taxonID	type	description	language	source
572A3C3B1276FFA351C7FF6DFE9BFF2B.taxon	description	(Figs 1, 5)	en	Min, Qiuyue, Li, Houhun, Wang, Shuxia (2026): Four species of the sakaiella group of Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from China. Zootaxa 5752 (1): 134-140, DOI: 10.11646/zootaxa.5752.1.6, URL: https://doi.org/10.11646/zootaxa.5752.1.6
572A3C3B1276FFA351C7FF6DFE9BFF2B.taxon	materials_examined	Type material. Holotype ♂, CHINA, Guangxi: Lanma, Jiuwanshan Nature Reserve, Rongshui County (25.25 ºN, 108.81 ºE), 357 m, 26. VII. 2022, leg. H Sun, ZX Xu and D Zhang, slide No. MQY 24243.	en	Min, Qiuyue, Li, Houhun, Wang, Shuxia (2026): Four species of the sakaiella group of Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from China. Zootaxa 5752 (1): 134-140, DOI: 10.11646/zootaxa.5752.1.6, URL: https://doi.org/10.11646/zootaxa.5752.1.6
572A3C3B1276FFA351C7FF6DFE9BFF2B.taxon	diagnosis	Diagnosis. The new species can be distinguished from its allies by the forewing with a white costal spot on the outer margin of the median fascia anteriorly; in the male genitalia by the broadly U-shaped uncus, the distal 1 / 3 of the valva curved upward, and the flagellate juxta exceeding the apex of the uncus and with dense spines in distal 1 / 3.	en	Min, Qiuyue, Li, Houhun, Wang, Shuxia (2026): Four species of the sakaiella group of Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from China. Zootaxa 5752 (1): 134-140, DOI: 10.11646/zootaxa.5752.1.6, URL: https://doi.org/10.11646/zootaxa.5752.1.6
572A3C3B1276FFA351C7FF6DFE9BFF2B.taxon	description	Description. Adult (Fig. 1) with wingspan 10.0 mm. Head. Vertex white, frons and occiput deep greenish brown. Labial palpus with second segment pale orange yellow on inner surface, yellowish brown on outer surface; third segment black, with yellowish brown scales. Antenna with scape white; flagellum white in basal 1 / 3, black alternating with white in distal 2 / 3 dorsally, dark brown ventrally. Thorax. Mesonotum and tegula dark brown. Forewing orange yellow, with dark brown scales from base to median fascia along costal margin; dark reddish-brown stripe from base of fold obliquely outward to dorsum, its outer margin edged with black scales and a white line adjacent to black scales; median fascia dark reddish brown, inner margin edged with black scales, a white streak adjacent to black scales running from dorsum to below anterior margin of cell, outer margin concave inward in anterior 1 / 2, edged with black scales; costal spot white, subovate, placed beyond median fascia, extending to posterior angle of cell; distal patch dark brown, diffused along termen to median fascia; fringe orange yellow except dark brown on distal part of costal margin and on extension of median fascia. Hindwing and fringe grey. Abdomen. Male genitalia (Fig. 5) with uncus broadly U-shaped; lateral arms uniform from base to basal 2 / 3, inflated from distal 1 / 3 to obtuse apex. Gnathos semicircular, 1 / 4 length of uncus. Tegumen furcate from posterior end, laterally subparallel to anterior end. Valva narrow at base, widened to middle, abruptly narrowed from middle to beyond distal 1 / 4, then subparallel to rounded apex, distal 1 / 3 curved upward, densely setose; costa roundly produced medially; sacculus narrow, subparallel basally, setose and fused with valva distally. Saccus elongate triangular, broad at base, narrowed to apex, approx. 2.5 times length of uncus. Juxta flagellate, slightly narrowed to acute apex, distal 1 / 3 with dense fine spines, wheatear-like, far exceeding apex of uncus; basal lobe clubbed, slightly shorter than saccus. Phallus tubular, approx. twice length of saccus; cornutus absent. Female unknown.	en	Min, Qiuyue, Li, Houhun, Wang, Shuxia (2026): Four species of the sakaiella group of Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from China. Zootaxa 5752 (1): 134-140, DOI: 10.11646/zootaxa.5752.1.6, URL: https://doi.org/10.11646/zootaxa.5752.1.6
572A3C3B1276FFA351C7FF6DFE9BFF2B.taxon	distribution	Distribution. China (Guangxi).	en	Min, Qiuyue, Li, Houhun, Wang, Shuxia (2026): Four species of the sakaiella group of Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from China. Zootaxa 5752 (1): 134-140, DOI: 10.11646/zootaxa.5752.1.6, URL: https://doi.org/10.11646/zootaxa.5752.1.6
572A3C3B1276FFA351C7FF6DFE9BFF2B.taxon	etymology	Etymology. The specific epithet is derived from the Latin bi- and folium, referring to the two lateral lobes of the uncus.	en	Min, Qiuyue, Li, Houhun, Wang, Shuxia (2026): Four species of the sakaiella group of Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from China. Zootaxa 5752 (1): 134-140, DOI: 10.11646/zootaxa.5752.1.6, URL: https://doi.org/10.11646/zootaxa.5752.1.6
572A3C3B1274FFA351C7FEF8FB77F933.taxon	description	(Figs 2, 6, 8)	en	Min, Qiuyue, Li, Houhun, Wang, Shuxia (2026): Four species of the sakaiella group of Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from China. Zootaxa 5752 (1): 134-140, DOI: 10.11646/zootaxa.5752.1.6, URL: https://doi.org/10.11646/zootaxa.5752.1.6
572A3C3B1274FFA351C7FEF8FB77F933.taxon	materials_examined	Type material. Holotype ♂, CHINA, Sichuan: Songpinggou, Maoxian County (32.09 ºN, 103.64 ºE), 2385 m, 4. VII. 2021, leg. S Yu, XJ Zhu and D Zhang, slide No. MQY 25159. Paratypes. 16 ♂ 1 ♀, 3 – 6. VII. 2021, same data as holotype, slide Nos. MQY 25155 ♂, MQY 25156 ♀, MQY 25636 ♂, MQY 25637 ♂.	en	Min, Qiuyue, Li, Houhun, Wang, Shuxia (2026): Four species of the sakaiella group of Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from China. Zootaxa 5752 (1): 134-140, DOI: 10.11646/zootaxa.5752.1.6, URL: https://doi.org/10.11646/zootaxa.5752.1.6
572A3C3B1274FFA351C7FEF8FB77F933.taxon	diagnosis	Diagnosis. The new species is similar to P. subquadrata Wang, 2019 in male genitalia. It can be distinguished by the triangular uncus about 2 / 3 the length of the gnathos, the gnathos produced to a small process apically, and the right valva with apex concave above the ventral margin. In P. subquadrata, the semicircular uncus is about 1 / 3 the length of the gnathos, the gnathos lacks an apical process, and the apex of the right valva is straight.	en	Min, Qiuyue, Li, Houhun, Wang, Shuxia (2026): Four species of the sakaiella group of Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from China. Zootaxa 5752 (1): 134-140, DOI: 10.11646/zootaxa.5752.1.6, URL: https://doi.org/10.11646/zootaxa.5752.1.6
572A3C3B1274FFA351C7FEF8FB77F933.taxon	description	Description. Adult (Fig. 2) with wingspan 12.0 – 14.0 mm. Head. Vertex white, frons greyish brown, occiput dark brown. Labial palpus with second segments yellow on inner surface, orange yellow on outer surface; third segment black except white at apex. Antenna with scape white dorsally, pale yellowish brown ventrally; flagellum black alternating with white dorsally, black ventrally. Thorax. Mesonotum and tegula blackish brown. Forewing orange yellow; basal fascia dark brown, outer margin edged with black scales, a white line adjacent to black scales; median fascia dark grey, edged with black scales, narrowed medially, widened to costal margin between basal 2 / 5 and before distal 1 / 3, widened to ventral margin between end of fold and tornus; distal patch greyish black, edged with black scales on inner margin, narrowed along termen to tornus; fringe orange yellow except greyish black on extension of distal patch and around tornus. Hindwing and fringe greyish brown. Male genitalia. (Fig. 6). Uncus weakly sclerotized, broadly triangular, gradually narrowed from base to before papillary apex. Gnathos sublingulate, produced to a papillary process apically. Tegumen furcate from posterior 1 / 3, narrowed to anterior end laterally. Valva broad subrectangular, asymmetrical: left valva subparallel to sclerotized, flat apex, costa broadly banded, with dense spines in distal 1 / 3, sacculus wide and parallel to middle, notched obliquely outward beyond middle; right valva parallel to apex, apex concave above ventral margin, forming two lobes: dorsal lobe large, rounded, with dense setae and spines on rounded apical margin, ventral lobe much smaller, subquadrate, costa banded, roundly produced distally, sacculus wide at base, narrowed to basal 2 / 5, concave in a small notch on dorsal margin near middle, uniformly narrow distally. Saccus subtriangular, distinctly narrowed to apex, longer than uncus. Juxta weakly sclerotized in basal 2 / 5, strongly sclerotized in distal 3 / 5, widened from base to basal 2 / 5, then narrowed to rounded apex; basal lobe slender, clavate, reaching 3 / 5 length of saccus. Phallus slender in basal 1 / 4, thereafter uniform to apex, curved at distal 1 / 4, with a small sclerotized plate near apex (6 a); cornutus absent. Female genitalia. (Fig. 8). Papillae anales subrectangular, sparsely setose. Apophyses anteriores about half length of apophyses posteriores. Eighth sternum with sparse setae, slightly concave at middle on posterior margin. Lamella antevaginalis subpentagonal, strongly sclerotized. Ductus bursae narrow posteriorly, widened toward corpus bursae, with a sclerotized rectangular plate extending into corpus bursae anteriorly. Corpus bursae ovate, membranous; signum linear, arched medially, with dense denticles.	en	Min, Qiuyue, Li, Houhun, Wang, Shuxia (2026): Four species of the sakaiella group of Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from China. Zootaxa 5752 (1): 134-140, DOI: 10.11646/zootaxa.5752.1.6, URL: https://doi.org/10.11646/zootaxa.5752.1.6
572A3C3B1274FFA351C7FEF8FB77F933.taxon	distribution	Distribution. China (Sichuan).	en	Min, Qiuyue, Li, Houhun, Wang, Shuxia (2026): Four species of the sakaiella group of Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from China. Zootaxa 5752 (1): 134-140, DOI: 10.11646/zootaxa.5752.1.6, URL: https://doi.org/10.11646/zootaxa.5752.1.6
572A3C3B1274FFA351C7FEF8FB77F933.taxon	etymology	Etymology. The specific epithet is from the type locality, Maoxian County, Sichuan Province.	en	Min, Qiuyue, Li, Houhun, Wang, Shuxia (2026): Four species of the sakaiella group of Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from China. Zootaxa 5752 (1): 134-140, DOI: 10.11646/zootaxa.5752.1.6, URL: https://doi.org/10.11646/zootaxa.5752.1.6
572A3C3B1274FFA551C7F880FEEBFBC3.taxon	description	(Figs 3, 7, 9)	en	Min, Qiuyue, Li, Houhun, Wang, Shuxia (2026): Four species of the sakaiella group of Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from China. Zootaxa 5752 (1): 134-140, DOI: 10.11646/zootaxa.5752.1.6, URL: https://doi.org/10.11646/zootaxa.5752.1.6
572A3C3B1274FFA551C7F880FEEBFBC3.taxon	materials_examined	Type material. Holotype ♂, CHINA, Hainan: Jianfeng Town, Ledong County (18.70 ºN, 108.79 ºE), 40 m, 27. IV. 2013, leg. YH Sun, W Guan and TT Liu, slide No. MQY 25210. Paratypes. Hainan: 1 ♀, same data as holotype, slide No. MQY 25211; 4 ♂ 4 ♀, Bawangling, Changjiang County (19.11 ºN, 109.09 ºE), 146 – 225 m, 13 – 15. VIII. 2017, leg. X Bai, P Liu and S Yu, slide Nos. MQY 25206 ♂, MQY 25207 ♀; 3 ♂, Datian Nature Reserve, Dongfang City (19.11 ºN, 108.79 ºE), 56 m, 4 – 5. VI. 2018, leg. P Liu, X Bai and S Yu, slide No. MQY 24322.	en	Min, Qiuyue, Li, Houhun, Wang, Shuxia (2026): Four species of the sakaiella group of Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from China. Zootaxa 5752 (1): 134-140, DOI: 10.11646/zootaxa.5752.1.6, URL: https://doi.org/10.11646/zootaxa.5752.1.6
572A3C3B1274FFA551C7F880FEEBFBC3.taxon	diagnosis	Diagnosis. The new species can be distinguished from its congeners in the male genitalia by the valva distally divided into a large ovate dorsal lobe and a furcate ventral lobe, and the juxta with strongly sclerotized lateral process extending along costa and apically exceeding the dorsal ovate process of the valva; in the female genitalia by the eighth sternum concave at middle on the posterior margin, the short antrum slightly narrowed to the ductus bursae, and the large paired signa extending from below the entrance to anterior 1 / 3 of the corpus bursae.	en	Min, Qiuyue, Li, Houhun, Wang, Shuxia (2026): Four species of the sakaiella group of Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from China. Zootaxa 5752 (1): 134-140, DOI: 10.11646/zootaxa.5752.1.6, URL: https://doi.org/10.11646/zootaxa.5752.1.6
572A3C3B1274FFA551C7F880FEEBFBC3.taxon	description	Description. Adult (Fig. 3) with wingspan 5.5 – 6.0 mm. Head. Vertex white, frons white with greyish brown scales, occiput white with black scales. Labial palpus with second segments creamy white mixed with brown scales on inner surface, deep brown except distal 1 / 3 white on outer surface; third segment creamy white except black at base and beyond middle. Antenna with scape creamy white mixed with brown scales dorsally, pale yellowish brown ventrally; flagellum black alternating with yellowish grey dorsally, dark brown ventrally. Thorax. Mesonotum and tegula black, mixed with white. Forewing golden yellow; basal fascia black, edged with white scales on both inner and outer margins, with orange yellow scales in posterior area; postmedian fascia black, with a few white scales, extending from distal 1 / 3 of costal margin to between end of fold and tornus, slightly narrowed posteriorly; distal patch black, narrowed along termen to tornus, where it meets postmedian fascia, with a white dot at apex; fringe golden yellow mixed with greyish black except grey around tornus. Hindwing and fringe grey. Male genitalia (Fig. 7). Uncus wide at base, narrowed to basal 2 / 3, thereafter subparallel to before rounded apex. Gnathos very short, digitate; basal arm wider and longer than gnathos. Tegumen furcate from posterior 2 / 5, with a triangular anterior emargination, narrowed to anterior end laterally. Valva parallel sided basally, slightly widened from basal 2 / 5 to apex; apex concave at middle, forming two apex-rounded lobes, dorsoapical lobe large, wider than ventroapical lobe; sacculus wide and parallel from base to basal 2 / 3, then slightly narrowed to apex, distally setose, free from valva, being the narrower ventroapical lobe, with a subtriangular process arising from before apex, directed dorsad to dorsal lobe. Saccus triangular, approx. as long as uncus, narrowly rounded at apex. Juxta fused and weakly sclerotized basally, bilobed distally, each lobe fusiform, widened medially, narrowed to an acute apex, approx. as long as uncus; lateral process snake-like, arising from middle, strongly sclerotized, extending along costa to beyond dorsoapical lobe of valva, with long spine-like setae at apex; basal lobe conic, slender, nearly reaching middle of saccus. Phallus tubular, about 3 times length of saccus; cornutus absent. Female genitalia (Fig. 9). Papillae anales subrectangular, sparsely setose. Apophyses anteriores about half length of apophyses posteriores. Eighth sternum concave at middle on posterior margin. Ostium bursae large, round; antrum short, slightly narrowed toward ductus bursae. Ductus bursae membranous, almost as long as corpus bursae. Corpus bursae membranous, elliptical; two signa very large, elongate elliptical, with dense denticles, extending from below entrance to anterior 1 / 3 of corpus bursae.	en	Min, Qiuyue, Li, Houhun, Wang, Shuxia (2026): Four species of the sakaiella group of Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from China. Zootaxa 5752 (1): 134-140, DOI: 10.11646/zootaxa.5752.1.6, URL: https://doi.org/10.11646/zootaxa.5752.1.6
572A3C3B1274FFA551C7F880FEEBFBC3.taxon	distribution	Distribution. China (Hainan).	en	Min, Qiuyue, Li, Houhun, Wang, Shuxia (2026): Four species of the sakaiella group of Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from China. Zootaxa 5752 (1): 134-140, DOI: 10.11646/zootaxa.5752.1.6, URL: https://doi.org/10.11646/zootaxa.5752.1.6
572A3C3B1274FFA551C7F880FEEBFBC3.taxon	etymology	Etymology. The specific epithet is derived from the Latin serpentinus, referring to the shape of the lateral process of the juxta.	en	Min, Qiuyue, Li, Houhun, Wang, Shuxia (2026): Four species of the sakaiella group of Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from China. Zootaxa 5752 (1): 134-140, DOI: 10.11646/zootaxa.5752.1.6, URL: https://doi.org/10.11646/zootaxa.5752.1.6
572A3C3B1272FFA551C7FBECFB9EF961.taxon	description	(Figs 4, 10)	en	Min, Qiuyue, Li, Houhun, Wang, Shuxia (2026): Four species of the sakaiella group of Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from China. Zootaxa 5752 (1): 134-140, DOI: 10.11646/zootaxa.5752.1.6, URL: https://doi.org/10.11646/zootaxa.5752.1.6
572A3C3B1272FFA551C7FBECFB9EF961.taxon	materials_examined	Material examined. CHINA, Guangxi: 2 ♂, Shaoping Forest Farm, Pingxiang City, 280 m, 27. IV. 2012, leg. XF Yang, slide Nos. MQY 25002, MQY 25633; 1 ♀, Shaoping Forest Farm, Pingxiang City, 280 m, 25. III. 2013, leg. XF Yang, slide No. MQY 25001.	en	Min, Qiuyue, Li, Houhun, Wang, Shuxia (2026): Four species of the sakaiella group of Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from China. Zootaxa 5752 (1): 134-140, DOI: 10.11646/zootaxa.5752.1.6, URL: https://doi.org/10.11646/zootaxa.5752.1.6
572A3C3B1272FFA551C7FBECFB9EF961.taxon	description	Description. Adult (Fig. 4). Wingspan length 6.0 – 7.0 mm. Head and thorax. As previously described for male (see Wang & Jia 2019). Female genitalia (Fig. 10). Papillae anales setose, rounded on posterior margin. Apophyses anteriores about half length of apophyses posteriores. Eighth sternum with wrinkles, sparsely setose on posterior margin. Ostium bursae large, round; antrum short, weakly sclerotized. Ductus bursae membranous, almost as long as corpus bursae. Corpus bursae membranous, elliptical; two elongate subrectangular signa as long as 2 / 3 length of corpus bursae, with dense denticles arranged in longitudinal lines.	en	Min, Qiuyue, Li, Houhun, Wang, Shuxia (2026): Four species of the sakaiella group of Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from China. Zootaxa 5752 (1): 134-140, DOI: 10.11646/zootaxa.5752.1.6, URL: https://doi.org/10.11646/zootaxa.5752.1.6
572A3C3B1272FFA551C7FBECFB9EF961.taxon	distribution	Distribution. China (Guangxi). Note. The female of P. spiniflagellata is described for the first time herein.	en	Min, Qiuyue, Li, Houhun, Wang, Shuxia (2026): Four species of the sakaiella group of Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from China. Zootaxa 5752 (1): 134-140, DOI: 10.11646/zootaxa.5752.1.6, URL: https://doi.org/10.11646/zootaxa.5752.1.6
