taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
5F7987D6FFB5B070FF51F111FAE367AA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226438/files/figure.png	https://doi.org/10.5281/zenodo.21226438	FIGURE 1. A–E, Epsilon achterbergi Giordani Soika, female paratype. F–J, Epsilon bertschi sp. nov., female holotype. A, F, habitus, lateral view. B, G, head, frontal view. C, H, mesosoma, dorsolateral view. D, I, head and mesosoma, lateral view. E, J, T1–T2, dorsal view.	FIGURE 1. A–E, Epsilon achterbergi Giordani Soika, female paratype. F–J, Epsilon bertschi sp. nov., female holotype. A, F, habitus, lateral view. B, G, head, frontal view. C, H, mesosoma, dorsolateral view. D, I, head and mesosoma, lateral view. E, J, T1–T2, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFB5B070FF51F111FAE367AA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226478/files/figure.png	https://doi.org/10.5281/zenodo.21226478	FIGURE 13. Distribution maps of Epsilon species in the Malay Archipelago.	FIGURE 13. Distribution maps of Epsilon species in the Malay Archipelago.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFB5B075FF51F685FB0867CA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226438/files/figure.png	https://doi.org/10.5281/zenodo.21226438	FIGURE 1. A–E, Epsilon achterbergi Giordani Soika, female paratype. F–J, Epsilon bertschi sp. nov., female holotype. A, F, habitus, lateral view. B, G, head, frontal view. C, H, mesosoma, dorsolateral view. D, I, head and mesosoma, lateral view. E, J, T1–T2, dorsal view.	FIGURE 1. A–E, Epsilon achterbergi Giordani Soika, female paratype. F–J, Epsilon bertschi sp. nov., female holotype. A, F, habitus, lateral view. B, G, head, frontal view. C, H, mesosoma, dorsolateral view. D, I, head and mesosoma, lateral view. E, J, T1–T2, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFB5B075FF51F685FB0867CA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226452/files/figure.png	https://doi.org/10.5281/zenodo.21226452	FIGURE 6. Epsilon latefasciatum sp. nov. A–B, male holotype. C–E, female. A, habitus, lateral view. B–C, head, frontal view. D, mesosoma, dorsal view. E, T1–T2, dorsal view.	FIGURE 6. Epsilon latefasciatum sp. nov. A–B, male holotype. C–E, female. A, habitus, lateral view. B–C, head, frontal view. D, mesosoma, dorsal view. E, T1–T2, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFB5B075FF51F685FB0867CA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226454/files/figure.png	https://doi.org/10.5281/zenodo.21226454	FIGURE 7. Epsilon manifestum (Smith). A, C–D, F, male. B, E, G–H, female. D, F, anomalous specimens. A, habitus of holotype, lateral view (credits: James Hogan, OUMNH). B–D, head, frontal view. E–F, mesosoma, dorsal view. G, T1–T2, dorsal view. H, apex of T2, lateral view.	FIGURE 7. Epsilon manifestum (Smith). A, C–D, F, male. B, E, G–H, female. D, F, anomalous specimens. A, habitus of holotype, lateral view (credits: James Hogan, OUMNH). B–D, head, frontal view. E–F, mesosoma, dorsal view. G, T1–T2, dorsal view. H, apex of T2, lateral view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFB5B075FF51F685FB0867CA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226478/files/figure.png	https://doi.org/10.5281/zenodo.21226478	FIGURE 13. Distribution maps of Epsilon species in the Malay Archipelago.	FIGURE 13. Distribution maps of Epsilon species in the Malay Archipelago.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFB0B075FF51F778FDF06469.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226441/files/figure.png	https://doi.org/10.5281/zenodo.21226441	FIGURE 2. Epsilon crassipunctatum Gusenleitner, stat. nov., female paratype. A, habitus, lateral view. B, head, frontal view. C, head, dorsal view. D, mesosoma, dorsal view. E, T1–T2, dorsal view.	FIGURE 2. Epsilon crassipunctatum Gusenleitner, stat. nov., female paratype. A, habitus, lateral view. B, head, frontal view. C, head, dorsal view. D, mesosoma, dorsal view. E, T1–T2, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFB0B075FF51F778FDF06469.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226478/files/figure.png	https://doi.org/10.5281/zenodo.21226478	FIGURE 13. Distribution maps of Epsilon species in the Malay Archipelago.	FIGURE 13. Distribution maps of Epsilon species in the Malay Archipelago.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFB1B076FF51F762FD94618A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226443/files/figure.png	https://doi.org/10.5281/zenodo.21226443	FIGURE 3. Epsilon dyscherum (de Saussure). A–B, D–E, female syntype of Odynerus chartergoides von Schulthess. C, nontypical male. F–H, non-typical female. A, habitus, lateral view. B–C, head, frontal view. D, head, dorsal view. E, mesosoma, dorsal view. F, T1–T2 from Luzon, dorsal view. G, T1–T2 from Mindanao, dorsal view. H, T1–T2 from Tawi-Tawi, dorsal view.	FIGURE 3. Epsilon dyscherum (de Saussure). A–B, D–E, female syntype of Odynerus chartergoides von Schulthess. C, nontypical male. F–H, non-typical female. A, habitus, lateral view. B–C, head, frontal view. D, head, dorsal view. E, mesosoma, dorsal view. F, T1–T2 from Luzon, dorsal view. G, T1–T2 from Mindanao, dorsal view. H, T1–T2 from Tawi-Tawi, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFB1B076FF51F762FD94618A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226476/files/figure.png	https://doi.org/10.5281/zenodo.21226476	FIGURE 12. Male genitalia of Epsilon species; from left to right: aedeagus in ventral view, aedeagus in lateral view, digitus+volsella. A, Epsilon dyscherum (de Saussure). B, Epsilon latefasciatum sp. nov. C, Epsilon manifestum (Smith). D, Epsilon rufipes Selis. E, Epsilon tigrinum sp. nov.	FIGURE 12. Male genitalia of Epsilon species; from left to right: aedeagus in ventral view, aedeagus in lateral view, digitus+volsella. A, Epsilon dyscherum (de Saussure). B, Epsilon latefasciatum sp. nov. C, Epsilon manifestum (Smith). D, Epsilon rufipes Selis. E, Epsilon tigrinum sp. nov.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFB1B076FF51F762FD94618A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226478/files/figure.png	https://doi.org/10.5281/zenodo.21226478	FIGURE 13. Distribution maps of Epsilon species in the Malay Archipelago.	FIGURE 13. Distribution maps of Epsilon species in the Malay Archipelago.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFB3B076FF51F0A5FAC760D2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226445/files/figure.png	https://doi.org/10.5281/zenodo.21226445	FIGURE 4. Epsilon incola Giordani Soika, female holotype. A, habitus, lateral view. B, head, frontal view. C, head, dorsal view. D, mesosoma, dorsolateral view. E, T1–T2, dorsal view.	FIGURE 4. Epsilon incola Giordani Soika, female holotype. A, habitus, lateral view. B, head, frontal view. C, head, dorsal view. D, mesosoma, dorsolateral view. E, T1–T2, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFB3B076FF51F0A5FAC760D2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226478/files/figure.png	https://doi.org/10.5281/zenodo.21226478	FIGURE 13. Distribution maps of Epsilon species in the Malay Archipelago.	FIGURE 13. Distribution maps of Epsilon species in the Malay Archipelago.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFBCB079FF51F2F9F86B6484.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226450/files/figure.png	https://doi.org/10.5281/zenodo.21226450	FIGURE 5. Epsilon laboriosum (Smith). A–B, male holotype (credits: James Hogan, OUMNH). C–H, female. A, habitus, lateral view. B–C, head, frontal view. D, scutellum and metanotum, lateral view. E, scutellum and metanotum, dorsal view. F, mid and hind legs, lateral view. G, T1–T2 from New Guinea, dorsal view. H, T1–T2 from Bismarck Archipelago, dorsal view.	FIGURE 5. Epsilon laboriosum (Smith). A–B, male holotype (credits: James Hogan, OUMNH). C–H, female. A, habitus, lateral view. B–C, head, frontal view. D, scutellum and metanotum, lateral view. E, scutellum and metanotum, dorsal view. F, mid and hind legs, lateral view. G, T1–T2 from New Guinea, dorsal view. H, T1–T2 from Bismarck Archipelago, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFBCB079FF51F2F9F86B6484.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226478/files/figure.png	https://doi.org/10.5281/zenodo.21226478	FIGURE 13. Distribution maps of Epsilon species in the Malay Archipelago.	FIGURE 13. Distribution maps of Epsilon species in the Malay Archipelago.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFBDB07DFF51F4A3FAD162FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226452/files/figure.png	https://doi.org/10.5281/zenodo.21226452	FIGURE 6. Epsilon latefasciatum sp. nov. A–B, male holotype. C–E, female. A, habitus, lateral view. B–C, head, frontal view. D, mesosoma, dorsal view. E, T1–T2, dorsal view.	FIGURE 6. Epsilon latefasciatum sp. nov. A–B, male holotype. C–E, female. A, habitus, lateral view. B–C, head, frontal view. D, mesosoma, dorsal view. E, T1–T2, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFBDB07DFF51F4A3FAD162FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226476/files/figure.png	https://doi.org/10.5281/zenodo.21226476	FIGURE 12. Male genitalia of Epsilon species; from left to right: aedeagus in ventral view, aedeagus in lateral view, digitus+volsella. A, Epsilon dyscherum (de Saussure). B, Epsilon latefasciatum sp. nov. C, Epsilon manifestum (Smith). D, Epsilon rufipes Selis. E, Epsilon tigrinum sp. nov.	FIGURE 12. Male genitalia of Epsilon species; from left to right: aedeagus in ventral view, aedeagus in lateral view, digitus+volsella. A, Epsilon dyscherum (de Saussure). B, Epsilon latefasciatum sp. nov. C, Epsilon manifestum (Smith). D, Epsilon rufipes Selis. E, Epsilon tigrinum sp. nov.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFBDB07DFF51F4A3FAD162FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226438/files/figure.png	https://doi.org/10.5281/zenodo.21226438	FIGURE 1. A–E, Epsilon achterbergi Giordani Soika, female paratype. F–J, Epsilon bertschi sp. nov., female holotype. A, F, habitus, lateral view. B, G, head, frontal view. C, H, mesosoma, dorsolateral view. D, I, head and mesosoma, lateral view. E, J, T1–T2, dorsal view.	FIGURE 1. A–E, Epsilon achterbergi Giordani Soika, female paratype. F–J, Epsilon bertschi sp. nov., female holotype. A, F, habitus, lateral view. B, G, head, frontal view. C, H, mesosoma, dorsolateral view. D, I, head and mesosoma, lateral view. E, J, T1–T2, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFBDB07DFF51F4A3FAD162FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226445/files/figure.png	https://doi.org/10.5281/zenodo.21226445	FIGURE 4. Epsilon incola Giordani Soika, female holotype. A, habitus, lateral view. B, head, frontal view. C, head, dorsal view. D, mesosoma, dorsolateral view. E, T1–T2, dorsal view.	FIGURE 4. Epsilon incola Giordani Soika, female holotype. A, habitus, lateral view. B, head, frontal view. C, head, dorsal view. D, mesosoma, dorsolateral view. E, T1–T2, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFBDB07DFF51F4A3FAD162FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226478/files/figure.png	https://doi.org/10.5281/zenodo.21226478	FIGURE 13. Distribution maps of Epsilon species in the Malay Archipelago.	FIGURE 13. Distribution maps of Epsilon species in the Malay Archipelago.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFB8B07FFF51F40DF882618A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226454/files/figure.png	https://doi.org/10.5281/zenodo.21226454	FIGURE 7. Epsilon manifestum (Smith). A, C–D, F, male. B, E, G–H, female. D, F, anomalous specimens. A, habitus of holotype, lateral view (credits: James Hogan, OUMNH). B–D, head, frontal view. E–F, mesosoma, dorsal view. G, T1–T2, dorsal view. H, apex of T2, lateral view.	FIGURE 7. Epsilon manifestum (Smith). A, C–D, F, male. B, E, G–H, female. D, F, anomalous specimens. A, habitus of holotype, lateral view (credits: James Hogan, OUMNH). B–D, head, frontal view. E–F, mesosoma, dorsal view. G, T1–T2, dorsal view. H, apex of T2, lateral view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFB8B07FFF51F40DF882618A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226476/files/figure.png	https://doi.org/10.5281/zenodo.21226476	FIGURE 12. Male genitalia of Epsilon species; from left to right: aedeagus in ventral view, aedeagus in lateral view, digitus+volsella. A, Epsilon dyscherum (de Saussure). B, Epsilon latefasciatum sp. nov. C, Epsilon manifestum (Smith). D, Epsilon rufipes Selis. E, Epsilon tigrinum sp. nov.	FIGURE 12. Male genitalia of Epsilon species; from left to right: aedeagus in ventral view, aedeagus in lateral view, digitus+volsella. A, Epsilon dyscherum (de Saussure). B, Epsilon latefasciatum sp. nov. C, Epsilon manifestum (Smith). D, Epsilon rufipes Selis. E, Epsilon tigrinum sp. nov.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFB8B07FFF51F40DF882618A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226478/files/figure.png	https://doi.org/10.5281/zenodo.21226478	FIGURE 13. Distribution maps of Epsilon species in the Malay Archipelago.	FIGURE 13. Distribution maps of Epsilon species in the Malay Archipelago.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFBAB060FF51F0B8FB4E618A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226457/files/figure.png	https://doi.org/10.5281/zenodo.21226457	FIGURE 8. Epsilon perforatum sp. nov., female holotype. A, habitus, lateral view. B, head, frontal view. C, mesosoma, dorsal view. D, T1–T2, dorsal view. E, S2, lateral view. F, S2, ventral view.	FIGURE 8. Epsilon perforatum sp. nov., female holotype. A, habitus, lateral view. B, head, frontal view. C, mesosoma, dorsal view. D, T1–T2, dorsal view. E, S2, lateral view. F, S2, ventral view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFBAB060FF51F0B8FB4E618A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226470/files/figure.png	https://doi.org/10.5281/zenodo.21226470	FIGURE 11. Epsilon vechti Giordani Soika, female paratype. A, habitus, lateral view. B, head, frontal view. C, vertex, dorsal view. D, head, lateral view. E, submarginal carina and propodeal valvula, lateral view. F, T1–T2, dorsal view.	FIGURE 11. Epsilon vechti Giordani Soika, female paratype. A, habitus, lateral view. B, head, frontal view. C, vertex, dorsal view. D, head, lateral view. E, submarginal carina and propodeal valvula, lateral view. F, T1–T2, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFBAB060FF51F0B8FB4E618A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226478/files/figure.png	https://doi.org/10.5281/zenodo.21226478	FIGURE 13. Distribution maps of Epsilon species in the Malay Archipelago.	FIGURE 13. Distribution maps of Epsilon species in the Malay Archipelago.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFA5B062FF51F0B8FDD763F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226459/files/figure.png	https://doi.org/10.5281/zenodo.21226459	FIGURE 9. Epsilon rufipes Selis. A–B, E, H, female holotype from Aru. C, G, I, female from New Guinea. D, F, male from New Guinea. A, habitus, lateral view. B–D, head, frontal view. E, mesosoma, lateral view. F, scutellum, posterior view. G, mid and hind legs, lateral view. H–I, T1–T2, dorsal view.	FIGURE 9. Epsilon rufipes Selis. A–B, E, H, female holotype from Aru. C, G, I, female from New Guinea. D, F, male from New Guinea. A, habitus, lateral view. B–D, head, frontal view. E, mesosoma, lateral view. F, scutellum, posterior view. G, mid and hind legs, lateral view. H–I, T1–T2, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFA5B062FF51F0B8FDD763F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226476/files/figure.png	https://doi.org/10.5281/zenodo.21226476	FIGURE 12. Male genitalia of Epsilon species; from left to right: aedeagus in ventral view, aedeagus in lateral view, digitus+volsella. A, Epsilon dyscherum (de Saussure). B, Epsilon latefasciatum sp. nov. C, Epsilon manifestum (Smith). D, Epsilon rufipes Selis. E, Epsilon tigrinum sp. nov.	FIGURE 12. Male genitalia of Epsilon species; from left to right: aedeagus in ventral view, aedeagus in lateral view, digitus+volsella. A, Epsilon dyscherum (de Saussure). B, Epsilon latefasciatum sp. nov. C, Epsilon manifestum (Smith). D, Epsilon rufipes Selis. E, Epsilon tigrinum sp. nov.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFA5B062FF51F0B8FDD763F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226478/files/figure.png	https://doi.org/10.5281/zenodo.21226478	FIGURE 13. Distribution maps of Epsilon species in the Malay Archipelago.	FIGURE 13. Distribution maps of Epsilon species in the Malay Archipelago.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFA7B064FF51F324FA7664B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226465/files/figure.png	https://doi.org/10.5281/zenodo.21226465	FIGURE 10. Epsilon tigrinum sp. nov. A–B, D–E, G, female holotype. C, male paratype. F, female paratype. A, habitus, lateral view. B–C, head, frontal view. D, scutellum and metanotum, lateral view. E, scutellum and metanotum, dorsal view. F, legs and metasoma, lateral view. G, T1–T2, dorsal view.	FIGURE 10. Epsilon tigrinum sp. nov. A–B, D–E, G, female holotype. C, male paratype. F, female paratype. A, habitus, lateral view. B–C, head, frontal view. D, scutellum and metanotum, lateral view. E, scutellum and metanotum, dorsal view. F, legs and metasoma, lateral view. G, T1–T2, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFA7B064FF51F324FA7664B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226476/files/figure.png	https://doi.org/10.5281/zenodo.21226476	FIGURE 12. Male genitalia of Epsilon species; from left to right: aedeagus in ventral view, aedeagus in lateral view, digitus+volsella. A, Epsilon dyscherum (de Saussure). B, Epsilon latefasciatum sp. nov. C, Epsilon manifestum (Smith). D, Epsilon rufipes Selis. E, Epsilon tigrinum sp. nov.	FIGURE 12. Male genitalia of Epsilon species; from left to right: aedeagus in ventral view, aedeagus in lateral view, digitus+volsella. A, Epsilon dyscherum (de Saussure). B, Epsilon latefasciatum sp. nov. C, Epsilon manifestum (Smith). D, Epsilon rufipes Selis. E, Epsilon tigrinum sp. nov.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFA7B064FF51F324FA7664B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226441/files/figure.png	https://doi.org/10.5281/zenodo.21226441	FIGURE 2. Epsilon crassipunctatum Gusenleitner, stat. nov., female paratype. A, habitus, lateral view. B, head, frontal view. C, head, dorsal view. D, mesosoma, dorsal view. E, T1–T2, dorsal view.	FIGURE 2. Epsilon crassipunctatum Gusenleitner, stat. nov., female paratype. A, habitus, lateral view. B, head, frontal view. C, head, dorsal view. D, mesosoma, dorsal view. E, T1–T2, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFA7B064FF51F324FA7664B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226443/files/figure.png	https://doi.org/10.5281/zenodo.21226443	FIGURE 3. Epsilon dyscherum (de Saussure). A–B, D–E, female syntype of Odynerus chartergoides von Schulthess. C, nontypical male. F–H, non-typical female. A, habitus, lateral view. B–C, head, frontal view. D, head, dorsal view. E, mesosoma, dorsal view. F, T1–T2 from Luzon, dorsal view. G, T1–T2 from Mindanao, dorsal view. H, T1–T2 from Tawi-Tawi, dorsal view.	FIGURE 3. Epsilon dyscherum (de Saussure). A–B, D–E, female syntype of Odynerus chartergoides von Schulthess. C, nontypical male. F–H, non-typical female. A, habitus, lateral view. B–C, head, frontal view. D, head, dorsal view. E, mesosoma, dorsal view. F, T1–T2 from Luzon, dorsal view. G, T1–T2 from Mindanao, dorsal view. H, T1–T2 from Tawi-Tawi, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFA7B064FF51F324FA7664B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226450/files/figure.png	https://doi.org/10.5281/zenodo.21226450	FIGURE 5. Epsilon laboriosum (Smith). A–B, male holotype (credits: James Hogan, OUMNH). C–H, female. A, habitus, lateral view. B–C, head, frontal view. D, scutellum and metanotum, lateral view. E, scutellum and metanotum, dorsal view. F, mid and hind legs, lateral view. G, T1–T2 from New Guinea, dorsal view. H, T1–T2 from Bismarck Archipelago, dorsal view.	FIGURE 5. Epsilon laboriosum (Smith). A–B, male holotype (credits: James Hogan, OUMNH). C–H, female. A, habitus, lateral view. B–C, head, frontal view. D, scutellum and metanotum, lateral view. E, scutellum and metanotum, dorsal view. F, mid and hind legs, lateral view. G, T1–T2 from New Guinea, dorsal view. H, T1–T2 from Bismarck Archipelago, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFA7B064FF51F324FA7664B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226459/files/figure.png	https://doi.org/10.5281/zenodo.21226459	FIGURE 9. Epsilon rufipes Selis. A–B, E, H, female holotype from Aru. C, G, I, female from New Guinea. D, F, male from New Guinea. A, habitus, lateral view. B–D, head, frontal view. E, mesosoma, lateral view. F, scutellum, posterior view. G, mid and hind legs, lateral view. H–I, T1–T2, dorsal view.	FIGURE 9. Epsilon rufipes Selis. A–B, E, H, female holotype from Aru. C, G, I, female from New Guinea. D, F, male from New Guinea. A, habitus, lateral view. B–D, head, frontal view. E, mesosoma, lateral view. F, scutellum, posterior view. G, mid and hind legs, lateral view. H–I, T1–T2, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFA7B064FF51F324FA7664B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226478/files/figure.png	https://doi.org/10.5281/zenodo.21226478	FIGURE 13. Distribution maps of Epsilon species in the Malay Archipelago.	FIGURE 13. Distribution maps of Epsilon species in the Malay Archipelago.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFACB069FF51F2F9FACE62EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226470/files/figure.png	https://doi.org/10.5281/zenodo.21226470	FIGURE 11. Epsilon vechti Giordani Soika, female paratype. A, habitus, lateral view. B, head, frontal view. C, vertex, dorsal view. D, head, lateral view. E, submarginal carina and propodeal valvula, lateral view. F, T1–T2, dorsal view.	FIGURE 11. Epsilon vechti Giordani Soika, female paratype. A, habitus, lateral view. B, head, frontal view. C, vertex, dorsal view. D, head, lateral view. E, submarginal carina and propodeal valvula, lateral view. F, T1–T2, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFACB069FF51F2F9FACE62EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226478/files/figure.png	https://doi.org/10.5281/zenodo.21226478	FIGURE 13. Distribution maps of Epsilon species in the Malay Archipelago.	FIGURE 13. Distribution maps of Epsilon species in the Malay Archipelago.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFACB06BFF51F69DFCE96085.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226470/files/figure.png	https://doi.org/10.5281/zenodo.21226470	FIGURE 11. Epsilon vechti Giordani Soika, female paratype. A, habitus, lateral view. B, head, frontal view. C, vertex, dorsal view. D, head, lateral view. E, submarginal carina and propodeal valvula, lateral view. F, T1–T2, dorsal view.	FIGURE 11. Epsilon vechti Giordani Soika, female paratype. A, habitus, lateral view. B, head, frontal view. C, vertex, dorsal view. D, head, lateral view. E, submarginal carina and propodeal valvula, lateral view. F, T1–T2, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFACB06BFF51F69DFCE96085.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226438/files/figure.png	https://doi.org/10.5281/zenodo.21226438	FIGURE 1. A–E, Epsilon achterbergi Giordani Soika, female paratype. F–J, Epsilon bertschi sp. nov., female holotype. A, F, habitus, lateral view. B, G, head, frontal view. C, H, mesosoma, dorsolateral view. D, I, head and mesosoma, lateral view. E, J, T1–T2, dorsal view.	FIGURE 1. A–E, Epsilon achterbergi Giordani Soika, female paratype. F–J, Epsilon bertschi sp. nov., female holotype. A, F, habitus, lateral view. B, G, head, frontal view. C, H, mesosoma, dorsolateral view. D, I, head and mesosoma, lateral view. E, J, T1–T2, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFACB06BFF51F69DFCE96085.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226441/files/figure.png	https://doi.org/10.5281/zenodo.21226441	FIGURE 2. Epsilon crassipunctatum Gusenleitner, stat. nov., female paratype. A, habitus, lateral view. B, head, frontal view. C, head, dorsal view. D, mesosoma, dorsal view. E, T1–T2, dorsal view.	FIGURE 2. Epsilon crassipunctatum Gusenleitner, stat. nov., female paratype. A, habitus, lateral view. B, head, frontal view. C, head, dorsal view. D, mesosoma, dorsal view. E, T1–T2, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFACB06BFF51F69DFCE96085.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226443/files/figure.png	https://doi.org/10.5281/zenodo.21226443	FIGURE 3. Epsilon dyscherum (de Saussure). A–B, D–E, female syntype of Odynerus chartergoides von Schulthess. C, nontypical male. F–H, non-typical female. A, habitus, lateral view. B–C, head, frontal view. D, head, dorsal view. E, mesosoma, dorsal view. F, T1–T2 from Luzon, dorsal view. G, T1–T2 from Mindanao, dorsal view. H, T1–T2 from Tawi-Tawi, dorsal view.	FIGURE 3. Epsilon dyscherum (de Saussure). A–B, D–E, female syntype of Odynerus chartergoides von Schulthess. C, nontypical male. F–H, non-typical female. A, habitus, lateral view. B–C, head, frontal view. D, head, dorsal view. E, mesosoma, dorsal view. F, T1–T2 from Luzon, dorsal view. G, T1–T2 from Mindanao, dorsal view. H, T1–T2 from Tawi-Tawi, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFACB06BFF51F69DFCE96085.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226445/files/figure.png	https://doi.org/10.5281/zenodo.21226445	FIGURE 4. Epsilon incola Giordani Soika, female holotype. A, habitus, lateral view. B, head, frontal view. C, head, dorsal view. D, mesosoma, dorsolateral view. E, T1–T2, dorsal view.	FIGURE 4. Epsilon incola Giordani Soika, female holotype. A, habitus, lateral view. B, head, frontal view. C, head, dorsal view. D, mesosoma, dorsolateral view. E, T1–T2, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFACB06BFF51F69DFCE96085.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226450/files/figure.png	https://doi.org/10.5281/zenodo.21226450	FIGURE 5. Epsilon laboriosum (Smith). A–B, male holotype (credits: James Hogan, OUMNH). C–H, female. A, habitus, lateral view. B–C, head, frontal view. D, scutellum and metanotum, lateral view. E, scutellum and metanotum, dorsal view. F, mid and hind legs, lateral view. G, T1–T2 from New Guinea, dorsal view. H, T1–T2 from Bismarck Archipelago, dorsal view.	FIGURE 5. Epsilon laboriosum (Smith). A–B, male holotype (credits: James Hogan, OUMNH). C–H, female. A, habitus, lateral view. B–C, head, frontal view. D, scutellum and metanotum, lateral view. E, scutellum and metanotum, dorsal view. F, mid and hind legs, lateral view. G, T1–T2 from New Guinea, dorsal view. H, T1–T2 from Bismarck Archipelago, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFACB06BFF51F69DFCE96085.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226452/files/figure.png	https://doi.org/10.5281/zenodo.21226452	FIGURE 6. Epsilon latefasciatum sp. nov. A–B, male holotype. C–E, female. A, habitus, lateral view. B–C, head, frontal view. D, mesosoma, dorsal view. E, T1–T2, dorsal view.	FIGURE 6. Epsilon latefasciatum sp. nov. A–B, male holotype. C–E, female. A, habitus, lateral view. B–C, head, frontal view. D, mesosoma, dorsal view. E, T1–T2, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFACB06BFF51F69DFCE96085.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226454/files/figure.png	https://doi.org/10.5281/zenodo.21226454	FIGURE 7. Epsilon manifestum (Smith). A, C–D, F, male. B, E, G–H, female. D, F, anomalous specimens. A, habitus of holotype, lateral view (credits: James Hogan, OUMNH). B–D, head, frontal view. E–F, mesosoma, dorsal view. G, T1–T2, dorsal view. H, apex of T2, lateral view.	FIGURE 7. Epsilon manifestum (Smith). A, C–D, F, male. B, E, G–H, female. D, F, anomalous specimens. A, habitus of holotype, lateral view (credits: James Hogan, OUMNH). B–D, head, frontal view. E–F, mesosoma, dorsal view. G, T1–T2, dorsal view. H, apex of T2, lateral view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFACB06BFF51F69DFCE96085.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226457/files/figure.png	https://doi.org/10.5281/zenodo.21226457	FIGURE 8. Epsilon perforatum sp. nov., female holotype. A, habitus, lateral view. B, head, frontal view. C, mesosoma, dorsal view. D, T1–T2, dorsal view. E, S2, lateral view. F, S2, ventral view.	FIGURE 8. Epsilon perforatum sp. nov., female holotype. A, habitus, lateral view. B, head, frontal view. C, mesosoma, dorsal view. D, T1–T2, dorsal view. E, S2, lateral view. F, S2, ventral view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFACB06BFF51F69DFCE96085.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226459/files/figure.png	https://doi.org/10.5281/zenodo.21226459	FIGURE 9. Epsilon rufipes Selis. A–B, E, H, female holotype from Aru. C, G, I, female from New Guinea. D, F, male from New Guinea. A, habitus, lateral view. B–D, head, frontal view. E, mesosoma, lateral view. F, scutellum, posterior view. G, mid and hind legs, lateral view. H–I, T1–T2, dorsal view.	FIGURE 9. Epsilon rufipes Selis. A–B, E, H, female holotype from Aru. C, G, I, female from New Guinea. D, F, male from New Guinea. A, habitus, lateral view. B–D, head, frontal view. E, mesosoma, lateral view. F, scutellum, posterior view. G, mid and hind legs, lateral view. H–I, T1–T2, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
5F7987D6FFACB06BFF51F69DFCE96085.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21226465/files/figure.png	https://doi.org/10.5281/zenodo.21226465	FIGURE 10. Epsilon tigrinum sp. nov. A–B, D–E, G, female holotype. C, male paratype. F, female paratype. A, habitus, lateral view. B–C, head, frontal view. D, scutellum and metanotum, lateral view. E, scutellum and metanotum, dorsal view. F, legs and metasoma, lateral view. G, T1–T2, dorsal view.	FIGURE 10. Epsilon tigrinum sp. nov. A–B, D–E, G, female holotype. C, male paratype. F, female paratype. A, habitus, lateral view. B–C, head, frontal view. D, scutellum and metanotum, lateral view. E, scutellum and metanotum, dorsal view. F, legs and metasoma, lateral view. G, T1–T2, dorsal view.	2026-07-06	Selis, Marco		Zenodo	biologists	Selis, Marco			
