identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
B78D812BE7E9525F86C0363CEB0FF7C2.text	B78D812BE7E9525F86C0363CEB0FF7C2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tylototriton wufengensis Li & Shi & Liu & Luo & Wang & Liao & Wang & Gong & Wu & Wang 2026	<div><p>Tylototriton wufengensis sp. nov.</p><p>Holotype.</p><p>CIB WF 20240502001 adult male, collected by Shize Li on 2 May 2024 in Wufeng County (30.187 ° N, 110.931 ° E; elevation 1045 m a. s. l.), Hubei Province, China.</p><p>Paratypes.</p><p>Three females, CIB WF 20240502004, CIB WF 20240502007, and CIB WF 20240502008, and four males, CIB WF 20240502002, CIB WF 20240502003, CIB WF 20240502005, and CIB WF 20240502006, were collected by Jing Liu on 2 May 2024. All of the paratypes were collected from the same place as the holotype.</p><p>Diagnosis.</p><p>Tylototriton wufengensis sp. nov. is assigned to the genus Tylototriton based upon molecular phylogenetic analyses and the following morphological characters: presence of dorsal granules, dorsolateral bony ridges on the head, knob-like warts on dorsolateral body, and the absence of a quadrate spine.</p><p>Tylototriton wufengensis sp. nov. can be distinguished from other congeners by the following characters: (1) body size moderate (SVL 63.7–77.2 mm in males and 74.9–88.1 mm in females); (2) head width larger than head length; (3) gular fold present; (4) the tail length shorter than the snout-vent length; (5) the distal ends and ventral surfaces of digits, peripheral area of cloaca, and the lower margin of tail orange; (6) the distal tips of the limbs just meeting when the fore and hindlimbs being pressed along the trunk; (7) fingertips reaching to the level beyond the snout when the forelimbs being stretched forward; (8) nodule-like warts on body sides that are continuous and not obvious.</p><p>Description of holotype.</p><p>Adult male (Figs 5, 6), body size moderate, SVL 74.8 mm; TOL 138.5 mm; head width larger than head length (HDW / HDL = 1.13); head slightly concave on the top; snout nearly square, extending beyond the lower lip; naris nearly snout; snout truncate in dorsal view; fabial fold absent; dorsolateral bony ridges on head prominent, moderately protruding, from above snout to above anterior end of parotoid, posterior ends relatively thick and scrolled inside; mid-dorsal ridge on head distinct and thin; parotoids enlarged, projecting backwards; tinter-nasal space is smaller than the inter-orbital space; eyes protrude from the dorsolateral portion of the head; the oral fissure flat; the joint of jaw articulation lies posterior to the caudal margin of the eyes; fine teeth present on the edge of jaw, vomerine teeth long and prominent, form a “ ∧ ” shape which separated from each other at the tip of the “ ∧ ” shaped; tongue oval and nearly entirely fixed at bottom but free at both lateral edges; neck is rounded and thick, gular fold present.</p><p>Body stout; skin on lateral body with no obvious presence of small glands, glandular vertebral ridge prominent, slightly rough, and segmented, extending from top of head to base of tail, rib nodules indistinct.</p><p>Four limbs slender, fingers from fore- and hindlimbs meeting when addressed along body; tips of fingers beyond snout when addressed along head; relative fingers length III&gt; II&gt; I&gt; IV, and relative toes length III&gt; IV&gt; II&gt; I&gt; V; both fingers and digits are with none webbed.</p><p>The tail length shorter than the snout-vent length; the tail laterally compressed, thin, and tip acuminated, distinctly thin and high; the tail height is greater than the width at the tail base, and the tip of the tail resembles a sword shape; the cloaca long and narrow, and the cloacal region slightly bulbous.</p><p>The skin rough, body covered with small tubercles; the labial margin, distal limbs, ventral limbs, and ventral edge of the tail smooth; on the lateral dorsum of the body, tubercles small, appearing to form lines that extend from the shoulder to the base of the tail, and the nodule-like tubercles continuously distributed with no clear separation between neighboring tubercles; the tubercles on pharyngeal small, distinct and sparse, and tubercles on on the abdomen relative large and dense.</p><p>Coloration in life.</p><p>In life, the overall body color is black. Orange coloration is present on the dorsal and ventral surfaces of the fingers and toes, the peripheral area of the cloaca, and the ventral edge of the tail (Fig. 6).</p><p>Color in preservative.</p><p>The specimen in preservative is blackish-brown. The orange coloration of the distal ends and ventral surface of digits, the peripheral area of the cloaca, and the ventral edge of the tail fades to light orange (Fig. 5).</p><p>Variation.</p><p>Measurements of all specimens are listed in Table 3 and Suppl. material 1. All specimens were very similar in morphology. But in CIB WF 20240502004, there is an orange-red protrusion behind the parotoid gland (Fig. 7 B); cloaca and the ventral edge of tail disconnected (Fig. 7 D).</p><p>Secondary sexual characters.</p><p>In male, the cloacal orifice long slit and has papilla on inner cloacal wall (Fig. 6 I) and in female, cloacal hole is short and its inner cloacal wall has no papilla (Fig. 7 D).</p><p>Distribution and habitats.</p><p>At present, Tylototriton wufengensis sp. nov. is known only from Wufeng County, Hubei Province, China. The population inhabits mountain ponds at an altitude of approximately 1000 m, where accumulated rocks are present. During the day, Tylototriton wufengensis sp. nov. hides under these rocks and becomes active at night. Near these ponds inhabited by Tylototriton wufengensis sp. nov., there are farmlands (Fig. 8). Polypedates megacephalus Hallowell, 1861, Pelophylax nigromaculatus (Hallowell, 1860), and Zhangixalus chenfui (Liu, 1945) were also found at the type locality of the new species.</p><p>Etymology.</p><p>The specific epithet “ wufeng ” refers to the distribution of this species, Wufeng County, Hubei Province, China. We propose the common name “ Wufeng Knobby Newt ” and its Chinese name as Wu Feng You Yuan (五峰疣螈).</p></div>	https://treatment.plazi.org/id/B78D812BE7E9525F86C0363CEB0FF7C2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Shize;Shi, Shengchao;Liu, Jing;Luo, Zhonghao;Wang, Jiajun;Liao, Linhong;Wang, Yeqing;Gong, Renhu;Wu, Jun;Wang, Bin	Li, Shize, Shi, Shengchao, Liu, Jing, Luo, Zhonghao, Wang, Jiajun, Liao, Linhong, Wang, Yeqing, Gong, Renhu, Wu, Jun, Wang, Bin (2026): Description of a new species of the Asian newt genus Tylototriton Anderson, 1871 (Urodela, Salamandridae) from central China. Zoosystematics and Evolution 102 (1): 181-197, DOI: 10.3897/zse.102.173283
