identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
503B2128FFFD65779A6F93EAFBD38707.text	503B2128FFFD65779A6F93EAFBD38707.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Austinograea alayseae Guinot 1990	<div><p>Austinograea alayseae Guinot, 1990:</p><p>Western Pacific • 1 ♂, 1 ♀; western Pacific: TUIM06MV cruise; dive PL142, slurp 1, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-176.13333&amp;materialsCitation.latitude=-20.316668" title="Search Plazi for locations around (long -176.13333/lat -20.316668)">Lau Basin</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-176.13333&amp;materialsCitation.latitude=-20.316668" title="Search Plazi for locations around (long -176.13333/lat -20.316668)">Cam Tow</a>; 2719 m; 19.V.2005; 20°19’S, 176°08’W; ZRC 2024.0727, ex MNHN-IU-2022-4049 .</p></div>	https://treatment.plazi.org/id/503B2128FFFD65779A6F93EAFBD38707	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guinot, Danièle;Ng, Peter K. L.	Guinot, Danièle, Ng, Peter K. L. (2025): Kallograea kulolasi n. gen., n. sp. and K. jolliveti (Guinot & Segonzac, 2018) from the hydrothermal Kulo Lasi Volcano Caldera, West Pacific; and a reappraisal of Austinograea Hessler & Martin, 1989 and Gandalfus McLay, 2007 (Decapoda, Brachyura, Bythograeoidea). Zoosystema 47 (33): 773-797, DOI: 10.5252/zoosystema2025v47a33, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2025v47a33_.pdf
503B2128FFFD65779A6F9348FCB086AD.text	503B2128FFFD65779A6F9348FCB086AD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Austinograea hourdezi Guinot & Segonzac 2018	<div><p>Austinograea hourdezi Guinot &amp; Segonzac, 2018:</p><p>Paratypes. Western Pacific • 1 ♂ (21.4 × 34.3 mm); dive 232, Tu’i Malila site; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-176.56816&amp;materialsCitation.latitude=-21.989" title="Search Plazi for locations around (long -176.56816/lat -21.989)">Lau Back-Arc Basin</a>; 1891 m; 11.IX.2006; 21°59.34’S, 176°34.09’W: MNHN-IU-2016-10740 • 1 ♀; dive 427, ABE site, Lau Back-Arc Basin; 2130 m; 7.IX.2009; 20°45.65’S, 176°11.45’W; MNHNIU-2016-10744 .</p></div>	https://treatment.plazi.org/id/503B2128FFFD65779A6F9348FCB086AD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guinot, Danièle;Ng, Peter K. L.	Guinot, Danièle, Ng, Peter K. L. (2025): Kallograea kulolasi n. gen., n. sp. and K. jolliveti (Guinot & Segonzac, 2018) from the hydrothermal Kulo Lasi Volcano Caldera, West Pacific; and a reappraisal of Austinograea Hessler & Martin, 1989 and Gandalfus McLay, 2007 (Decapoda, Brachyura, Bythograeoidea). Zoosystema 47 (33): 773-797, DOI: 10.5252/zoosystema2025v47a33, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2025v47a33_.pdf
503B2128FFFD65779A6F94CFFB61860F.text	503B2128FFFD65779A6F94CFFB61860F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Austinograea williamsi Hessler & Martin 1989	<div><p>Austinograea williamsi Hessler &amp; Martin, 1989:</p><p>Paratypes. Western North Pacific • 1♂ (21.6 × 34.2 mm), 1 ♀ (22.5 × 35.3 mm), 1 small ♂ (15.2 × 23.8 mm); dive 1845; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=144.70718&amp;materialsCitation.latitude=18.209984" title="Search Plazi for locations around (long 144.70718/lat 18.209984)">Alice Springs</a> vent field; 3640 m; 6.V.1987; 18°12.599’N, 144°42.431’E; MNHN-IU-2008-11121 (= MNHN-B20910) .</p></div>	https://treatment.plazi.org/id/503B2128FFFD65779A6F94CFFB61860F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guinot, Danièle;Ng, Peter K. L.	Guinot, Danièle, Ng, Peter K. L. (2025): Kallograea kulolasi n. gen., n. sp. and K. jolliveti (Guinot & Segonzac, 2018) from the hydrothermal Kulo Lasi Volcano Caldera, West Pacific; and a reappraisal of Austinograea Hessler & Martin, 1989 and Gandalfus McLay, 2007 (Decapoda, Brachyura, Bythograeoidea). Zoosystema 47 (33): 773-797, DOI: 10.5252/zoosystema2025v47a33, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2025v47a33_.pdf
503B2128FFFD65779A6F9240FB0C84DF.text	503B2128FFFD65779A6F9240FB0C84DF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gandalfus puia McLay 2007	<div><p>Gandalfus puia McLay, 2007:</p><p>Western Pacific • holotype ♂ (15.5 × 24.3 mm); stn TAN0107 /128; Rumble III; 35°44.22-44.04’S, 178°29.72-29.63’E; 270- 239 m; 21.V.2001; NIWA 27855 • 1 paratype ♂ (21.3 × 33.5 mm); Macauley Caldera, Kermadec Islands; 337 m; 12.IV.2005; 30°12.78’S, 181°33.04’E; NIWA 18017 • 1 paratype ♂ (22.9 × 36.4 mm); Brothers Seamount, dive KOK0506/32; 1647 m; 2.V.2005; 34°51.70’S, 179°3.58’E; NIWA 18019 • 1 ♂ (22.6 × 36.7 mm) [coated with a brown ferric deposits, slightly damaged], 1 ♂ (11.6 × 18 mm); Kermadec Islands, Macauley Caldera, stn KOK0506/22; 337 m; 12.IV.2005; 30°12.78’S, 181°33.04’E; NIWA 18018 • 1 ♂ (entirely white) (18.8 × 29.0 mm); stn TAN1213/59; 405.0- 408 m; 26.X.2012; NIWA 86453 • 1♂, 6 ♀ [1 ♀ 10.4 × 14.3 mm, ZRC 2024.0726]; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=179.62517&amp;materialsCitation.latitude=-32.617332" title="Search Plazi for locations around (long 179.62517/lat -32.617332)">Haungaroa Caldera</a>; HYDROTHERMADEC cruise; dive 413; stn 026; 707 m; 31.XII.2016; 32°37.04’S, 179°37.51’E; MNHNIU-2024-6553 (ex SH162-065) .</p></div>	https://treatment.plazi.org/id/503B2128FFFD65779A6F9240FB0C84DF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guinot, Danièle;Ng, Peter K. L.	Guinot, Danièle, Ng, Peter K. L. (2025): Kallograea kulolasi n. gen., n. sp. and K. jolliveti (Guinot & Segonzac, 2018) from the hydrothermal Kulo Lasi Volcano Caldera, West Pacific; and a reappraisal of Austinograea Hessler & Martin, 1989 and Gandalfus McLay, 2007 (Decapoda, Brachyura, Bythograeoidea). Zoosystema 47 (33): 773-797, DOI: 10.5252/zoosystema2025v47a33, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2025v47a33_.pdf
503B2128FFFD65779A6F919BFA6085FA.text	503B2128FFFD65779A6F919BFA6085FA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gandalfus yunohana Takeda, Hashimoto & Ohta 2000	<div><p>Gandalfus yunohana Takeda, Hashimoto &amp; Ohta, 2000:</p><p>Western Pacific • 1♀ (28.2 × 42.2 mm); off central Japan, Philippine Sea Plate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.07784&amp;materialsCitation.latitude=26.705833" title="Search Plazi for locations around (long 141.07784/lat 26.705833)">Kaikata Seamount</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.07784&amp;materialsCitation.latitude=26.705833" title="Search Plazi for locations around (long 141.07784/lat 26.705833)">Shinkai 2000</a>, dive #1014; 26°42.35’N, 141°04.67’E; 448 m deep; 18.V.1998; MNHN-IU-2024-6053 (= MNHN-B28759) (ex JAMSTEC) • 1 ♀ (43.4 × 28.4 mm); same data; MNHN-IU-2024-6054 (= MNHN-B28759) (ex JAMSTEC) • 1 ♂ (20.5 × 29.9 mm); same data; MNHN-IU-2024-6055 (= MNHN-B28759) (ex JAMSTEC) • paratype ♂ (20.5 × 29.9 mm); off central Japan, Philippine Sea Plate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=139.86734&amp;materialsCitation.latitude=32.10317" title="Search Plazi for locations around (long 139.86734/lat 32.10317)">Myojin Knoll</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=139.86734&amp;materialsCitation.latitude=32.10317" title="Search Plazi for locations around (long 139.86734/lat 32.10317)">Shinkai 2000</a>, dive # 1007, 32°06.19’N, 139°52.04’E, 1263 m deep, 5.V.1998; MNHN-IU-2008-11865 (= MNHN-B28419).</p></div>	https://treatment.plazi.org/id/503B2128FFFD65779A6F919BFA6085FA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guinot, Danièle;Ng, Peter K. L.	Guinot, Danièle, Ng, Peter K. L. (2025): Kallograea kulolasi n. gen., n. sp. and K. jolliveti (Guinot & Segonzac, 2018) from the hydrothermal Kulo Lasi Volcano Caldera, West Pacific; and a reappraisal of Austinograea Hessler & Martin, 1989 and Gandalfus McLay, 2007 (Decapoda, Brachyura, Bythograeoidea). Zoosystema 47 (33): 773-797, DOI: 10.5252/zoosystema2025v47a33, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2025v47a33_.pdf
503B2128FFFC6576982596E8FEC88470.text	503B2128FFFC6576982596E8FEC88470.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kallograea Peter & L. 2025	<div><p>Kallograea n. gen.</p><p>urn:lsid:zoobank.org:act: 4AA24DD0-6035-4ED5-8075-54ECFABE95E6</p><p>TYPE SPECIES. — Kallograea kulolasi n. sp., by present designation.</p><p>OTHER INCLUDED SPECIES. — Austinograea jolliveti Guinot &amp; Segonzac, 2018 (see below).</p><p>ETYMOLOGY. — The name is derived from the Greek “ kallo ” for “beautiful”, and the name for marine crabs “ graea ” (cf. Liddell &amp; Scott 1940); alluding to the slender and delicate chelipeds of the two constituent species. The gender is feminine.</p><p>DIAGNOSIS. — Carapace transversely elliptical. Dorsal surface of carapace almost flat; regions indistinct. Eyestalk absent, podophthalmite fused to floor of orbital region; cornea absent, no visible pigment. Antennules, antennae recessed under front. Mxp3 ischium distinctly elongate; merus subtriangular, distally produced. Adult male chelipeds prominently elongate; merus long, slender; palm elongated (especially in K. kulolasi n. sp.), with setal patch either exclusively on inner surface and between fingers of both chelipeds ( K. jolliveti n. comb.) or on either side of superior margin (including the superior margin itself), thus on both sides of palm and extending between fingers ( K. kulolasi n. sp.); crusher with thick blunt-tipped fingers. No spot on anterior portion of palm near base of dactylus, but coloured spot at base of fixed finger ( K. jolliveti n. comb.). Adult male ambulatory legs slender, elongate, especially merus, propodus. Thoracic sternum short, wide. Male sternopleonal cavity gently concave, relatively wide distally, but without obvious depression for G1 tips. Longitudinal median line along entire male and female thoracic sternite 8, deep. G1 almost straight, obliquely directed, not crossing each other along distal parts. G2 either rather long, with the flagellum slightly longer than basal part, or shorter and with a small flagellum. Vulvae relatively large, occupying most of surface of thoracic sternite 6.</p><p>REMARK</p><p>The genera closest to Kallograea n. gen. are Austinograea Hessler &amp; Martin, 1989 and Gandalfus McLay, 2007, for which we deem necessary to give detailed illustrations of the species for comparisons of these two genera with the new genus established here; see below.</p></div>	https://treatment.plazi.org/id/503B2128FFFC6576982596E8FEC88470	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guinot, Danièle;Ng, Peter K. L.	Guinot, Danièle, Ng, Peter K. L. (2025): Kallograea kulolasi n. gen., n. sp. and K. jolliveti (Guinot & Segonzac, 2018) from the hydrothermal Kulo Lasi Volcano Caldera, West Pacific; and a reappraisal of Austinograea Hessler & Martin, 1989 and Gandalfus McLay, 2007 (Decapoda, Brachyura, Bythograeoidea). Zoosystema 47 (33): 773-797, DOI: 10.5252/zoosystema2025v47a33, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2025v47a33_.pdf
503B2128FFFC657D987F91EFFB5C8193.text	503B2128FFFC657D987F91EFFB5C8193.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kallograea KULOLASI N.	<div><p>Kallograea kulolasi n. sp.</p><p>(Figs 4-7)</p><p>urn:lsid:zoobank.org:act: 8E69D842-1647-45A5-8EB9-AC968353F769</p><p>TYPE MATERIAL. — Holotype. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-177.00961&amp;materialsCitation.latitude=-15.16782" title="Search Plazi for locations around (long -177.00961/lat -15.16782)">Western Pacific</a> • ♂ (right-handed) 7.3 × 12.7 mm; CHUBACARC 2019 cruise, Leg 1; Futuna Volcanic Arc, Kulo Lasi Caldera; 15°10.0692’S, 14°42.9563, 177°28.4632’, 177°00.5768’W (latitude minimum/maximum and longitude minimum/maximum, respectively); PL729 - GBT1; 1472 m; MNHNIU-2024-6067.</p><p>Paratypes. Western Pacific • 1 ♂ (right-handed) 7.5 × 12.0 mm; same data as holotype; MNHN-IU-2024-6084 • 1♂ 9.7 × 15.4 mm (without chelipeds); MNHN-IU-2024-6035 [CHU 036] • 1 ♀ 8.1 × 12.6 mm (with detached chelipeds in tube); MNHN-IU-2024-6033 • 1 ♂ 9.6 × 16.4 mm; MNHN-IU-2024-6552 [CHU 035] • 1 ♂ (left-handed) 8.8 × 14.1 mm; ZRC 2024.0724 (ex MNHNIU-2024-6085) .</p><p>TYPE LOCALITY. — Kulo Lasi Caldera, on periphery.</p><p>ETYMOLOGY. — Named after the Kulo Lasi Volcano where the species was discovered. Kulo lasi is a common name meaning ‘big cauldron’ in Futunian. Used as a noun in apposition.</p><p>DESCRIPTION</p><p>Carapace</p><p>Small size (carapace width 12.0- 16.4 mm). Carapace elliptical, very short, very elongated transversely, width-to-length ratio 1.59-1.74, flat; regions indistinct (Figs 4A, C; 5A; 6A). Dorsal surface entirely smooth, glabrous (Figs 4A, C; 5A; 6A). Anterolateral margin regularly rounded, with minute, barely discernible granules; supra-orbital margin with more obvious granules (Figs 4A, C, E; 5A, C; 6A, E). Posterolateral margins convergent posteriad; posterior margin slightly concave (Figs 4A, C; 5A; 6A). Subhepatic regions entirely glabrous (Figs 4E; 5B, C; 6E). Front broad, not protruded, practically straight, not emarginate medially, barely pointed medially, with two indistinct lobes; margin without discernible granules (Figs 4A, C, E; 5A; 6A, E). Infra-orbital region with scattered small granules (Figs 4E; 6E). Eyes, antennules, antennae recessed below front (Figs 4E; 6E). Orbit not delimited; orbital region extending as groove, lateral to area with vestigial eyestalks and antennae (Figs 4E; 5A, C; 6E). Eyestalk absent; podophthalmite barely visible, as small fixed piece fused to floor of orbital region; cornea absent but a tiny dark pigment barely visible (Figs 4E; 5A, C; 6E). Antennules folded horizontally (Figs 4E; 5C; 6E). Antenna very small; urinary article fixed, recessed; basal article (2 + 3) cylindrical, moveable; article 4 slightly elongated, inclined; flagellum not long. Proepistome very thin (Figs 4E; 5C, E; 6E). Posterior margin of epistome wide, with lateral margins distinctly concave, median projection obtusely triangular (Figs 4E; 6E). Pterygostomial lobe with small granules; pterygostomial region smooth (except minute granules along lateral line), glabrous, except along lateral line (Figs 4E; 5B, C; 6E).</p><p>Mxp3</p><p>Mxp3 completely closing buccal cavity, on all parts, especially between antero-external margin of merus and pterygostomial lobe. Ischium long, external margin oblique; longitudinal internal groove weak.Merus: external margin proximally straight, then obliquely directed; distal and external margins almost touching pterygostomial lobe; merus distal part markedly narrow, produced; internal margin bluntly angled medially. Carpus inserted on distal part of antero-internal margin of merus; propodus thick, short; dactylus long, reaching about three-quarters length of ischium; inner margins of propodus and dactylus with brush-like setae. Mxp3 coxa with only proximal portion visible, lateral projection hidden by junction of thoracic sternum (sternite 4) with pterygostome. Exopod thick, longer than endopod ischium (Figs 4D, E; 5B; 6E).</p><p>Male chelipeds</p><p>Male chelipeds elongate, not clearly heteromorphic (weak heterochely in females, see below) (Figs 4 A-C; 5C-F). Merus very long, slender, extending far beyond margin of carapace, subcylindrical, triangular in cross section, anterior border straight (without expansion), with regularly spaced small granules. Carpus covered with thick patches of thin soft setae on whole external surface; propodus of both chelipeds very long, with superior margin covered by thick patches of thin soft setae, appearing as a white down-like pubescence (Fig. 4C) (more developed on major chela), partially extending on both outer and inner parts of palm (more on inner part) and extending partially to basal part of dactylus. Fingers distinctly shorter than palm, with dark colour extending along distal third- or quarter-length; no setae along occluding margins. No obvious spot at base of propodus. Dark pigmentation on fingers not extending to palm, confined to tips in males (as in females) (Fig. 5 C-E).</p><p>In right-handed males (like holotype), major cheliped (crusher) stouter, slightly shorter than minor chela (cutter); both fingers very thick, distinctly blunt-tipped, weakly gaping; dactylus with small tooth, then molariform tooth on occluding margin; fixed finger thick, with some small denticles on occluding margin; dark colour extending dorsally on only distal part of fingers. Minor cheliped (cutter) with finger tips pointed. Merus as in major chela; palm rather long, outer surface slightly convex, smooth; both fingers not gaping at occluding margin, both tips curved, crossed; dactylus elongate, with occluding margin lined with some minute proximal teeth; fixed finger very thick, with thin extension bearing nearly straight occluding margin, with some small proximal teeth (Figs 4A, B; 5C, D).</p><p>In left-handed male (e.g., ZRC 2024.0724, ex MNHNIU-2024-6085), chelae less developed; fingers of major chela much stouter than pointed fingers of minor chela; both fingers relatively thinner, only very weakly gaping, less blunt-tipped, slightly curved at tip; minor chela elongated, with less dense patch of thin soft setae on superior margin; fixed finger regularly teethed (medially granular in left-handed males) (Fig. 5E, F).</p><p>Female chelipeds</p><p>Female chelipeds (♀ 8.1 × 12.6 mm mm, MNHNIU-2024-6033) not as developed as those of males, with merus distinctly shorter and narrower than in males (Fig. 6A, B). Chelae slightly heteromorphic, not blunt-tipped, with both fingers not gaping, crossing at tips; major chela only a little stouter as minor chela. As in males, patches of thin soft setae, appearing as down-like pubescence, on superior margin of carpus, propodus, extending on both outer and inner part of palm. Dark pigmentation of fingers in most part confined to tips (as in males) (Fig. 6C, D).</p><p>Ambulatory legs</p><p>P2-P5, in particular meri, distinctly long, slender. P2 much shorter than chelipeds; P3, P4 longest.Merus of P2-P4 smooth, without patches of setae, only with a few sparse setae; carpus, propodus and dactylus of P2-P5 with sparse setae on ventral margins (Figs 4 A-C; 6A, B).</p><p>Thoracic sternum</p><p>Thoracic sternum very wide, short, extending laterally, elliptical; suture 2/3 complete; sutures 4/5-7/8 incomplete; sutures 6/7 and 7/8 medially less calcified (Fig. 4D, F). Junction of sternite 4 with pterygostome represented by only short juxtaposition, without patch of setae. Median line at level of sternite 8 in both sexes, reaching to junction with sternite 7 (Fig. 4F). In males, press-button of locking mechanism acute, on sternite 5 abutting against suture 5/6 (Fig. 4F). Entirely smooth, glabrous in both sexes (Figs 4D, F; 6B, F).</p><p>Pleon</p><p>Both sexes with six free somites and telson. Male pleon rather wide, regularly triangular; pleonal somite 3 widest; pleonal somite 6 longest; telson very short, bluntly semicircular (Fig. 4B, D). Press-button on narrow portion of sternite 5, absent in females (Fig. 4F).</p><p>Female pleon almost round, covering most of thoracic sternites; telson semicircular (Fig. 6B).</p><p>Gonopods</p><p>G1 almost straight, directed obliquely into sternopleonal cavity, never contiguous, and even distant throughout their entire length, including their tips which are apart, not recessed in medial depression at end of sternopleonal cavity. Only some setae along mid distal part (Figs 4G; 7 A-D). G2 relatively long, flagellum slightly longer than basal part, curved, blade-like, distal part slightly dilated (Figs 4H; 7E, F).</p><p>Vulvae</p><p>Vulvae distant from each other, subhemispherical; close to but separate from thoracic sternal suture 5/6, occupying 2/3 of surface of sternite 6 (Fig. 6F).</p><p>REMARKS</p><p>Five males and one female of Kallograea kulolasi n. gen., n. sp., all small specimens, have been collected by the French CHUBACARC 2019 cruise on the periphery of the caldera of Kulo Lasi Volcano, through a unique dive in the north sector (Figs 2; 3). Immediately suspected of being new by Stéphane Hourdez (pers. comm.), they are here confirmed to represent an undescribed species.</p><p>Kallograea kulolasi n. gen., n. sp. is a small-sized species, however, with well-developed gonopods and vulvae. It is also the smallest bythograeid known in the world. The smallest bythograeid previously was Bythograea microps Saint Laurent, 1984 (Saint Laurent 1984: 356) from the East Pacific Rise (between 21°N and 9 ° 50 ’ N), which is indicated as having a maximum carapace width 40 mm according to Guinot &amp; Segonzac (2006b: 468), but it seems that this size may be overestimated: the male collected during the Hot 96 mission measuring 13 × 24 mm likely represents the largest B. microps known to date (Guinot &amp; Segonzac 1997: 139; Guinot &amp; Hurtado 2003: 437). Kallograea kulolasi n. gen., n. sp. is distinctly smaller, with specimens measuring 12 mm in carapace width already mature (e.g., MNHN-IU-2024-6084).</p><p>Kallograea kulolasi n. gen., n. sp. is characterised by a flat carapace, the males possessing distinctly elongate chelipeds, with especially blunt fingers on the major chela, and a prominent setal patch on the propodus that appears as a white down-like pubescence on the superior margin of the cheliped palm and extending on the inner and outer surfaces. This pubescence is best observed in water or ethanol, as the setae collapse and clump together when the specimen is studied dried (compare Fig. 4C with Fig. 5E, F). Since in most Austinograea species the setal patch is exclusively located on the inside of the palm, its presence on either side of the upper palm margin is a unique feature of K. kulolasi n. gen., n. sp. A setal patch is also present over the entire external surface of the carpus, which is not found in other bythograeoids, which have a glabrous carpus instead. The markedly thick and blunt-tipped fingers of the crusher cheliped is also a remarkable character of K. kulolasi n. gen., n. sp. The condition of these finger tips is unusual and perhaps functions similarly to the spoon-tipped fingers of other crabs. As indicated by Davie et al. (2015), not much is known about the precise function of spoon-tipped fingers, but they seem to be generally used for feeding on detritus, scooping up mucus from corals or picking up other soft foods, scraping off encrusting algae, effective gripping of filamentous algae, or scraping epilithic algae off coral rock. The rounded finger tips of the major chelae of K. kulolasi n. gen., n. sp. may be adapted for harvesting the filamentous bacterial matter in its habitat.</p><p>The palm and fingers of male adult chelipeds appear very heavy relative to the long and slender merus, in which the apodemes and attached muscles have little room, and together with the flat carapace, gives the crab a very distinctive appearance. Male Kallograea kulolasi n. gen., n. sp. clearly prioritises investing in strong chelipeds but in females, the chelipeds are proportionately shorter and the chelae are less strongly developed. This marked sexual dimorphism suggests the male chelipeds may have a role in reproductive selection.</p><p>DISTRIBUTION</p><p>Known only from the Futuna volcanic Arc, on the periphery of the Kulo Lasi Caldera.</p></div>	https://treatment.plazi.org/id/503B2128FFFC657D987F91EFFB5C8193	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guinot, Danièle;Ng, Peter K. L.	Guinot, Danièle, Ng, Peter K. L. (2025): Kallograea kulolasi n. gen., n. sp. and K. jolliveti (Guinot & Segonzac, 2018) from the hydrothermal Kulo Lasi Volcano Caldera, West Pacific; and a reappraisal of Austinograea Hessler & Martin, 1989 and Gandalfus McLay, 2007 (Decapoda, Brachyura, Bythograeoidea). Zoosystema 47 (33): 773-797, DOI: 10.5252/zoosystema2025v47a33, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2025v47a33_.pdf
503B2128FFF765699A7B948AFACF8751.text	503B2128FFF765699A7B948AFACF8751.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kallograea jolliveti (Guinot & Segonzac 2018) Peter & L. 2025	<div><p>Kallograea jolliveti (Guinot &amp; Segonzac, 2018) n. comb.</p><p>(Figs 8; 9)</p><p>Austinograea jolliveti Guinot &amp; Segonzac, 2018: 89, figs 9A-H, 10A-E, 11A-E.</p><p>MATERIAL EXAMINED. — Holotype. Western Pacific • ♂ 12.8 × 20.0 mm (right-handed); North Fiji Basin; STARMER II cruise, dive 18 (PL 18); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=173.48334&amp;materialsCitation.latitude=-18.833334" title="Search Plazi for locations around (long 173.48334/lat -18.833334)">Mussel Valley site</a>; 18°50’S, 173°29’E; 2750 m; 3.VII.1989; MNHN-IU-2016-10769.</p><p>Paratypes. Western Pacific • 1 ♂ (slightly right-handed) 14.7 × 25.4 mm, 1♀ 12.4 × 21.6 mm; same data as for holotype; MNHNIU-2016-10770 .</p><p>OTHER MATERIAL. — Western Pacific • 1 ♀ 13.6 × 22.0 mm (left-handed); Lau Back-Arc Basin; MGLN07MV cruise, dive 237; ABE site; 20°45.65’S, 176°11.45’E; 2130 m; 25.IX.2006; MNHNIU-2016-10751 .</p><p>NEW MATERIAL. — Western Pacific • 1 ♂ 14.3 × 22.7 mm (slightly left-handed), Futuna 1 cruise, Kulo Lasi, PL1776-4 – aspi 4; 14°56.35’S, 177°15.57W; 1406 m; 12.IX.2010; L. Menot coll.; R/V Atalante; ROV Nautile; M. Segonzac 06.2011 det. Austinograea sp.; ZRC 2024.0725 (MNHN-IU-2009-4045) • 2 ♂ 8.0 × 13.4 mm, 9.0 × 12.4 mm, 1 ♀ 8.4 × 13.8 mm (most of legs and chelipeds detached), Futuna 3 cruise, PL06 – aspi 04; MNHN-IU-2024-6075 • 1 ♀ 13.5 × 20.0 mm, CHUBACARC cruise, PL734 - GBT2, Manus Basin, Pac Manus, Roman’s Ruins; 03°43 649, 151°40 861; 1725 m; MNHN-IU-2024-6026 [CHU 511] • 1 ♂ 12.2 × 16.8 mm (right-handed); CHUBACARC cruise, PL733 - GBT5; Manus Basin, Pac Manus, Fenway; MNHN-IU-2024-6551 .</p><p>REMARKS</p><p>See description by Guinot &amp; Segonzac (2018: 89, figs 9AH; 10A-E; 11A-E) and remarks on K. kulolasi n. gen., n. sp. In Kallograea jolliveti n. comb., the pigmented spot near the base of the fixed finger and more or less in continuity with the dark colour of the fixed finger, and present on both chelae in both sexes (Figs 8C; 9C, D), recalls the distinctly coloured, more or less purplish, oval spot of Bythograea microps Saint Laurent, 1984, located practically in the same place and also present in both sexes (Saint Laurent 1984: pl. 1F; Guinot &amp; Segonzac 2006b: fig. 5). In addition to the similarly coloured spot on the cheliped palm, these two species are both relatively small; but, whereas B. microps shows only a drastic reduction of the eyestalks (Saint Laurent 1984: 386; 1988; Guinot &amp; Segonzac 2006b: fig. 2), K. jolliveti n. comb. is entirely blind: the eyestalks are absent, the podophthalmite is fused to the floor of the orbital region, and the cornea is absent, without any visible pigment (Fig. 8B).</p><p>DISTRIBUTION</p><p>Kallograea jolliveti n. comb. has a wide distribution in the western Pacific: initially, North Fiji and Lau Basins, now Kulo Lasi Volcano and Manus Basin.</p><p>COMPARISON BETWEEN KALLOGRAEA KULOLASI N. GEN., N. SP. (Figs 4-7) AND K. JOLLIVETI N. COMB. (Figs 8, 9) Kallograea kulolasi n. gen., n. sp. can easily be distinguished from K. jolliveti n. comb. by the suborbital region being glabrous or almost so (Figs 4E; 5A; 6E) (vs densely setose in K. jolliveti n. comb.; Fig. 8B); median lobe of the posterior epistomial margin being obtusely triangular, with the lateral margins concave (Figs 4E; 5C; 6E) (vs median lobe more acutely triangular with lateral margins gently sinuous to almost straight in K. jolliveti n. comb.; Fig. 8B); the mxp3 completely closes the buccal cavity across the anterior part, with the distal and external margins of the merus almost touching the pterygostomial lobe (Figs 4E; 5C; 6E) (vs distinct gap between the distal and external margins of the merus and the pterygostomial lobe in K. jolliveti n. comb.; Fig. 8B); the P2-P5, in particular the meri, are distinctly longer and more slender (Figs 4 A-C; 6A, B) (vs shorter and stouter in K. jolliveti n. comb.; Figs 8A; 9A); the ventral margins of P2–P4 have only scattered setae (vs with dense tomentum in K. jolliveti n. comb.); in both sexes, the dorsal margin of palm as well as the upper third or half of the outer surface (including the superior margin) and the upper third of the inner surface of adult male palm are densely setose but the rest of the inner surface is glabrous (Figs 4 A-C; 5C-F; 6A-D) (vs dorsal and outer surfaces glabrous although the inner surface is densely setose in K. jolliveti n. comb.; Figs 8A, C-F; 9A, C-E); there is no spot at the base of the male pollex (Figs 5C, E; 6C) (vs with a visible pale-coloured spot in K. jolliveti n. comb.; Figs 8C, D; 9, D); the fingers of the male chela are relatively shorter than the palm, with the pigmentation extending along the distal third or quarter (Figs 5 C-F; 6C, D) (vs the fingers of the male chela are as long as the palm, with the pigmentation extending along the distal half in K. jolliveti n. comb.; Figs 8A, C-F; 9A, C-E); the male thoracic sternum is proportionately wider, in relation with the short body of the species (Fig. 4B, D, F) (vs thoracic sternum less wide in K. jolliveti n. comb.; Figs 8G; 9G); the longitudinal median line on sternite 8 is relatively deeper (Figs 4F; 6F) (vs median line less deep in K. jolliveti n. comb.; Figs 8H; 9G); the male pleon is distinctly wider, with the telson semicircular in form (Fig. 4B, D) (vs pleon less wide with the telson more triangular in K. jolliveti n. comb.; Fig. 8G); the G1 is more strongly curved outwards (Figs 4G; 7A, B) (vs distinctly straighter in K. jolliveti n. comb.; Fig. 8I, J); the G2 is proportionately much longer in K. kulolasi n. gen., n. sp., with the flagellum slightly longer than basal part (Figs 4H; 7E) (vs G2 relatively shorter, with a small flagellum in K. jolliveti; Fig. 8K); and the medially positioned subhemispherical vulva occupying slightly more than half the space of sternite 6 (Fig. 6F) (vs relatively larger, ovate and occupying most of the space of sternite 6, with the anterior edge touching sternal suture 5/ 6 in K. jolliveti n. comb.; Fig. 11G).</p><p>The differences in the structure of the posterior epistomial margin (i.e., whether the mxp3 completely closes the anterior part of the buccal cavity), shape of the male telson, the very different proportions of the flagellum and basal part of the G2, and the structures of the vulvae, however, are significant, and may suggest K. jolliveti n. comb. is not a member of Kallograea n. gen. That being said, the flat carapace, wide male anterior thoracic sternum, wide male pleon, and elongate male chelipeds with the rounded finger tips are important shared characters with K. kulolasi n. gen., n. sp., and indicate that placing Austinograea jolliveti in Kallograea n. gen. is the best decision for the time being.</p><p>It is interesting to find two distinct species in a narrow perimeter on the periphery of the circular caldera of the Kulo Lasi Vulcano, an area with a diameter of only 5 km. The biota in this area, however, is known to be very dense and varied. Kallograea kulolasi n. gen., n. sp. seems confined to the Kulo Lasi Volcano, whereas the geographic range of K. jolliveti now extends from west to east, up to the Manus Basin (Figs 1-3).</p><p>COMPARISON BETWEEN KALLOGRAEA N. GEN. (Figs 4-9) AND AUSTINOGRAEA HESSLER &amp; MARTIN, 1989 (Figs 10; 11)</p><p>The genus Austinograea is known from four species from the western Pacific: the type species A. williamsi Hessler &amp; Martin, 1989, A. alayseae Guinot, 1990 (note that genetic differences have been found between individuals of A. alayseae from the Tofua Arc and the Manus Basin, see Kim et al. 2014), A. hourdezi Guinot &amp; Segonzac, 2018, and A. chubacarc Guinot, 2025 . A fifth species, A. rodriguezensis Tsuchida &amp; Hashimoto, 2002, occurs in the western Indian Ocean.</p><p>Austinograea williamsi (see Hessler &amp; Martin 1989: figs 1, 2, 4, 5a, 6a, 7a, 8b, 9-11, 13a, 14a-d; Tsuchida &amp; Fujikura 2000: figs 3, 5, 6, 8; Segonzac 2006: figs 1-4; see also Desbruyères et al. 2006; Kojima &amp; Watanabe 2015: figs 25.1, 25.2), found in abundance in beds of the snail Alviniconcha hessleri Okutani &amp; Ohta, 1988 that is common at the vent openings, is endemic to the Mariana Trough in the north-western Pacific, in the Mariana Back-Arc Basin, just west of the Mariana Island Arc. It should be noted that the ocular region of adult A. alayseae is not significantly different from that of adult A. williamsi (Hessler &amp; Martin 1989: fig. 4): the orbital region is only more recessed in A. williamsi, but, likewise, the eye is vestigial, fused to the orbital floor, and replaced by a small oval region in the posterior orbital wall lateral to the antenna, and with a trace of cornea that is more or less discernible and virtually unpigmented (sometimes a small spot) (cf. Hessler &amp; Martin 1989: figs 4, 5a for A. williamsi; Guinot 1990: fig. 1A, B for A. alysaeae). A key to the species of Austinograea was provided by Guinot &amp; Segonzac (2018: 96). One species described from the western Pacific, Austinograea yunohana Takeda, Hashimoto &amp; Ohta, 2000, was subsequently transferred by McLay (2007) to his new genus Gandalfus (see below).</p><p>Kallograea n. gen. is markedly different from Austinograea in that the dorsal surface of the carapace is almost flat (Figs 4A, C, E; 5A; 6A, E) (vs surface gently but distinctly convex in frontal view in Austinograea; Figs 10A, C; 11A); the posterior margin of the epistome is relatively wider (Figs 4E; 6E) (vs transversely narrower in Austinograea; Fig. 10C); the ischium of the mxp3 is distinctly elongate (Fig. 5B) (vs shorter in Austinograea, except in A. williamsi; Fig. 10H); the merus of the mxp3 is subtriangular in shape, with the anterior part much produced (Figs 4E; 5B; 6E) (vs subquadrate in Austinograea, except in A. williamsi; Fig. 10H); the male anterior thoracic sternum is short and very wide (Fig. 4B, D, F) (vs transversely narrower in Austinograea; Fig. 10B, H, I); the adult male chelipeds are distinctly more elongate and slender, especially the merus that is longer, distinctly extending well beyond the carapace margin, and slender, narrow on its whole length, and regularly toothed on the dorsal margin (Figs 4 A-C; 5C, D) (vs distinctly shorter and stouter, of moderate length and width in Austinograea [the chelipeds are relatively longer in A. willliamsi but still distinctly shorter than in Kallograea n. gen.]; Fig. 10A, B); the adult male ambulatory legs are proportionately more slender and elongate, especially the merus and propodus (Fig. 4 A-C) (vs relatively shorter and stouter in Austinograea; Fig. 10A, B); the male sternopleonal cavity is gently concave, relatively wide distally and without a concavity for the G1 tip (Fig. 4F) (vs cavity relatively deeper, relatively narrowing distally Fig. 10I, sometimes with a distinct depression for the G1 tips, in Austinograea); and the almost straight G1 is positioned more or less longitudinally with the distal parts of each not overlapping (Figs 4F, G; 7 A-D) (vs distinctly sinuous or straight with the distal parts of each overlapping to some degree in Austinograea (Fig. 10I) [condition not clear for A. rodriguezensis, cf. Tsuchida &amp; Fujikura 2000: fig. 8].</p><p>The form of the median lobe on the posterior epistomial margin of K. kulolasi n. gen., n. sp., being obtusely triangular (Figs 4E; 6E), is different from that of Austinograea species (acutely triangular in form; Fig. 10C); and is more similar in condition to that in Gandalfus (Figs 12 B-D; 14B, C; 15B; 16B), although the overall margin of K. kulolasi n. gen., n. sp. is distinctly wider. On the other hand, the median lobe of the margin in K. jolliveti n. comb., being acutely triangular (Figs 8B; 9B), is more similar in form to that of Austinograea species, although the overall margin is still wider in K. jolliveti n. comb.</p><p>In Austinograea alayseae Guinot, 1990, the extremities of the G1 join at the tips at the end of the sternopleonal cavity in situ and are positioned in a small deep depression (cf. Guinot 1990: 884, 891, fig. 2C; Guinot &amp; Segonzac 2006a: fig. 5). In A. hourdezi Guinot &amp; Segonzac, 2018 (Guinot 1990: figs 4C, 6C) and A. williamsi (Fig. 10I), the G1 tips also cross distally and are positioned in a narrow channel at the anterior part of the sternopleonal cavity. In contrast, in Kallograea n. gen., the obliquely directed G1 have the tips well separated and the anterior part of the sternopleonal cavity is proportionally wider, flatter and without a special distal depression to receive the G1 tips (Figs 4F; 8H).</p><p>Another significant distinction between Austinograea and Kallograea kulolasi n. gen., n. sp. is observed in the length of the flagellum of G2. It is much shorter than the basal article in Austinograea species (Fig. 10I) and is as long as the basis in Kallograea kulolasi n. gen., n. sp. (Figs 4H; 7E). In K. jolliveti n. comb., the G2 is proportionately shorter and the flagellum much shorter than that in K. kulolasi n. gen., n. sp. (Fig. 8K); Guinot &amp; Segonzac 2018: figs 9D, 11C-E).</p><p>No other Austinograea species, except A. williamsi, in which the male crusher has rounded finger tips (Fig. 10B, D, G), has finger tips as blunt as those on the crusher of Kallograea n. gen. Their overall cheliped structures, however, differ significantly, with that of Kallograea n. gen. being elongate and slender (Figs 4 A-C; 5C-F; 8A, C-F) and that of Austinograea, short and stout (Fig. 8A, B, D, G).</p><p>The above-discussed differences justify the transfer of Austinograea jolliveti into Kallograea n. gen., albeit with some reservations (see previously: Comparison between Kallograea kulolasi n. gen., n. sp. and K. jolliveti n. comb.).</p><p>COMPARISON BETWEEN KALLOGRAEA N. GEN. (Figs 4-9) AND GANDALFUS MCLAY, 2007 (Figs 12-16)</p><p>The type species, Gandalfus puia, has been collected from hydrothermal sites of the Kermadec Arc in the south-west Pacific Ocean, the longest underwater volcanic ridge on the planet, particularly in three active volcanoes between 30°12’S- 35°44’S and 181°33’E- 178°29’E (at 270- 239 m, at 1604-1647 m, and at 337 m). The ridge, which forms the base of the Kermadec Islands, is linear for about 1000 km and is a prolongation of the Tonga ridge. The Tonga-Kermadec Ridge is underlying the Tonga-Kermadec island arc and, on its western side, it is flanked by the back-arc-basin, the Lau Basin, which is at the boundary of the Australian and Pacific Plates (De Ronde et al. 2001; Wright 2001; Wright et al. 2002; Hauff et al. 2021).</p><p>The carapace of adult Gandalfus species is distinctly higher, with the dorsal surface more convex (Figs 12A, C, D; 13A, D; 14A, B; 15A; 16A) compared to those of Kallograea spp. which are lower and flatter (Figs 4A, C, E; 5A; 6A, E; 8A, B; 9A). The chelipeds of G. puia and G. yunohana are relatively short, have a short merus that is irregularly serrated and triangular in cross section, and a granulous and glabrous palm ending in relatively acute fingers that are armed with several proximal teeth of variable size, including one larger tooth at midlength of fixed finger (Figs 12A, F, G; 13A; 14A, E-G; 15E-H; 16A, D, E) (vs adult male chelipeds distinctly more elongate, with the merus distinctly longer, slender along its entire length, with the dorsal margin dentate, and extending well beyond carapace margin; the stout palm possessing setal patches and ending in thick and blunt fingers, and on the major chela, there is only a molariform tooth on the fixed finger in Kallograea n. gen.; Figs 4 A-C; 5C-F; 8A, C-F). While adult Gandalfus spp. are generally larger (carapace width: 36.7 mm in G. puia; 42.2 mm in G. yunohana), we have subadult specimens of G. puia (e.g., female 10.4 × 14.3 mm, ZRC 2024.0726, ex MNHN-IU-2024-6553) which are comparable in size to the two Kallograea species, but these differences remain valid.</p><p>McLay (2007: fig. 2D-F, table 2) recognised a new genus for his new species, Gandalfus puia, arguing that its G2 was quite different from those of Austinograea s. str. In G. puia, the G2 is approximately as long as the G1, with the flagellum as long as or longer than the basal article. Because of this character, McLay (2007) transferred Austinograea yunohana Takeda, Hashimoto &amp; Ohta, 2000 to Gandalfus (see Takeda et al. 2000: 164, 168, figs 4I, 6c, e) and, in doing so, he restricted Austinograea for species in which the G2 is clearly shorter than the G1, and has a shorter flagellum as well. The G2 is distinctly less than half the length of the G 1 in A. williamsi (Fig. 10I; Hessler &amp; Martin 1989: fig. 14a; Tsuchida &amp; Fujikura 2000: fig. 6; Guinot &amp; Segonzac 2006c: fig. 4) and A. rodriguezensis (cf. Tsuchida &amp; Hashimoto 2002: fig. 8; Tsuchida 2006: fig. 3); about half the length of the G 1 in A. hourdezi Guinot &amp; Segonzac, 2018 (cf. Guinot &amp; Segonzac 2018: fig. 6D, E); and more than half the length of the G1 (which is curved and armed with spiniform setae along its whole length) in A. alayseae Guinot, 1990 (cf. Guinot 1990: fig. 3A-C; Guinot &amp; Segonzac 2006a: fig. 6).</p><p>The discovery of the relative lengths of G2/G 1 in Kallograea kulolasi n. gen., n. sp. and K. jolliveti n. comb. reinforces the value of this character as a key taxonomic index. In both species of Kallograea n. gen., the G2 is curved, and is as long as or shorter than the G1 (Figs 4H; 7E; 8K); with the flagellum slightly longer than the basal part. This is also one of the main characters that distinguishes Gandalfus from Austinograea .</p><p>Another distinctive character of Gandalfus documented by McLay (2007), the slightly sinuous posterior epistomial margin (vs the strongly sinuous margin in Austinograea; see Tsuchida &amp; Hashimoto 2002: fig. 3), was difficult to appreciate because, unfortunately, the shape of the epistome of G. puia was not clearly illustrated in McLay’s paper. Our examination of the two genera shows that the median lobe of the margin is obtusely triangular in Gandalfus (Figs 12 B-D; 14B, D; 15B; 16B, C) but more distinctly acutely triangular in Austinograea (Fig. 8C; Tsuchida &amp; Hashimoto 2002: fig. 3). The posterior epistomial margin in both Kallograea species is distinctly wider than those of Gandalfus or Austinograea . As discussed earlier under Austinograea, the median lobe of this margin is obtusely triangular in K. kulolasi n. gen., n. sp. (Figs 4E; 5C; 6E).</p><p>McLay (2007: 7) described the vulva in a female of Gandalfus puia (carapace width 36.5 mm) as “slit-like”, oriented along anterior posterior body axis, not operculate, but he did not give an illustration. Gandalfus puia has a rounded vulva, with a distinct pointed tubercle at the anterior edge, which may be interpreted as a narrow sternal cover (Fig. 13F), as that of G. yunohana illustrated by Takeda et al. (2000: 166, fig. 4 E, as Austinograea yunohana; Fig. 16G). The vulva of K. kulolasi n. gen., n. sp. (Fig. 6F) and K. jolliveti n. comb. (Fig. 9G; Guinot &amp; Segonzac 2018: fig. 10E, as Austinograea jolliveti) is large, occupying most of the surface of sternite 6, and rounded, without any trace of an anterior tubercle. The vulva of Austinograea species is rounded, closed by soft membrane: in A. williamsi (Fig. 9B) and A. hourdezi (Guinot &amp; Segonzac 2018: fig. 10E), without any trace of an anterior tubercle.</p><p>The larvae of Gandalfus yunohana were reported on by Nakajima et al. (2010) and Hamasaki et al. (2010). The complete mitogenome of G. yunohana, from the Nikko Seamount on the Izu-Ogasawara Ridge, was reported by Yang et al. (2010: fig. 3) who showed that it had highly conserved characteristics and appeared to be related to brachyurans such as Pseudocarcinus gigas (Lamarck, 1818) . Pseudocarcinus gigas is at present in its own family (see Ng &amp; Davie 2020), and this supposed relationship needs to be revisited. The complete mitochondrial genome of G. puia, from the Tonga Arc, was obtained by Kim et al. (2015: fig. 1), showing that the genetic data of this taxon is distinct from that of Austinograea s. str. The phylogenetic tree of Wang et al. (2019: fig. 1) includes Gandalfus puia, G. yunohana, Austinograea alayseae, A. rodriguezensis and Segonzacia mesatlantica (Williams, 1988), and they show the same separation, indicating Gandalfus and Austinograea are separate genera (see also the Bayesian inference tree in Ma et al. 2024). Barcoding sequences of the mitochondrial COI gene of G. yunohana, from four vent fields (including two on the Izu Arc, one on the northern Mariana Arc and one in the Okinawa Trough), have shown a similar genetic diversity of populations on the Izu and northern Mariana Arcs and a sharing of the dominant haplotypes, without genetic subdivision regardless of the habitat depth, the whole suggesting a high dispersal capability for G. yunohana (cf. Watanabe et al. 2020).</p></div>	https://treatment.plazi.org/id/503B2128FFF765699A7B948AFACF8751	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guinot, Danièle;Ng, Peter K. L.	Guinot, Danièle, Ng, Peter K. L. (2025): Kallograea kulolasi n. gen., n. sp. and K. jolliveti (Guinot & Segonzac, 2018) from the hydrothermal Kulo Lasi Volcano Caldera, West Pacific; and a reappraisal of Austinograea Hessler & Martin, 1989 and Gandalfus McLay, 2007 (Decapoda, Brachyura, Bythograeoidea). Zoosystema 47 (33): 773-797, DOI: 10.5252/zoosystema2025v47a33, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2025v47a33_.pdf
