identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
5B7A156CFFB8D1257A49FDFD606AFBAD.text	5B7A156CFFB8D1257A49FDFD606AFBAD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Wormaldia McLachlan 1865	<div><p>Genus Wormaldia McLachlan, 1865</p><p>Key to the Vietnamese male species of Wormaldia McLachlan, 1865</p><p>1. Harpagones are clearly shorter than coxopodites ............................................................................. 2</p><p>– Harpagones about as long as, or longer than coxopodites ................................................................ 9</p><p>2. Harpagones about straight in lateral view ........................................................................................ 3</p><p>– Harpagones curving ventrally in lateral view ................................................................................... 6</p><p>3. Tergite X exceeding coxopodites posteriorly ............................ Wormaldia sinocornuta Mey, 1996</p><p>– Tergite X not exceeding coxopodites posteriorly ............................................................................. 4</p><p>4. Harpagones almost parallel-sided in lateral view ......................... Wormaldia caobangensis sp. nov.</p><p>– Harpagones triangular in lateral view ............................................................................................... 5</p><p>5. Tergite X curving dorsally at apex; harpagones with rounded mesal projection in ventral view ....... .................................................................................................... Wormaldia meosorum (Mey, 1996)</p><p>– Tergite X curving pointing distally at apex; harpagones with mesal margin straight in ventral view ................................................................................................ Wormaldia muoinam (Malicky, 1995)</p><p>6. In lateral view, harpagones as high at mid-length as at base ............................................................ 7</p><p>– In lateral view, harpagones clearly lower at mid-length than at base ............................................... 8</p><p>7. In lateral view, coxopodites with straight ventral margin; dorsal margin with triangular projection oriented dorsally .............................................................................. Wormaldia piadenensis sp. nov.</p><p>– In lateral view, coxopodites with convex ventral margin; dorsal margin smoothly convex, without triangular projection ................................................................... Wormaldia muoihai Malicky, 1995</p><p>8. In lateral view, harpagones uniformly curving venrally along their length ........................................ ................................................................................................. Wormaldia nyctimon (Schmid, 1991)</p><p>– In lateral view, harpagones sharply bending ventrally before apex .................................................... ....................................................................................... Wormaldia sonlama Oláh &amp; Malicky, 2010</p><p>9. Harpagones curving ventrally in lateral view ................................................................................. 10</p><p>– Harpagones about straight in lateral view .......................................................................................11</p><p>10. Each posterodorsal corner of segment strongly prolonged into slender, ventrally curving posterior projection ................................................................................ Wormaldia muoisan (Malicky, 1995)</p><p>– Each posterodorsal corner of segment not prolonged posteriorly Wormaldia muoiba (Malicky, 1995)</p><p>11. Tergite VIII produced posteriorly above segment IX ..................................................................... 12</p><p>– Tergite VIII not produced posteriorly above segment IX ............................................................... 16</p><p>12. In lateral view, harpagones narrowing dorsally and ventrally at mid-length .................................. 13</p><p>– In lateral view, harpagones as high at mid-length as distally of mid-length .................................. 14</p><p>13. In lateral view, segment IX longer than coxopodites ................. Wormaldia muoimot Malicky, 1995</p><p>– In lateral view, segment IX as long as coxopodites .................. Wormaldia relicta (Martynov, 1935)</p><p>14. In lateral view, harpagones narrowing along their length ................................................................... ......................................................................................... Wormaldia hephoa Oláh &amp; Malicky, 2010</p><p>– In lateral view, harpagones equally high along their length .......................................................... 15.</p><p>15. In ventral view, harpagones narrowing along their length ...... Wormaldia muoibon (Malicky, 1995)</p><p>– In ventral view, harpagones parallel-sided along their length ...................... Wormaldia nui sp. nov.</p><p>16. In lateral view, harpagones clearly longer than coxopodites .......................................................... 17</p><p>– In lateral view, harpagones about as long as coxopodites .............................................................. 18</p><p>17. In lateral view, harpagones almost equally high along their length .................................................... ............................................................... Wormaldia montuosa Arefina-Armitage &amp; Armitage, 2015</p><p>– In lateral view, harpagones shallower at mid-length than subapically ............................................... ............................................................................................................ Wormaldia clavella Mey, 1995</p><p>18. Phallus with three spines shorter than high of posterior part of phallus ............................................. ........................................................................................... Wormaldia daona Oláh &amp; Malicky, 2010</p><p>– Phallus with one spine almost as long as phallus ..................... Wormaldia longicornuta Mey, 1996</p></div>	https://treatment.plazi.org/id/5B7A156CFFB8D1257A49FDFD606AFBAD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johanson, Kjell Arne;Pham, Hong-Thai	Johanson, Kjell Arne, Pham, Hong-Thai (2025): Description of three new species of Wormaldia McLachlan, 1865 from Vietnam (Trichoptera: Philopotamidae). European Journal of Taxonomy 1007: 24-34, DOI: 10.5852/ejt.2025.1007.2991, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2991/13451
5B7A156CFFBFD1277A56FB8E61B4FD30.text	5B7A156CFFBFD1277A56FB8E61B4FD30.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Wormaldia piadenensis Johanson & Pham 2025	<div><p>Wormaldia piadenensis sp. nov.</p><p>urn:lsid:zoobank.org:act: A47E096C-BDC2-4038-97C3-82309676D9E3</p><p>Figs 1–2, 5–8</p><p>Diagnosis</p><p>The male genitalia of Wormaldia piadenensis sp. nov. exhibit a notable similarity to those of Wormaldia muoihai, particularly in the presence of downwardly curving harpagones, which are of equal height at mid-length and at the base, and are each shorter than the coxopodites. However, W. piadenensis can be distinguished from W. muoihai by several key characteristics in the male genitalia: segment IX is shorter, segment IX possesses a distinct posteroventral process, the dorsal margin of the coxopodites is more sharply produced dorsally, and the phallus features a single posterior curved sclerite and a long, needleshaped anterior sclerite, in contrast to the two short, straight sclerites present in W. muoihai .</p><p>Etymology</p><p>Wormaldia piadenensis sp. nov. was named after Phia Dén Village, near the type locality of the species.</p><p>Type material</p><p>Holotype VIETNAM • ♂ (in alcohol); Cao Bang Province, N Nui Pia Oac Nature Reserve, about 9.5 km (rd) N of Phia Dén Village, 100 m upstream of bridge; 10 Apr. 2012; Johanson and Pham leg.; light trap at stream with mixed stream bottom; loc#VN028; VNMN.</p><p>Description (holotype)</p><p>Forewing (Fig. 2) 5.5 mm (N = 1), membrane light brown, veins dark brown. Forks 1–5 present; fork 1 originates at distal corner of Dc; basis of M1+2 and Ms hyaline; arc located opposite to base of fork 5. Abdominal segments VII and VIII with long posteroventral lobes (Figs 5, 7). In lateral view (Fig. 5), segment IX anteriorly triangular and posteroventral part produced posteriorly; coxopodites with almost straight ventral margin, dorsal margin slightly produced into triangular; harpagones almost uniformly curving ventrad along their length, about ½× as long as coxopodites, each with rounded apex. In dorsal view (Fig. 6), segment IX with wide U-shaped anterior margin; tergum X triangular, with few setae. In ventral view (Fig. 7), with sternum IX produced into small, narrow triangle; basal half of coxopodites fused, distal half with irregularly shaped mesal margin bearing stout setae at posteromesal corner; harpagones almost straight, with small mesal process at mid-length. Phallus (Fig. 8) with one pair of very short sclerites running along its length, and one sickle-shaped sclerite near apex.</p></div>	https://treatment.plazi.org/id/5B7A156CFFBFD1277A56FB8E61B4FD30	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johanson, Kjell Arne;Pham, Hong-Thai	Johanson, Kjell Arne, Pham, Hong-Thai (2025): Description of three new species of Wormaldia McLachlan, 1865 from Vietnam (Trichoptera: Philopotamidae). European Journal of Taxonomy 1007: 24-34, DOI: 10.5852/ejt.2025.1007.2991, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2991/13451
5B7A156CFFBDD1267A84FD0D6428FE32.text	5B7A156CFFBDD1267A84FD0D6428FE32.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Wormaldia nui Johanson & Pham 2025	<div><p>Wormaldia nui sp. nov.</p><p>urn:lsid:zoobank.org:act: CB2E57C6-4B85-4619-8D95-F40DC1A02ECB</p><p>Figs 1, 3, 9–12</p><p>Diagnosis</p><p>The male genitalia of Wormaldia nui sp. nov. closely resemble those of Wormaldia muoibon, particularly in lateral view, where the harpagones are almost straight and parallel-sided, and of equal length as the coxopodites.Additionally, tergite VIII is produced above segment IX. However, W. nui is distinguishable from W. muoibon based on several male genital features: in W. nui, the harpagones are parallel-sided in ventral view, whereas they slightly taper in W. muoibon; further, the phallus of W. nui has two short, straight sclerites at the posterior end, in contrast to the three longer, straight sclerites found in W. muoibon .</p><p>Etymology</p><p>Wormaldia nui sp. nov. was named after the Nui Pia Oac Nature Reserve, the type locality of the species.</p><p>Type material</p><p>Holotype VIETNAM • ♂ (in alcohol); Cao Bang Province, N Nui Pia Oac Nature Reserve, about 9.5 km (rd) N of Phia Dén Village, 100 m upstream of bridge; 10 Apr. 2012; Johanson and Pham leg.; light trap at stream with mixed stream bottom; loc#VN028; VNMN.</p><p>Paratype VIETNAM • 1 ♂ (in alcohol); Cao Bang Province, Nui Pia Oac Nature Reserve, about 200 m downstream on rd to radio tower; 10 Apr. 2012; Johanson and Pham leg.; light trap at village center; loc#VN030; NHRS .</p><p>Description (holotype)</p><p>Forewing (Fig. 3) 4.5 mm (N = 1), membrane light brown, veins dark brown. Forks 1–5 present; fork 1 originates shortly before distal corner of Dc; basis of M1+2 and Ms hyaline; arc located slightly distally of base of fork 5. Abdominal segments VII and VIII with long posteroventral lobes (Figs 9, 11). In lateral view (Fig. 8), segment IX forming triangular anteriorly, posteroventral corner not produced posteriorly; coxopodites with slightly convex ventral margin, dorsal margin strongly convex; harpagones straight, with almost parallel-sided dorsal and ventral margins, about as long as coxopodites, each with rounded apex. In dorsal view (Fig. 10), segment IX with wide U-shaped anterior margin; tergum X with basal three-quarters with parallel-sided lateral margins, with two pairs of short setae before narrowing apex. In ventral view (Fig. 11), with sternum IX with small central incision at posterior margin; basal twothirds of coxopodites fused, each with distal half of mesal margin straight, bearing few stout setae near posteromesal corner; harpagones almost straight, without mesal process. Phallus (Fig. 12) with thin sclerites in group before mid-length and one pair of short, triangular sclerites situated close to each other near apex.</p></div>	https://treatment.plazi.org/id/5B7A156CFFBDD1267A84FD0D6428FE32	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johanson, Kjell Arne;Pham, Hong-Thai	Johanson, Kjell Arne, Pham, Hong-Thai (2025): Description of three new species of Wormaldia McLachlan, 1865 from Vietnam (Trichoptera: Philopotamidae). European Journal of Taxonomy 1007: 24-34, DOI: 10.5852/ejt.2025.1007.2991, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2991/13451
5B7A156CFFB3D1287A5BFEEA6792FD57.text	5B7A156CFFB3D1287A5BFEEA6792FD57.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Wormaldia caobangensis Johanson & Pham 2025	<div><p>Wormaldia caobangensis sp. nov.</p><p>urn:lsid:zoobank.org:act: 412B7D98-4ED4-4DAB-BA79-3F7F50EAED53</p><p>Figs 1, 4, 13–16</p><p>Diagnosis</p><p>The male genitalia of Wormaldia caobangensis sp. nov. exhibit similarities to those of Wormaldia muoihai, particularly in the presence of the downwardly curving harpagones that are of equal height at mid-length and at the base, and are shorter than the coxopodites. However, in lateral view, W. caobangensis is distinguishable from W. muoihai by the more prominently produced anterior segment IX.</p><p>Etymology</p><p>Wormaldia caobangensis sp. nov. was named after the Vietnamese Cao Bang Province, the type locality of the species.</p><p>Type material</p><p>Holotype VIETNAM • ♂ (in alcohol); Cao Bang Province, Nui Pia Oac Nature Reserve, N Phia Dén Village, upstream road to the radio tower; 8–10 Apr. 2012; Johanson and Pham leg.; Malaise trap across stream with large stones; loc#VN024; VNMN.</p><p>Description (holotype)</p><p>Forewing (Fig. 4) 5.5 mm (N = 1), membrane light brown, veins dark brown. Forks 1–5 present; fork 1 originates shortly before distal corner of Dc; basis of M1+2 and Ms hyaline; arc located slightly distally of base of fork 5. Abdominal segments VII and VIII with long posteroventral lobes (Figs 13, 15). In lateral view (Fig. 13), segment IX anteriorly triangular, posteroventral corner not produced posteriorly; coxopodites with slightly undulating ventral margin, dorsal margin strongly convex; harpagones with weakly convex dorsal margin and weakly concave margins, about 3 / 5 as long as coxopodites, each with rounded apex. In dorsal view (Fig. 14), segment IX with wide U-shaped anterior margin; tergum X hidden below triangular tergite VIII. In ventral view (Fig. 15), with sternum IX with small central incision at posterior margin; basal half of coxopodites fused, each with distal half of mesal margin almost straight, bearing scattered setae; harpagones almost straight, without mesal process. Phallus (Fig. 16) with one long, straight sclerite and one distally hook-shaped, slender sclerite, both originating approximately at mid-length of phallus.</p></div>	https://treatment.plazi.org/id/5B7A156CFFB3D1287A5BFEEA6792FD57	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johanson, Kjell Arne;Pham, Hong-Thai	Johanson, Kjell Arne, Pham, Hong-Thai (2025): Description of three new species of Wormaldia McLachlan, 1865 from Vietnam (Trichoptera: Philopotamidae). European Journal of Taxonomy 1007: 24-34, DOI: 10.5852/ejt.2025.1007.2991, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2991/13451
