identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
4CF7A1ED2C865C8787729A5891CF21AE.text	4CF7A1ED2C865C8787729A5891CF21AE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dactylolabis (Coenolabis) posthabita (Bergroth 1888)	<div><p>Dactylolabis (Coenolabis) posthabita (Bergroth, 1888)</p><p>Comments.</p><p>Dactylolabis (Coenolabis) posthabita is a very rare and poorly known species, originally described by Bergroth (1888) from Slovenia (Ljubljana) as Limnophila posthabita . Savchenko (1982) examined the type series (three males), redescribed the male genitalia, and transferred the species to Dactylolabis, tentatively placing it in the subgenus Coenolabis . However, this placement remained uncertain because the subgenus is primarily defined by female characters, while only males were known at that time.</p><p>Erhan-Dincă (1984) later reported the species from Romania (Eastern Carpathians), providing records of 16 males and 10 females and schematically illustrating the male terminalia. She referred to the species as Coenolabis without question and without further discussion. Savchenko (1986) later noted that he had received specimens from Erhan-Dincă, including females, and briefly mentioned that the ovipositor of the Romanian D. (C.) posthabita females is similar to that of D. (C.) aberrans, characterised by short hypogynial valves, although this was not illustrated. The species was later reported from Hungary by Kolcsár et al. (2021) based on five males. In addition, the first author located further material in the HNHM, including female specimens from Hungary. Erhan-Dincă (1984) reported 14 males and 10 females deposited in the Grigore Antipa National Museum of Natural History (Bucharest, Romania). However, the Romanian material now appears to be either lost or remains uncatalogued in the Diptera collection of the Grigore Antipa Museum (Melania Stan pers. comm.).</p><p>Comparison of three males and one female from Serbia with the male genitalia illustrations provided by Savchenko (1982, repeated in 1986), as well as those of Erhan-Dincă (1984), and with photographs of Hungarian specimens published by Kolcsár et al. (2021), together with additional Hungarian material, revealed several minor differences. The most notable difference is observed in the Hungarian and Serbian females, which possesses well-developed, elongate hypogynial valves reaching approx. half-length of the cerci, in contrast to the condition mentioned by Savchenko (1986) for the Romanian material. Unfortunately, due to the unavailability of Romanian specimens, direct comparison with the original material was not possible and the illustration provided by Erhan-Dincă (1984) is schematic and cannot be used for detailed comparison in case of males.</p><p>We compared the type material deposited in the MZH with specimens from Hungary and Serbia and found clear but relatively minor differences among the three populations, most notably in the Hungarian specimens (see discussion below). However, the examined material shows no distinct differences in habitus, coloration, wing venation, or antennal segmentation, a degree of morphological uniformity also observed in other species of Coenolabis (Savchenko 1969) . Therefore, we treat the Hungarian population as a new subspecies rather than a distinct species, pending future genetic analyses that may clarify its taxonomic status and degree of differentiation.</p></div>	https://treatment.plazi.org/id/4CF7A1ED2C865C8787729A5891CF21AE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Kolcsár, Levente-Péter;d’Oliveira, Micha Camiel;Graf, Wolfram;Kovács, Tibor;Murányi, Dávid;Ivković, Marija	Kolcsár, Levente-Péter, d’Oliveira, Micha Camiel, Graf, Wolfram, Kovács, Tibor, Murányi, Dávid, Ivković, Marija (2026): A new subspecies of Dactylolabis (Coenolabis) posthabita (Bergroth, 1888) from Hungary and new faunistic records of crane flies (Diptera, Tipuloidea) from the Balkans. ZooKeys 1291: 303-324, DOI: 10.3897/zookeys.1291.199961
059DDC9068905E6AB9F0A4479C30C5C2.text	059DDC9068905E6AB9F0A4479C30C5C2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dactylolabis (Coenolabis) posthabita subsp. posthabita (Bergroth 1888)	<div><p>Dactylolabis (Coenolabis) posthabita posthabita (Bergroth, 1888)</p><p>Figs 1 A – E, 2 A – G, 3 A – F, 4 A – F</p><p>Type material examined.</p><p>Slovenia • Holotype ♂; Laibach (Ljubljana); 1870; leg. Palmén, Johan Axel; MZH • Paratypes 2 ♂♂; same data as for holotype; MZH; http://id.luomus.fi/GV.28617 .</p><p>Additional material.</p><p>Serbia • 3 ♂♂, 1 ♀; Raška district, Kraljevo city, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=20.5447&amp;materialsCitation.latitude=43.762" title="Search Plazi for locations around (long 20.5447/lat 43.762)">Samaila</a>, open oak forest and cemetery in a church garden; 43.7620°N, 20.5447°E; alt. 235 m a. s. l.; 7 May 2023; leg. LPK, TK, DM; CKLP .</p><p>Diagnosis.</p><p>Fused median part of interbase situated at two-thirds length of aedeagal complex (at half-length in p. tothi Kolcsár, ssp. nov.). Ventral arm of interbase drop-shaped, with a triangular apex in caudal view (rectangular, with a narrow, pointed apex in p. tothi Kolcsár, ssp. nov.); narrow, band-like, with an evenly curved posterior margin in lateral view (as wide as tip of aedeagus and bent almost perpendicularly at mid-length in p. tothi Kolcsár, ssp. nov.).</p><p>Redescription.</p><p>A detailed general description of the species was provided by Savchenko (1982, 1986) and is not repeated here; only the characters of the male and female terminalia are discussed below. The general appearance of the female is similar to that of the male.</p><p>Male terminalia. Epandrium fused with hypandrium, approx. as wide as long; posterior margin slightly convex, with small lateral lobe (Fig. 2 A, B, H). Posterior margin of hypandrium variable in shape: in type material rounded and bearing several setae (Fig. 2 E, G), whereas Serbian specimens possess a distinct, finger-like lobe with long setae (Figs 2 C, 3 C). In well-dissected specimens, the proctiger is fully inflated and roughly quadrate, with lateral lobes triangular, with obtuse apices directed approx. 45 ° inward (Fig. 2 A, F). If median part of proctiger is slightly compressed, the lateral lobes may appear more perpendicularly directed inward (as in Fig. 2 B). Gonocoxite (Fig. 2 A – G) 1.7–1.9 × longer than wide, slightly longer than epandrium. Gonostyli broadly fused (Fig. 2); lobe of gonostylus (outer gonostylus) balloon-shaped, clasper of gonostylus (inner gonostylus) flamingo-beak-shaped, arising near apex of fused gonostyli. Interbase (Fig. 3 B, D) with median portion fused into a narrow band, situated at approx. 2 / 3 of length of aedeagal complex; lateral arm widened near gonocoxal apodeme, sinuate in lateral view (Fig. 3 E). Ventral arm partially covering lateral arm in caudal view (Fig. 3 F), short, drop-shaped, with ventral part directed inward at nearly a right angle; dorsal apex triangular. Ventral arm in lateral view (Fig. 3 E) narrow, band-like, with posterior margin evenly curved. Paramere (Fig. 3 B, D, E) slender, anterior end weakly curved outward. Aedeagus robust, dorsoventrally flattened, with well-visible median tube (Fig. 3 B, D), slightly curved dorsally (Fig. 3 E). Sperm pump drop-shaped, approx. half the length of aedeagus (Fig. 3 B, D) and wider than aedeagus in lateral view (Fig. 3 E). Ejaculatory apodeme (Fig. 3 B, D, E) with arms widened outward and directed ventrolaterally.</p><p>Female terminalia. Tergite 10 roughly rectangular in dorsal view (Fig. 4 A). Cercus (Fig. 4 B) longer than tergite 10, basally widened, distally narrowing, curving upward in lateral view; apex blunt, without teeth or saw-tooth-like structures (Fig. 4 B). Sternite 8 (Fig. 4 C) slightly longer than wide. Hypogynial valves well developed, approx. as long as sternite 8; apex reaching approx. to mid-length of cercus (Fig. 4 B). A small additional lobe near base of lateral margin of hypogynial valve, triangular in ventral view (Fig. 4 C). Sternite 9 with anterior margin deeply notched medially; posterior margin with a central anteriorly directed lobe, posterolateral parts evenly curved, directed posteriorly (Fig. 4 D). Genital fork widening anteriorly, sinuate in lateral view, as long as sternite 8 (Fig. 4 C). Genital opening weakly sclerotised; common spermathecal duct as long as genital fork, bifurcating into two branches; no sclerotised spermathecae visible. Lateral arms of genital frame with central part distinctly produced, sombrero-shaped in ventral view (Fig. 4 D), roughly circular in caudal view (Fig. 4 F).</p><p>Comments.</p><p>Previously reported from Slovenia, the Hungarian record reported by Kolcsár et al. (2021) is transferred to D. posthabita tothi Kolcsár, ssp. nov., whereas the Romanian records remain tentative (see discussion below). First record from Serbia.</p></div>	https://treatment.plazi.org/id/059DDC9068905E6AB9F0A4479C30C5C2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Kolcsár, Levente-Péter;d’Oliveira, Micha Camiel;Graf, Wolfram;Kovács, Tibor;Murányi, Dávid;Ivković, Marija	Kolcsár, Levente-Péter, d’Oliveira, Micha Camiel, Graf, Wolfram, Kovács, Tibor, Murányi, Dávid, Ivković, Marija (2026): A new subspecies of Dactylolabis (Coenolabis) posthabita (Bergroth, 1888) from Hungary and new faunistic records of crane flies (Diptera, Tipuloidea) from the Balkans. ZooKeys 1291: 303-324, DOI: 10.3897/zookeys.1291.199961
48D35C0603CF5409B374F7C5D733FB4B.text	48D35C0603CF5409B374F7C5D733FB4B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dactylolabis (Coenolabis) posthabita subsp. tothi Kolcsar 2026	<div><p>Dactylolabis (Coenolabis) posthabita tothi Kolcsár ssp. nov.</p><p>Figs 1 F, 1 G, 2 H, 2 I, 3 G – K</p><p>Dactylolabis (Coenolabis) posthabita (Bergroth, 1888) in Kolcsár et al. 2021.</p><p>Type material.</p><p>Holotype: Hungary • ♂ (pinned); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.64&amp;materialsCitation.latitude=45.93" title="Search Plazi for locations around (long 17.64/lat 45.93)">Potony</a>; 45.93°N, 17.64°E (approx.; not stated on the label); 03 May 1977; leg. S. Tóth; HOLOTYPE Dactylolabis (Coenolabis) posthabita tothi Kolcsár, ssp. nov. [red label]; HNHM .</p><p>Paratypes: Hungary • 6 ♂♂, 1 ♀ (pinned); same locality and date as holotype; HNHM • 3 ♂♂ (pinned); Darány, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.563&amp;materialsCitation.latitude=45.98" title="Search Plazi for locations around (long 17.563/lat 45.98)">Barcsi-Ősborókás</a>; 45.98°N, 17.563°E (approx.; not stated on the label); 03 May 1977; leg. S. Tóth; HNHM • 2 ♀♀ (pinned); Pécs, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.33&amp;materialsCitation.latitude=46.13" title="Search Plazi for locations around (long 18.33/lat 46.13)">Bányatelep</a>; 46.13°N, 18.33°E (approx.; not stated on the label); 05–07 June 1957; leg. F. Mihály; det. as Dactylolabis tergestina Egger, 1863 by Mannheims 1965; HNHM • 1 ♀ (pinned); Pécs, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.33&amp;materialsCitation.latitude=46.13" title="Search Plazi for locations around (long 18.33/lat 46.13)">Bányatelep</a>, gesztenyés felett; 46.13°N, 18.33°E (approx.; not stated on the label); 05–07 June 1957; leg. F. Mihály; det. as Dactylolabis tergestina Egger, 1863 by Mannheims 1965; HNHM • 1 ♂ (pinned); Pécs, Mecsek-hg., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.14&amp;materialsCitation.latitude=46.09" title="Search Plazi for locations around (long 18.14/lat 46.09)">Jakob hg., tölgyes</a>; 46.09°N, 18.14°E (approx.; not stated on the label); 05–07 June 1957; leg. F. Mihály; det. as Dactylolabis tergestina Egger, 1863 by Mannheims 1965; HNHM .</p><p>Diagnosis.</p><p>Fused median part of interbase situated at approx. half-length of aedeagal complex (at two-thirds length in p. posthabita). Ventral arm of interbase rectangular, with a narrow, pointed apex in caudal view (drop-shaped, with a triangular apex in p. posthabita); as wide as tip of aedeagus and bent almost perpendicularly at mid-length in lateral view (narrow, band-like, with an evenly curved posterior margin in p. posthabita).</p><p>Description.</p><p>General appearance and size as nominate subspecies in both sexes (Fig. 1).</p><p>Male terminalia. In general (Fig. 2 H, I, 3 G, H), similar to the nominate subspecies, differing as follows: interbase (Fig. 3 I) with fused median part situated at approx. half-length of aedeagal complex; in caudal view (Fig. 3 K) ventral arm of interbase longer than in nominate subspecies, nearly rectangular, directed more ventrally at approx. 45 ° with dorsal apex narrow, pointed; in lateral view (Fig. 3 J), as wide as apex of aedeagus, posterior margin bent almost perpendicularly at mid-length. Sperm pump as wide as aedeagus in lateral view (Fig. 3 J).</p><p>Female. General appearance as male.</p><p>Female terminalia. In general, similar to the nominate subspecies, differing as follows: lateral arms of genital frame elongate, approx. twice as long as wide in caudal view (Fig. 4 G).</p><p>Etymology.</p><p>The subspecies name is dedicated to the memory of the recently deceased Sándor Tóth (1930–2024), a Hungarian entomologist, who collected the majority of the types. The name is a noun in genitive case and masculine form.</p><p>Discussion.</p><p>The male genitalia of D. posthabita are very similar to those of D. imeretica, suggesting a closer morphological affinity between these taxa than between D. imeretica and D. aberrans . However, according to Savchenko (1969), females of D. imeretica cannot be distinguished from those of D. aberrans and therefore possess short hypogynial valves. In contrast, females of both subspecies of D. posthabita have distinctly elongate hypogynial valves. The two subspecies mainly differ in the shape and position of the ventral arm of the interbase. While the interbase appears identical in the paratype of p. posthabita from Slovenia and the specimen from Serbia, the posterior margin of the hypandrium differs; in the type material it is rounded in dorsal view (Fig. 2 E, G), whereas in the Serbian specimens it is produced into a finger-like lobe, clearly visible in lateral view (Fig. 3 C), which is also characteristic of D. imeretica . The Hungarian p. tothi Kolcsár, ssp. nov., shows an intermediate condition, with the posterior margin slightly produced (Fig. 3 H). Other observed differences visible in the figures, such as the shape of the proctiger, appear to result largely from deformation or slight differences in orientation. The wing venation of the examined specimens of both subspecies shows variation in the position of m – cu relative to the fork of M (Fig. 1 B, D, G). However, this character is highly variable among specimens and may even vary between the left and right wings of the same individual, suggesting that it is not associated with subspecific or population-level differences.</p><p>The Hungarian specimens were collected in Potony, most likely along the Drava River or in a relatively natural forest patch in 1977 (Sándor Tóth pers. comm. 2018), with additional specimens collected a few kilometres further in an old natural Juniperus - dominated, open, sandy steppe nature reserve. Further specimens were collected in the Mecsek Mountains in oak and sweet chestnut forests. The Serbian specimens were collected near a small graveyard with large oak trees in the surrounding area; one male was observed resting on mosses at the base of a concrete fence. Both the Hungarian p. tothi Kolcsár, ssp. nov., and Serbian p. posthabita specimens were collected from early May to early June. The exact collection date of the Slovenian type material of p. posthabita is unknown; only the year is available.</p><p>Erhan-Dincă (1984) reported that the Romanian specimens were collected in a wet, well-forested montane habitat between mid-September and late October. If the hypogynial valves of these specimens are indeed short, as stated by Savchenko (1986), then, considering both their occurrence at higher elevations and their markedly different flight period (autumn rather than late spring to early summer, as observed in other populations), the Romanian population could represent a distinct species and warrants further investigation.</p></div>	https://treatment.plazi.org/id/48D35C0603CF5409B374F7C5D733FB4B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Kolcsár, Levente-Péter;d’Oliveira, Micha Camiel;Graf, Wolfram;Kovács, Tibor;Murányi, Dávid;Ivković, Marija	Kolcsár, Levente-Péter, d’Oliveira, Micha Camiel, Graf, Wolfram, Kovács, Tibor, Murányi, Dávid, Ivković, Marija (2026): A new subspecies of Dactylolabis (Coenolabis) posthabita (Bergroth, 1888) from Hungary and new faunistic records of crane flies (Diptera, Tipuloidea) from the Balkans. ZooKeys 1291: 303-324, DOI: 10.3897/zookeys.1291.199961
B58F754A48B75EADB73001E714E20D05.text	B58F754A48B75EADB73001E714E20D05.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dactylolabis (Coenolabis) Savchenko 1969	<div><p>Subgenus Dactylolabis (Coenolabis) Savchenko, 1969</p><p>Type species.</p><p>Dactylolabis (Coenolabis) aberrans Savchenko, 1963 by original designation in Savchenko (1969).</p><p>Remarks.</p><p>Prior to the present study, Dactylolabis (Coenolabis) comprised only four recognised species. Dactylolabis arsianensis Savchenko, 1969 and D. imeretica Savchenko, 1969 were originally described as subspecies of D. aberrans but are now treated as distinct species, due to their partial sympatry (Oosterbroek 2026). The subgenus is restricted to the Western Palearctic.</p></div>	https://treatment.plazi.org/id/B58F754A48B75EADB73001E714E20D05	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Kolcsár, Levente-Péter;d’Oliveira, Micha Camiel;Graf, Wolfram;Kovács, Tibor;Murányi, Dávid;Ivković, Marija	Kolcsár, Levente-Péter, d’Oliveira, Micha Camiel, Graf, Wolfram, Kovács, Tibor, Murányi, Dávid, Ivković, Marija (2026): A new subspecies of Dactylolabis (Coenolabis) posthabita (Bergroth, 1888) from Hungary and new faunistic records of crane flies (Diptera, Tipuloidea) from the Balkans. ZooKeys 1291: 303-324, DOI: 10.3897/zookeys.1291.199961
