taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
48D8371928B75D86B68EE960A1D40603.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1788958	https://doi.org/10.3897/phytokeys.279.181697.figure1	Figure 1. Illustration of Elymus chaobotryus Y. C. Zhang, sp. nov. (1) habit; (2) pendulous compound spike showing lateral spike-bearing branchlets; (3) spikelet; (4) caryopsis, dorsal view; (5) caryopsis, ventral view; (6) lemma awn; (7) lemma; (8) palea. Drawn by Yuan Liu.	Figure 1. Illustration of Elymus chaobotryus Y. C. Zhang, sp. nov. (1) habit; (2) pendulous compound spike showing lateral spike-bearing branchlets; (3) spikelet; (4) caryopsis, dorsal view; (5) caryopsis, ventral view; (6) lemma awn; (7) lemma; (8) palea. Drawn by Yuan Liu.	2026-09-24	Zhang, Yong-Chao;Xiao, Sa;Ma, Xin-Yun;Chen, Ji-Gui;Qin, Yan;Liu, Yong;Guo, Zhi-Ying;Zhou, Yu-Qing;Zhao, Dian-Zhi;Shan, Qing-Sheng;Lu, Hai-Zhang;Wei, Xiao-Xing		Zenodo	biologists	Zhang, Yong-Chao;Xiao, Sa;Ma, Xin-Yun;Chen, Ji-Gui;Qin, Yan;Liu, Yong;Guo, Zhi-Ying;Zhou, Yu-Qing;Zhao, Dian-Zhi;Shan, Qing-Sheng;Lu, Hai-Zhang;Wei, Xiao-Xing			
48D8371928B75D86B68EE960A1D40603.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1788959	https://doi.org/10.3897/phytokeys.279.181697.figure2	Figure 2. Holotype specimen of Elymus chaobotryus (389653), deposited in the Qinghai-Tibetan Plateau Museum of Biology (HNWP).	Figure 2. Holotype specimen of Elymus chaobotryus (389653), deposited in the Qinghai-Tibetan Plateau Museum of Biology (HNWP).	2026-09-24	Zhang, Yong-Chao;Xiao, Sa;Ma, Xin-Yun;Chen, Ji-Gui;Qin, Yan;Liu, Yong;Guo, Zhi-Ying;Zhou, Yu-Qing;Zhao, Dian-Zhi;Shan, Qing-Sheng;Lu, Hai-Zhang;Wei, Xiao-Xing		Zenodo	biologists	Zhang, Yong-Chao;Xiao, Sa;Ma, Xin-Yun;Chen, Ji-Gui;Qin, Yan;Liu, Yong;Guo, Zhi-Ying;Zhou, Yu-Qing;Zhao, Dian-Zhi;Shan, Qing-Sheng;Lu, Hai-Zhang;Wei, Xiao-Xing			
48D8371928B75D86B68EE960A1D40603.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1788960	https://doi.org/10.3897/phytokeys.279.181697.figure3	Figure 3. Elymus chaobotryus in the wild. A. Habit of a clump-forming plant; B, C. Close-ups of the pendulous compound spikes, showing short lateral branchlets bearing spikelets; D. Culms and leaves. Photographs by Yong-Chao Zhang.	Figure 3. Elymus chaobotryus in the wild. A. Habit of a clump-forming plant; B, C. Close-ups of the pendulous compound spikes, showing short lateral branchlets bearing spikelets; D. Culms and leaves. Photographs by Yong-Chao Zhang.	2026-09-24	Zhang, Yong-Chao;Xiao, Sa;Ma, Xin-Yun;Chen, Ji-Gui;Qin, Yan;Liu, Yong;Guo, Zhi-Ying;Zhou, Yu-Qing;Zhao, Dian-Zhi;Shan, Qing-Sheng;Lu, Hai-Zhang;Wei, Xiao-Xing		Zenodo	biologists	Zhang, Yong-Chao;Xiao, Sa;Ma, Xin-Yun;Chen, Ji-Gui;Qin, Yan;Liu, Yong;Guo, Zhi-Ying;Zhou, Yu-Qing;Zhao, Dian-Zhi;Shan, Qing-Sheng;Lu, Hai-Zhang;Wei, Xiao-Xing			
48D8371928B75D86B68EE960A1D40603.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1788961	https://doi.org/10.3897/phytokeys.279.181697.figure4	Figure 4. Pendulous compound spikes of Elymus chaobotryus from field-grown (cultivated) plants. A. Compound spike at anthesis; B. Compound spike at maturity; C. Excised compound spike with ruler (cm) showing overall size; D. Excised compound spike with lateral branchlets gently splayed to display the branching pattern and their insertion along the main rachis (ruler in cm).	Figure 4. Pendulous compound spikes of Elymus chaobotryus from field-grown (cultivated) plants. A. Compound spike at anthesis; B. Compound spike at maturity; C. Excised compound spike with ruler (cm) showing overall size; D. Excised compound spike with lateral branchlets gently splayed to display the branching pattern and their insertion along the main rachis (ruler in cm).	2026-09-24	Zhang, Yong-Chao;Xiao, Sa;Ma, Xin-Yun;Chen, Ji-Gui;Qin, Yan;Liu, Yong;Guo, Zhi-Ying;Zhou, Yu-Qing;Zhao, Dian-Zhi;Shan, Qing-Sheng;Lu, Hai-Zhang;Wei, Xiao-Xing		Zenodo	biologists	Zhang, Yong-Chao;Xiao, Sa;Ma, Xin-Yun;Chen, Ji-Gui;Qin, Yan;Liu, Yong;Guo, Zhi-Ying;Zhou, Yu-Qing;Zhao, Dian-Zhi;Shan, Qing-Sheng;Lu, Hai-Zhang;Wei, Xiao-Xing			
48D8371928B75D86B68EE960A1D40603.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1788962	https://doi.org/10.3897/phytokeys.279.181697.figure5	Figure 5. Structure of the compound spike in Elymus chaobotryus. A. Portion of a compound spike showing a node with a lateral branchlet; B, C. Close-up views of lateral branchlets bearing several spikelets; D. An excised node from the main rachis, showing one elongated lateral branchlet together with two sessile spikelets that lack branchlet elongation.	Figure 5. Structure of the compound spike in Elymus chaobotryus. A. Portion of a compound spike showing a node with a lateral branchlet; B, C. Close-up views of lateral branchlets bearing several spikelets; D. An excised node from the main rachis, showing one elongated lateral branchlet together with two sessile spikelets that lack branchlet elongation.	2026-09-24	Zhang, Yong-Chao;Xiao, Sa;Ma, Xin-Yun;Chen, Ji-Gui;Qin, Yan;Liu, Yong;Guo, Zhi-Ying;Zhou, Yu-Qing;Zhao, Dian-Zhi;Shan, Qing-Sheng;Lu, Hai-Zhang;Wei, Xiao-Xing		Zenodo	biologists	Zhang, Yong-Chao;Xiao, Sa;Ma, Xin-Yun;Chen, Ji-Gui;Qin, Yan;Liu, Yong;Guo, Zhi-Ying;Zhou, Yu-Qing;Zhao, Dian-Zhi;Shan, Qing-Sheng;Lu, Hai-Zhang;Wei, Xiao-Xing			
48D8371928B75D86B68EE960A1D40603.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1788963	https://doi.org/10.3897/phytokeys.279.181697.figure6	Figure 6. Spikelet morphology of Elymus chaobotryus. A. Spikelet, lateral view, showing the paired glumes enclosing the florets and an awned lemma; B. Glumes, abaxial (outer) faces; C. Glumes, adaxial (inner) faces; D. Palea; E. Lemma with distal awn; F. Awn, enlarged detail.	Figure 6. Spikelet morphology of Elymus chaobotryus. A. Spikelet, lateral view, showing the paired glumes enclosing the florets and an awned lemma; B. Glumes, abaxial (outer) faces; C. Glumes, adaxial (inner) faces; D. Palea; E. Lemma with distal awn; F. Awn, enlarged detail.	2026-09-24	Zhang, Yong-Chao;Xiao, Sa;Ma, Xin-Yun;Chen, Ji-Gui;Qin, Yan;Liu, Yong;Guo, Zhi-Ying;Zhou, Yu-Qing;Zhao, Dian-Zhi;Shan, Qing-Sheng;Lu, Hai-Zhang;Wei, Xiao-Xing		Zenodo	biologists	Zhang, Yong-Chao;Xiao, Sa;Ma, Xin-Yun;Chen, Ji-Gui;Qin, Yan;Liu, Yong;Guo, Zhi-Ying;Zhou, Yu-Qing;Zhao, Dian-Zhi;Shan, Qing-Sheng;Lu, Hai-Zhang;Wei, Xiao-Xing			
48D8371928B75D86B68EE960A1D40603.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1788964	https://doi.org/10.3897/phytokeys.279.181697.figure7	Figure 7. Caryopsis (grain) of Elymus chaobotryus at two developmental stages. A, B. Grain-filling stage: A. Dorsal side; B. Ventral side showing the ventral groove; C, D. Mature stage: C. Dorsal side; D. Ventral side with a conspicuous ventral groove.	Figure 7. Caryopsis (grain) of Elymus chaobotryus at two developmental stages. A, B. Grain-filling stage: A. Dorsal side; B. Ventral side showing the ventral groove; C, D. Mature stage: C. Dorsal side; D. Ventral side with a conspicuous ventral groove.	2026-09-24	Zhang, Yong-Chao;Xiao, Sa;Ma, Xin-Yun;Chen, Ji-Gui;Qin, Yan;Liu, Yong;Guo, Zhi-Ying;Zhou, Yu-Qing;Zhao, Dian-Zhi;Shan, Qing-Sheng;Lu, Hai-Zhang;Wei, Xiao-Xing		Zenodo	biologists	Zhang, Yong-Chao;Xiao, Sa;Ma, Xin-Yun;Chen, Ji-Gui;Qin, Yan;Liu, Yong;Guo, Zhi-Ying;Zhou, Yu-Qing;Zhao, Dian-Zhi;Shan, Qing-Sheng;Lu, Hai-Zhang;Wei, Xiao-Xing			
48D8371928B75D86B68EE960A1D40603.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1788965	https://doi.org/10.3897/phytokeys.279.181697.figure8	Figure 8. Florets of Elymus chaobotryus during grain filling. A. Floret with the palea removed (lemma retained), exposing the developing caryopsis (ovary) and basal lodicules; parts of the pistil and stamens visible; B. A different floret with both lemma and palea removed, showing the pistil (ovary with two plumose stigmas) and three stamens with slender filaments; C. Anther, close-up, dorsifixed and dehiscing by longitudinal slits, with pollen grains visible.	Figure 8. Florets of Elymus chaobotryus during grain filling. A. Floret with the palea removed (lemma retained), exposing the developing caryopsis (ovary) and basal lodicules; parts of the pistil and stamens visible; B. A different floret with both lemma and palea removed, showing the pistil (ovary with two plumose stigmas) and three stamens with slender filaments; C. Anther, close-up, dorsifixed and dehiscing by longitudinal slits, with pollen grains visible.	2026-09-24	Zhang, Yong-Chao;Xiao, Sa;Ma, Xin-Yun;Chen, Ji-Gui;Qin, Yan;Liu, Yong;Guo, Zhi-Ying;Zhou, Yu-Qing;Zhao, Dian-Zhi;Shan, Qing-Sheng;Lu, Hai-Zhang;Wei, Xiao-Xing		Zenodo	biologists	Zhang, Yong-Chao;Xiao, Sa;Ma, Xin-Yun;Chen, Ji-Gui;Qin, Yan;Liu, Yong;Guo, Zhi-Ying;Zhou, Yu-Qing;Zhao, Dian-Zhi;Shan, Qing-Sheng;Lu, Hai-Zhang;Wei, Xiao-Xing			
48D8371928B75D86B68EE960A1D40603.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1788966	https://doi.org/10.3897/phytokeys.279.181697.figure9	Figure 9. Chromosome counting, GISH and FISH analyses of Elymus chaobotryus. A. DAPI staining showing 2 n = 42; B. GISH identifying St-genome chromosomes; C. Sequential FISH distinguishing chromosome groups interpreted as H and Y genomes. Scale bars: 10 μm.	Figure 9. Chromosome counting, GISH and FISH analyses of Elymus chaobotryus. A. DAPI staining showing 2 n = 42; B. GISH identifying St-genome chromosomes; C. Sequential FISH distinguishing chromosome groups interpreted as H and Y genomes. Scale bars: 10 μm.	2026-09-24	Zhang, Yong-Chao;Xiao, Sa;Ma, Xin-Yun;Chen, Ji-Gui;Qin, Yan;Liu, Yong;Guo, Zhi-Ying;Zhou, Yu-Qing;Zhao, Dian-Zhi;Shan, Qing-Sheng;Lu, Hai-Zhang;Wei, Xiao-Xing		Zenodo	biologists	Zhang, Yong-Chao;Xiao, Sa;Ma, Xin-Yun;Chen, Ji-Gui;Qin, Yan;Liu, Yong;Guo, Zhi-Ying;Zhou, Yu-Qing;Zhao, Dian-Zhi;Shan, Qing-Sheng;Lu, Hai-Zhang;Wei, Xiao-Xing			
