identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
E02A9F0DC67654DE84EEFF46F64562B3.text	E02A9F0DC67654DE84EEFF46F64562B3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomechusoides umbrosus Kjærby & Bek Craig & Thomsen & Hansen 2026	<div><p>Lomechusoides umbrosus sp. nov.</p><p>Figs 2 A, 2 C, 2 E, 2 F, 2 I, 2 K, 7 A</p><p>Type locality.</p><p>Hammarskog, Uppsala, Uppland, Sweden. 59.774586, 17.578827.</p><p>Material examined.</p><p>Holotype: Sweden • 1 ♂; Hammarskog, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.578827&amp;materialsCitation.latitude=59.774586" title="Search Plazi for locations around (long 17.578827/lat 59.774586)">Uppland</a>; 59.774586, 17.578827; 20. IX. 2023; leg. H. Kjærby; w. Formica polyctena; NHMD . Paratypes: Belarus • 4 ♂♂, 1 ♀; Vitebsk reg., Polotsk distr., 3 km SW of Novopolotsk, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=28.6318&amp;materialsCitation.latitude=55.5168" title="Search Plazi for locations around (long 28.6318/lat 55.5168)">near the lake Luechovo</a>, 55.5168, 28.6318, 152 m, in the nest of Formica polyctena, fir-wood, sorrel, 30. V. 2013, leg. E. Pliskevich; cSem • 1 ♂; Vitebsk reg., Polotsk distr., 3 km SW of Novopolotsk, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=28.6325&amp;materialsCitation.latitude=55.5167" title="Search Plazi for locations around (long 28.6325/lat 55.5167)">near the lake Luechovo</a>; 55.5167, 28.6325; 152 m; 10. VI. 2015; leg. E. Pliskevich; in the nest of Formica polyctena, fir-wood, sorrel; cSem . Russia • 2 ♀♀; Smolensk distr., near Vysokoye; 54.73, 32.21; 23. V. – 11. VI. 2022; leg. O. Semionenkov; window trap near dead spruce and nest of Formica polyctena, coniferous-broad-leaved forest; cSem • 1 ♂, 3 ♀♀; Svedlovsk Reg., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=59.7734&amp;materialsCitation.latitude=57.3734" title="Search Plazi for locations around (long 59.7734/lat 57.3734)">Visimskij Nature Res.</a>; 57.3734, 59.7734; 570 m; 18. VI. 2019; leg. A. Solodovnikov, A. K. Hansen, A. Tokareva; spruce dominated forest, in Formica nests; NHMD . Sweden • 1 ♀; Hacksta, Uppland; 59.55, 17.38; 30. IV. 2022; leg. H. - E. Wanntorp; w. Formica polyctena; cWann • 1 ♂; Hållvik, Uppland; 59.93, 18.41; 12. V. 2025; leg. H. - E. Wanntorp; cWann • 3 ♂♂, 4 ♀♀; Hammarskog, Uppland; 59.77, 17.57; 20. IX. 2023; leg. H. Kjærby; w. Formica polyctena; cKjær • 6 ♂♂, 3 ♀♀; Skåne, Vittsjö; 56.34, 13.66; Formica rufa nest; 24. V. 1980; leg. A. Carlson; MZLU .</p><p>Additional material.</p><p>Estonia • 1 ♂; Välgi; 58.55, 26.92; 18. V. – 7. VII. 2024; forest edge window trap on dead spruce; TAMZ . Finland • 1 ♀; Karelia australis, Joutseno, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=28.729&amp;materialsCitation.latitude=61.073" title="Search Plazi for locations around (long 28.729/lat 61.073)">Kuurmanpohja</a>; 61.073, 28.729; 71 m; 8. VI. 2015 – 11. VII. 2015; leg. Eero Helve; window trap, mushroomed log pile (aspen, spruce, birch); ZMUO . Poland • 1 ♀; Bialowieza, Czerlonka; 52.69, 23.72; 19. VI. 1991; leg. L. Borowiec; DBET .</p><p>Differential diagnosis.</p><p>The new species can be separated from all other members of the Lomechusoides strumosus group sensu Jászay et al. (2023) by the following combination of characters: 1) widest point of the pronotum positioned at ~ ¼ – ⅓ of the length of the pronotal lateral margin from the posterior corners, making the posterior corners distinctly obtuse. 2) pronotal lateral margins smoothly rounded in their entire length 3) a visible sharp margin on the posterior curved part of the pronotal margins (best viewed laterally and slightly from behind) (Fig. 2 G). 4) pronotal disc glossy with very limited, uneven microsculpture, but with very well-defined tubercles. 5) head, sternites and tergites predominantly black. 6) clypeus and frontal median impression of the head shiny with limited, uneven microsculpture. 7) two transverse carinae on the anterior part of visible tergites II to V connected in the centre of the tergite by a broad septum, occasionally two or more broad septa, rarely absent.</p><p>See Table 2 for a comprehensive list of identified traits that separate L. umbrosus sp. nov. from its close congener L. strumosus .</p><p>Description.</p><p>Female. Measurements (mm): TL 6.81; FL 3.07; AL 3.34; HW 0.91; HL 0.94; PW 1.99; PL 1.20; EW 2.24; EL 0.96; FTL 1.30; MTL 1.66; HTL 1.95. Ratios: HW / HL 0.96; PW / PL 1.66; EW / EL 2.33.</p><p>Male. Measurements (mm). TL 6.46; FL 2.88; AL 2.90; HW 0.84; HL 1.08; PW 1.72; PL 1.04; EW 2.07; EL 0.94; FTL 1.25; MTL 1.47; HTL 1.75. Ratios: HW / HL 0.78; PW / PL 1.65; EW / EL 2.2.</p><p>Colouration dark. Legs and antennae dark reddish-brown. Head entirely black. Pronotal disc deep bordeaux reddish-brown, with a semicircular blackish pattern along the posterior margin; the pronotal lateral margins are paler reddish-brown than the pronotal disc. Elytra almost entirely bright reddish-brown, blackish-brown along anterior margins. Thorax with sterna black. Tergites and sternites almost entirely black, each with a thin reddish-brown transverse stripe along the posterior margins.</p><p>Head rectangular, with deep V-shaped frontal median impression very weakly microsculptured, shiny, very sparsely and finely punctate (Fig. 2 H). A central spot at the hind part of the head without microsculpture. Clypeus finely and unevenly microsculptured, shiny. The rest of the head is evenly microsculptured and finely punctate, dull. Eyes slightly to strongly protuberant. Temples overall concave, usually being strongly narrowed immediately behind the eyes, subparallel posteriad.</p><p>Antennae (Fig. 2 D) relatively gracile and long (Fig. 3 B). Male with dense tufts of golden pubescence on antennal segments III – IV.</p><p>Male. Scape 1.52 × as long as wide, and 1.8 × as long as pedicel; pedicel 1.31 × as long as wide; antennomere III 1.89 × as long as wide; antennomere IV 1.4 × as long as wide; antennomeres V – VIII all ~ 1.5 × as long as wide; terminal antennomere slender and acute, 3.25 × as long as wide, 1.36 × as long as scape.</p><p>Female. Scape 1.6 × as long as wide, and 1.68 × as long as pedicel; pedicel 1.26 × as long as wide; antennomere III 1.86 × as long as wide; antennomere IV 1.41 × as long as wide; antennomeres V – VIII all ~ 1.4–1.6 × as long as wide; terminal antennomere slender and acute, 2.9 × as long as wide, 1.37 × as long as scape.</p><p>Pronotum (Fig. 2 J) with concave anterior margin; anterior corners obtuse; with visible microsculpture along anterior and posterior margins, and in depressions. Pronotal lateral margins are smoothly rounded, semicircular convex and with strong microsculpture. The widest point of the pronotum ~ ¼ – ⅓ of the lateral marginal length from the posterior corners. As the pronotum narrows from the widest point towards the anterior corners, the lateral margins are evenly rounded without angular bends. Behind the widest point, the pronotum narrows slightly posteriorly, making the posterior corners distinctly obtuse. This results in the pronotal lateral margins being smoothly rounded in their entire length.</p><p>In lateral view, the pronotal lateral margins are thick in the anterior half, then abruptly thinning in the posterior half. At posterior corners the margin is strongly curved and very thin with a visible sharp edge (Fig. 2 G). The length of sharp edge-bearing margin varies but is usually rather short and constrained to the curved part at posterior corners.</p><p>The pronotal lateral margins are separated from the pronotal disc by shallow longitudinal depressions with the lateral area with visible microsculpture. Pronotal disc (Fig. 2 J) with either very limited, unevenly distributed fine microsculpture, or no microsculpture at all, giving the central part of the pronotum a very glossy appearance. Pronotal disc with well-defined tubercles and median line. Pronotal tubercles are very noticeable due to the lack of surrounding microsculpture. Pronotal tubercles bearing long, thin erect golden pubescence. Lateral margins of pronotum with 5–7 black macrosetae, anterior margin with 1–4 black macrosetae.</p><p>Metaventral process microsculptured and sparsely punctate with concave anterior and lateral margins, possessing lateral lines that converge anteriorly. Anterior part with no / very weakly defined transverse carina, posterior part of median line lacking lentil-like depression. Metaventral disc microsculptured and sparsely punctate with weakly defined longitudinal shallow impressions on either side of the posterior part of the median line.</p><p>Elytra finely and evenly punctate and microsculptured. Each puncture bears a short golden hair. Anterior part bearing eight or nine black macrosetae, humeral part with two black macrosetae.</p><p>Abdomen with visible tergite I densely punctate and evenly microsculptured; II and III densely punctate along posterior margin, otherwise finely and sparsely punctate, finely, and unevenly microsculptured. Visible tergites IV and VII finely and sparsely punctate and finely and unevenly microsculptured; visible tergites V – VI equally finely and sparsely punctate in median part, lateral parts bearing slightly larger and denser punctures and more visible microsculpture. Visible tergites I – IV with dense golden trichomes laterally, visible tergites I – III additionally with shorter, golden trichomes along posterior margin, thinning towards the centre. Visible tergites V – VII without trichomes. Anterior part of visible tergites II – V bearing two transverse carinae. The two carinae are joined near the centre of the tergite by a septum (Fig. 2 L). Occasionally, two or more connecting septa may be present, and rarely they may be absent. Area around the two transverse carinae with wavy microsculpture. Posterior parts of visible tergites I – III, especially II, with numerous black macrosetae. Posterior margins of visible tergites IV – VII with fewer, smaller dark setae.</p><p>Sternites I – IV finely microsculptured and sparsely punctate, rather glossy. Sternite V and especially VI more strongly microsculptured and moderately punctate. Max distance between punctures on sternite VI is ~ 4 × the diameter of punctures. Sternites I – III with dense, flat-laying, golden trichomes laterally in the anterior part. All sternites with numerous black macrosetae.</p><p>Male genitalia.</p><p>Aedeagus (Fig. 4 B, D) with robust median lobe, apical lobe shorter than basal bulb. In lateral view, apex is thin, with a subapical expansion followed by a notch (Fig. 4 D). In lateral view, the entire dorsal margin of the chitinised part of the median lobe is concave. In dorsoventral view, apex is relatively thin, roughly shaped like an acute triangle, sides of median lobe softly concave. In ventral view, lateral sides of apical lobe narrow apically along entire length, most strongly narrowing in basal part. Apex of parameres with a short, backwards-curving tooth.</p><p>Female genitalia.</p><p>Spermatheca (Fig. 4 F) with basal and apical parts approximately equal in length; basal part 0.88–1.16 × the length of apical part; tip of apical part rounded. The entire spermatheca is uniformly dark brown.</p><p>Etymology.</p><p>The species epithet umbrosus, meaning shadowy in Latin (nominative masculine), refers both to the generally very dark colouration, and its habitat being nests of Formica polyctena, who are often found in relatively shaded forest environments.</p><p>Distribution.</p><p>Lomechusoides umbrosus sp. nov. is known from a few sites in Sweden, Finland, Estonia, Poland, Belarus, and Northwestern Russia (Fig. 5 A). In Sweden it is found in the region of Uppland on several sites between Stockholm and Uppsala (Hammarskog, Hacksta, and others), as well as Vittsjö in Scania in southernmost Sweden. In Finland it has been collected on a site in South Karelia. In Estonia it has been found near Tartu. In Poland it is known from Bialowieza Primeval Forest. In Belarus it has been collected near Vitebsk (Novopolotsk), and nearby in Russia it is known from Smolensk (Vysokoye). The second known Russian site is in the Urals (Visimskij Nature Reserve) and marks the easternmost known site for the species.</p><p>Bionomics.</p><p>Lomechusoides umbrosus sp. nov. is only known from the mounds of Formica polyctena (Fig. 6 A – D). On several occasions during both spring and autumn, individuals of L. umbrosus sp. nov. have been collected on the surface of the F. polyctena host anthills, sometimes whilst being carried by workers (HSK pers. obs.; Artportalen. se). On other occasions, L. umbrosus sp. nov. has been found by sifting nest material of F. polyctena (AKH pers. obs.; Artportalen. se).</p><p>Due to its association with F. polyctena, L. umbrosus sp. nov. is most commonly found in coniferous or mixed forests, often in rather shaded environments (Fig. 6 A). It is not strictly associated with this habitat, however. At Hacksta (Vallentuna, Uppland, Sweden), it has been recorded in a F. polyctena nest located on a forest-adjacent open pasture (Artportalen. se).</p><p>The myrmecophile behaviour of L. umbrosus sp. nov. has never been studied. However, it is thought to be similar to the closely related L. strumosus, that has been subject to many behavioural studies since the mid 1800 ’ s. Lomechusoides strumosus infiltrates host F. sanguinea nests through an elaborate adoption process using the pheromone-producing adoption glands located under the abdominal trichomes. Inside the nest, it lives as a socially integrated kleptoparasite, being fed by the host through trophallaxis. The beetle larvae are also fed through trophallaxis, but in addition to this also predate on ant brood. It is assumed that all L. strumosus - group species are similar to L. strumosus in the relationship to their respective Formica hosts. See Wasmann (1915) and Hölldobler et al. (2018) for detailed accounts of L. strumosus ’ myrmecophilous habits.</p><p>Notes.</p><p>The larva is unknown but can safely be assumed to be similar to the well-known larva of L. strumosus, as all known larvae of both Lomechusoides and Lomechusa are generally very similar (Wasmann 1915).</p><p>Detailed comparison.</p><p>Lomechusoides umbrosus sp. nov. belongs to the L. strumosus complex within the L. strumosus species group sensu Jászay et al. (2023).</p><p>Although L. umbrosus sp. nov. has until now been confused with L. strumosus, it does not resemble it particularly much in comparison to other Lomechusoides species. Lomechusoides umbrosus sp. nov. shares more characteristics with the East Siberian L. zerchei and L. primoricus than with the European L. strumosus, L. folgaricus Jászay, Hlaváč &amp; Baňař, 2023 and L. wellenii . These two East Siberian species, with L. sibiricus and L. zeyai, are the only other described members of the strumosus complex with a visible sharp edge on the posterior curved part of the pronotal lateral margins in lateral view. Lomechusoides zeyai is easily differentiated by its unique, strongly concave pronotal lateral margins and acute hind corners, a very different spermatheca, and smaller size (~ 5 mm). Lomechusoides sibiricus is easily differentiated by its acute hind corners of the pronotum, slightly concave lateral margins of the pronotum, and longer antennomeres (Jászay et al. 2023). However, L. zerchei and L. primoricus are more similar to L. umbrosus sp. nov.</p><p>Lomechusoides zerchei (Fig. 3 D) is the only described member of the L. strumosus complex besides L. umbrosus sp. nov. where the pronotum is widest well in front of the posterior corners and narrows significantly posteriorly. It is easily differentiated from L. umbrosus sp. nov. by its distinctly trapezoidal pronotum shape, with the lateral margins being diverging and straight in the part anterior of the widest point, by having a much lighter reddish-brown head, lacking lateral lines on the metaventral process, and the transverse carinae on anterior parts of tergites II – V being disconnected in their entire length. The host of the examined L. zerchei specimens is a species in the F. rufa group.</p><p>Lomechusoides primoricus (Fig. 3 C) is differentiated from L. umbrosus sp. nov. by having a dark brown head. Furthermore, it is differentiated by having 90 - degree posterior corners of the pronotum, and widest point of the pronotum being at the posterior corners. The pronotal lateral margins of L. primoricus are much more parallel (sometimes slightly concave posteriorly) and less rounded and convex than in L. umbrosus sp. nov., making the pronotum appear much more square in L. primoricus . The length of the sharp edge on the lateral pronotal margin is also much longer in L. primoricus, where it covers the posterior third of the lateral margin. In L. umbrosus sp. nov. it is shorter and usually constrained to the curved part near the posterior corners. The host of the examined L. primoricus specimens is F. sanguinea .</p><p>Lomechusoides zerchei, L. primoricus, L. sibiricus, and L. zeyai are only known from regions east of Lake Baikal (Jászay et al. 2023), and all are generally lighter in colouration than the very dark L. umbrosus sp. nov.</p><p>European species.</p><p>In Europe, described members of the L. strumosus complex consist of L. strumosus (much of Europe), L. wellenii (Fennoscandia), L. folgaricus (The Alps), L. rossii Jászay, Hlaváč &amp; Baňař, 2023 (Abruzzi), L. siculus Fiori, 1914 (Sicily), L. teres (Caucasus), L. dudkorum Jászay, Hlaváč &amp; Baňař, 2023 (Urals and eastwards) and L. richteri Jászay, Hlaváč &amp; Baňař, 2023 (Saratovsk, Volga). None of these species share pronotum shape with L. umbrosus sp. nov., and few overlap in range (Jászay et al. 2023). The Italian L. folgaricus and L. rossii share with L. umbrosus sp. nov. a glossy pronotal disc with very limited microsculpture but have either very weakly defined tubercles or entirely lack them. In L. umbrosus sp. nov., they are very well defined.</p><p>Northern European species.</p><p>In the Nordic Countries, L. strumosus (Figs 2 A, 2 C, 2 E, 2 F, 2 I, 2 K, 7 A) and L. wellenii (Fig. 3 B) of the strumosus complex, and L. inflatus (Fig. 3 A) of the inflatus complex are the only other known representatives of the genus. Lomechusoides umbrosus sp. nov. is a large species, equal in size to L. strumosus, whereas the other known species L. wellenii and L. inflatus are slightly smaller to much smaller, respectively. In general, L. umbrosus sp. nov. has a much darker appearance than other Nordic Lomechusoides species, with both tergites and sternites being predominantly black, each bearing a thin brown stripe at the posterior margins. This is very apparent in life, when the abdomen is held in an upright position when the beetle is at rest. The jet black, glossy sternites of L. umbrosus sp. nov. (Fig. 7 B) gives it a very different appearance than L. strumosus, who has a conspicuously vibrant deep red abdominal apex (Fig. 7 A). This colour difference matches the general colour difference between their specific host ants, with F. sanguinea being more brightly red than the darker F. rufa group. On museum specimens the colour difference is often less clear than on living specimens, and colour characteristics should always be used together with morphological characteristics.</p><p>Among Nordic members of the genus, the spermatheca of L. umbrosus sp. nov. is unique in its apical and basal parts being approximately equal in length (basal part 0.88–1.08 × the length of apical part) (Fig. 4 F), where the basal part is significantly longer than the apical part in L. strumosus (Fig. 4 E), and significantly shorter in L. wellenii and L. inflatus . The spermatheca is usually also darker than in L. strumosus, which is especially apparent in the basal part.</p><p>Comparative material examined.</p><p>Lomechusoides strumosus: Austria • 1 ♀; Hainburger Berge; 48.13, 16.93; leg. Scheerpeltz; ZMUH • 3 ♂♂, 5 ♀♀; Linz; 48.29, 14.31; 29. III. 1913; leg. J. Kloiber; NHMW • 2 ♂♂, 2 ♀♀; Marzer Kogel, n. Marz-Rohrbach; 47.72, 16.45; I. IV. 1926, leg. Scheerpeltz; NHMW [Pinned host: Formica sanguinea] • 5 ♂♂, 2 ♀♀; n. Graz; 47.07, 15.44; leg. Scheerpeltz; NHMW • 4 ♂♂, 6 ♀♀; Rekawinkel n. Vienna; 48.18, 16.03; leg. Scheerpeltz; NHMW [Pinned host: Formica sanguinea]. Bosnia and Herzegovina • 1 ♀; Nevesinje; 43.26, 18.11; leg. Schuster; NHMW. Czechia • 5 ♂♂, 4 ♀♀; Hannsdorf [Hanušovice]; 50.07, 16.93; leg. V. Zoufal; NHMW • 2 ♀♀; Jíloviště; 49.93, 14.34; 24. IV. 1910; leg. K. Klenka; SMNS • 2 ♂♂, 5 ♀♀; Písecké Hory; 49.28, 14.23; 1917; leg. Scheerpeltz; NHMW [Pinned host: Formica sanguinea] • 1 ♀, 1 ♂; Pollauer Berge [Vrchy Pálavské]; 48.86, 16.64; 18. – 19. IV. 1924; leg. Scheerpeltz; NHMW [Pinned host: Formica sanguinea] • 1 ♂; Příbram; 49.67, 13.99; leg. J. Roubal; NHMW • 1 ex.; Tabor; 49.41, 14.68; leg. Luse; NHMW • 4 ex.; Zvole n. Prague; 49.93, 14.42; leg. Skalitzky; NHMW. Denmark • 2 ♂♂, 1 ♀; Feldborg Nørreskov; 56.36, 8.94; 22. IV. 2025; leg. H. Kjærby; w. Formica sanguinea; cKjær • 1 ♀; Gludsted Plantage; 56.08, 9.36; 8. V. 2024; leg. H. Kjærby; w. Formica sanguinea; cKjær • 2 ♂♂, 2 ♀♀; Hønning Plantage; 55.18, 8.95; 11. V. 2025; leg. H. Kjærby; w. Formica sanguinea; cKjær • 1 ♀; Høstemark Skov; 56.93, 10.22; 17. VIII. 2020; leg. H. Kjærby; w. Formica sanguinea; cKjær • 2 ♂♂, 4 ♀♀; Kompedal Plantage; 56.23, 9.26; 9. IV. 2025; leg. H. Kjærby; w. Formica sanguinea; cKjær • 1 ♀; Melby Overdrev; 56.01, 11.99; 27. V. 2025; leg. H. Kjærby; w. Formica sanguinea; cMart • 1 ♂; Nørlund Plantage; 56.05, 9.21; 24. III. 2025; leg. H. Kjærby; w. Formica sanguinea; cKjær • 2 ♂♂; Nørlund Plantage; 56.05, 9.21; 3. IV. 2025; leg. H. Kjærby; w. Formica sanguinea; cKjær • 1 ♂; Store Hjøllund Plantage; 56.05, 9.37; 17. VIII. 2021; leg. H. Kjærby; w. Formica sanguinea; cKjær • 1 ♂, 1 ♀; Store Hjøllund Plantage; 56.05, 9.37; 18. IV. 2024; leg. H. Kjærby; w. Formica sanguinea; cKjær • 1 ♀; Store Hjøllund Plantage, 56.05, 9.37, 21. III. 2024, leg. H. Kjærby, w. Formica sanguinea; cKjær • 3 ♀♀; Store Hjøllund Plantage; 56.05, 9.37; 7. IV. 2024; leg. H. Kjærby; Formica sanguinea; cKjær • 1 ♀; Store Hjøllund Plantage; 56.05, 9.37; 11. VI. 2024; leg. H. Kjærby; Formica sanguinea; cKjær • 1 ♀; Tingskov; 57.08, 9.42; 22. IV. 2023; leg. H. Kjærby; Formica sanguinea; cKjær • 1 ♂, 6 ♀♀; Tisvilde; 56.05, 12.06; 31. V. 1908; leg. Rye; NHMA. Estonia • 1 ♀ Kõrkküla; 58.15, 22.37; 25. IV. – 6. VI. 2023; forest cut area window trap on aspen stand; TAMZ • 1 ♂; Aegviidu Pillapalu; 59.28, 25.61; 25. V. 1994; TAMZ. Finland • 1 ♂; Helsinge [Helsinki]; 60.23, 24.99; leg. Karvonen; MZLU • 1 ♂; Helsingfors; 60.23, 24.99; leg. O. Wellenius; MZLU • 1 ♀; Joutseno; 61.12, 28.51; leg. E. Thuneberg; MZLU • 1 ♂, 3 ♀♀; Nylandia, Helsinki; 60.23, 24.99; 24. IV. 1919; leg. Karvonen; ZMUO • 1 ♀; Pirkkala; 61.46, 23.64; leg. Grönblom; MZLU • 1 ♂, 2 ♀♀; Regio aboënsis, Raisio; 60.48, 22.16; 1940 ’ s; leg. R. Linnavuori. ZMUO • 1 ♂; Regio aboënsis, Rymättylä; 60.37, 21.94; 1940 ’ s; leg. R. Linnavuori. ZMUO • 1 ♂; Tampere; 61.49, 23.76; leg. Soyrinki; MZLU • 1 ♂, 1 ♀; Vehkalaliti; 60.57, 27.14; 2. VI. 1962; leg. L. Tiensuu; MZLU. France • 1 ♀; Cote d’Or; 47.50, 4.64; leg. Rouget; SMNS • 1 ♀; Hautes Pyrénées; 42.99, 0.14; leg. Pandellé; NHMW. Germany • 4 ♂♂, 4 ♀♀; Asselfingen; 48.53, 10.19; 13. V. 1952; SMNS • 1 ♀; Düsseldorf; 51.22, 6.78; leg. Dr. Friis; NHMW • 1 ♀; Eifel, 2 km ENE Schönecken; 50.17, 6.49; 7. IX. 1991; leg. Schulz; Clearing in Pine Forest; NHMW • 1 ♀; Hannover, Helstorf Reiterheide; 52.58, 9.61; 30. V. 1991; leg. V. Assing; NHMW • 1 ♂, 1 ♀; Kyffhauser; 51.11, 11.08; 2. V. 1941; leg. Folwaczny; SMNS • Peutenhausen; 48.52, 11.23; 23. III. 1974; leg. R Papperitz; SMNS • 2 ♂♂; Taubertal; 49.69, 9.63; 20. IV. 1981; leg. R. Bickel; SMNS. Greece • 1 ♂; Parnassus, 38.55, 22.66; NHMW [Pinned host: Formica sanguinea]. Italy • 1 ♀; Val di Laux; 45.04, 7.03; 10. VI. 1908; leg. Pinker; NHMW [Pinned host: Formica sanguinea] • 1 ♀; Valle di Saviore; 46.07, 10.43; VII. 1855; leg. Cadamuro; NHMW. Netherlands • 1 ♂, 1 ♀; Exaten; 52.18, 6.41; V. 1890; leg. Wasmann; NHMW [Pinned host: Formica sanguinea]. Norway • 1 ♀; Akershus, Vestby, Kroken; 59.56, 10.67; 9. V. 1996; leg. S. Ligaard; cØde • 1 ♀; Buskerud, Kongsberg; 59.66, 9.65; 22. IV. 1990; leg. B. Sagvolden; cØde • 1 ♀; Oppland, Nord fron Stordalsberge; 61.58, 9.81; 10. VII. 1988; leg. O. Hanssen; In nest of Formica sanguinea at the root of fallen birch; cHans • 1 ♂; Oppland, Nord fron Stordalsberge; 61.61, 9.83; 4. VI. – 3. IV. 1998; leg. F. Ødegaard; pitfall trap with Formica sanguinea; cØde • 1 ♂; Oslo, Sogn; 59.95, 10.74; 15. V. 1961; leg. J. Andersen; UiT. Russia • 1 ♀; Demidov distr., near Nikitenki; 55.49, 31.78; 22. – 26. VII. 2015; leg. O. Semionenkov; near a dead birch with Formica sanguinea; cSem • 1 ♀; Demidov distr., near Poboistche, SW part of “ Kolpitski Mokh ”; 55.46, 31.63; 1. – 5. VII. 2017; leg. O. Semionenkov; in nest of Formica uralensis, oligotrophic moor; cSem • 1 ♀; Ulyanovo Distr., “ Kaluzhskiye Zaseki ” Nature Reserve, Novaya Dyeryevnya; 53.59, 35.80; 10. – 20. VI. 2009; leg. S. Alekseev; window trap on a willow; cSem • 1 ♀; Vladimir area, Meshchera National Park, near Tasino; 55.52, 40.19; 13. VI. 2008; leg. V. B. Semenov; cSem. Slovakia • 3 ♂♂, 4 ♀♀; Nyitra County; 48.31, 18.08; leg. H. Raffenberg; NHMW. Sweden • 1 ♀; Dalarna, St. Kopparberg; 60.61, 15.63; leg. Klefbeck; MZLU • 2 ♀♀; Halland, Släp; 57.51, 11.98; leg. Agren; MZLU • 2 ♂♂, 1 ♀; Skåne, Hässleholm; 56.16, 13.77; 7. V. 1977; leg. F. Olsson; MZLU • 1 ♀; Skåne, Klostersågen; 55.64, 13.59: 15. V. 1982; leg. H. Andersson; MZLU • 1 ♀, 1 ♂; Skåne, Vankiva; 56.19, 13.74; 9. V. 1961; leg. F. Olsson; MZLU • 1 ♀, 1 ♂; Småland, Kalmar; 56.67, 16.32; 1. V. 1980; leg. B. Anderson; MZLU • 1 ♀; Västra Götaland, Kråkhult; 57.71, 13.09; 29. VIII. 1959; leg. H. - E. Wanntorp; cWann.</p><p>Lomechusoides wellenii: Finland • 1 ♀; Helsinge [Helsinki]; 60.23, 24.99; 30. IV. 1940; leg. O. Wellenius; w. Formica uralensis; NHMD • 1 ♀; Helsinge [Helsinki]; 30. IV. 1940; leg. O. Wellenius; w. Formica uralensis; ZMUO. Sweden • 1 ♂ Dalarna, Norr Oradtjärnberg; 60.95, 14.18; 2006; leg. H. - E. Wanntorp; det. H. Kjærby; cWann • 1 ♀; unknown locality, leg. Gyllenhall; det. H. Kjærby; NHMD.</p><p>Lomechusoides inflatus: Norway • 2 ♀♀; Tana, Torhop; 70.48, 27.98; 29. V. 2024; leg. A. Staverløkk; w. Formica gagatoides; cKjær.</p><p>Lomechusoides folgaricus: Italy • 3 ♂♂; Folgaria; 45.91, 11.18; leg. Winkler; NHMW • 1 ♂; Serraia; 46.13, 11.25; NHMW • 1 ♀; Colle St. Lucia; 46.45, 12.01; leg. Kaiser; NHMW.</p><p>Lomechusoides teres: Georgia • 3 ♂♂; Caucasus, Lomis Mts.; 41.87 43.25; 2100 m; leg. König, NHMW [Pinned host: Formica sanguinea]. Turkey • 1 ♂; Trabzon, Doğu Karadeniz Dağları, Maçka, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=39.6866&amp;materialsCitation.latitude=40.6285" title="Search Plazi for locations around (long 39.6866/lat 40.6285)">Fırınoba</a> plateau environs; 2200 m; 40.6285, 39.6866, 14. V. 2025; leg. A. Solodovnikov; alpine zone; under stones; NHMD.</p><p>Lomechusoides primoricus: Paratypes Russia • 1 ♂ 1 ♀; Primorskij kray [= Primorsky Kray], Sikhote-Alin, Meteorol. St., 28 km SE Chuguyevka; 43.59, 134.08; 900 m; 1. VI. 1993; leg. Zerche; SDEI [Pinned host: Formica sanguinea].</p><p>Lomechusoides zerchei: Paratypes Russia • 2 ♂♂ 2 ♀♀; Primorsky kray, Anisimovka (= Kangaus), Krinichnaya Mt., 70 km E Vladivostok; 43.07, 132.41; 500 m; 8. VI. 1993; leg. L. Zerche; SDEI [Pinned host: Formica rufa group].</p><p>Lomechusoides dudkorum: Holotype Russia • ♂; 6 км к югу от г. Тюмени [6 km k yugu ot g. Tyumeni (= 6 km to the south from town Tyumen)]; 57.07, 65.55; 3. IV. 1995; leg. A. et. R. Dudko; NHMD [Pinned host: Formica sanguinea] Paratypes • 2 ♀♀; 6 км к югу от г. Тюмени [= 6 km to the south from town Tyumen]; 57.07, 65.55; 3. IV. 1995; leg. A. et. R. Dudko; NHMD.</p><p>Lomechusoides mongolicus: Russia • 1 ♀; Buryatia Rep., Selenga R. 8 km N Tarbagataj; 51.68, 108.41; 28. V .. 2002; leg. A. Anistschenko; NHMD • 1 ♀; Даурия [Chita area], оз. Барун-Торей, протока Утыча на [Lake Barun-Torey, Utych Strait]; 50.02, 115.39; 1. VI. 2000; O. Корсун [O. Korsun] leg.; NHMD.</p><p>Lomechusoides fallax: Paratype Russia • 1 ♀; Primorsky kray, Anisimovka (= Kangaus), Krinichnaya Mt., 70 km E Vladivostok; 43.07, 132.41; 500 m; 8. VI. 1993; leg. L. Zerche; SDEI [Pinned host: Formica rufa group].</p></div>	https://treatment.plazi.org/id/E02A9F0DC67654DE84EEFF46F64562B3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Kjærby, Hjalte Sebastian;Bek Craig, Sean Birk;Thomsen, Philip Francis;Hansen, Aslak Kappel	Kjærby, Hjalte Sebastian, Bek Craig, Sean Birk, Thomsen, Philip Francis, Hansen, Aslak Kappel (2026): A new species of Lomechusoides (Coleoptera, Staphylinidae) from Formica polyctena anthills in Northeastern Europe. ZooKeys 1275: 229-257, DOI: 10.3897/zookeys.1275.182781
