identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
556987F9FF9FFFDC5A3F0CB7FE627387.text	556987F9FF9FFFDC5A3F0CB7FE627387.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cerceros Kraatz 1879	<div><p>Subgenus  Cerceros Kraatz, 1879</p><p>The  Macrolycus gracilis group</p><p>Diagnosis</p><p>Male antennomeres III usually triangular, seldom lamellate or cylindrical. Elytra elongate, with lateral margins feebly to moderately, but never strongly diverging posteriorly. Aedeagus with phallus never notched at apex, but attached with a circular lamellar structure (e.g., Figs 2A–C, 3).</p><p>Included species</p><p>M. aurantiacus Kazantsev, 2001,  M. baihualingensis Li, Bocak &amp; Pang, 2015,  M. gracilis Pic, 1923,  M. lizipingensis Li, Bocak &amp; Pang, 2015,  M. mucronatus Li, Bocak &amp; Pang, 2012,  M. multicostatus</p><p>Kazantsev, 2002,  M. muyuensis Li, Bocak &amp; Pang, 2012,  M. phoeniceus Li, Bocak &amp; Pang, 2015,  M. rhodoneurus Li, Bocak &amp; Pang, 2015, and  M. rosaceus Li, Bocak &amp; Pang, 2015 .</p><p>Distribution (Fig. 1)</p><p>China, Laos, and Vietnam.</p></div>	https://treatment.plazi.org/id/556987F9FF9FFFDC5A3F0CB7FE627387	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Du, Ruolan;Yang, Yuxia;Yang, Xingke;Liu, Haoyu	Du, Ruolan, Yang, Yuxia, Yang, Xingke, Liu, Haoyu (2025): Taxonomic study of the Macrolycus gracilis group (Coleoptera: Lycidae), with descriptions of four new species from China. European Journal of Taxonomy 981: 265-279, DOI: 10.5852/ejt.2025.981.2825, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2825/12885
556987F9FF9EFFDB59930FD1FB9D715F.text	556987F9FF9EFFDB59930FD1FB9D715F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrolycus mucronatus Li, Bocak & Pang 2012	<div><p>Macrolycus mucronatus Li, Bocak &amp; Pang, 2012</p><p>Figs 1–2</p><p>Macrolycus mucronatus Li, Bocak &amp; Pang, 2012: 53, figs 27, 37–38, 50.</p><p>Macrolycus mucronatus – Li 2015: 32.</p><p>Material examined</p><p>CHINA • 2 ♂♂; Guangxi, Longsheng, Huaping,  Anjiangping; 2 Jun. 2023; H.Q. Lin and S.L. Yuan leg.; MHBU  •   1 ♂; Hubei, Shennongjia,  Wenshui Forestry; 17 Jul. 2003; F. Lin leg.; IZAS  .</p><p>Descriptive notes</p><p>Male (Fig. 2D)</p><p>AEDEAGUS. Phallus slender, nearly parallel-sided at basal part in dorsal and ventral views (Fig. 2A–B), subapical part strongly and almost symmetrically inflated laterally, about 1.8× as wide as basal part, with ventral, fusiform cavity, apical part abruptly narrowed distad, apex about 0.22 × as wide as subapical part; curved ventrally at basal ¼ portion, and bent dorsally at apical 3 / 8 portion in lateral view (Fig. 2C), subapical part moderately inflated ventrally, apical part nearly straight and feebly inflated ventrally near apex.</p><p>Distribution</p><p>China (Guangxi, Hubei, Hunan).</p><p>Remarks</p><p>The external appearance of this species was described well in the original publication (Li et al. 2012), but the aedeagus was simply described and illustrated only in lateral and ventral views. Here, the material of this species is available for us, which makes it possible to provide a detailed description and illustrations of the aedeagus, to make better comparison with the new species below.</p></div>	https://treatment.plazi.org/id/556987F9FF9EFFDB59930FD1FB9D715F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Du, Ruolan;Yang, Yuxia;Yang, Xingke;Liu, Haoyu	Du, Ruolan, Yang, Yuxia, Yang, Xingke, Liu, Haoyu (2025): Taxonomic study of the Macrolycus gracilis group (Coleoptera: Lycidae), with descriptions of four new species from China. European Journal of Taxonomy 981: 265-279, DOI: 10.5852/ejt.2025.981.2825, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2825/12885
556987F9FF99FFD959B50DA7FE7C7427.text	556987F9FF99FFD959B50DA7FE7C7427.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrolycus nigricollis Y. Yang, Liu & X. Yang 2025	<div><p>Macrolycus nigricollis Y. Yang, Liu &amp; X. Yang sp. nov.</p><p>urn:lsid:zoobank.org:act: 23F1E034-0B3F-4520-A49D-02B8790A8617</p><p>Figs 1, 3A–C, 4A–B</p><p>Diagnosis</p><p>This species can be separated from other species of the  M. gracilis group by a characteristic aedeagus with moderately robust phallus, which is nearly parallel-sided in the basal part in all views (Fig. 3A–C). The species most closely resembles  M. aurantiacus Kazantsev, 2001 in the triangular antennomere III, but can be easily distinguished from the latter by the black pronotum and dark red elytra, while both pronotum and elytra are orange in  M. aurantiacus; phallus almost even in width in apical part in lateral view (Fig. 3C), and abruptly narrowed distad in  M. aurantiacus (Kazantsev 2001: fig. 9).</p><p>Etymology</p><p>The specific epithet is derived from the Latin ‘ niger ’ (‘black’) and ‘ collum ’ (‘neck’), referring to its uniformly black pronotum.</p><p>Type material</p><p>Holotype</p><p>CHINA • ♂; Zhejiang, Fengyangshan,  Ludai; 1123 m a.s.l.; 3 May 2021; Y. Kang and X.X. Wang leg.; HBU(E)410056; MHBU.</p><p>Paratypes CHINA • 1 ♂; Zhejiang, Tianmushan,  Dahenglu; 1200 m a.s.l.; 6 Jun. 1998; M.S. Zhao leg.; IOZ(E)2058406; IZAS  •   1 ♂; Fujian, Chongan, Xingcun,  Road of Sangang to  Jianyang,  Aotou; 740– 1170 m a.s.l.; 20 May 1960; C.L. Ma leg.; IOZ(E)1119994; IZAS  •  1 ♀; Fujian, Chongan, Xingcun,</p><p>Sangang; 740 m a.s.l.; 25 May 1960; Y. R. Zhang leg.; IOZ(E)1120123; IZAS.</p><p>Description</p><p>Male (Fig. 4A)</p><p>MEASUREMENTS. Length 7.8–10.0 mm (8.7 mm in holotype), width at humeri 1.7–2.3 mm (1.9 mm in holotype).</p><p>HABITUS AND COLORATION. Body black brown. Pronotum and scutellum black, elytra dark red. Surface covered with decumbent red pubescence.</p><p>HEAD. Eyes small, interocular distance about 1.7 × as long as eye diameter. Antennae flabellate starting from antennomeres IV, overlapping basal ⅔ length of elytra when inclined. Antennomere III triangular and feebly longer than wide, IV‒XI lamellate, lamellae pointed at apices, lamella of IV as half long as joint itself, lamella of IX longest, 3.3× as long as joint itself (Fig. 4A).</p><p>PRONOTUM. Trapezoidal, 1.2 × as wide as long. Anterior margin widely rounded, lateral margins feebly sinuate and posterior margin bisinuate; anterior angles rounded, posterior angles sharp and feebly projected. Scutellum trapezoidal, emarginate at apex (Fig. 4A).</p><p>ELYTRA. 3.8 × as long as humeral width. Costa I weak but visible in whole length, costa IV as strong as II and both separated terminally, costa III visible only at the basal half part (Fig. 4A).</p><p>AEDEAGUS. Phallus slender, nearly parallel-sided at basal part in dorsal and ventral views (Fig. 3A–B), subapical part moderately and asymmetrically inflated laterally, about 1.45× as wide as basal part, with ventral, fusiform cavity, apical part gradually narrowed distad, apex about 0.38 × as wide as subapical part; curved ventrally at basal 2 /7 portion, and bent dorsally at apical 5 /14 portion in lateral view (Fig. 3C), subapical part moderately inflated ventrally, apical part nearly straight and flat near apex.</p><p>Female (Fig. 4B)</p><p>Similar to males, but larger in body size. Antennae serrate and shorter, reaching elytral midlength when inclined, antennomere III feebly widened apically, about 2.5 × as long as wide. Pronotum narrower, 1.1× as wide as long. Elytral costa II stronger than IV and both fused then split terminally at apical part of elytra.</p><p>Variation within type series</p><p>Pronotum of some specimens has the posterior margin straight.</p><p>Distribution</p><p>China (Zhejiang, Fujian).</p></div>	https://treatment.plazi.org/id/556987F9FF99FFD959B50DA7FE7C7427	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Du, Ruolan;Yang, Yuxia;Yang, Xingke;Liu, Haoyu	Du, Ruolan, Yang, Yuxia, Yang, Xingke, Liu, Haoyu (2025): Taxonomic study of the Macrolycus gracilis group (Coleoptera: Lycidae), with descriptions of four new species from China. European Journal of Taxonomy 981: 265-279, DOI: 10.5852/ejt.2025.981.2825, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2825/12885
556987F9FF9BFFD759B80858FE1F7360.text	556987F9FF9BFFD759B80858FE1F7360.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrolycus graciliramus Y. Yang, Du & Liu 2025	<div><p>Macrolycus graciliramus Y. Yang, Du &amp; Liu sp. nov.</p><p>urn:lsid:zoobank.org:act: 4DB1D779-6B9E-4291-8609-CA259A12D17D</p><p>Figs 1, 3D–F, 4D–E</p><p>Diagnosis</p><p>This species can be easily distinguished from all others of the  M. gracilis group by the male antennae with extremely slender branches on antennomeres III‒X (Fig. 4D). The species more resembles  M. gracilis Pic, 1923 in the presence of lamella on antennomere III, but differs in the following characters: longer antennae, overlapping basal ¾ length of elytra when inclined, lamella of antennomere III shorter, 0.7× as long as joint itself; black pronotum; apical part of phallus gradually narrowed distad in ventral view (Fig. 3E). In comparison, in  M. gracilis, the antennae are shorter, at most reaching basal ⅔ length of elytra, lamella of antennomere III is longer, as long as joint itself; pronotum is orange; apical part of the phallus is parallel-sided at apical 1 /5 portion in ventral view (Li 2015: fig. 7u).</p><p>Etymology</p><p>The specific epithet is derived from the Latin  ‘ gracilis ’ (‘slender’) and ‘ ramus ’ (‘branch’), referring to the slender lamellae on male antennomeres III‒X.</p><p>Type material</p><p>Holotype</p><p>CHINA • ♂; Guangxi, Longsheng, Huaping,  Anjiangping; 4 Jun. 2023; H.Q. Lin and S.L. Yuan leg.; HBU(E)410057; MHBU.</p><p>Paratypes</p><p>CHINA • 2 ♂♂; same data as for holotype; HBU(E)410059 to 410060; MHUB •  2 ♀♀; same data as for holotype; HBU(E)410061 to 410062; MHUB •  1 ♀; same data as for holotype; 2 Jun. 2023; HBU(E)410063; MHBU .</p><p>Description</p><p>Male (Fig. 4D)</p><p>MEASUREMENTS. Length 10.0– 10.6 mm (10.6 mm in holotype), width at humeri 2.1–2.4 mm (2.4 mm in holotype).</p><p>HABITUS AND COLORATION. Body black brown. Pronotum and scutellum black, elytra dark red. Surface covered with decumbent red pubescence.</p><p>HEAD. Eyes small, interocular distance about 1.8 × as long as eye diameter. Antennae flabellate starting from antennomeres III, overlapping basal ¾ length of elytra when inclined. Antennomeres III‒XI lamellate, lamellae pointed at apices, lamella of III 0.7× as long as joint itself, lamella of VIII longest, 3.4× as long as joint itself (Fig. 4D).</p><p>PRONOTUM. Trapezoidal, 1.1× as wide as long. Anterior margin widely rounded and projecting anteriad, lateral margins feebly sinuate and posterior margin straight; anterior angles rounded, posterior angles sharp and moderately projected. Scutellum trapezoidal, nearly straight at apex (Fig. 4D).</p><p>ELYTRA. 3.8 × as long as humeral width. Costa I weak but visible at basal ⅔ length, costa II stronger than IV and both separated terminally at apical part of elytra, costa III visible only at the basal ¼ part (Fig. 4D).</p><p>AEDEAGUS. Phallus slender, nearly parallel-sided at basal part in dorsal and ventral views (Fig. 3D–E), subapical part strongly and almost symmetrically inflated laterally, about twice as wide as basal part, with an oval ventral-cavity, apical part gradually narrowed distad, apex about 0.25 × as wide as subapical part; curved ventrally at basal ⅓ portion, and bent dorsally at apical 4 / 7 portion in lateral view (Fig. 3F), subapical part feebly inflated ventrally, apical part slightly inclined dorsally and flat near apex.</p><p>Female (Fig. 4E)</p><p>Similar to males, but larger in body size. Antennae serrate and shorter, reaching elytral midlength when inclined. Pronotum wider, 1.2× as wide as long, anterior angles obtuse-angled. Elytra wider and 3.6× as long as humeral width, costa II much stronger than IV and both fused terminally at apical part of elytra.</p><p>Distribution</p><p>China (Guangxi).</p></div>	https://treatment.plazi.org/id/556987F9FF9BFFD759B80858FE1F7360	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Du, Ruolan;Yang, Yuxia;Yang, Xingke;Liu, Haoyu	Du, Ruolan, Yang, Yuxia, Yang, Xingke, Liu, Haoyu (2025): Taxonomic study of the Macrolycus gracilis group (Coleoptera: Lycidae), with descriptions of four new species from China. European Journal of Taxonomy 981: 265-279, DOI: 10.5852/ejt.2025.981.2825, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2825/12885
556987F9FF95FFD6598F0F83FEF9746C.text	556987F9FF95FFD6598F0F83FEF9746C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrolycus breviramus Y. Yang, Du & Liu 2025	<div><p>Macrolycus breviramus Y. Yang, Du &amp; Liu sp. nov.</p><p>urn:lsid:zoobank.org:act: 042B1564-6E58-4915-BF4C-A5C933F5040F</p><p>Figs 1, 3G–I, 4C</p><p>Diagnosis</p><p>This species can be easily distinguished from all others of the  M. gracilis group by the cylindrical antennomere III, orange red margins of pronotum and elytra (Fig. 4C), and very robust but uneven phallus (Fig. 3G–I).</p><p>Etymology</p><p>The specific epithet is derived from the Latin ‘ brevis ’ (‘short’) and ‘ ramus ’ (‘branch’), referring to its short antennal lamellae on male antennomeres VI‒X.</p><p>Type material</p><p>Holotype</p><p>CHINA • ♂; Hubei, Shennongjia, Pingqian,  Baiwawu; 1350 m a.s.l.; 2 Jul. 2022; J.B. Tong and X.Y. Ge leg.; HBU(E)410058; MHBU.</p><p>Description</p><p>Male (Fig. 4C)</p><p>MEASUREMENTS. Length 8.7 mm, width at humeri 2.0 mm.</p><p>HABITUS AND COLORATION. Body black brown. Pronotum and elytra dark red, with red margins. Scutellum black. Surface covered with decumbent red pubescence.</p><p>HEAD. Eyes small, interocular distance about 1.6 × as long as eye diameter. Antennae flabellate starting from antennomeres VI, overlapping basal ⅔ length of elytra when inclined.Antennomere III cylindrical, IV long-triangular, about 2× as long as wide, V wide-triangular, feebly longer than wide, VI‒XI lamellate, with lamellae acute at apices, lamella of IX longest, 2× as long as joint itself (Fig. 4C).</p><p>PRONOTUM. Square, 1.1× as wide as long. Anterior margin widely rounded and projecting anteriad, lateral margins feebly sinuate and posterior margin nearly straight; anterior angles obtuse-angled, posterior angles sharp and moderately projected. Scutellum trapezoidal, nearly straight at apex (Fig. 4C).</p><p>ELYTRA. 3.8 × as long as humeral width. Costa I weak but visible at basal ⅔ length; costa III visible only at the base; costa II stronger than IV and both fused terminally at apical part of elytra (Fig. 4C).</p><p>AEDEAGUS. Phallus stout, expanded to one side at basal part in dorsal and ventral views (Fig. 3G–H), subapical part moderately and asymmetrically inflated laterally, about 1.25× as wide as basal part, with an oblong ventral-cavity, apical part gradually narrowed distad, apex about 0.20 × as wide as subapical part; curved ventrally at basal 5 / 14 portion, and bent dorsally at apical ⅓ portion in lateral view (Fig. 3I), subapical part flat, apical part slightly inclined dorsally and feebly inflated ventrally near apex.</p><p>Female</p><p>Unknown.</p><p>Distribution</p><p>China (Hubei).</p></div>	https://treatment.plazi.org/id/556987F9FF95FFD6598F0F83FEF9746C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Du, Ruolan;Yang, Yuxia;Yang, Xingke;Liu, Haoyu	Du, Ruolan, Yang, Yuxia, Yang, Xingke, Liu, Haoyu (2025): Taxonomic study of the Macrolycus gracilis group (Coleoptera: Lycidae), with descriptions of four new species from China. European Journal of Taxonomy 981: 265-279, DOI: 10.5852/ejt.2025.981.2825, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2825/12885
556987F9FF94FFD359B30896FAE37074.text	556987F9FF94FFD359B30896FAE37074.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrolycus acutiapex Y. Yang, Liu & X. Yang 2025	<div><p>Macrolycus acutiapex Y. Yang, Liu &amp; X. Yang sp. nov.</p><p>urn:lsid:zoobank.org:act: 28C36CE6-34D4-4FBE-84A8-0151F8ACF9DC</p><p>Figs 1, 3J–L, 4F</p><p>Diagnosis</p><p>This species can be distinguished from the remaining species of the  M. gracilis group by the antennomere III with slender and short lamella, 0.7 × as long as joint itself. It is more similar to  M. mucronatus Li, Bocak &amp; Pang, 2012, but can be differentiated by the antennomere III with slender lamella, 0.7 × as long as joint itself; apical part of the phallus gradually narrowed distad in ventral view (Fig. 3K), and subapical part inflated ventrally in lateral view (Fig. 3L). Unlikely, in  M. mucronatus, the antennomere III is stouter, with lamella about half length of joint itself; apical part of the phallus is abruptly narrowed distad in ventral view (Li et al., 2012: fig. 38), and subapical part flat in lateral view (Li et al., 2012: fig. 37).</p><p>Etymology</p><p>The specific epithet is derived from the Latin ‘ acutus ’ (‘sharp’) and ‘apex ’ (‘tip’), referring to its lamella of antennomere III with pointed apex.</p><p>Type material</p><p>Holotype</p><p>CHINA • ♂; Hubei, Xingshan,  Longmenhe; 2200 m a.s.l.; 13 Jun. 1995; S.Y. Wang leg.; IOZ(E)1119901; IZAS.</p><p>Description</p><p>Male (Fig. 4F)</p><p>MEASUREMENTS. Length 7.3 mm, width at humeri 1.6 mm.</p><p>HABITUS AND COLORATION. Body black brown. Pronotum, elytra and scutellum dark red. Elytral costae IV orange-red. Surface covered with decumbent red pubescence.</p><p>HEAD. Eyes small, interocular distance about 1.7 × as long as eye diameter. Antennae flabellate, overlapping basal ⅔ length of elytra when inclined. Antennomeres III‒XI lamellate, lamellae pointed at apices, lamella of III extremely slender, about 0.7 × as long as joint itself, lamella of IX longest, 4.2× as long as joint itself (Fig. 4F).</p><p>PRONOTUM. Trapezoidal, 1.3 × as wide as long. Anterior margin widely rounded, lateral margins obviously sinuate and posterior margin nearly straight; anterior angles confluent with anterior margin, posterior angles sharp and moderately projected. Scutellum trapezoidal, nearly straight at apex (Fig. 4F).</p><p>ELYTRA. Slender and subparallel-sided, 3.9 × as long as humeral width. Costae I and III weak but visible at whole length, costa IV weaker than II and both separated at apical part of elytra (Fig. 4F).</p><p>AEDEAGUS. Phallus slender, with basal part narrowed to middle in dorsal and ventral views (Fig. 3J–K), subapical part strongly and almost symmetrically inflated laterally, about 1.67× as wide as basal part, with an oval ventral-cavity, apical part gradually narrowed distad, apex about 0.25× as wide as subapical part; curved ventrally at basal 5 /16 portion, and bent dorsally at apical 7 /16 portion in lateral view (Fig. 3L), subapical part feebly inflated ventrally, apical part nearly straight and flat ventrally.</p><p>Female</p><p>Unknown.</p><p>Distribution</p><p>China (Hubei).</p><p>Key to the species of the  Macrolycus gracilis group</p><p>1. Antennomere III lamellate in male, lamella at least 0.7 × as long as joint itself (e.g., Fig. 4D, F; Li 2015: fig. 3e) ..................................................................................................................................... 2</p><p>– Antennomere III triangular or cylindrical in male, without any lamella (e.g., Fig. 4A, C; Kazantsev 2001: fig. 8; 2002: fig. 9; Li et al. 2012: figs 50–51; 2015: figs 2–6) ............................................... 4</p><p>2. Pronotum black (Fig. 4D); phallus with apical part slightly bent dorsally in lateral view (Fig. 3F) .. .....................................................................................................................  M. graciliramus sp. nov.</p><p>– Pronotum dark red or orange (e.g., Fig. 4F; Li 2015: fig. 3e); phallus with apical part straight in lateral view (e.g., Fig. 4L; Li 2015: fig. 10e) .................................................................................... 3</p><p>3. Lamella of antennomere III as long as joint itself (Li 2015: fig. 7u); apical part of phallus parallel-sided in ventral view (Li 2015: fig. 10f) ............................................................  M. gracilis Pic, 1923</p><p>– Lamella of antennomere III 0.7× as long as joint itself (Fig. 4F); apical part of phallus narrowed distad in ventral view (Fig. 4K) ........................................................................  M. acutiapex sp. nov.</p><p>4. Antennomere III cylindrical in male (Fig. 4C); basal part of phallus obviously asymmetrical, expanded to one side in ventral view (Fig. 3H) .............................................  M. breviramus sp. nov.</p><p>– Antennomere III triangular in male (e.g., Fig. 4A; Kazantsev 2001: fig. 8; 2002: fig. 9; Li et al. 2012: figs 50–51; 2015: figs 2–6); basal part of phallus nearly symmetrical (e.g., Fig. 3B; Kazantsev 2001: fig. 9; 2002: fig. 10; Li et al. 2012: figs 38, 40; 2015: figs 15, 17, 19, 21, 23) ................................. 5</p><p>5. Apical part of phallus moderately inflated distad in ventral view (Li et al. 2015: fig. 15) ................ ......................................................................................  M. baihualingensis Li, Bocak &amp; Pang, 2015</p><p>– Apical part of phallus narrowed distad in ventral view (e.g., Fig. 3B; Kazantsev 2001: fig. 9; 2002: fig. 10; Li et al. 2012: figs 38, 40; 2015: figs 17, 19, 21, 23) ........................................................... 6</p><p>6. Pronotum black (Fig. 4A); subapical part of phallus moderately inflated ventrally in lateral view (Fig. 3C) ...........................................................................................................  M. nigricollis sp. nov.</p><p>– Pronotum dark red or orange; subapical part of phallus flat ventrally in lateral view (e.g., Kazantsev 2001: fig. 10; 2002: fig. 11; Li et al. 2012: figs 37, 39; 2015: figs 16, 18, 20, 22) ........................... 7</p><p>7. Phallus with apical part curved dorsally in lateral view (e.g., Li et al. 2015: figs 20, 22) ............... 8</p><p>– Phallus with apical part straight in lateral view (e.g., Kazantsev 2001: fig. 10; 2002: fig. 11; Li et al. 2012: figs 37, 39; 2015: figs 16, 18) ................................................................................................. 9</p><p>8. Elytral costa II orange-red; phallus with basal part narrowed to middle in ventral view (Li et al. 2015: fig. 21), and apical part moderately curved dorsally in lateral view (Li et al. 2015: fig. 20) ...............................................................................  M. rhodoneurus Li, Bocak &amp; Pang, 2015</p><p>– Elytral costa II dark red; phallus with basal part parallel-sided in ventral view (Li et al. 2015: fig. 23), and apical part strongly curved dorsally in lateral view (Li et al. 2015: fig. 22) ............................... ..................................................................................................  M. rosaceus Li, Bocak &amp; Pang, 2015</p><p>9. Phallus with apical part gradually narrowed distad (e.g., Kazantsev 2001: fig. 9; 2002: fig. 10; Li et al. 2015: fig. 19) .......................................................................................................................... 10</p><p>– Phallus with apical part abruptly narrowed distad (e.g., Li et al. 2012: figs 38, 40; Li et al. 2015: fig. 17) ............................................................................................................................................. 12</p><p>10. Pronotum and elytra orange; apex of phallus about ¼ as wide as subapical part (Kazantsev 2001: fig. 9) ...............................................................................................  M. aurantiacus Kazantsev, 2001</p><p>– Pronotum and elytra dark red (e.g., Li et al. 2015: fig. 4); apex of phallus at most 1 /5 as wide as subapical part (e.g., Kazantsev 2002: fig. 10; Li et al. 2015: fig. 19) ..............................................11</p><p>11. Pronotum and elytra orange red (Li et al. 2015: fig. 4); basal part of phallus uneven in width in ventral view (Li et al. 2015: fig. 19); apical part moderately inflated ventrally near apex in lateral view (Li et al. 2015: fig. 18) ...............................................  M. phoeniceus Li, Bocak &amp; Pang, 2015</p><p>– Pronotum and elytra dark red; basal part of phallus parallel-sided in ventral view (Kazantsev 2002: fig. 10), apical part flat ventrally in lateral view (Kazantsev 2002: fig. 11) ....................................... .......................................................................................................  M. multicostatus Kazantsev, 2002</p><p>12. Phallus with apical part abruptly narrowed distad in lateral view (Li et al. 2015: fig. 16) ................ ............................................................................................  M. lizipingensis Li, Bocak &amp; Pang, 2015</p><p>– Phallus with apical part nearly parallel-sided in lateral view (e.g., Li et al. 2012: figs 37, 39) ..... 13</p><p>13. Antennae relatively long, overlapping basal 4 /5 length of elytra when inclined (Li et al. 2012: fig. 50); phallus bent dorsally at basal ⅓ portion in lateral view (Li et al. 2012: fig. 37) ................................ ............................................................................................  M. mucronatus Li, Bocak &amp; Pang, 2012</p><p>– Antennae relatively short, overlapping basal ½ length of elytra when inclined (Li et al. 2012: fig. 51); phallus bent dorsally at basal 1 /6 portion in lateral view (Li et al. 2012: fig. 39) ............................... ..............................................................................................  M. muyuensis Li, Bocak &amp; Pang, 2012</p></div>	https://treatment.plazi.org/id/556987F9FF94FFD359B30896FAE37074	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Du, Ruolan;Yang, Yuxia;Yang, Xingke;Liu, Haoyu	Du, Ruolan, Yang, Yuxia, Yang, Xingke, Liu, Haoyu (2025): Taxonomic study of the Macrolycus gracilis group (Coleoptera: Lycidae), with descriptions of four new species from China. European Journal of Taxonomy 981: 265-279, DOI: 10.5852/ejt.2025.981.2825, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2825/12885
