taxonID	type	description	language	source
5B7A156CFFBFD1277A56FB8E61B4FD30.taxon	description	urn: lsid: zoobank. org: act: A 47 E 096 C-BDC 2 - 4038 - 97 C 3 - 82309676 D 9 E 3 Figs 1 – 2, 5 – 8	en	Johanson, Kjell Arne, Pham, Hong-Thai (2025): Description of three new species of Wormaldia McLachlan, 1865 from Vietnam (Trichoptera: Philopotamidae). European Journal of Taxonomy 1007: 24-34, DOI: 10.5852/ejt.2025.1007.2991, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2991/13451
5B7A156CFFBFD1277A56FB8E61B4FD30.taxon	diagnosis	Diagnosis The male genitalia of Wormaldia piadenensis sp. nov. exhibit a notable similarity to those of Wormaldia muoihai, particularly in the presence of downwardly curving harpagones, which are of equal height at mid-length and at the base, and are each shorter than the coxopodites. However, W. piadenensis can be distinguished from W. muoihai by several key characteristics in the male genitalia: segment IX is shorter, segment IX possesses a distinct posteroventral process, the dorsal margin of the coxopodites is more sharply produced dorsally, and the phallus features a single posterior curved sclerite and a long, needleshaped anterior sclerite, in contrast to the two short, straight sclerites present in W. muoihai.	en	Johanson, Kjell Arne, Pham, Hong-Thai (2025): Description of three new species of Wormaldia McLachlan, 1865 from Vietnam (Trichoptera: Philopotamidae). European Journal of Taxonomy 1007: 24-34, DOI: 10.5852/ejt.2025.1007.2991, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2991/13451
5B7A156CFFBFD1277A56FB8E61B4FD30.taxon	etymology	Etymology Wormaldia piadenensis sp. nov. was named after Phia Dén Village, near the type locality of the species.	en	Johanson, Kjell Arne, Pham, Hong-Thai (2025): Description of three new species of Wormaldia McLachlan, 1865 from Vietnam (Trichoptera: Philopotamidae). European Journal of Taxonomy 1007: 24-34, DOI: 10.5852/ejt.2025.1007.2991, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2991/13451
5B7A156CFFBFD1277A56FB8E61B4FD30.taxon	materials_examined	Type material Holotype VIETNAM • ♂ (in alcohol); Cao Bang Province, N Nui Pia Oac Nature Reserve, about 9.5 km (rd) N of Phia Dén Village, 100 m upstream of bridge; 10 Apr. 2012; Johanson and Pham leg.; light trap at stream with mixed stream bottom; loc # VN 028; VNMN.	en	Johanson, Kjell Arne, Pham, Hong-Thai (2025): Description of three new species of Wormaldia McLachlan, 1865 from Vietnam (Trichoptera: Philopotamidae). European Journal of Taxonomy 1007: 24-34, DOI: 10.5852/ejt.2025.1007.2991, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2991/13451
5B7A156CFFBFD1277A56FB8E61B4FD30.taxon	description	Description (holotype) Forewing (Fig. 2) 5.5 mm (N = 1), membrane light brown, veins dark brown. Forks 1 – 5 present; fork 1 originates at distal corner of Dc; basis of M 1 + 2 and Ms hyaline; arc located opposite to base of fork 5. Abdominal segments VII and VIII with long posteroventral lobes (Figs 5, 7). In lateral view (Fig. 5), segment IX anteriorly triangular and posteroventral part produced posteriorly; coxopodites with almost straight ventral margin, dorsal margin slightly produced into triangular; harpagones almost uniformly curving ventrad along their length, about ½ × as long as coxopodites, each with rounded apex. In dorsal view (Fig. 6), segment IX with wide U-shaped anterior margin; tergum X triangular, with few setae. In ventral view (Fig. 7), with sternum IX produced into small, narrow triangle; basal half of coxopodites fused, distal half with irregularly shaped mesal margin bearing stout setae at posteromesal corner; harpagones almost straight, with small mesal process at mid-length. Phallus (Fig. 8) with one pair of very short sclerites running along its length, and one sickle-shaped sclerite near apex.	en	Johanson, Kjell Arne, Pham, Hong-Thai (2025): Description of three new species of Wormaldia McLachlan, 1865 from Vietnam (Trichoptera: Philopotamidae). European Journal of Taxonomy 1007: 24-34, DOI: 10.5852/ejt.2025.1007.2991, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2991/13451
5B7A156CFFBDD1267A84FD0D6428FE32.taxon	description	urn: lsid: zoobank. org: act: CB 2 E 57 C 6 - 4 B 85 - 4619 - 8 D 95 - F 40 DC 1 A 02 ECB Figs 1, 3, 9 – 12	en	Johanson, Kjell Arne, Pham, Hong-Thai (2025): Description of three new species of Wormaldia McLachlan, 1865 from Vietnam (Trichoptera: Philopotamidae). European Journal of Taxonomy 1007: 24-34, DOI: 10.5852/ejt.2025.1007.2991, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2991/13451
5B7A156CFFBDD1267A84FD0D6428FE32.taxon	diagnosis	Diagnosis The male genitalia of Wormaldia nui sp. nov. closely resemble those of Wormaldia muoibon, particularly in lateral view, where the harpagones are almost straight and parallel-sided, and of equal length as the coxopodites. Additionally, tergite VIII is produced above segment IX. However, W. nui is distinguishable from W. muoibon based on several male genital features: in W. nui, the harpagones are parallel-sided in ventral view, whereas they slightly taper in W. muoibon; further, the phallus of W. nui has two short, straight sclerites at the posterior end, in contrast to the three longer, straight sclerites found in W. muoibon.	en	Johanson, Kjell Arne, Pham, Hong-Thai (2025): Description of three new species of Wormaldia McLachlan, 1865 from Vietnam (Trichoptera: Philopotamidae). European Journal of Taxonomy 1007: 24-34, DOI: 10.5852/ejt.2025.1007.2991, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2991/13451
5B7A156CFFBDD1267A84FD0D6428FE32.taxon	etymology	Etymology Wormaldia nui sp. nov. was named after the Nui Pia Oac Nature Reserve, the type locality of the species.	en	Johanson, Kjell Arne, Pham, Hong-Thai (2025): Description of three new species of Wormaldia McLachlan, 1865 from Vietnam (Trichoptera: Philopotamidae). European Journal of Taxonomy 1007: 24-34, DOI: 10.5852/ejt.2025.1007.2991, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2991/13451
5B7A156CFFBDD1267A84FD0D6428FE32.taxon	materials_examined	Type material Holotype VIETNAM • ♂ (in alcohol); Cao Bang Province, N Nui Pia Oac Nature Reserve, about 9.5 km (rd) N of Phia Dén Village, 100 m upstream of bridge; 10 Apr. 2012; Johanson and Pham leg.; light trap at stream with mixed stream bottom; loc # VN 028; VNMN. Paratype VIETNAM • 1 ♂ (in alcohol); Cao Bang Province, Nui Pia Oac Nature Reserve, about 200 m downstream on rd to radio tower; 10 Apr. 2012; Johanson and Pham leg.; light trap at village center; loc # VN 030; NHRS.	en	Johanson, Kjell Arne, Pham, Hong-Thai (2025): Description of three new species of Wormaldia McLachlan, 1865 from Vietnam (Trichoptera: Philopotamidae). European Journal of Taxonomy 1007: 24-34, DOI: 10.5852/ejt.2025.1007.2991, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2991/13451
5B7A156CFFBDD1267A84FD0D6428FE32.taxon	description	Description (holotype) Forewing (Fig. 3) 4.5 mm (N = 1), membrane light brown, veins dark brown. Forks 1 – 5 present; fork 1 originates shortly before distal corner of Dc; basis of M 1 + 2 and Ms hyaline; arc located slightly distally of base of fork 5. Abdominal segments VII and VIII with long posteroventral lobes (Figs 9, 11). In lateral view (Fig. 8), segment IX forming triangular anteriorly, posteroventral corner not produced posteriorly; coxopodites with slightly convex ventral margin, dorsal margin strongly convex; harpagones straight, with almost parallel-sided dorsal and ventral margins, about as long as coxopodites, each with rounded apex. In dorsal view (Fig. 10), segment IX with wide U-shaped anterior margin; tergum X with basal three-quarters with parallel-sided lateral margins, with two pairs of short setae before narrowing apex. In ventral view (Fig. 11), with sternum IX with small central incision at posterior margin; basal twothirds of coxopodites fused, each with distal half of mesal margin straight, bearing few stout setae near posteromesal corner; harpagones almost straight, without mesal process. Phallus (Fig. 12) with thin sclerites in group before mid-length and one pair of short, triangular sclerites situated close to each other near apex.	en	Johanson, Kjell Arne, Pham, Hong-Thai (2025): Description of three new species of Wormaldia McLachlan, 1865 from Vietnam (Trichoptera: Philopotamidae). European Journal of Taxonomy 1007: 24-34, DOI: 10.5852/ejt.2025.1007.2991, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2991/13451
5B7A156CFFB3D1287A5BFEEA6792FD57.taxon	description	urn: lsid: zoobank. org: act: 412 B 7 D 98 - 4 ED 4 - 4 DAB-BA 79 - 3 F 7 F 50 EAED 53 Figs 1, 4, 13 – 16	en	Johanson, Kjell Arne, Pham, Hong-Thai (2025): Description of three new species of Wormaldia McLachlan, 1865 from Vietnam (Trichoptera: Philopotamidae). European Journal of Taxonomy 1007: 24-34, DOI: 10.5852/ejt.2025.1007.2991, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2991/13451
5B7A156CFFB3D1287A5BFEEA6792FD57.taxon	diagnosis	Diagnosis The male genitalia of Wormaldia caobangensis sp. nov. exhibit similarities to those of Wormaldia muoihai, particularly in the presence of the downwardly curving harpagones that are of equal height at mid-length and at the base, and are shorter than the coxopodites. However, in lateral view, W. caobangensis is distinguishable from W. muoihai by the more prominently produced anterior segment IX.	en	Johanson, Kjell Arne, Pham, Hong-Thai (2025): Description of three new species of Wormaldia McLachlan, 1865 from Vietnam (Trichoptera: Philopotamidae). European Journal of Taxonomy 1007: 24-34, DOI: 10.5852/ejt.2025.1007.2991, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2991/13451
5B7A156CFFB3D1287A5BFEEA6792FD57.taxon	etymology	Etymology Wormaldia caobangensis sp. nov. was named after the Vietnamese Cao Bang Province, the type locality of the species.	en	Johanson, Kjell Arne, Pham, Hong-Thai (2025): Description of three new species of Wormaldia McLachlan, 1865 from Vietnam (Trichoptera: Philopotamidae). European Journal of Taxonomy 1007: 24-34, DOI: 10.5852/ejt.2025.1007.2991, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2991/13451
5B7A156CFFB3D1287A5BFEEA6792FD57.taxon	materials_examined	Type material Holotype VIETNAM • ♂ (in alcohol); Cao Bang Province, Nui Pia Oac Nature Reserve, N Phia Dén Village, upstream road to the radio tower; 8 – 10 Apr. 2012; Johanson and Pham leg.; Malaise trap across stream with large stones; loc # VN 024; VNMN.	en	Johanson, Kjell Arne, Pham, Hong-Thai (2025): Description of three new species of Wormaldia McLachlan, 1865 from Vietnam (Trichoptera: Philopotamidae). European Journal of Taxonomy 1007: 24-34, DOI: 10.5852/ejt.2025.1007.2991, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2991/13451
5B7A156CFFB3D1287A5BFEEA6792FD57.taxon	description	Description (holotype) Forewing (Fig. 4) 5.5 mm (N = 1), membrane light brown, veins dark brown. Forks 1 – 5 present; fork 1 originates shortly before distal corner of Dc; basis of M 1 + 2 and Ms hyaline; arc located slightly distally of base of fork 5. Abdominal segments VII and VIII with long posteroventral lobes (Figs 13, 15). In lateral view (Fig. 13), segment IX anteriorly triangular, posteroventral corner not produced posteriorly; coxopodites with slightly undulating ventral margin, dorsal margin strongly convex; harpagones with weakly convex dorsal margin and weakly concave margins, about 3 / 5 as long as coxopodites, each with rounded apex. In dorsal view (Fig. 14), segment IX with wide U-shaped anterior margin; tergum X hidden below triangular tergite VIII. In ventral view (Fig. 15), with sternum IX with small central incision at posterior margin; basal half of coxopodites fused, each with distal half of mesal margin almost straight, bearing scattered setae; harpagones almost straight, without mesal process. Phallus (Fig. 16) with one long, straight sclerite and one distally hook-shaped, slender sclerite, both originating approximately at mid-length of phallus.	en	Johanson, Kjell Arne, Pham, Hong-Thai (2025): Description of three new species of Wormaldia McLachlan, 1865 from Vietnam (Trichoptera: Philopotamidae). European Journal of Taxonomy 1007: 24-34, DOI: 10.5852/ejt.2025.1007.2991, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2991/13451
