identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
5C5987E2FFA8D94FFF40704AB8090DF6.text	5C5987E2FFA8D94FFF40704AB8090DF6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tillancoccus Ben-Dov 1989	<div><p>Tillancoccus Ben-Dov, 1989 .</p><p>Type species: Tillancoccus tillandsiae Ben-Dov 1989 .</p><p>Generic diagnosis. Body of adult female almost circular; dorsum becoming sclerotised and slightly convex at maturity. Anal cleft fused. Dorsal setae robust and either lanceolate or conical. Dorsal setae set in radial lines, with either 19 or 25 lines around body. Preopercular pores distinct, present in a medial band or in small groups extending from anal plates to thorax. Other dorsal pores of possibly two types present, one a microduct. Dorsal tubular ducts absent. Submarginal tubercles present or absent. Anal plates about as long as their combined widths, each plate with a discal seta. Anogenital fold with 2 setae on anterior margin and 6 or 7 on each lateral margin. Marginal setae either slender and pointed or of two sizes, one capitate. Each stigmatic cleft with 3 stigmatic spines, one much longer than other two. Ventral microducts minute, frequent, probably present throughout. Multilocular disc-pores with mainly either 6, 7 or 10 loculi, present in a large group on either side of genital opening. Spiracular disc-pores with 3-6 loculi in a narrow band between each spiracle and margin. Ventral tubular ducts absent. Antennae well developed, with 6 or 8 segments. Legs well developed; tibio-tarsal segmentation obscure or absent; tarsal digitules both narrow; claw digitules broader than tarsal digitules; claw without a denticle.</p><p>Note. Caballero et al. (2017), when describing their new species T. koreguajae, state that it has 2 types of dorsal setae and 2 types of marginal setae. This is likely to be incorrect. Tillancoccus tillandsiae Ben-Dov and T. mexicanus Ben-Dov, do not have these setae and nor does the new species described below. The ventral setae on the new species come close to the margin but are definitely all ventral and there are no small setose setae on the dorsum or on the actual margin. It is here considered likely that T. koreguajae is the same and that the setae described by Caballero et al. are in fact all ventral but appear to be dorsal because the specimen is very flattened. It is notable that, in Caballero et al. ’s description, the size of all these setae is the same. Caballero et al. also suggested (their Table 1) that dorsal and ventral microducts are absent from T. tillandsiae and T. mexicanus but, in fact, they are present (see Ben-Dov 1989 and Hodgson 1994 - although these two latter authors only reported one type of dorsal pore).</p></div>	https://treatment.plazi.org/id/5C5987E2FFA8D94FFF40704AB8090DF6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hodgson, Chris	Hodgson, Chris (2025): A new species of the soft scale insect genus Tillancoccus Ben-Dov (Hemiptera: Coccomorpha: Coccidae) on coconut palm in Trinidad. Zootaxa 5637 (1): 175-179, DOI: 10.11646/zootaxa.5637.1.9, URL: https://doi.org/10.11646/zootaxa.5637.1.9
5C5987E2FFA9D94DFF4074E8B9180AD7.text	5C5987E2FFA9D94DFF4074E8B9180AD7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tillancoccus bendovi Hodgson 2025	<div><p>Tillancoccus bendovi Hodgson sp. nov. (Fig. 1)</p><p>Material examined: One slide labelled: left label: TRINIDAD / coconut palm / CIE 5886-17304; right label: Tillancoccus / bendovi / Hodgson / Holotype ♀ (1/1 adf, good but with body partially split lengthwise) (NHML).</p><p>Unmounted material. Not seen.</p><p>Slide-mounted material. Body almost round, 2.75 mm long, 2.6 mm at widest; length of anal cleft (without anal plates) approximately 1/5 th of total body length. Probably slightly convex in life as derm split on mounting.</p><p>Dorsum. Derm fairly heavily sclerotised throughout but degree of sclerotisation varying as follows: (i) with a less sclerotised, fairly broad marginal band about 40‒100 µm wide; (ii) with 19 radial rays of less sclerotised derm extending medially from near margin, each reaching middle of dorsum which is also rather less sclerotised (these rays probably not clear on immature, unsclerotised adult females): with 3 rays on head and, on each side, 1 extending from each stigmatic cleft, 1 between clefts and 6 on abdomen; (iii) derm between radial rays with numerous small nodules throughout (Fig. 1a), these absent from each ray and from the marginal band; and (iv) a narrow area of stronger sclerotization around anal plates. Dorsal setae restricted to a line within each radial ray, each seta large, spinose and lanceolate, 23‒37 µm long and about 11 µm wide, with a sharp apex and narrow base, and a basal socket 11‒15 µm wide; with 4‒8 setae in each ray plus an occasional 1 on submargin; each basal socket with 3‒6 closely associated pores. Dorsal pores of 3 types present: preopercular pores, each round and 3.5‒5.5 µm wide, in a non-sclerotised area of derm 8‒12 µm wide; with 54 pores extending anteriorly from anal plates to about metathorax, more or less in two medial bands; and two types of smaller pores: (i) very small pores, probably dorsal microducts, represented by minute pale unsclerotised spots, each about 1 µm wide, in sparse loose groups or short radial lines within the marginal band, with a group either at marginal end of each ray or in a small group near margin between rays; and (ii) slightly larger pores represented by unsclerotised pale spots, each 1.5‒3.5 µm wide, which may be dorsal simple pores, present throughout, including medially and within rays (Fig. 1b), and in groups of 3‒6 associated with each dorsal setal socket but absent from marginal band. Submarginal tubercles present, each about 10 µm wide with a broad outer orifice; with 1 on each side of head, 1 on each side between clefts and 2 on each abdominal margin. Pocket-like sclerotizations possibly present submarginally but small and indistinct. Cribriform plate-like structures (Fig. 1c) present submedially, mainly on antero-lateral margin of each ray; each with 4‒8 celllike bodies in a cluster, with derm slightly more sclerotised around each group; with 10 on each side. Anal plates as long as their combined widths, each about 143 µm long and combined widths 143 µm; anterior margin slightly concave; apex slightly attenuated; each plate with a sclerotised posterior border and with an area of thickening underneath, running from each outer corner apparently to each anal cleft margin; each plate with a discal seta (missing) and perhaps 3 other small setae (obscure). Ano-genital fold with 2 setae on anterior margin, each outer seta 23 µm long, each inner seta 11‒12 µm long; anterior margins of each anal plate apodeme without setae but with 6 or 7 setae along each posterior margin. Anal ring with 8 setae. Eyespots present just dorsad to margin, each about 22 µm wide.</p><p>Margin. Smooth, not crenulated. Marginal setae of 2 types present: (i) a larger seta with a capitate apex, each slightly curved and 28‒30 µm long, and (ii) a smaller, narrow seta with a blunt apex and parallel sides, mainly straight, each 10‒12 µm long; with 11 large setae and 12 smaller setae on head between stigmatic clefts, and, on each side, 4‒6 large setae and 5 small setae between clefts and 15 or 16 large setae and 17 or 18 small setae on abdomen. Stigmatic clefts each distinct and strongly indented but somewhat asymmetrical, each with 3 stigmatic spines clearly set on posterior margin of cleft: 2 small, rather parallel-sided, spines with a blunt apex and 1 much larger stout, but more sharply pointed, and set further back from cleft; larger spines difficult to measure as mostly vertical but perhaps 25 µm long, each smaller spine about 5 µm long.</p><p>Venter. Derm membranous. Multilocular disc-pores present in abdominal segment VII only, each with mainly 10 loculi; with a group of about 70 on either side of anogenital area. Spiracular disc-pores present in narrow band 1‒2 pores wide between each spiracle and margin; each pore small with 3‒5 loculi (3 loculi quite common); with 19‒27 pores in each band. Ventral microducts very small, each perhaps 1.0‒1.5 µm wide, probably present throughout. Setae fairly sparse; only 1 pair of pregenital setae present, on abdominal segment VII, each seta about 40 µm long; with 2 pairs of inter-antennal setae: each pair with a long seta perhaps 50 µm long and a small setae about 7 µm long; submarginal setae very small and sparse, many rather close to margin; those close to margin 3‒5 µm long, those situated more medially up to 8 µm long. Antennae 6 segmented, third segment much the longest; total length 225‒235 µm; setal distribution showing nothing distinctive. Clypeolabral shield about 126 µm long; clypeal setae present. Spiracles: anterior pair each 33‒35 µm wide, posterior pair each about 40 µm wide. Legs well developed, each without a tibio-tarsal articulation and with tibio-tarsal segmentation absent or unclear; lengths (in µm) of metathoracic legs: coxa 122‒138, trochanter + femur 185‒195, tibia 113‒118, tarsus 88; setae very few; claw 20‒22 µm long, without a denticle; tarsal digitules both narrow, claw digitules both with broad apices.</p><p>Comments. Three species of Tillancoccus were known prior to this study, two on Tillandsia sp. ( Bromeliaceae), namely T. tillandsiae Ben-Dov from Guatemala, Honduras and Mexico and T. mexicanus Ben-Dov from Guatemala and Mexico. The third species, T. koreguajae Caballero &amp; Ramos, is known only from Colombia on sugarcane ( Poaceae). The new species, T. bendovi Hodgson, was on coconut palm ( Cocos nucifera) from Trinidad. The genus appears to be restricted to the geographic area around the Caribbean.</p><p>Name derivation. The new species name, bendovi, is in recognition of Dr Yair Ben-Dov, the Volcani Center, Bet Dagan, Israel, who introduced this genus and is an acknowledged authority on Coccomorpha from southern Africa and the Middle East. The four species can be separated by the following key.</p></div>	https://treatment.plazi.org/id/5C5987E2FFA9D94DFF4074E8B9180AD7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hodgson, Chris	Hodgson, Chris (2025): A new species of the soft scale insect genus Tillancoccus Ben-Dov (Hemiptera: Coccomorpha: Coccidae) on coconut palm in Trinidad. Zootaxa 5637 (1): 175-179, DOI: 10.11646/zootaxa.5637.1.9, URL: https://doi.org/10.11646/zootaxa.5637.1.9
5C5987E2FFABD94DFF4073C9B8C708E6.text	5C5987E2FFABD94DFF4073C9B8C708E6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tillancoccus Ben-Dov 1989	<div><p>Key to adult female Tillancoccus species, based on adult female morphology</p><p>1. Submarginal tubercles present on dorsum. Dorsal setae lanceolate, each with pores associated with their setal socket...... 2</p><p>- Submarginal tubercles absent from dorsum. Dorsal setae not lanceolate but conical, without pores associated with their setal sockets............................................................................................. 3</p><p>2. With 2 sizes of marginal setae, one capitate and the other smaller and parallel-sided with a blunt apex. Multilocular disc-pores each with mainly 10 loculi. Each stigmatic cleft relatively deep, with stigmatic spines placed on posterior border....................................................................................... T. bendovi Hodgson sp. nov.</p><p>- With only 1 size of marginal setae, each sharply spinose. Multilocular disc-pores each with mainly 7 loculi. Each stigmatic cleft shallow with stigmatic spines placed centrally............................................ T. mexicanus Ben-Dov.</p><p>3. Antennae 8 segmented. With about 25 radial lines of dorsal setae.................... T. koreguajae Caballero &amp; Ramos.</p><p>- Antennae 6 segmented. With 19 radial lines of with dorsal setae.............................. T. tillandsiae Ben-Dov.</p></div>	https://treatment.plazi.org/id/5C5987E2FFABD94DFF4073C9B8C708E6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hodgson, Chris	Hodgson, Chris (2025): A new species of the soft scale insect genus Tillancoccus Ben-Dov (Hemiptera: Coccomorpha: Coccidae) on coconut palm in Trinidad. Zootaxa 5637 (1): 175-179, DOI: 10.11646/zootaxa.5637.1.9, URL: https://doi.org/10.11646/zootaxa.5637.1.9
