apex of each mesial margin squared, with brown spot. Laterotergites VIII with black margins; apex of each developed as a spine. Gonocoxites IX trapezoid, about three times wider than long, anterior, posterior and lateral margins straight. Laterotergites IX rounded apically, mesial margins forming a right angle, lateral margins convex, posterior margins projected, slightly surpassing tergite VIII; segment X rectangular. Internal genitalia: Ring sclerites elliptical; thickening of vaginal intima triangular posteriorly, rounded anteriorly; ductus receptaculi proximally slightly enlarged near vesicular area, shorter than vesicular area; median wall of vesicular area straight, slightly enlarged at apex; ductus receptaculi distally straight, shorter than pars intermedialis, with rounded dilation apically; proximal and distal annular flanges convergent; pars intermedialis convoluted, sclerotized basally, apical half straight, membranous; capsula seminalis thumb-like, dilated anteriorly.	en	Bianchi, Filipe Michels, Krein, Verônica, Rider, David, Grazia, Jocelia (2021): Finding Padaeus bovillus (Hemiptera: Pentatomidae): A Phylogenetic Placement and the Description of Its Sister Species. Zoological Studies 60 (11), No. 11: 1-13, DOI: 10.6620/ZS.2021.60-11, URL: http://dx.doi.org/10.5281/zenodo.12825774
622E6F09DD6FFFA5FC861178F3577E88.taxon	distribution	Distribution: Mitripus bovillus is distributed throughout southern Central America, with records from Costa Rica and Panama (Fig. 5).	en	Bianchi, Filipe Michels, Krein, Verônica, Rider, David, Grazia, Jocelia (2021): Finding Padaeus bovillus (Hemiptera: Pentatomidae): A Phylogenetic Placement and the Description of Its Sister Species. Zoological Studies 60 (11), No. 11: 1-13, DOI: 10.6620/ZS.2021.60-11, URL: http://dx.doi.org/10.5281/zenodo.12825774
622E6F09DD6FFFA5FC861178F3577E88.taxon	discussion	Comments: Distant (1900) originally speculated that Padaeus bovillus resembled Sibaria armata. More recently, Rolston (1976) called attention to the “ arcuately truncate termination of the bucculae well before the distal end of the first rostral segment ” that should remove P. bovillus from Padaeus and suggested that the species was close to Mormidea, and that further investigation was needed. A phylogenetic hypothesis previously placed Padaeus within Euschistus group, close to Proxys Spinola, although Padaeus bovillus was not sampled (Bar „ o et al. 2020).	en	Bianchi, Filipe Michels, Krein, Verônica, Rider, David, Grazia, Jocelia (2021): Finding Padaeus bovillus (Hemiptera: Pentatomidae): A Phylogenetic Placement and the Description of Its Sister Species. Zoological Studies 60 (11), No. 11: 1-13, DOI: 10.6620/ZS.2021.60-11, URL: http://dx.doi.org/10.5281/zenodo.12825774
