taxonID	type	description	language	source
5857225FFFF95C15A390FCD2FEE0FA37.taxon	description	Eurydoxa advena Filipjev, 1930. Doklady Akademii Nauk SSSR (A) (1930): 374. Figs 1 - 3. TL: Russia, “ auf dem Gipfel des Berges Chualasa (ungefähr 1400 m hoch) in Sutchan Gebiet ” (Far East, Partizansk [Sutchan] District, Khalaza [Chualasa] Mountain). Types. Holotype ♀: “ Уссур. край, Сучан, верш. ХудлаЗы, Куренцов, 11. VIII. 28 [Ussuri Region, Sutschan, peak of Khudlazy, Kurentsov, 11. VIII. 28] ”, gold circle, “ Holotypus Eurydoxa advena Filipjev, 1930 ”, “ ZISP INS _ LEP _ 0001403 ”, Praep. micr. SVN 24188 Eurydoxa advena Filipjev, 1930 S. Nedoshivina pr. 2024 ”. Present status. Eurydoxa advena Filipjev, 1930.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFF95C15A390FCD2FEE0FA37.taxon	discussion	Comments. The venation of the forewing and hindwing was illustrated in the original description. Taxonomic notes. Eurydoxa advena is the type species of the genus Eurydoxa Filipjev. The female genitalia of the holotype are figured here for the first time. In Gilligan et al. (2018) orbimaculata Kurentsov, 1956, and ussuriensis Kurentsov, 1956 were listed as junior synonyms of advena with the mention that their types are deposited in ZISP. However, the types of species described by Kurentsov were initially kept in the Federal Scientific Center of the East Asia Terrestrial Biodiversity in Vladivostok, Russia. Unfortunately, Kurentsov’s collection is no longer deposited in Vladivostok and it is most likely impossible to determine whether it was transferred or lost. As a result, the type specimens of orbimaculata Kurentsov, 1956, and ussuriensis Kurentsov, 1956 have not been located and are presumed lost.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFF95C15A390F9CEFB21F818.taxon	description	Present status. Clepsis altitudinarius (Filipjev, 1962).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFF95C15A390F9CEFB21F818.taxon	discussion	Comments. In Gilligan et al. (2018), the type locality was erroneousely listed as “ Russia, Caucasus ”. The genitalia of the holotype were illustrated in the original description and by Razowski (1979).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFFF5C13A390FF5BFB20FB81.taxon	description	Cnephasia atlantis Filipjev, 1934. Bulletin de l’Académie des Sciences de l’URSS 9: 1404. TL: Morocco, “ Высокий Атлас [High Atlas], Tachdirt ”. Types. Paratypes: 4 ♂ – “ Marokko, Gr. Atlas Tachdirt 2200 – 2700 m, 2. – 10. VII. 33. Zerny ”, red label “ 7 / m 34 N I м. г. ”, “ Paratype Cnephasia atlantis Fil ”; “ Marokko, Gr. Atlas Tachdirt 2200 – 2700 m, 2. – 10. VII. 33. Zerny ”, “ praep. micr. № 2600 | Cneph. atlantis Fil. ♂ Marokko ”, “ Praep. micr. SVN 24200 Cnephasia stephensiana atlantis (Filipjev, 1934) S. Nedoshivina praep. 2024 ”, “ Paratype Cnephasia atlantis Fil ”; “ Marokko, Gr. Atlas Tachdirt 2200 – 2700 m, 2. – 10. VII. 33. Zerny ”, “ Paratype Cnephasia atlantis Fil ”; “ H. Atlas Maroc. Tachdirt 2300 – 2700 m 3. – 25. VII. 33 Schwingenschuss ”, “ Paratype Cnephasia atlantis Fil ”. Present status. Cnephasia stephensiana atlantis (Filipjev, 1934).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFFF5C13A390FF5BFB20FB81.taxon	discussion	Comments. The holotype is deposited in NHMV. In the original description, Filipjev mentioned three male specimens from ZISP: 2 males collected 2 – 10. VII 1933 by Zerny and one male collected 25. VII 1933 by Schwingenschuss. However, an additional male specimen is present in ZISP with the same label as the first two by Zerny, though it lacks labels explicitly designating it as a part of the type series. Despite this omission, I consider it likely to belong to the type series, as it is improbable that a specimen sharing identical labels and morphological characteristics would have been excluded intentionally by the author. Genitalia of both sexes were illustrated by Razowski (1965). Taxonomic notes. Razowski (1965) initially listed atlantis Filipjev, 1934 as a bona species, but later, in his work of 1989, he reclassified it as one of three subspecies of stephensiana Doubleday, 1850, alongside anatolica Obraztsov, 1950, and stolidana Walker, 1863. This subspecies is distributed in Morocco (Razowski 1989) and differs from the nominate subspecies by its shorter sacculus. We follow this criterium and keep the taxon as a subspecies. However, given the variability of the male genitalia of stephensiana Doubleday, 1850, a deep taxonomic study of the whole group, including molecular analysis, is in need to clarify the real status of atlantis.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFFF5C12A390FB94FB9BFC6B.taxon	description	Paratypes: 2 ♂ and ♀ – ♂ “ Lagodechi 2. V. 885 Млокосевич [Mlokosevich] ”, “ колл. б. Вел. Кн. НиколаЯ Михайловича [coll. of the former Grand Duke Nikolay Mihailovich] ”, white circle, “ praep. micr. № 3151 Peronea caucasica Fil. ♂ Лагодехи [Lagodekhi] ”, “ Paratypus Peronea caucasica Fil. ”; ♂ “ Лагодехи. Зака-тал окр. Тифл. г. Млокосъвич 15. VIII. 96 [Lagodekhi. Outskirts of Zaqatala, Tbilisi Region, Mlokosevich 15. VIII. 96] ”, white circle, “ praep. micr. № 3145 Peronea caucasica Fil. ♂ Лагодехи [Lagodekhi] ”, “ Paratypus Peronea caucasica Fil. ”; ♀ “ Lagodechi ♀ | 9.10.87 Tripunctana ”, “ колл. б. Вел. Кн. НиколаЯ Михайловича [coll. of the former Grand Duke Nikolay Mihailovich] ”, gold circle, “ praep. micr. № 3145 Peronea caucasica Fil. ♀ Лагодехи [Lagodekhi] ”, “ Paratypus Peronea caucasica Fil. ”, “ Gemalt von Mgr. R. Zawada-Skrzyński für Micr. Pal. ”, “ Praep. micr. SVN 24229 Acleris caucasica (Filipjev, 1962) S. Nedoshivina pr. 2024 ”. Present status. Acleris caucasica (Filipjev, 1962).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFFF5C12A390FB94FB9BFC6B.taxon	discussion	Comments. In the original description, Filipjev stated that the female was collected on 3. X. 1887. However, the only female specimen bearing a gold circle, which indicates its type status, was collected on 9. X. 1887. Therefore, I conclude that the collection date provided in the original description was likely erroneous. Taxonomic notes. The species caucasica was originally described in the genus Peronea and was subsequently transferred to Acleris by Razowski (1966) as a bona species. Later, in his works of 2001, 2002 and 2008, the same author considered it as a subspecies of Acleris lacordairana (Duponchel, 1836). Genitalia of both sexes were illustrated in the original description; however, Razowski (2008) noted that female genitalia of caucasica were not re-examined and from the original drawing one cannot correctly determine the species. Unfortunately, in the vial containing genitalia preparation № 3145, corresponding to the female paratype, only bursa copulatrix was preserved. The shape of corpus bursae and signum matches that of lacordairana according to Razowski (2008). On the other hand, I examined two females from the ZISP collection that are externally similar to the holotype of caucasica and lacordairana sensu Razowski (2008): one originates from the type locality of caucasica (Fig. 6), and the other is from Tiflis (now Tbilisi), a city located approximately 120 km west of the type locality. Both specimens have a bursa copulatrix of the same shape as that of the caucasica paratype but lack a signum (Fig. 82). In general, the female genitalia of these Georgian specimens are close to those of the nominotypical subspecies of lacordairana and its other subspecies, roxana Razowski et Yasuda, 1964, from Japan (according to Razowski 2008). However, they show clear differences: antrum broad, its length equals width, whereas in lacordairana, it is 1.5 times the width; proximal lobes of sterigma almost as broad as in roxana; proximal 2 / 3 of ductus bursae folded and maximum 4 times broader than basal third (in lacordairana and roxana, maximum 3 times broader). I preliminarily conclude that these two females most likely belong to caucasica, and a discrimination based on the female genitalia is provided above. However, additional material is needed to confirm this assumption. In the male genitalia of caucasica ventral incision of sacculus before subterminal lobe shorter than in two other subspecies and directed backward toward valva base, overlapping notch of lower valva edge. In Gilligan et al. (2018) caucasica and roxana are interpreted as bona species and longulana Eversmann, 1844, is listed as a junior synonym of caucasica. Based on my investigation, the specific status of caucasica can be supported by the differences in male and female genitalia discussed above. However, the synonymy of longulana is entirely incorrect. In the male genitalia of the longulana lectotype (Nedoshivina 2007), sacculus concave before end as in lacordairana whereas in caucasica, it is slightly curved; subterminal lobe of sacculus in longulana is long and distinct as in lacordairana, while in caucasica, it is relatively small, and ventral incision before the lobe is shorter than in longulana and lacordairana. Therefore, longulana Eversmann, 1844, should be considered as a junior synonym of lacordairana Duponchel, 1836, as proposed by Kyrki (1982).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFFE5C12A390FBF3FB5EF9C0.taxon	description	Laspeyresia coeruleosparsa Filipjev, 1924. Jahrbuch des Martjanov’schen Staatsmuseums in Minussinsk 2 (3): 26. TL: Russia, “ Сад МуЗеЯ [Museum garden] ” (Krasnoyarsk Region, Outskirts of Minusinsk). Types. Holotype ♀: “ Окр. Минусинска 10 / VI 1917 Кожанчиковъ | Сад при муЗее [Outskirts of Minusinsk 10 / VI 1917 Kozhantshikov | Museum garden] ”, gold circle, “ 213 ”, “ микр. преп. [praep. micr.] N 7768 ”, “ Laspeyresia coeruleosparsa Fil. type [underlined] ”, “ Photograph Marek Kopeć ”, “ Holotypus Laspeyresia coeruleosparsa Fil. ”, “ ZISP INS _ LEP _ 0001523 ”, “ Praep. micr. SVN 24197 Cydia exquisitana coeruleosparsa (Filipjev, 1931) S. Nedoshivina pr. 2024 ”. Present status. Cydia exquisitana coeruleosparsa (Filipjev, 1924).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFFE5C12A390FBF3FB5EF9C0.taxon	discussion	Comments. The collecting date of the holotype, 23.6.1917, mentioned in the original description, is likely an error. The only specimen in ZISP bearing the label “ Laspeyresia coeruleosparsa n. sp. Fil. type [underlined], ” handwritten by Filipjev, was collected on 10. VI. 1917. Taxonomic notes. This Siberian subspecies is currently known only from the type specimen.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFFE5C11A390F954FB3EFD9F.taxon	description	Paratype: ♀ — “ Алма-Ата, Заповедник, ниж. теч. р. Батарейка прав. склон 27. VII 1936 г. В. Шайдуров [Almaty Nature Reserve, lower river Batareyka (= Bedel’bay) right slope 27. VII 1936 V. Shaydurov] ”, gold circle, “ praep. micr. № 3105 | Peronea coniferarum Fil. ♀ Алма-Ата [Almaty] ”, “ Allotypus Peronea coniferarum Fil. ”, “ Praep. micr. SVN 24232 Acleris coniferarum (Filipjev, 1962) S. Nedoshivina pr. 2024 ”. Present status. Acleris coniferarum (Filipjev, 1962).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFFE5C11A390F954FB3EFD9F.taxon	discussion	Comments. Genitalia of both sexes were illustrated in the original description. The genitalia of the male holotype were divided into parts, and in Microlepidoptera Palaearctica (Razowski 1984), the drawing of the tuba analis was placed separately from other genitalia structures. This is likely the reason why, in a subsequent work (Razowski 2008), this structure was omitted from the corresponding plate. However, the tuba analis, with its strong, pointed thorn, is a diagnostic feature and should be illustrated. Taxonomic notes. Since its description, the validity of this name has not been questioned.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFFD5C11A390FBE8FB00FA65.taxon	description	Euxanthis disjuncta Filipjev, 1924. Jahrbuch des Martjanov’schen Staatsmuseums in Minussinsk 2 (3): 54. TL: Russia, “ ГрЯды [Ridges] ” (Krasnoyarsk Region, mountain ridges in outskirts of Minusinsk). Types. Holotype ♂: “ Euxanthis disjuncta Fil. ♂ Type ”, “ Окр. Минусинска 11. VII 1923 Кожанчиковъ [Outskirts of Minusinsk 11. VII 1923 Kozhantschikov] ”, “ к. Н. Филипьева [coll. of N. Filipjev] ”, gold circle, “ Holotypus Euxanthis disjuncta Filipjev, 1924 ”, “ ZISP INS _ LEP _ 0001575 ”, “ Praep. micr. SVN 24221 Aphelia disjuncta (Filipjev, 1924) S. Nedoshivina pr. 2024 ”. Present status. Aphelia disjuncta (Filipjev, 1924)	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFFD5C11A390FBE8FB00FA65.taxon	discussion	Comments. In Gilligan et al. (2018), the type locality was given erroneously as “ Tomsk District, Teleckoe Lake ”. Taxonomic notes. This taxon has been treated as a bona species since its description.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFFD5C11A390FDAFFC3AFC5C.taxon	description	Phalonia costignata Filipjev, 1926. Archiv für Naturgeschichte 91 (1) (1925): 115. TL: Russia, “ Grushevka ” (Stavropol Region, Grushevskoye). Types. Holotype ♂: “ Ставрополь Кавк. Грушевка 25. V. 1921 [Stavropol Caucas. Grushevka 25. V. 1921] ”, gold circle, “ G. S. – 8034 ♂ J. Razowski ”, “ Phalonia costisignata Fil. ♂ Type ”, “ costisignata ”, “ Holotypus Phalonia costignata Filipjev, 1926 ”, “ ZISP INS _ LEP _ 0001536 ”. Present status. Aethes williana (Brahm, 1791).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFFD5C11A390FDAFFC3AFC5C.taxon	discussion	Comments. Genitalia of the male holotype were illustrated by Razowski (1970). Taxonomic notes. Synonymy was established by Razowski (1970).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFFD5C10A390F9E1FD80FE2B.taxon	description	Semasia elongata Filipjev, 1924. Jahrbuch des Martjanov’schen Staatsmuseums in Minussinsk 2 (3): 24. TL: Russia, “ Джойское оЗ. [Dzhoyskoye Lake] ” (Krasnoyarsk Region, Dzhoyka Lake). Types. Lectotype ♂ (here designated): “ Окр. Минусинска 17. V 1917 Кожанчиковъ [Outskirts of Minusinsk 17. V 1917 Kozhantschikov] ”, “ Epiblema elongata Fil. ♂ Type ”, “ Cotypus Semasia elongata Fil. ”, “ Lectotypus Semasia elongata Filipjev, 1924 ”, “ ZISP INS _ LEP _ 0001593 ”, “ Praep. micr. SVN 24233 Eucosma lignana (Snellen, 1883) S. Nedoshivina pr. 2024 ”. Paralectotypes 2 ♂: “ Окр. Минусинска 20. V 1917 Кожанчиковъ [Outskirts of Minusinsk 17. V 1917 Kozhantschikov] ”, “ Epiblema elongata Fil. ♂ Type ”, “ Cotypus Semasia elongata Fil. ”. Present status. Eucosma lignana (Snellen, 1883).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFFD5C10A390F9E1FD80FE2B.taxon	discussion	Comments. In the original description, on page 24 (in Russian), the author noted that the type specimens were collected on 30. V and 2. VI. However, in the same publication on page 38, the author translated the description of elongata into German, where the collecting dates were given as 17. V and 20. V. In the ZISP collection, there are only three specimens of elongata designated as types by Filipjev, with the collecting dates 17. V and 20. V. Therefore, it can be assumed that the dates 30. V and 2. VI were provided erroneously. In Gilligan et al. (2018), the type locality was imprecisely listed as “ Siberia. ” Taxonomic notes. The synonymy was likely established by Kuznetzov (e. g., 1978), but the syntypes were not dissected. My examination of the type material confirms this synonymy, and the genitalia of the male lectotype are illustrated here for the first time.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFFC5C10A390FE3AFEB9FA04.taxon	description	Peronea expressa Filipjev, 1931. Annuaire du Musée Zoologique de l’Académie des Sciences de l’URSS 1931 (1930): 517. Plate XXIV, fig. 3; Plate XXXI, fig. 2. TL: Russia, “ Jakovlevka, Lazarevka-Weg ” (Far East, Yakovlevka). Types. Holotype ♂: “ Яковлевка Спас. у. Уссур. кр. 1. V. 926 ДьЯконов Филипьев [Yakovlevka, Ussuri Region, 1. V. 1926 Djakonov, Filipjev] ”, “ ЛаЗаревскаЯ дорога 1. V. 926 [Lazarevka Road, 1. V. 1926] ”, “ Pe. expressa Fil. ♂ N. Filipjev det. ”, “ praep. micr. № 988 ”, gold circle, “ Holotypus Peronea expressa Fil. ”, “ ZISP INS _ LEP _ 0001614 ”, “ Praep. micr. SVN 24206 Acleris expressa (Filipjev, 1931) S. Nedoshivina pr. 2024 ”. Paratype ♀: “ Виноградовка Уссур. кр. 13. V. 929 ДьЯконов Филип. [Vinogradovka Ussuri Region 13. V. 929 Djakonov, Filipjev] ”, “ Peronea expressa Fil. ♀ Filipjev det. ”, “ praep. micr. 988 Ac. expressa Fil. ♀ Uss. ”, “ Allotypus Peronea expressa Fil. ”, “ Praep. micr. SVN 24207 Acleris expressa (Filipjev, 1931) S. Nedoshivina pr. 2024 ”. Present status. Acleris expressa (Filipjev, 1931).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFFC5C10A390FE3AFEB9FA04.taxon	discussion	Comments. Razowski (1984) illustrated the genitalia of both male and female type specimens in Microlepidoptera Palaearctica. He accurately listed the labels of the male holotype in the text; however, the captions for the genitalia images were incorrect. Specifically, he noted that the male genitalia depicted on Plate 63, originating from genitalia preparation 988, belonged to a specimen labeled “ Okr. [estnosti] Wladiwostoka, p. [ol] - ostr. [ow] De-Fris, 27. VI. 1957 ” [Outskirts of Vladivostok] from the ZISP collection. However, genitalia preparation 988 contains only the male and female genitalia of the type specimens, none of which originated from the outskirts of Vladivostok. It is now impossible to determine whether a type specimen is illustrated here. Additionally, in the caption for the female genitalia drawing on Plate 98, the labels of the male holotype listed in the text were erroneously provided. These errors were repeated in Razowski (2008), with an additional mistake: the caption for the female genitalia states “ holotype, Russia, Amur District, ” whereas both Yakovlevka and Vinogradovka are located in Primorsky Krai (= Russian Far East), which is near the Amur Region but not identical to it. In Gilligan et al. (2018), the type locality was incorrectly listed as “ Japan, ” and the holotype sex was marked as “ unknown. ”	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFFC5C1FA390FA00FBBEFEC6.taxon	description	Paratypes: 2 ♂ and ♀ – ♀ “ Уссур. край, Сучан, ХуалаЗа, Куренцов, 21. VII. 28 [Ussuri Region, Sutschan, Khualaza, Kurentsov, 21. VII. 28] ”, “ Praep. micr. № 2992 | Latiunca fulva Fil. ♀ Ussuri ”, “ Paratypus Cacoecia fulva Fil. ”; ♂ “ Уссур. край, Сучан, Куренцов, 26. VII. 28 [Ussuri Region, Sutschan, Kurentsov, 26. VII. 28] ”, white circle, “ брюшко потерЯно [abdomen lost], “ Paratypus Cacoecia fulva Fil. ”; ♂ “ Сучанский район Уссур. кр. Васильев [Sutschan district, Ussuri Region, Vasilyev] ”, white circle, “ Praep. micr. № 2992 | Latiunca fulva Fil. ♂ Ussuri ”, “ Paratypus Cacoecia fulva Fil. ”, “ Praep. micr. SVN 24217 Daemilus fulva (Filipjev, 1962) S. Nedoshivina pr. 2024 ”. Present status. Daemilus fulva (Filipjev, 1962).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFFC5C1FA390FA00FBBEFEC6.taxon	discussion	Comments. The collection date of the male paratype collected by Kurentsov was mentioned in the original description as 28. VII. 28 probably by mistake. The specimen deposited in ZISP was collected 26. VII. 28 and undoubtedly belongs to the type series. Genitalia of both sexes were illustrated in the original description. In Gilligan et al. (2018), type locality was imprecisely listed as “ Russia, Pri-morski Territory ”.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFF35C1FA390FE5EFDF3FAEB.taxon	description	Piercea fulvimixta Filipjev, 1940. Travaux de l’Institut Zoologique de l’Académie des Sciences de l’URSS 6: 181. Fig. 17. TL: Russia, “ Jakovlevka ” (Far East, Yakovlevka). Types. Lectotype ♂ (here designated): “ Яковлевка Спас. у. Уссур. кр. 8. VIII. 926 ДьЯконв Филипьев [Yakovlevka, Ussuri Region, 8. VIII. 1926 Djakonov, Filipjev] ”, “ fulvomixta ”, gold circle, “ Gemalt von Dr. F. Gregor für Micr. Pal. ”, “ Lectotypus Piercea fulvimixta Filipjev, 1940 ”, “ ZISP INS _ LEP _ 0001635 ”, “ Praep. micr. SVN 24220 Phalonidia fulvimixta (Filipjev, 1940) S. Nedoshivina pr. 2024 ”. Paralectotypes: 7 ♂ – “ Виноградовка Уссур. кр. 7. VIII. 929 ДьЯконов Филип [Vinogradovka Ussuri Region 7. VIII. 929 Djakonov Filipjev] ”, “ Paratypus ”; “ Виноградовка Уссур. кр. 1 - 2. VIII. 929 ДьЯконов Филип [Vinogradovka Ussuri Region 1 - 2.929 Djakonov Filipjev] ”, white circle, gold circle, “ praep. micr. № 2281 Ph. fulvimixta Fil. ♂ Ussuri ”, “ Paratypus ”; “ Уссур. край Сучан Куренцов 27. VII 28 | Истоки р. Сицы [Ussuri Region Sutchan Kurentsov 27. VII 28 | Upper river Sitsa] ”, “ Paratypus ”; “ Уссур. край Сучан Куренцов 8. VIII 28 | Истоки р. Сицы [Ussuri Region Sutchan Kurentsov 8. VIII 28 | Upper river Sitsa] ”, “ praep. micr. № 2282 Ph. fulvimixta Fil. ♂ Ussuri ”, white circle, “ Paratypus ”; “ Уссур. край Сучан Куренцов 19. VIII 28 | Истоки р. Сицы [Ussuri Region Sutchan Kurentsov 19. VIII 28 | Upper river Sitsa] ”, “ praep. micr. № 2280 Ph. fulvimixta Fil. ♂ Ussuri ”, white circle, “ Paratypus ”; “ Уссур. край Сучан ист. р. Сицы Куренцов 25. VII 28 [Ussuri Region Sutchan upper river Sitsa Kurentsov 25. VII 28] ”, white circle, “ Paratypus ”; “ ОкеанскаЯ бл. Владивостока Траншель 10. VIII. 928 [Okeanskaya nearby Vladivostok Transhel 10. VIII. 928] ”, white circle, “ Paratypus ”. Present status. Phalonidia fulvimixta (Filipjev, 1940).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFF35C1FA390FE5EFDF3FAEB.taxon	discussion	Comments. Instead of the three males from Vinogradovka mentioned in the original description, only two males were found in the ZISP collection. Taxonomic notes. Razowski (1970) indicated type specimen collected in Yakovlevka 8. VIII 1926 as a holotype and illustrated the genitalia of a paratype from Vinogradovka. However, in the original description, holotype was not designated and the male genitalia were illustrated without important diagnostic features, such as the shape of the valva and socii. Therefore, the lectotype of fulvimixta Filipjev, 1940, is designated here, and its genitalia are illustrated in high quality for the first time.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFF35C1FA390FA73FE5BF86D.taxon	description	Semasia (Steganoptycha) funesta Filipjev, 1931. Annuaire du Musée Zoologique de l’Académie des Sciences de l’URSS 1931 (1930): 341. Figs 1 - 3, 7. TL: Russia, “ Ussuri-Gebiet, Chabarovsk ” (Far East, Khabarovsk Region, Khabarovsk). Types. Holotype ♂: “ Хабаровск 1926 [Khabarovsk 1926] ”, gold circle, “ Holotypus Semasia (Steganoptycha) funesta Filipjev, 1931 ”, “ ZISP INS _ LEP _ 0001637 ”, “ Praep. micr. SVN 24222 Zeiraphera funesta (Filipjev, 1931) S. Nedoshivina pr. 2024 ”. Paratypes: 2 ♂ and 2 ♀ – ♂ and ♀ “ Хабаровск 1926 [Khabarovsk 1926] ”, “ Paratypus Semasia (Steganoptycha) funesta Fil. ”; ♂ and ♀ “ г. Хабаровск 12. VII. 26 оп. поле [Khabarovsk, 12. VII. 26, experimental field] ”, “ гусеницы в бутонах Яблонь и груш [larvae in buds of apple and pear trees] ”, praep. micr. № 1089 Steg. funesta Fil. ♀ Ussuri ”, “ Paratypus Semasia (Steganoptycha) funesta Fil. ”, “ Praep. micr. SVN 24209 Zeiraphera funesta (Filipjev, 1931) S. Nedoshivina pr. 2024 ”. Present status. Zeiraphera funesta (Filipjev, 1931).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFF35C1FA390FA73FE5BF86D.taxon	discussion	Comments. Genitalia of both sexes were illustrated in the original description. Gilligan et al. (2018) noted that the holotype sex is unknown.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFF25C1EA390FB25FC71F977.taxon	description	Peronea grisea Filipjev, 1931. Annuaire du Musée Zoologique de l’Académie des Sciences de l’URSS 1931 (1930): 519. Plate A, fig. 4, Plate XXVII, fig. 1. TL: Russia, “ Baranovka, Suifun ” (Far East, Baranovskiy, Razdol’naya [= Suifun] River). Types. Holotype ♂: “ ♂ Suifun | Hedem [ann]. ”, “ 78 78 ”, 2 gold circles, “ praep. micr. № 1246 ”, “ колл. б. Вел. Кн. НиколаЯ Михайловича [coll. of the former Grand Duke Nikolay Mihailovich] ”, “ filipjevi ”, “ T. parisiana Gn. (Boscana F. var. [unreadable] Snell. ”, “ ZISP INS _ LEP _ 0001660 ”, “ Praep. micr. SVN 24216 Acleris filipjevi (Obraztsov, 1956) S. Nedoshivina pr. 2024 ”. Present status. Acleris filipjevi (Obraztsov, 1956).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFF25C1EA390FB25FC71F977.taxon	discussion	Comments. Genitalia of the male holotype were figured in the original description and by Razowski (2008). Genitalia preparation is divided into parts and right valva is lost. Gilligan et al. (2018) listed the type locality imprecisely as “ Ussuri ” and marked the holotype sex as “ unknown ”. Taxonomic notes. The species name grisea was preoccupied by Acalla hastiana f. grisea Müller-Rutz, 1924 (Razowski 2008). Obraztsov (1956) provided the replacement name filipjevi.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFF25C1EA390FF5BFB26FB11.taxon	description	Semasia gracilis Filipjev, 1924. Jahrbuch des Martjanov’schen Staatsmuseums in Minussinsk 2 (3): 55. TL: Russia, “ Джойское оЗ. [Dzhoyskoye Lake] ” (Krasnoyarsk Region, Dzhoyka Lake). Types. Lectotype ♂ (here designated): “ Окр. Минусинска 10. VI 1923 Кожанчиковъ [Outskirts of Minusinsk 10. VI 1923 Kozhantschikov] ”, “ Semasia gracilis Fil. ♂ Type ”, “ praep. micr. № 9195 ♂ ”, gold circle, “ к. Н. Филипьева [coll. of N. Filipjev] ”, “ Lectotypus Semasia gracilis Filipjev, 1924 ”, “ ZISP INS _ LEP _ 0001658 ”, “ Praep. micr. SVN 24199 Eucosma gracilis (Filipjev, 1924) S. Nedoshivina pr. 2024 ”. Paralectotype ♂: “ Окр. Минусинска 10. VI 1923 Кожанчиковъ [Outskirts of Minusinsk 10. VI 1923 Kozhantschikov] ”, gold circle, “ Semasia gracilis Fil. ♂ Type ”, “ к. Н. Филипьева [coll. of N. Filipjev] ”, “ Paralectotypus Semasia gracilis Fil. ”. Present status. Eucosma gracilis (Filipjev, 1924), stat. rev.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFF25C1EA390FF5BFB26FB11.taxon	discussion	Comments. In Gilligan et al. (2018), the type locality was given imprecisely as “ Siberia ” and the holotype sex was marked as “ unknown ”. Taxonomic notes. Gilligan et al. (2018) referred to the holotype as the type specimen; however, no holotype was designated in the original description. The species was synonymized with wimmerana Treitschke, 1835, likely by Kuznetzov (e. g., 1978), and subsequent works followed this synonymy. My examination of the type and non-type material of gracilis, as well as a series of specimens of wimmerana from Siberia in the ZISP collection, reveals that although the male genitalia of gracilis are generally very similar to those of wimmerana (e. g., as illustrated in Razowski 2003), consistent differences can be observed in the shape of the valva and wing coloration. These characters are considered here as the basis for rejecting the previously established synonymy and treating gracilis as a distinct species (stat. rev.). However, molecular data are needed to confirm this status for gracilis. Forewing ground color of gracilis dark ochreous, lighter nearby costal edge, whereas in wimmerana ground colour grey with whitish scales forming thin longitudinal lines. Costal strigulae in gracilis short, distinct only on costa; in wimmerana costal strigulae longer, merged distally. Median fascia not defined in gracilis wing pattern, but in wimmerana it forms dorsal whitish spot. In male genitalia of gracilis ventral edge of sacculus forms obtuse angle, valvae neck shorter than sacculus, arched ventrally. In wimmerana ventral edge of sacculus forms almost 90 - degree angle, valvae neck almost as long as sacculus with parallel edges and less rounded ventrally.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFF25C1DA390F88EFE67FDE7.taxon	description	Paratype ♂: “ Уссур. край, Сучан, Тигровое, 24. VII. 28, Куренцов [Ussuri Region, Suchan, Tigrovoye, 7. VII. 28, Kurentsov] ”, white circle, “ praep. micr. № 2557 Cn. kurentzovi ”, “ 2557 kurentzovi ”, Cnephasia alternella Stph. опр. Razowski в 1960 г. [Cnephasia alternella Stph. det. Razowski in 1960], “ Paratypus Cnephasia kurentzovi Fil. ”. Present status. Cnephasia stephensiana stolidana (Walker, 1863).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFF25C1DA390F88EFE67FDE7.taxon	discussion	Comments. In Gilligan et al. (2018), the species was listed erroneously as C. kurentsovi. Taxonomic notes. Cnephasia kurentzovi Filipjev, 1962 was considered as a junior synonym of Cnephasia alternella (Stephens, 1852) by Razowski (1965) and later synonymized with Cnephasia stephensiana stolidana (Walker, 1863) (Razowski 1989). Its subspecies status needs re-examination with more material involved (see also taxonomic notes for atlantis).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFF15C1DA390FD7EFBDAFB58.taxon	description	Phalonia lanceolata Filipjev, 1924. Jahrbuch des Martjanov’schen Staatsmuseums in Minussinsk 2 (3): 52. TL: Russia, “ Протока [Protoka River] ” (Krasnoyarsk Region, Minusinsk, Protoka River). Types. Lectotype ♂ (here designated): “ Окр. Минусинска 25. V 1923 Кожанчиковъ [Outskirts of Minusinsk 25. V 1923 Kozhantschikov] ”, “ Phalonia lanceolata Fil. ♂ Type ”, “ praep. micr. № 6572 ♂ ”, gold circle, “ Lectotypus Phalonia lanceolata Filipjev, 1924 ”, “ ZISP INS _ LEP _ 0001745 ”, “ Praep. micr. SVN 24208 Aethes moribundana (Staudinger, 1859) S. Nedoshivina pr. 2024 ”. Paralectotype ♂: “ Окр. Минусинска 31. V 1923 Кожанчиковъ | КладбиЩе [Outskirts of Minusinsk 31. V 1923 Kozhantschikov | Cemetery] ”, gold circle, “ Phalonia lanceolata Fil. ♂ Type ”, “ Paralectotypus Phalonia lanceolata Fil. ”. Present status. Aethes moribundana (Staudinger, 1859).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFF15C1DA390FD7EFBDAFB58.taxon	discussion	Comments. The collecting date for the paralectotype was listed in the original description as 21. V, however, the only male specimen marked as a type by Filipjev and bearing a gold circle in the ZISP collection was collected on 31. V. In Gilligan et al. (2018), the type locality was listed imprecisely as “ Siberia ”.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFF15C1BA390FAFCFDFCFE73.taxon	description	Piercea lydiae Filipjev, 1940. Travaux de l’Institut Zoologique de l’Académie des Sciences de l’URSS 6: 180. Fig. 16. TL: Russia, “ Voroshilov-Ussurijsk ” (Far East, Ussuriysk). Types. Lectotype ♂ (here designated): “ Никольск-Уссур., на свет, Л. Токарева, 14. VIII. 31 [Nikol’sk-Ussuriyskiy, on light, L. Tokareva, 14. VIII. 31] ”, gold circle, “ praep. micr. № 2259 ”, “ Lectotypus Piercea lydiae Filipjev, 1940 ”, “ ZISP INS _ LEP _ 0001770 ”. Paralectotypes: 2 ♂ and 2 ♀ – ♂ “ Никольск-Уссур., на свет, Л. Токарева, 16. VIII. 31 [Nikol’sk-Ussuriyskiy, on light, L. Tokareva, 16. VIII. 31] ”, white circle, “ praep. micr. № 2263 | Ph. lydiae Fil. ♂ Ussuri ”, “ Paratypus ”, “ Praep. micr. SVN 24225 Phalonidia lydiae (Filipjev, 1940) S. Nedoshivina pr. 2024 ”; ♂ “ Никольск-Уссур., на свет, Л. Токарева, 17. VIII. 31 [Nikol’sk-Ussuriyskiy, on light, L. Tokareva, 17. VIII. 31] ”, white circle, “ praep. micr. № 2264 | Ph. lydiae Fil. ♂ Ussuri ”, “ Paratypus ”; ♀ “ Яковлевка Спас. у. Уссур кр. 12. VII 926 ДьЯконв Филипьев [Yakovlevka Spasskoye District Ussuri Region 12. VII 926 Djakonov Filipjev] ”, white circle, “ Пасека Квашука [Kvashuk’s bee-garden] ”, “ praep. micr. № 2263 | Phal. lydiae Fil. ♀ Ussuri ”, “ Paratypus ”, “ Praep. micr. SVN 24226 Phalonidia lydiae (Filipjev, 1940) S. Nedoshivina pr. 2024 ”; ♀ “ Никольск-Уссур., на свет, Л. Токарева, 16. VIII. 31 [Nikol’sk-Ussuriyskiy, on light, L. Tokareva, 16. VIII. 31] ”, gold circle, “ praep. micr. № 2259 | Ph. lydiae Fil. ♀ Ussuri ”, “ lydiae ”, “ Gemalt von Dr. F. Gregor für Micr. Pal. ”, “ Paratypus ”. Present status. Phalonidia lydiae (Filipjev, 1940).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFF15C1BA390FAFCFDFCFE73.taxon	discussion	Comments. A vial containing genitalia preparation № 2259 of the male lectotype is empty, and the genitalia are presumably lost. Therefore, I provide here an image of the male paralectotype genitalia instead. Taxonomic notes. The genitalia of the male holotype from “ E Russia, Ussuri District ” were illustrated by Razowski (2009). However, since the species was described from multiple specimens and no holotype was designated in the original description, the specimen illustrated by Razowski should be designated as the lectotype. Female genitalia were not illustrated in the original description. In his paper, Filipjev only mentioned that the female genitalia of the new species differ from all other species by the presence of chitinous wrinkles on the ductus bursae. Razowski (2009) noted that the female genitalia of lydiae Fil. remain unknown. Thus, an image of the female genitalia is provided here for the first time.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFF75C1BA390FD8AFB2EFBAC.taxon	description	Laspeyresia marginestriana Filipjev, 1925. Revue Russe d’Entomologie [Russkoe Entomologicheskoe Obozrenie] 19: 50. TL: Georgia, “ Bolshoe Pozharistshe, westlich von Borzhom, Westtranskaukasien ” (outskirts of Borjomi). Types. Holotype ♀: “ Б. ПожариЩе, 17. VI. 16, И. Филипьев [Bolshoe Pozharistshe, 17. VI. 16, I. Filipjev] ”, “ Grapholitha marginestriana Filipyev, type ”, “ praep. micr. № 7655 ”, “ 157 ”, “ Holotypus Laspeyresia marginestriana Fil. ”, “ ZISP INS _ LEP _ 0001784 ”, “ Praep. micr. SVN 24198 Dichrorampha marginestriana (Filipjev, 1931) S. Nedoshivina pr. 2024 ”. Present status. Dichrorampha marginestriana (Filipjev, 1925).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFF75C1BA390FD8AFB2EFBAC.taxon	discussion	Comments. Filipjev designated the only type specimen in the original description as a male; however, in the ZISP collection, there is only a single female of marginestriana bearing the type labels by Filipjev. The genitalia of the holotype were illustrated by Danilevsky & Kuznetzov (1968). Taxonomic notes. This species name has the status of a valid taxon since its description.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFF75C1BA390FBB0FDD8F86D.taxon	description	Peronea nigriradix Filipjev, 1931. Annuaire du Musée Zoologique de l’Académie des Sciences de l’URSS 1931 (1930): 513. Plate A, figs 4 - 6; Plate XXIII, fig. 1; Plate XXIX, fig. 5. TL: Russia, “ Jakovlevka, Daubiche-Wald ” (Far East, Yakovlevka). Types. Holotype ♂: “ Яковлевка Спас. у. Уссур. кр. 14. IX. 926 ДьЯконов, Филипьев [Yakovlevka, Ussuri Region, 14. IX. 1926 Djakonov, Filipjev] ”, “ Лес по раЗливу р. ДаубихЭ [Forest nearby the river Daubikhe (= Arsen’yevka)] ”, gold circle, “ Holotypus, Sihe Foto! ”, “ Holotypus Peronea nigriradix Filipjev, 1931 ”, “ ZISP INS _ LEP _ 0001821 ”, “ Praep. micr. SVN 24192 Acleris nigriradix (Filipjev, 1931) S. Nedoshivina pr. 2024 ”. Paratypes: 32 ♂ and 12 ♀ – 21 ♂ and 7 ♀ “ Яковлевка Спас. у. Уссур. кр. ДьЯконов, Филипьев [Yakovlevka, Ussuri Region, Djakonov, Filipjev] ”, “ Лес по раЗливу р. ДаубихЭ [Forest nearby the river Daubikhe (= Arsen’yevka)] ”, white circle, “ Paratypus Peronea nigriradix Fil. ” (♀ 10. IX. 926; ♂ 10. V. 926; 4 ♂, ♀ 24. V. 926; 4 ♂, 3 ♀ 8. IX. 926; 7 ♂ 10. IX. 926; ♂ 14. IX. 926; ♂ 16. IX. 926; 3 ♂, 2 ♀ 17. IX. 926); 8 ♂ and ♀ “ Яковлевка Спас. у. Уссур. кр. ДьЯконов, Филипьев [Yakovlevka, Ussuri Region, Djakonov, Filipjev] ”, “ Пасека И. Лешко [I. Leshko’s apiary] ”, white circle, “ Paratypus Peronea nigriradix Fil. ”; 3 ♂ and 4 ♀ “ Виноградовка Спас. у. Уссур. кр. ДьЯконов, Филипьев [Vinogradovka, Ussuri Region, Djakonov, Filipjev] ”, white circle, “ Paratypus Peronea nigriradix Fil. ” (♀ 13. V. 929; ♀ 16. V. 929; 2 ♂ 19. V. 929; ♂, ♀ 20. V. 929; ♀ 3. VI. 929). Present status. Acleris nigriradix (Filipjev, 1931).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFF75C1BA390FBB0FDD8F86D.taxon	discussion	Comments. Gilligan et al. (2018) listed the type locality imprecisely as “ Ussuri ” and marked the holotype sex as “ unknown. ” Genitalia of both sexes were illustrated in the original description. Several probable errors exist in the list of type material provided in the original description. Two additional males, collected on 24. V. 1926 and 8. IX. 1929 and bearing a white circle, were found in the ZISP collection. At the same time, 10 males and 8 females mentioned in the original description were not located in the collection and are presumably lost. A single male from Yakovlevka, collected on 1. V. 1926 and bearing a white circle, was not mentioned in the original description and is considered here as a non-type specimen.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFEB5C07A390FF5BFBBFFD67.taxon	description	Paratypes: ♂ and ♀ – ♂ “ Уссур. край, Сучан, ХуалаЗа, Куренцов, 21. VII. 28 [Ussuri Region, Sutschan, Khualaza, Kurentsov, 21. VII. 28] ”, white circle, “ Praep. micr. № 2904 | Epag. orientis ♂ Ussuri ”, “ Paratypus Epagoge orientis Fil. ”, “ Praep. micr. SVN 24211 Gnorismoneura orientis (Filipjev, 1962) S. Nedoshivina pr. 2024 ”; ♀ “ Уссур. край, Сучан, ТигроваЯ, Куренцов, 24. VII. 28 [Ussuri Region, Sutschan, Tigrovaya, Kurentsov, 24. VII. 28] ”, gold circle, “ Praep. micr. № 2904 | Epag. orientis ♀ Ussuri ”, “ Paratypus Epagoge orientis Fil. ”. Present status. Gnorismoneura orientis (Filipjev, 1962).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFEB5C07A390FF5BFBBFFD67.taxon	discussion	Comments. Genitalia of both sexes were illustrated in the original description.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFEB5C07A390FCFFFB19FB2C.taxon	description	Euxanthis rectifascia Filipjev, 1924. Jahrbuch des Martjanov’schen Staatsmuseums in Minussinsk 2 (3): 53. TL: Russia, “ Джойское ос. [Dzhoyskoye Lake] ” (Krasnoyarsk Region, Dzhoyka Lake). Types. Holotype ♂: “ Euxanthis rectifascia Fil. ♂ Type ”, “ Окр. Минусинска 10. VI 1923 Кожанчиковъ [Outskirts of Minusinsk 10. VI 1923 Kozhantschikov] ”, gold circle, “ Holotypus Euxanthis rectifascia Filipjev, 1924 ”, “ ZISP INS _ LEP _ 0001914 ”, “ Praep. micr. SVN 24219 Cochylimorpha hedemanniana (Snellen, 1883) S. Nedoshivina pr. 2024 ”. Present status. Cochylimorpha hedemanniana (Snellen, 1883).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFEB5C07A390FCFFFB19FB2C.taxon	discussion	Comments. In Gilligan at al. (2018), the type locality was given erroneously as Minsk.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFEB5C07A390FB30FBA4F95C.taxon	description	Olethreutes retortimacula Filipjev, 1929. Annuaire du Musée Zoologique de l’Académie des Sciences de l’URSS: 4. Plate I, fig. 4; plate II, fig. 1. TL: “ Munku-Sardyk, Ost-Sayan-Gebirge ” (Border between Russia and Mongolia, East Sayan Mountains, Munku-Sardyk). Types. Lectotype ♀ (designated by Nedoshivina, 2016): “ Munko Sardyk, Sajan mont. ”, “ praep. micr. № 448 ”, gold circle, “ Phiaris retortimacula Fil. ♀ N. Filipjev det. ”, “ Lectotypus Olethreutes retortimacula Filipjev, 1929 ”. Paralectotypes: 2 ♀ – “ Munko Sardyk, Sajan mont. ”, “ A Bang-Haas ”, “ praep. micr. № 449 ”, gold circle, “ Phiaris retortimacula Fil. ♀ N. Filipjev det. ”, “ ZISP INS _ LEP _ 0001920 ”, “ Paralectotypus Olethreutes retortimacula Filipjev, 1929 ”; “ Munko Sardyk, Sajan mont. ”, “ A Bang-Haas ”, “ praep. micr. № 451 ”, “ Paralectotypus Olethreutes retortimacula Filipjev, 1929 ”. Present status. Olethreutes inquietana (Walker, 1863).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFEB5C07A390FB30FBA4F95C.taxon	discussion	Comments. The female genitalia were illustrated in the original description.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFEB5C06A390F8E0FA9FFEC7.taxon	description	Phalonia roseotincta Filipjev, 1924. Jahrbuch des Martjanov’schen Staatsmuseums in Minussinsk 2 (3): 51. TL: Russia, “ Тагарский остр. [Tagarskiy Island] ” (Krasnoyarsk Region, Tagarskiy Island). Types. Holotype ♂: “ Phalonia roseotincta Fil. ♂ Type ”, “ Окр. Минусинска 11. VI 1923 Кожанчиковъ [Outskirts of Minusinsk 11. VI 1923 Kozhantschikov] ”, gold circle, “ Holotypus Phalonia roseotincta Filipjev, 1924 ”, “ ZISP INS _ LEP _ 0001934 ”, “ Praep. micr. SVN 24228 Falseuncaria degreyana (McLachlan, 1869) S. Nedoshivina pr. 2024 ”. Present status. Falseuncaria degreyana (McLachlan, 1869).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFEB5C06A390F8E0FA9FFEC7.taxon	discussion	Comments. In Gilligan et al. (2018), the type locality was given erroneously as Okresnosti Minsk.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFEA5C05A390FC20FE64FF53.taxon	description	Pamene [sic!] seminotata Filipjev, 1926. Archiv für Naturgeschichte 91 (1) (1925): 116. TL: Russia, “ Stavropol ”. Types. Lectotype ♀: “ Ставрополь – К. 29 / III 14 г. [Stavropol – K. 29. III 14] ”, “ Pamene seminotata Fil. ♀ Type ”, gold circle, “ микр. преп. [praep. micr.] N 7010 ♀ ”, “ Lectotypus Pammene seminotata Fil. ”, “ ZISP INS _ LEP _ 0001955 ”, “ Praep. micr. SVN 24194 Pammene regiana (Zeller, 1849) S. Nedoshivina pr. 2024 ”. Paralectotype: ♂ – “ Х. Романовский Куб. обл. лес Н. Филипьев 23 / IV 20 [Romanovsky farm, Kuban Region, forest 23 / IV 20 N. Filipjev] ”, gold circle, “ Pammene seminotata Fil. ♂ Type ”, “ микр. преп. N 7010 ♂ | Pammene seminotata Fil. [praep. micr. № 7010 ♂ | Pammene seminotata Fil.] ”, “ Paralectotypus Pammene seminotata Fil. ”, “ Praep. micr. SVN 24193 Pammene regiana (Zeller, 1849) S. Nedoshivina pr. 2024 ”. Present status. Pammene regiana (Zeller, 1849) syn. nov.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFEA5C05A390FC20FE64FF53.taxon	discussion	Comments. The genitalia of the male paralectotype were figured by Danilevsky & Kuznetzov (1968). They also considered the type specimens as a holotype and a paratype. Since holotype was not designated in the original description, the specimen mentioned by Danilevsky & Kuznetzov (1968) as the holotype is designated here to be a lectotype. Taxonomic notes. Danilevsky & Kuznetzov (1968) treated seminotata as a subspecies of Pammene trauniana (Dennis et Schiffermüller, 1775), distributed in Crimea, the Caucasus, Iran, and mountainous Turkmenistan. They noted that high variability and the presence of intermediate forms do not justify recognizing seminotata as a separate species. Razowski (2003) questioned this opinion and suggested that these specimens require re- consideration. In addition to the previously mentioned types, the ZISP collection includes the following specimens of seminotata: a female from Crimea, a specimen without an abdomen from Georgia, eight specimens from Armenia, and one female from Turkmenistan. My examination of the type specimens of seminotata and the material mentioned above reveals that seminotata does not correspond to any specimens of trauniana in the ZISP collection or those illustrated in Razowski (2003). The male and female genitalia of seminotata are identical to those of regiana Zeller, 1849, sensu Razowski (2003). At the same time, regiana includes another subspecies described from Georgia – Pammene regiana caucasica Kuznetzov, 1968 – which does not correspond to either regiana sensu Razowski (2003) or seminotata. Since type and non-type specimens of seminotata show no clear differences from European specimens of regiana, seminotata is considered here as a junior synonym of regiana (syn. nov.). The taxonomic status of caucasica remains unclear and requires further research.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFEA5C06A390FE5EFD3EFC1C.taxon	description	Paratype: ♀ – “ Алма-Ата Высота 1800 metr 18. VII 1935 Е. Самойлович | Малина [Almaty 1800 m 18. VII 1935 E. Samoylovich | on Malus] ”, white circle, “ praep. micr. № 3106 ”, “ Paratypus Peronea rubivorella Fil. ”. Present status. Acleris rubivorella (Filipjev, 1962).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFEA5C06A390FE5EFD3EFC1C.taxon	discussion	Comments. Genitalia of the female holotype were illustrated in the original description. The species was treated by Razowski (2008) as valid. Gilligan et al. (2018) misspelled it as “ rubrivorella ” (lapsus calami). Photograph of the holotype was published by Razowski (2008).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFE95C05A390FB05FBBFF897.taxon	description	Paratypes: 3 ♀ – “ Уссур. Край верх р. Супутинки Ник. Усс. р-на Куренцов 5. VII. 1933 [Ussuri Region, upper Suputinka River] ”, white circle, “ Praep. micr. № 2907 | Anacron seriepuncta Fil. ♂ Ussuri ”, “ Paratypus Paratorna seriepuncta Fil. ”, “ Praep. micr. SVN 24213 Paratorna catenulella (Christoph, 1882) S. Nedoshivina pr. 2024 ”; “ Уссур. Край верх р. Супутинки Ник. Усс. р-на Куренцов 5. VII. 1933 [Ussuri Region, upper Suputinka River] ”, white circle, “ Paratypus Paratorna seriepuncta Fil. ”; “ Уссур. Край верх р. Супутинки Ник. Усс. р-на Куренцов 5. VII. 1933 [Ussuri Region, upper Suputinka River] ”, gold circle, “ Paratypus Paratorna seriepuncta Fil. ”. Present status. Paratorna catenulella (Christoph, 1882).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFE95C05A390FB05FBBFF897.taxon	discussion	Comments. Genitalia of both sexes were illustrated in the original description.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFE95C04A390F897FBBFFE73.taxon	description	Peronea similis Filipjev, 1931. Annuaire du Musée Zoologique de l’Académie des Sciences de l’URSS 1931 (1930): 515. Plate A, figs 2 - 3, Plate XXIII, figs 2, 2 a, 2 b, Plate XXIX, fig. 3. TL: Russia, “ Jakovlevka, Daubiche-Wald ” (Far East, Yakovlevka). Types. Holotype ♂: “ Яковлевка Спас. у. Уссур. кр. 17. IX. 926 ДьЯконов, Филипьев [Yakovlevka, Ussuri Region, 17. IX. 1926 Djakonov, Filipjev] ”, “ Лес по раЗливу р. ДаубихЭ [Forest nearby the river Daubikhe (= Arsen’yevka)] ”, gold circle, “ praep. micr. № 1001 ”, “ Holotypus Peronea similis Filipjev, 1931 ”, “ ZISP INS _ LEP _ 0001963 ”, “ Praep. micr. SVN 24196 Acleris similis (Filipjev, 1931) S. Nedoshivina pr. 2024 ”. Paratype: ♀ – “ ♀ Anitino Amur | 19.5.77 Hed. [emann] ab. Buringerana ”, gold circle, “ Кол. Вел. Кн. НиколаЯ Михайловича [coll. of the Grand Duke Nikolay Mihailovich] ”, “ praep. micr. № 1239 | Ac. species ♀ Amur ”, “ Allotypus Peronea similis Fil. ”, “ Praep. micr. SVN 24195 Acleris similis (Filipjev, 1931) S. Nedoshivina pr. 2024 ”. Present status. Acleris similis (Filipjev, 1931).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFE95C04A390F897FBBFFE73.taxon	discussion	Comments. Genitalia of both sexes were illustrated in the original description.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFE85C04A390FD8AFBF9FC64.taxon	description	Present status. Stenopteron stenoptera (Filipjev, 1962).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFE85C04A390FD8AFBF9FC64.taxon	discussion	Comments. Genitalia of the male holotype were illustrated in the original description. Later, in his 2001 monograph, Kuznetzov reproduced the same drawing, slightly modified in his own style.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFE85C04A390FBF8FAF3F905.taxon	description	Peronea strigifera Filipjev, 1931. Annuaire du Musée Zoologique de l’Académie des Sciences de l’URSS 1931 (1930): 518. Plate B, figs 1 - 3; Plate XXVI, fig. 2; Plate XXXII, fig. 2. TL: Russia, “ Vinogradovka ” (Far East, Vinogradovka). Types. Holotype ♂: “ Виноградовка Уссур. кр. 28. V. 929 ДьЯконов, Филип. [Vinogradovka, Ussuri Region, 28. V. 1929 Djakonov, Filipjev] ”, gold circle, “ praep. micr. № 1778 ”, “ Holotypus Peronea strigifera Fil. ”, “ ZISP INS _ LEP _ 0001977 ”, “ Praep. micr. SVN 24190 Acleris strigifera (Filipjev, 1931) S. Nedoshivina pr. 2024 ”. Paratypes: ♂ and ♀ – ♂ “ Виноградовка Уссур. кр. 20. V. 929 ДьЯконов Филип. [Vinogradovka Ussuri Region 20. V. 929 Djakonov, Filipjev] ”, gold circle, “ praep. micr. 1773 | Per. strigifera Fil. ”, “ Paratypus Peronea strigifera Fil. ”; ♀ “ Виноградовка Уссур. кр. 22. V. 929 ДьЯконов Филип. [Vinogradovka Ussuri Region 22. V. 929 Djakonov, Filipjev] ”, gold circle, “ praep. micr. 1773 | Per. strigifera Fil. ”, “ Gemalt von Mgr. R. Zawada-Skrzyński für Micr. Pal. ”, “ Allotypus Peronea strigifera Fil. ”. Present status. Acleris strigifera (Filipjev, 1931).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFE85C04A390FBF8FAF3F905.taxon	discussion	Comments. Gilligan et al. (2018) listed the type locality imprecisely as “ Ussuri ” and marked the holotype sex as “ unknown ”. Genitalia of both sexes were illustrated in the original description. The genitalia of the male holotype were also illustrated in Razowski (2008), and those of the female paratype were illustrated in Razowski (1984).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFE85C03A390F919FAD0FE73.taxon	description	Polychrosis subherculeana Filipjev, 1924. Jahrbuch des Martjanov’schen Staatsmuseums in Minussinsk 2 (3): 55. TL: Russia, “ Джойское оЗ. [Dzhoyskoye Lake] ” (Krasnoyarsk Region, Dzhoyka Lake). Types. Lectotype ♂ (here designated): “ Окр. Минусинска 15. VI 1923 Кожанчиковъ [Outskirts of Minusinsk 15. VI 1923 Kozhantschikov] ”, “ Polychrosis subherculeana Fil. ♂ Type ”, “ praep. micr. № 329 ”, “ к. Н. Филипьева [coll. of N. Filipjev] ”, “ Holotypus Polychrosis subherculeana Fil. ♂ ”, “ ZISP INS _ LEP _ 0001982 ”. Paralectotypes: 2 ♂ – “ Окр. Минусинска 10. VI 1923 Кожанчиковъ [Outskirts of Minusinsk 10. VI 1923 Kozhantschikov] ”, gold circle, “ Polychrosis subherculeana Fil. ♂ Type ”, “ к. Н. Филипьева [coll. of N. Filipjev] ”, “ Paralectotypus Polychrosis subherculeana Fil. ♂ ”; “ Окр. Минусинска 14. VI 1923 Кожанчиковъ [Outskirts of Minusinsk 14. VI 1923 Kozhantschikov] ”, gold circle, “ Polychrosis subherculeana Fil. ♂ Type ”, “ к. Н. Филипьева [coll. of N. Filipjev] ”, “ Paralectotypus Polychrosis subherculeana Fil. ♂ ”, “ Photograph Marek Kopeć ”. Present status. Lobesia subherculeana (Filipjev, 1924).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFE85C03A390F919FAD0FE73.taxon	discussion	Comments. In Gilligan at al. (2018), the type locality was given erroneously as “ Siberia, Minsk ” and the holotype sex was marked as unknown. Taxonomic notes. The genitalia slide of the male lectotype was prepared by Falkovitsh, and the label “ Holotypus Polychrosis subherculeana Fil. ♂ ” was likely written by him. However, since the holotype of subherculeana was not designated in the original description, this specimen should be treated as the lectotype (here designated).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFEF5C03A390FD8AFAADFBF4.taxon	description	Paratype: ♂ – “ Култукъ, ю. - З. Байкалъ Чекановскiй 14 V 78 [Kultuk, southwest Baikal, Chekanovskiy 14 V 1878] ”, “ praep. micr. № 3125 | Peronea submaccana Fil. ♂ Байкал [Baikal] ”, “ Paratypus Peronea submaccana Fil. ”. Present status. Acleris submaccana (Filipjev, 1962).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFEF5C03A390FD8AFAADFBF4.taxon	discussion	Comments. In Gilligan et al. (2018), the type locality was imprecisely listed as “ Siberia ”. Another male specimen in the ZISP collection, labeled “ Минусинск 20. V. 24, Филипьев [Minusinsk, 20. V. 24, Filipjev] ”, with a gold circle, “ praep. micr. № 1404 | Ac. submaccana Fil. ♂ Минус. [Minus.] ”, “ Paratypus Peronea submaccana Fil. ” should not be treated as a type specimen, despite the gold circle, as it was not mentioned in the original description.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFEF5C02A390FB08FF7CFE2B.taxon	description	Ancylis transientana Filipjev, 1926. Archiv für Naturgeschichte 91 (1) (1925): 117. TL: Russia, “ Archierejskij Wald ” (Stavropol). Types. Lectotype ♂ (here designated): “ Ставрополь Кавк. Арх. л. 16. V. 1921 [Stavropol Cauc. [asus] Archiyereyskiy forest 16. V. 1921], white circle, gold circle, “ Ancylis transientana Fil. ♂ type N. Filipjev det. ”, “ к Н. Филипьева [collection of N. Filipjev] ”, “ Photograph Marek Kopeć ”, “ Lectotypus Ancylis transientana Filipjev, 1926 ”, ” Praep. micr. SVN 24224 Ancylis transientana Filipjev, 1926 S. Nedoshivina pr. 2024 ”. Paralectotypes: 5 ♂ and 3 ♀ – 5 ♂: “ Б. ПожариЩе [Bolshoe Pozharitshe] 6000 ’ 17. VI. 16 [unreadable] ”, “ An. transientana Fil. ♂ N. Filipjev det. ”, “ к Н. Филипьева [collection of N. Filipjev] ”, “ Paralectotypus Ancylis transientana Fil. ”; same labels, “ ZISP INS _ LEP _ 0002035 ”; “ Mangl. [is] ♂ | 19.5.84 Hed. transientana ”, “ колл. б. Вел. Кн. НиколаЯ Михайловича [coll. of the former Grand Duke Nikolay Mihailovich] ”, “ Anc. transientana ♂ Fil. type N. Filipjev det. ”, blue circle, gold circle, “ Paralectotypus Ancylis transientana Fil. ”,; “ Ставрополь Кавк. Майский лес 13. V. 21 [Stavropol Caucas. Mayskiy forest 13. V. 21] ”, gold circle, “ Anc. transientana ♂ Fil. type N. Filipjev det. ”, “ к Н. Филипьева [collection of N. Filipjev] ”, “ Paralectotypus Ancylis transientana Fil. ”; “ Borshom ”, blue circle, gold circle, “ praep. micr. № 1132 | Anc. transientana Fil. ♂ ♀ ”, “ Paralectotypus Ancylis transientana Fil. ”; 3 ♀: “ Suchum | 12.5.80 / 30.4 Chr. transientana ”, blue circle, gold circle, “ Anc. transientana Fil. ♀ type N. Filipjev det. ”, “ колл. б. Вел. Кн. НиколаЯ Михайловича [coll. of the former Grand Duke Nikolay Mihailovich] ”, “ Photograph Marek Kopeć ”, “ Paralectotypus Ancylis transientana Fil. ”; “ Mangl. [is] ♀ | Hed. 10.5.84 transientana ”, blue circle, gold circle, “ колл. б. Вел. Кн. НиколаЯ Михайловича [coll. of the former Grand Duke Nikolay Mihailovich] ”, “ Anc. transientana Fil. ♀ type N. Filipjev det. ”, “ Paralectotypus Ancylis transientana Fil. ”, “ Praep. micr. SVN 24223 Ancylis transientana Filipjev, 1926 S. Nedoshivina pr. 2024 ”; “ Borshom. ♀ | 30.6.80 Chr. ”, blue circle, gold circle, “ колл. б. Вел. Кн. НиколаЯ Михайловича [coll. of the former Grand Duke Nikolay Mihailovich] ”, “ Anc. transientana Fil. ♂ ♀ typen N. Filipjev det. ”, “ брюшко потерЯно [abdomen lost] ”, “ Paralectotypus Ancylis transientana Fil. ”. Present status. Ancylis transientana Filipjev, 1926.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFEF5C02A390FB08FF7CFE2B.taxon	discussion	Comments. Genitalia of both sexes were illustrated in the original description. Gilligan et al. (2018) listed the type locality imprecisely as “ Russia ” and noted that the holotype is unknown. The specific name transientana is misspelled in the original description as transienta (p: 117, lapsus calami); corrected directly in the same publication (p: 183).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFEE5C02A390FBDCFE9FF969.taxon	description	Paratypes: 5 ♂ – “ Уссур. край, Сучан, Цамо-ДынЗа, 6. VII. 28, Куренцов [Ussuri Region, Suchan, Tsamo-Dynza, 6. VII. 28, Kurentsov] ”, “ praep. micr. № 2555 | Cn. ussurica Fil. ♂ Ussuri ”, “ Paratypus Cnephasia ussurica Fil. ”; “ Уссур. край, Сучан, Тигровое, 8. VII. 28, Куренцов [Ussuri Region, Suchan, Tigrovoye, 8. VII. 28, Kurentsov] ”, “ Paratypus Cnephasia ussurica Fil. ”; “ Уссур. край, Сучан, 6. VII. 28, Куренцов [Ussuri Region, Suchan, 6. VII. 28, Kurentsov] ”, “ praep. micr. № 2553 | Cn. ussurica Fil. ♂ Ussuri ”, “ N 2553 ussurica ”, “ Paratypus Cnephasia ussurica Fil. ”; “ Уссур. край, Сучан, Тигровое, 24. VII. 28, Куренцов [Ussuri Region, Suchan, Tigrovoye, 24. VII. 28, Kurentsov] ”, “ Paratypus Cnephasia ussurica Fil. ”. Present status. Cnephasia ussurica Filipjev, 1962.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFEE5C02A390FBDCFE9FF969.taxon	discussion	Comments. In the original description, Filipjev noted Tigrovoye as the type locality; however, the label on the holotype specifies Tsamo-Dynza. Since Tsamo-Dynza Mountain is located near Tigrovoye, we consider Tigrovoye as the type locality in accordance with the original description. Male genitalia were illustrated in the original description.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFEE5C02A390FE32FC77FC38.taxon	description	Peronea uniformis Filipjev, 1931. Annuaire du Musée Zoologique de l’Académie des Sciences de l’URSS 1931 (1930): 512. Plate A, fig. 1; Plate XXIX, fig. 2. TL: Russia, “ Jakovlevka, Leško’s Bienenstand ” (Far East, Yakovlevka). Types. Holotype ♀: “ Яковлевка Спас. у. Уссур. кр. 12. IX. 926 ДьЯконов, Филипьев [Yakovlevka, Ussuri Region, 12. IX 1926 Djakonov, Filipjev] ”, “ Пасека И. Лешко [I. Leshko’s bee-garden] ”, “ praep. micr. № 1003 ”, gold circle, ” Gemalt fon Mgr. Zawada-Skrzyński für Micr. Pal. ”, “ Holotypus Peronea uniformis Fil. ”, “ ZISP INS _ LEP _ 0002035 ”. Present status. Acleris uniformis (Filipjev, 1931).	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
5857225FFFEE5C02A390FE32FC77FC38.taxon	discussion	Comments. Genitalia of the female holotype were illustrated in the original description. Genitalia preparation was probably lost, but it was figured also by Razowski (2008) in good quality. Gilligan et al. (2018) listed the type locality imprecisely as “ Ussuri ” and marked the holotype sex as “ unknown ”.	en	Nedoshivina, Svetlana V. (2025): An annotated list of the type specimens of Tortricidae (Lepidoptera) in the collection of Zoological Institute, St. Petersburg, Russia. 2. Types of species described by N. N. Filipjev. Zootaxa 5627 (3): 480-502, DOI: 10.11646/zootaxa.5627.3.4, URL: https://doi.org/10.11646/zootaxa.5627.3.4
