taxonID	type	description	language	source
50613239FF8C6C45FF0AFC55FEBEFE05.taxon	description	(Figs 1, 4 – 6, 13, 30)	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF8C6C45FF0AFC55FEBEFE05.taxon	materials_examined	Type material examined. Holotype ♂: CHINA: SICHUAN: ‘ CHINA: SICHUAN PROV., | Emeishan Mt., 9. VI. 2014, | LEIDONGPING env., 2410 m, | 29 ° 32´40 ´´ 103 ° 20´02 ´´ E, ’ <printed>, ‘ sift # 8, mixed forest with Acer, | Abies, Picea, Rhododendron, | around limestone rocks, | J. Hájek & J. Růžička leg. ’ <printed>, ‘ HOLOTYPE | Omalium | anguliferum sp. nov. | Shavrin A. V. des. 2025 ’ <printed> (NMPC). Paratypes (12 ♂♂, 11 ♀♀): CHINA: SICHUAN: 1 ♀: same data as the holotype (cSh); 2 ♂♂ (one specimen dissected): ‘ CHINA: SICHUAN PROV., | Emeishan Mt., 2420 m | JIEYINGDIAN TEMPLE, | 29 ° 32´17 ´´ 103 ° 19´59 ´´ E, ’ <printed>, ‘ 10. VI. 2014, sift # 12, | secondary mixed forest | above temple, | J. Hájek & J. Růžička leg. ’ <printed> (NMPC); 1 ♀ (dissected): ‘ CHINA: SICHUAN PROV., | Emeishan Mt., 2480 m | JIEYINGDIAN TEMPLE, | 29 ° 32´01 ´´ 103 ° 19´57 ´´ E, ’ <printed>, ‘ 7. VI. 2014, sift # 6, | bamboo and mixed forest | below temple near road, | J. Hájek & J. Růžička leg. ’ <printed> (NMPC); 1 ♂ (dissected), 1 ♀: ‘ CHINA: Sichuan | Emei Shan, Wannian | 1050 m, 19. – 30. III. 1999 | leg. P. JÄGER’ <printed> (SMNS); 3 ♂♂ (one specimen dissected), 3 ♀♀: ‘ P. R. CHINA, Sichuan, | EmeiShan, N 29 ° 33´36.3 ´´ | E 103 ° 20´38.0 ´´, | 15. vi. 2010, 1947 m, | sifting 33, V. Grebennikov’ <printed> (1 ♂, 1 ♀: cSh; 2 ♂♂, 2 ♀♀: CNC); 1 ♂, 3 ♀♀: ‘ P. R. CHINA, Sichuan, | Emei Shan, N 29 ° 33.6´| E 103 ° 20.6´, 27. vi. – 5. vii. | 2009, 1800 – 2400 m, sifti- | ng 11 – 17, V. Grebennikov’ <printed> (1 ♂, 1 ♀: cSh; 2 ♀♀: CNC); 2 ♂♂, 2 ♀♀: ‘ P. R. CHINA. Sichuan, | EmeiShan, N 29 ° 33´36.3 ´´ | E 103 ° 20´38.0 ´´, | 22. vi. 2010, 1947 m, | sifting 39, V. Grebennikov’ <printed> (1 ♂, 1 ♀: cSh; 1 ♂, 1 ♀: CNC); 1 ♂: ‘ CHINA: Sichuan (2) | Qingcheng-Shan, | Rückseite, 650 – 700 m | 30.53.56 N, 103.33.01 E | 18.05.1997, M. Schülke’ <printed>, ‘ Omalium [handwritten] | spec. [handwritten] | det. M. Schülke 1999 ’ <printed>, ‘ Museum für Naturkunde | Berlin | Sammlung M. Schülke’ <printed> (cSch); CHINA: SHAANXI: 2 ♂♂ (one specimen dissected): ‘ CHINA: S-Shaanxi (Qinling Shan) | pass on rd. Zhouzhi — Foping, | 105 km SW Xi`an, N-slope, | 1990 m, 33 ° 44´N, 107 ° 59´E | leg. M. Schülke [C 01 - 01] ’ <printed>, ‘ 2. / 4. VII. 2001, | small creek valley, mixed | deciduous forest, bamboo, | small meadows, dead wood, | mushrooms (sifted) [C 01 - 01] ’ <printed>, ‘ Sammlung | M. Schülke | Berlin’ <printed> (cSch). All paratypes with additional red printed label: ‘ PARATYPE | Omalium | anguliferum sp. nov. | Shavrin A. V. des. 2025 ’.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF8C6C45FF0AFC55FEBEFE05.taxon	description	Description. Measurements (n = 24): HW: 0.51 – 0.57; HL: 0.34 – 0.41; OL: 0.12 – 0.17; TL: 0.07 – 0.10; AL (holotype): 1.00; PL: 0.44 – 0.52; PWmax: 0.61 – 0.72; PWmin: 0.59 – 0.65; ESL: 0.76 – 0.92; EW: 0.84 – 0.97; MTbL (holotype): 0.42; MTrL (holotype): 0.19 (MTrL 1 – 4: 0.07; MTrL 5: 0.12); AW: 0.84 – 0.90; AedL: 0.55 – 0.60; BL: 2.45 – 3.35 (holotype). Habitus as in Fig. 1. Body brown to reddish-brown, with distinctly paler lateral and basal margins of pronotum and sometimes with slightly paler elytra and abdomen; antennomeres 5 – 11 or 6 – 11 brown; mouthparts, antennomeres 1 – 4 or 1 – 5 and legs yellow-brown; tarsi yellowish. Punctation of head dense, coarse and deep, sparser and finer in frontal and larger in middle portions, with interspaces between punctures in middle about as long as diameter of one nearest puncture or slightly shorter; neck with irregular, sparse and moderately fine punctation; punctation of pronotum regular, about as that in middle of head, distinctly finer and slightly sparser in mediobasal and significantly sparser in laterobasal portions; scutellum without punctures; punctation of elytra distinctly sparser, but larger and deeper than that on pronotum, coarser in medioapical part, finer and sparser along suture; abdominal tergites with fine and relatively dense punctation, indistinct on abdominal tergites VI – VIII. Forebody glossy; anterior portion of clypeus with fine transverse microreticulation, supra-antennal elevations with fine and sometimes indistinct diagonal meshes; lateral sides of neck with diagonal sculpture; pronotum and elytra without microsculpture; scutellum without or with indistinct fine transverse sculpture; abdomen with dense microsculpture, somewhat transverse on abdominal tergites IV – V and isodiametric on tergites VI – VIII. Anterior part of head with several moderately long and erect setae; lateral margins of pronotum with several short setae; abdominal tergites with sparse and short setation; posterior margin of pronotum with row of short cuticular fringe. Head 1.3 – 1.5 times as broad as long, with broad clypeus and distinctly elevated supra-antennal elevations, with relatively narrow and deep anteriomedian depressions, almost reaching level of anterior margins or anterior third of eyes; posteriolateral margins of clypeus stretching posteriad toward level of anterior third of eyes. Latero-apical margin between anterior margin of eyes and clypeus without semicircular notch. Dorsal surface with strong longitudinal and diagonal elevations between punctures in middle and on infraorbital portions; irregular elevations on postocular portions sometimes forming three to four longitudinal wrinkles. Anteocellar foveae moderately narrow and deep, distinctly convergent latero-apicad toward level of middle or anterior third of eyes. Temples long, 1.7 times as long as longitudinal length of eye, from posterior margins of eyes slightly narrowed posteriad, each with subacute angle distinctly protruded basad; postocular ridges missing. Apical part of neck narrowly and moderately deeply depressed, with several short and narrow elevations; each lateral margin with acute small projections, slightly protruded laterad. Eyes large and convex. Ocelli large, located significantly behind level of posterior margins of eyes; distance between ocelli short, 1.6 – 1.8 times as long as distance between ocellus and posterior margin of eye. Maxillary palpi long, apical palpomere about four times as long as small and transverse penultimate segment, from widest basal portion gradually narrowed toward acute apex. Antenna reaching basal portion of elytra when reclined, with slightly elongate antennomeres 4 – 6 and distinctly transverse antennomeres 7 – 10; antennomere 3 slightly shorter and distinctly narrower than 2, 4 1.4 times as long as and narrower than 3, 5 distinctly longer than 4, 6 slightly longer and broader than 5, 7 slightly broader than 6, 8 distinctly broader than 7, 9 – 10 slightly broader than 8, apical antennomere 1.3 – 1.4 times as long as preceding segment, from middle gradually narrowed toward subacute apex. Pronotum convex, 1.3 times as broad as long, 1.1 – 1.2 times as broad as head, from widest middle distinctly more narrowed posteriad than anteriad. Apical angles slightly protruded anteriad. Anterior margin widely rounded, slightly protruded anteriad. Laterobasal portions sometimes slightly concave in front of obtuse hind angles. Lateral portions relatively deeply and narrowly impressed, deeper in middle. Lateral margins narrowly marginated, sometimes indistinctly crenulate in latero-apical part. Surface of disc with two long and wide longitudinal depressions, broadened basad; some paratypes with indistinct oval depression in medioapical portion; each laterobasal portion widely and deeply depressed; surface between all pronotal depressions distinctly elevated. Middle portion with irregular longitudinal or diagonal elevations between punctures. Elytra slightly broader than long, 1.7 times as long as pronotum, from basal portion slightly broadened posteriad; lateral margins marginated; lateral portions narrowly impressed; hind margins somewhat straight or widely rounded. Dorsal surface of each elytron with strong irregular elevations between punctures: transverse and subdiagonal in mediobasal and longitudinal in apical portions. Wings fully developed. Metatarsi slightly more than twice shorter than metatibia. Abdomen convex, about as wide as elytra, with two small oval wing-folding patches in middle of abdominal tergite IV and distinct narrow palisade fringe on apical margin of abdominal tergite VII, sometimes with narrow to wide intersegmental membranes between abdominal tergites III – V. Male. Posterior margin of abdominal tergite VIII slightly and sternite VIII widely and moderately deeply sinuate. Aedeagus with wide basal portion, gradually narrowed toward median lobe; median lobe wide, with deeply sinuate lateral parts in preapical portion and widely rounded apex; mediolateral portions with narrow and relatively short accessory plates rounded apically; parameres moderately narrow, distinctly broadened apically, reaching preapical part of median lobe, each with four moderately long apical setae; internal sac wide and relatively short, with two small oval structures in basal part (Fig. 4). Lateral aspect of the aedeagus as in Fig. 5; apical portion of median lobe (lateral view) with three ventrolateral obtuse teeth (Fig. 6). Female. Posterior margin of abdominal tergite VIII truncate. Posterior margin of abdominal sternite VIII truncate or rounded. Accessory sclerite with moderately narrow basal portion and relatively elongate median part, rounded apically (Fig. 9). Spermatheca not recognized.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF8C6C45FF0AFC55FEBEFE05.taxon	discussion	Comparative notes. Based on the shape of temples, O. anguliferum sp. nov. belong to the acutangulum group, recently established for two species (O. acutangulum Shavrin, 2023 and O. amicorum Shavrin, 2023) from Nepal (Shavrin 2023 a). Besides that, it related to Chinese O. rutilum sp. nov. and O. sectum sp. nov. Regarding the shape of postocular portions of head slightly narrowed posteriad and forming distinct hind angles, the shape of the pronotum with apical angles slightly protruded apicad, O. anguliferum sp. nov. is more similar to O. acutangulum and O. rutilum sp. nov. It can be distinguished from O. acutangulum by the finer elevations between punctures in basal part of the head, and slightly narrower pronotum with finer and smaller depressions on the pronotal disc. From O. rutilum sp. nov. it differ by the shape of transverse antennomere 8 and the shape of narrower pronotum, widest in middle and with less pronounced dorsal impressions. From both species it can be distinguished by the darker body, significantly denser and coarser punctation of the pronotum, different shape of the broader median lobe and other details of internal and external morphology of the aedeagus, and different shape of the female accessory sclerite. From O. sectum sp. nov. it differs by the widely rounded anterior angles of the pronotum and different morphology of the aedeagus.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF8C6C45FF0AFC55FEBEFE05.taxon	distribution	Distribution. Omalium acutangulum sp. nov. is known from several localities in Qinling Shan mountain range in Shaanxi and in Emeishan and Qingcheng-Shan mountain ranges in Sichuan, China (Fig. 30).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF8C6C45FF0AFC55FEBEFE05.taxon	etymology	Etymology. The specific epithet (Latin adjective: that carries angles) refers to the shape of temples strongly narrowed and protruding posteriad.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF8C6C45FF0AFC55FEBEFE05.taxon	biology_ecology	Bionomics. Specimens were collected at elevations from 650 to 2480 m a. s. l. and were sifted from litter of mixed coniferous (Acer, Abies, Picea, Rhododendron), and deciduous forest with bamboo, dead wood and mushrooms (C 01 - 01).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF8B6C48FF0AFDE6FC97FD00.taxon	description	(Figs 2, 7 – 9, 14 – 15, 30)	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF8B6C48FF0AFDE6FC97FD00.taxon	materials_examined	Type material examined. Holotype ♂: CHINA: YUNNAN: ‘ CHINA: N-Yunnan [C 2005 - 09] | Diqing Tibet. Aut. Pref., | Deqin Co., Meili Xue Shan, | E-side, 14 km W Deqin, 2580 m, ’ <printed>, ‘ 28 ° 27.47´N, 98 ° 46.35´E, creek | valley below glacier, mixed forest, | leaf litter, moss, dead wood, sifted, | 11. VI. 2005, M. Schülke [C 2005 - 09] ’ <printed>, ‘ Museum für Naturkunde | Berlin | Sammlung M. Schülke’ <printed>, ‘ HOLOTYPE | Omalium | rutilum sp. nov. | Shavrin A. V. des. 2025 ’ <red, printed> (cSch). Paratypes (1 ♂, 2 ♀♀): YUNNAN: 1 ♀ (dissected; both antennes missing): same data as the holotype (cSh); 1 ♀: ‘ CHINA: N-Yunnan Diqing Tibet | Aut. Pr. Deqin Co. Meili Xue | Shan E-side 14 km W Deqin | 28 ° 27.47´N, 98 ° 46.35´E 2580 m | 11. VI. 2005 A. Smetana [C 158] ’ (CNC); 1 ♂ (dissected; right elytron missing): ‘ CHINA (Yunnan) | Dali Bai Auton. Pref., | Diancang Shan W Dali | 25 ° 41´20 ´´ N / 100 ° 06´12 ´´ E | 3160 m (small creek valley, | litter and debris sifted | 27. V. 2007 D. W. Wrase [02] ’ <printed>, ‘ Museum für Naturkunde | Berlin | Sammlung M. Schülke’ <printed> (cSch). All paratypes with additional red printed label: ‘ PARATYPE | Omalium | rutilum sp. nov. | Shavrin A. V. des. 2025 ’.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF8B6C48FF0AFDE6FC97FD00.taxon	description	Description. Measurements (n = 4): HW: 0.56 – 0.62; HL: 0.37 – 0.42; OL: 0.15 – 0.16; TL: 0.10 – 0.12; AL (holotype): 1.15; PL: 0.37 – 0.49; PWmax: 0.66 – 0.76; PWmin: 0.60 – 0.72; ESL: 0.83 – 0.90; EW: 0.87 – 1.10; MTbL (holotype): 0.47; MTrL (holotype): 0.20 (MTrL 1 – 4: 0.09; MTrL 5: 0.11); AW: 0.85 – 1.11; AedL: 0.47; BL: 2.57 – 3.10 (holotype: 2.70). Habitus as in Fig. 2. Body reddish-brown, with distinctly darker head and abdomen; antennomeres 7 – 11 brown; mouthparts and antennomeres 1 – 6 yellow-brown; lateral portions of pronotum, lateral and apical portions of elytra, paratergites of abdomen and legs yellowish (tarsi slightly paler). Punctation of head dense, coarse and deep, denser and coarser in middle and on infraorbital portions, with interspaces between punctures in middle slightly less than diameter of nearest puncture; punctation of pronotum about as that in middle portion of head, but slightly larger, more irregular, slightly finer and sparser in middle and finer and sparser in mediobasal portion, with interspaces between punctures in middle about as long as diameter of nearest puncture; punctation of elytra slightly sparser and distinctly larger and deeper than that on pronotum, denser in mediobasal and mediolateral part, finer and sparser along suture and in medioapical portion. Anterior portion with distinct transverse microreticulation, middle part of clypeus with fine transverse and supra-antennal elevations with diagonal sculpture; mediobasal part of neck with transverse meshes; scutellum with dense and fine transverse microsculpture; abdomen with dense isodiametric sculpture. Apical and basal margins of pronotum with row of short cuticular fringe. Head 1.4 – 1.5 times as broad as long, with strongly elevated supra-antennal elevations and wide and deep anteriomedian depressions. Dorsal surface with strong longitudinal elevations between punctures, distinctly diagonal in laterobasal parts of clypeus and on infraorbital portions; postocular parts with irregular longitudinal wrinkles. Anteocellar foveae narrow and deep, somewhat subparallel, almost reaching level of middle length of eyes. Temples 1.3 – 1.5 times as long as longitudinal length of eyes, each with subacute posterior angle protruded basad. Medioapical part of neck narrowly and slightly depressed, with short and narrow elevations. Distance between ocellus 1.5 – 1.7 times as long as distance between ocellus and posterior margin of eye. Apical palpomere about three and a half times as long as penultimate segment. Antenna with slightly elongate antennomeres 7 – 8 and distinctly transverse 9 – 10; antennomere 3 about as long as and distinctly narrower than 2, 4 distinctly shorter and slightly narrower than 3, 5 slightly longer than 4, 6 distinctly longer than 5, 7 indistinctly broader than 6, 8 slightly broader than 7, 9 – 10 distinctly broader than 8, apical antennomere slightly longer than preapical antennomere, from middle gradually narrowed toward rounded apex. Pronotum transverse, 1.5 – 1.7 times as broad as long, widest in anterior third, gradually narrowed posteriad. Laterobasal portions not concave in front of obtuse hind angles. Lateral and laterobasal portions deeply and widely impressed. Surface of disc with two long and deep longitudinal depression, distinctly broadened and deepened basad; medioapical part with deep oval depression; surface between all pronotal depressions strongly elevated. Elytra slightly less than twice or slightly more than twice as long as pronotum, lateral margins narrowly impressed; hind margins straight. Dorsal surface of each elytron with strong irregular elevations, finer in middle. Male. Posterior margin of abdominal tergite VIII somewhat straight. Posterior margin of abdominal sternite VIII sinuate. Aedeagus with wide basal portion, slightly narrowed toward median lobe; median lobe moderately narrow, from widest middle gradually narrowed toward relatively small rounded apex; accessory plates narrow, moderately short and slightly curved apically; parameres wide, broadened in preapical part, each with two long and two short apical setae; internal sac moderately wide, long, without elongate sclerotized structures (Fig. 7). Lateral aspect of the aedeagus as in Fig. 8; apical portion of median lobe elongate (lateral view), without ventrolateral teeth (Fig. 9). Female. Posterior margin of abdominal tergite VIII truncate. Posterior margin of abdominal sternite VIII widely rounded. Accessory sclerite with wide basal portion and short median part, strongly narrowed toward acute apex (Fig. 14). Spermatheca as in Fig. 15.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF8B6C48FF0AFDE6FC97FD00.taxon	discussion	Comparative notes. Based on the shape of postocular portions of head and the shape of the apical part of the pronotum with apical angles slightly protruded apicad, O. rutilum sp. nov. is similar to Nepalese O. acutangulum and Chinese O. anguliferum sp. nov. Based on general shapes of the body, the aedeagus and the female accessory sclerite, it is more similar to O. acutangulum, from which it can be distinguished by the paler body, finer longitudinal elevations of median part of the head, denser punctation of the pronotum and the elytra, slightly shorter median lobe, broader parameres, and other details of the external and internal morphology of the aedeagus. From O. anguliferum sp. nov., it differs by the narrower antennomere 8, the shape of more transverse pronotum, widest in anterior third and with more pronounced dorsal (including medioapical) impressions, and different morphology of the aedeagus and the female accessory sclerite.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF8B6C48FF0AFDE6FC97FD00.taxon	distribution	Distribution. Omalium rutilum sp. nov. is known from three localities in Qinling Shan mountain range in Shaanxi and in Meili Xue Shan and Diancang Shan mountain ranges in Yunnan, China (Fig. 30).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF8B6C48FF0AFDE6FC97FD00.taxon	etymology	Etymology. The specific epithet is the Latin adjective rutilus, - a, - um (reddish). It alludes to the coloration of the body.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF8B6C48FF0AFDE6FC97FD00.taxon	biology_ecology	Bionomics. Specimens were collected at elevations from 2580 to 3160 m a. s. l. and were sifted from leaf litter, debris and mosses. One specimen was collected by sifting of wet leaf litter, debris and moss on and around small seepages and on edges of a small creek in a mixed broadleaved forest (C 158). It was collected in Meili Xue Shan together with O. sectum sp. nov. and O. tibetanum sp. nov.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF866C49FF0AFCE2FBFCFC39.taxon	description	(Figs 3, 10 – 12, 30)	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF866C49FF0AFCE2FBFCFC39.taxon	materials_examined	Type material examined. Holotype ♂ (Fig. 3; dissected): CHINA: YUNNAN: ‘ CHINA: N-Yunnan [C 2005 - 09] | Diqing Tibet. Aut. Pref., | Deqin Co., Meili Xue Shan, | E-side, 14 km W Deqin, 2580 m, ’ <printed>, ‘ 28 ° 27.47´N, 98 ° 46.35´E, creek | valley below glacier, mixed forest, | leaf litter, moss, dead wood, sifted, | 11. VI. 2005, M. Schülke [C 2005 - 09] ’ <printed>, ‘ Museum für Naturkunde | Berlin | Sammlung M. Schülke’ <printed>, ‘ HOLOTYPE | Omalium | sectum sp. nov. | Shavrin A. V. des. 2025 ’ <red, printed> (cSch).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF866C49FF0AFCE2FBFCFC39.taxon	description	Description. Measurements: HW: 0.62; HL: 0.43; OL: 0.17; TL: 0.12; AL: 1.35; PL: 0.55; PWmax: 0.72; PWmin: 0.67; ESL: 1.01; EW: 1.16; MTbL: 0.35; MTrL: 0.17 (MTrL 1 – 4: 0.07; MTrL 5: 0.10); AW: 1.02; AedL: 0.70; BL: 3.54. Habitus as in Fig. 3. Body reddish-brown, with slightly paler abdomen; antennomeres 6 – 11 brown; mouthparts, antennomeres 1 – 5 and legs yellow-brown. Punctation of head dense, denser and coarser in middle and on infraorbital portions; neck with moderately fine and sparse punctation; punctation of pronotum fine and sparse, sparser in medioapical, lateral and mediobasal portions; punctation of pronotum distinctly denser than that on pronotum, coarser in middle; abdominal tergites without punctation. Clypeus with fine transverse microsculpture; scutellum with fine isodiametric microreticulation; abdomen with dense isodiametric sculpture. Head 1.4 times as broad as long, with broad and deep anteriomedian depressions; posteriolateral margins of clypeus stretching toward level of middle length of eyes. Latero-apical margin between anterior margin of eyes and clypeus with small semicircular notch. Dorsal surface with distinct transverse and longitudinal elevations between punctures in middle and irregular elongate elevations on postocular parts. Anteocellar foveae very wide and deep, somewhat elongate, reaching level of posterior third length of eyes. Temples 1.4 times as long as longitudinal length of eye, from posterior margin of eyes gradually narrowed posteriad, each with distinct subacute angle protruded basad. Apical part of neck relatively deeply depressed, with short irregular and narrow longitudinal elevations. Distance between ocelli 1.7 times as long as distance between ocellus and posterior margin of eye. Apical maxillary palpomere about three times as long as penultimate segment. Antenna with distinctly elongate antennomeres 2 – 8 and slightly elongate 9 – 10; antennomere 3 slightly longer and narrower than 2, 4 distinctly shorter than 3, 5 longer than 4, 6 – 7 longer and distinctly broader than 5, 8 slightly shorter than 7, 9 – 10 slightly shorter and distinctly broader than 8, apical antennomere slightly longer than 10. Pronotum 1.3 times as broad as long, 1.1 times as broad as head, from widest anterior portion gradually narrowed posteriad toward widely rounded hind angles. Anterior angles acute and protruded anteriad. Anterior margin rounded, distinctly protruded anteriad. Each laterobasal portion not concave. Surface of disc with two wide and deep longitudinal and oval and deep anteriomedian depressions; surface between all depressions strongly elevated. Middle portion with irregular indistinct elevations. Elytra 1.1 times as broad as long, 1.8 times as long as pronotum, from basal portion distinctly broadened posteriad; hind margins straight. Dorsal surface of each elytron with irregular elevations, diagonal in medioapical and median, and longitudinal in apical portions. Metatarsi slightly less than twice as long as metatibia. Abdomen slightly narrower than elytra, with wide intersegmental membranes between abdominal tergites V – VI. Male. Posterior margin of abdominal tergite VIII straight. Posterior margin of abdominal sternite VIII sinuate. Aedeagus with wide basal portion, gradually narrowed toward median lobe; median lobe moderately long, from narrow basal part significantly broadened slightly in front middle, strongly narrowed in preapical part and broadened in apical portion, with widely rounded apex; accessory plates narrow, curved and relatively long, rounded apically; parameres wide and long, reaching apical part of median lobe, from widest preapical parts gradully narrowed apically, each with two moderately short apical and two preapical setae; internal sac narrow and long, without sclerotized structures (Fig. 10). Lateral aspect of the aedeagus as in Fig. 11; apical portion of median lobe (lateral view) with crenulate ventrolateral margin (Fig. 12). Female unknown.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF866C49FF0AFCE2FBFCFC39.taxon	discussion	Comparative notes. Omalium sectum sp. nov. differs from the remaining species of the acutangulum group by the shape of the pronotum with acute anterior angles protruded anteriad, and the shape of the median lobe.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF866C49FF0AFCE2FBFCFC39.taxon	distribution	Distribution. Omalium sectum sp. nov. is known from one locality in Meili Xue Shan and Diancang Shan mountain ranges in Yunnan, China (Fig. 30).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF866C49FF0AFCE2FBFCFC39.taxon	etymology	Etymology. The specific epithet is the Latin sectus, - a, - um (divided). It alludes to the shape of the median lobe.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF866C49FF0AFCE2FBFCFC39.taxon	biology_ecology	Bionomics. Specimens were collected at elevation 2580 m a. s. l. and was sifted from leaf litter, debris and mosses. It was collected together with O. rutilum sp. nov. and O. tibetanum sp. nov.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF846C4DFF0AFF7FFE66FD3D.taxon	description	(Figs 31, 34 – 36, 39)	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF846C4DFF0AFF7FFE66FD3D.taxon	materials_examined	Type material examined. Holotype ♂ (Fig. 31; dissected; left apical part of paramere missing): CHINA: SHAANXI: ‘ CHINA (S-Shaanxi) Qinling Shan | pass on r [oa] d Zhouzhi-Foping | 105 km SW Xi`an, N slope | 1990 m, 33 ° 44´N / 107 ° 59´E | (small creek vall [ey]. / mix [ed]. decid [uous]. | for [est]. / bamboo / small meadows) | 2. / 4. VII. 2001 Wrase [01] ’ <printed>, ‘ HOLOTYPE | Omalium | pratense sp. nov. | Shavrin A. V. des. 2025 ’ <red, printed> (cSch).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF846C4DFF0AFF7FFE66FD3D.taxon	description	Description. Measurements: HW: 0.50; HL: 0.35; OL: 0.17; TL: 0.03; AL: 0.85; PL: 0.41; PWmax: 0.62; PWmin: 0.42; ESL: 0.80; EW: 0.90; MTbL (holotype): 0.40; MTrL: 0.19 (MTrL 1 – 4: 0.07; MTrL 5: 0.12); AW: 0.82; AedL: 0.60; BL: 2.80. Habitus as in Fig. 31. Head, pronotum and abdomen dark-brown (laterobasal and basal margins of pronotum slightly and apical part of abdomen and paratergites distinctly paler); antennomeres 6 – 11 and elytra brown; mouthparts, antennomeres 1 – 5 and legs yellowish. Head with irregular and moderately large punctation, slightly denser in middle and on infraorbital portions; neck with sparse and fine punctation; punctation of pronotum irregular, larger and deeper than that in middle portion of head, finer and sparser in lateral and mediobasal portions; punctation of elytra about as that on pronotum, denser around scutellum, finer and sparser in middle; abdomen without visible punctation. Anterior portion of clypeus with fine transverse microreticulation, laterobasal portions of clypeus with fine diagonal meshes, portions between apical margins of eyes and clypeus with distinct elongate microsculpture; scutellum without meshes; abdominal tergites with dense transverse sculpture. Head 1.4 times as broad as long, with broad distinctly elevated clypeus and moderately wide and deep anteriomedian depressions, reaching level of apical margins of eyes; posteriolateral margins of clypeus relatively strongly narrowed posteriad and reaching level of anterior third of eyes. Latero-apical margin between anterior margin of eyes and clypeus with small semicircular notch. Dorsal surface with irregular transverse elevations between punctures in middle; postocular portions with two indistinct, narrow longitudinal wrinkles. Anteocellar foveae narrow, linear, reaching level of middle length of eyes. Temples very short, from posterior margins of eyes strongly narrowed toward neck. Apical part of neck without depression. Ocelli located slightly below level of posterior margins of eyes; distance between ocelli 1.4 times as long as distance between ocellus and posterior margin of eye. Apical maxillary palpomere about three and a half times as long as penultimate segment. Antenna with elongate antennomeres 4 – 7 and transverse 8 – 10; 3 distinctly narrower than 2, 4 distinctly shorter than 3, 5 broader than 4, 6 slightly longer and broader than 5, 7 indistinctly shorter and broader than 6, 8 broader than 7, 9 – 10 broader than 8, apical antennomere about twice as long as 10. Pronotum transverse, 1.5 times as broad as long, 1.2 times as broad as head, from widest middle distinctly more narrowed posteriad than anteriad. Anterior margin rounded, not protruded anteriad, indistinctly concave in middle. Laterobasal portions not concave in front of subacute hind angles. Laterobasal portions widely impressed and relatively deeply impressed. Surface of disc with two indistinct longitudinal depressions, broadened in mediobasal portion; medioapical portion without depression. Middle portion without longitudinal elevations between punctures; each latero-apical portion with distinct longitudinal elevations. Elytra 1.1 times as broad as long, 1.9 times as long as pronotum; hind margins straight. Dorsal surface of each elytron with irregular and moderately fine longitudinal elevations, more distinct in mediolateral and apical portions. Abdomen slightly narrower than elytra. Male. Posterior margin of abdominal tergite VIII truncate. Posterior margin of abdominal sternite VIII slightly sinuate. Aedeagus with wide and moderately short basal portion, slightly narrowed toward median lobe; median lobe elongate, from slightly broadened basal part indistinctly narrowed apically toward widely rounded apex; accessory plates relatively short and narrow; parameres moderately narrow, slightly narrowed apically, reaching preapical part of median lobe, each with three moderately short apical setae; internal sac wide and relatively short, with two small oval structures in basal part (Fig. 34). Lateral aspect of the aedeagus as in Fig. 35; apical portion of median lobe (lateral view) relatively wide, gradually narrowed toward subacute apex (Fig. 36). Female unknown.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF846C4DFF0AFF7FFE66FD3D.taxon	discussion	Comparative notes. Based on the general shapes of the body and the the aedeagus, O. pratensum sp. nov. is similar to the Palaearctic O. littorale Kraatz, 1857, widely distributed from Europe to Central Asia (Shavrin & Khachikov 2023; Zanetti 1987, 2002, 2006, 2011). It can be distinguished from it by the significantly shorter temples and slightly broader apical part of the median lobe.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF846C4DFF0AFF7FFE66FD3D.taxon	distribution	Distribution. Omalium pratense sp. nov. is known only from the type locality in Qinling Shan mountain range, Shaanxi, China (Fig. 39).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF846C4DFF0AFF7FFE66FD3D.taxon	etymology	Etymology. The specific epithet is the Latin adjective pratensis, - is, - e (living in meadows). It alludes to the holotype having been collected in a meadow.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF846C4DFF0AFF7FFE66FD3D.taxon	biology_ecology	Bionomics. The holotype was collected at elevation 1990 m a. s. l. by sifting in mixed deciduous forest with bamboo and small meadows.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF836C4DFF0AFCC6FEBEFB29.taxon	description	(Figs 37 – 38)	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF836C4DFF0AFCC6FEBEFB29.taxon	materials_examined	Material examined. CHINA: SICHUAN: 1 ♂: ‘ Chung King Szechwan China centralis’ (MHNG).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF836C4DFF0AFCC6FEBEFB29.taxon	discussion	Remarks. The Himalayan O. alticola is known from India (Kashmir, Uttarakhand) and western Nepal (Shavrin 2023 a). Aedeagus as in Figs 37 – 38 and in Figs 43 – 45 in Shavrin (2023 a). It is here recorded from China for the first time.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF836C51FF0AFA8AFB79FE79.taxon	description	(Figs 16 – 17, 32, 40 – 43, 47, 49)	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF836C51FF0AFA8AFB79FE79.taxon	materials_examined	Type material examined. Holotype ♂ (Fig. 32): CHINA: YUNNAN: ‘ CHINA: Yunnan, Dali | Cangshan ca 2500 m | 10 .. IV. 2003 in moss | G. de. Rougemont leg. ’ <printed>, ‘ Omalium | n. sp. | det. 2002 | G. de Rougemont [printed] ’ <handwritten>, ‘ HOLOTYPE | Omalium | cocleare sp. nov. | Shavrin A. V. des. 2025 ’ <printed> (OUMNH). Paratypes (16 ♂♂, 9 ♀♀): CHINA: YUNNAN: 2 ♂♂ (one specimen dissected), 1 ♀: ‘ CHINA: Yunnan, Dali | Cangshan ca 2500 m | 10. IV. 2003 stream moss | G. de. Rougemont leg. ’ <printed> (1 ♂: cSh; 1 ♂, 1 ♀: OUMNH); 1 ♂ (dissected; left paramere missing): ‘ P. R. CHINA, Yunnan, E | slope Cangshan at Dali, | N 25 ° 40´15.1 ´´ E 100 ° 07´| 39.9 ´´, 10. v. 2010, 2711 m, | sifting 04, V. Grebennikov’ <printed> (CNC); 2 ♂♂ (one specimen dissected), 5 ♀♀: (two specimens dissected): ‘ CHINA: Yunnan: Lijiang | Yushuizhai ca 2600 m | 14. IV. 2003 stream moss | G. de Rougemont leg. ’ <printed> (1 ♂, 3 ♀♀: cSh; 1 ♂, 2 ♀♀: OUMNH); 1 ♂: ‘ CHINA: Yunnan, Lijiang | Yushuizhai ca 2500 m | 14. IV. 2003 stream moss | G. de Rougemont leg. ’ <printed> (OUMNH); 1 ♂: ‘ CHINA X. 1986 | Yunnan: Kunming | G. de Rougemont’ <printed>, ‘ Omalium sp. [handwritten] | O. malaisei [handwritten] | det. 198 [printed] Scheerp? [handwritten] | G. de. Rougemont’ <printed> (OUMNH); 1 ♂: ‘ CHINA: Yunnan, Lijiang | Yushuizai ca 2500 m | 14. IV. 2003 stream moss | G. de Rougemont’ <printed> (OUMNH); 1 ♂ (dissected): ‘ CHINA: N-Yunnan [C 03 - 19 B] | Dali Bai Nat. Aut. Pref., Diancang Shan, | 3 km W Dali old town, | pine forest at “ Cloud Road ”, | right upper chairlift station, | 25 ° 41.1´N, 100 ° 06.8´E, 2650 – 2750 m, ’ <printed>, ‘ [C 03 - 19 B] pine needles, moss | (dry) in ditches, mushrooms, tra [?] | 1. IX. 2003, leg. M. Schülke’ <printed>, ‘ Museum für Naturkunde | Berlin | Sammlung M. Schülke’ (cSch); 1 ♂: ‘ CHINA: N-Yunnan [C 2005 - 11] | Dali Bai Nat. Aut. Pref., Diancang | Shan, 3 km W Dali old town, pine | forest at “ Cloud Road ”, 25 ° 41.1´N, | 100 ° 06.8´E, 2650 – 2750 m, | 17. VI. 2005, M. Schülke’ <printed> (cSch); 1 ♂ (dissected): ‘ CHINA (Yunnan) Dali Bai Aut. Pref., | mount range E Weishan, 12 km NE | Weishan, 2630 – 2660 m (scrub with | pines and bamboo, litter sifted) | 25 ° 17´02 - 15 ´´ N / 100 ° 22´23 - 30 ´´ E | 15. IX. 2009 D. W. Wrase [54 A] ’ <printed>, ‘ Museum für Naturkunde | Berlin | Sammlung M. Schülke’ <printed> (cSh); 1 ♀ (right antennomeres 3 – 11 missing): ‘ CHINA: N-Yunnan Diqing Tibet | Aut. Pr. Bitai Hai Lake area | 29 km ESE Zhongdian 27 ° 43.65´N | 99 ° 58.97´E 3540 m 1. VI. 2005 | A. Smetana [C 147] ’ <printed> (cSh); 1 ♀: (left antennomeres 7 – 11 missing): ‘ CHINA: N-Yunnan Diqing Tibet. | Aut. Pr. Zhongdian Co. 35 km ESE | Zhongdian 27 ° 41.00´N | 100 ° 01.47´E 3450 m | 3. VI. 2005 A. Smetana [C 150] ’ (CNC); SICHUAN: 1 ♂ (dissected): ‘ CHINA X. 1986 | Sichuan: Emei Shan | G. de Rougemont’ <printed>, ‘ Omalium sp. [handwritten] | det. 198 | G. de Rougemont’ <printed> (OUMNH); 1 ♂: ‘ China S Sichuan | S Xichang Mt. Luoji | 2300 – 2500 litter | 16 – 24.07. [19] 96 Kurbatov’ <printed> (MHNG); 1 ♀: ‘ CHINA: Sichuan, | 50 km W Dayi, Xiling | mts., 2300 m, 5. – 8. V. 2006 | S. Murzin & I. Shokhin’ <printed>, ‘ Museum für Naturkunde | Berlin | Sammlung M. Schülke’ <printed> (cSch); 1 ♂: ‘ CHINA: Yunnan | above Dali, 2000 – 2200 m | 4. – 17. IV. 1999 | leg. W. SCHAWALLER’ <printed> (SMNS); 2 ♂♂ (one specimen dissected): ‘ CHINA: Yunnan | above Dali, 2500 – 2700 m | 8. – 18. IV. 1999 | leg. W. SCHAWALLER’ <printed> (1 ♂: cSh; 1 ♂: SMNS). All paratypes with additional red printed label: ‘ PARATYPE | Omalium | cocleare sp. nov. | Shavrin A. V. des. 2025 ’.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF836C51FF0AFA8AFB79FE79.taxon	description	Description. Measurements (n = 26): HW: 0.49 – 0.52; HL: 0.36 – 0.40; OL: 0.13 – 0.15; TL: 0.07; AL (holotype): 1.10; PL: 0.39 – 0.46; PWmax: 0.64 – 0.70; PWmin: 0.55 – 0.61; ESL: 0.85 – 0.95; EW: 0.88 – 0.96; MTbL (holotype): 0.42; MTrL (holotype): 0.25 (MTrL 1 – 4: 0.08; MTrL 5: 0.17); AW: 0.88 – 0.94; AedL: 0.58 – 0.62; BL: 2.70 – 3.56 (holotype: 3.55). Habitus as in Fig. 32. Body brown to reddish-brown, with slightly darker head and abdomen (lateral and basal portions of abdomen, lateral and apical portions of elytra and paratergites of abdomen sometimes yellowish); antennomeres 6 – 11 or 7 – 11 brown; mouthparts, antennomeres 1 – 5 or 1 – 6 and legs yellowish. Punctation of head moderately dense, usually larger and deeper in middle, sometimes slightly finer in laterobasal portions of clypeus and on infraorbital portions, with interspaces between punctures in middle about as long as diameter of one-two nearest punctures; neck with dense and fine punctation, sparser in middle in several paratypes; pronotum with dense punctation about as that in middle part of head, distinctly sparser in mediobasal portion; punctation about as that on pronotum, sometimes slightly denser and coarser in basal portion and finer and sparser in middle, particularly along suture; abdomen with indistinct, fine and sparse or without visible punctation. Clypeus with distinct transverse microreticulation; scutellum with dense and fine isodiametric meshes; abdominal tergites with dense isodiametric microsculpture. Anterior and posterior margins of pronotum with indistinct short cuticular fringe. Head 1.3 times as broad as long, with strongly convex supra-antennal elevations and deep and relatively narrow anteriomedian depressions, reaching level of apical margins of eyes; posteriolateral margins of clypeus subdiagonally stretching posteriad and reaching level of anterior third of eyes. Latero-apical margin between anterior margin of eyes and clypeus with small semicircular notch. Dorsal surface with distinct diagonal elevations between punctures in basal portion of clypeus, with irregular elevations in middle and on postocular portions, sometimes forming indistinct four to six longitudinal wrinkles. Anteocellar foveae deep and moderately wide, suboval, indistinctly convergent latero-apicad toward level of posterior third or middle length of eyes. Temples 1.8 times to twice as long as longitudinal length of eye, from posterior margins of eyes gradually narrowed toward widely rounded hind angles. Apical part of neck sometimes narrowly depressed, with several short longitudinal elevations between punctures. Distance between ocelli about 1.8 times to twice as long as distance between ocellus and posterior margin of eye. Antenna with elongate antennomeres 5 – 7 and distinctly transverse 8 – 10; 4 short, about as long as broad, significantly shorter and slightly narrower than 3, 5 longer than 4, 6 slightly longer and broader than 5, 7 indistinctly broader than 6, 8 slightly shorter and distinctly broader than 7, 9 – 10 slightly broader than 8, apical antennomere about 1.3 times as long as 10. Pronotum 1.5 – 1.6 times as broad as long, 1.3 times as broad as head, widest in anterior third portion, distinctly more narrowed posteriad than anteriad. Apical angles not or slightly protruded anteriad. Lateral and laterobasal portions deeply and widely impressed. Surface of disc with two longitudinal depressions, significantly broadened basad; medioapical portion with indistinct or distinct and moderately shallow oval depression. Middle portions with indistinct or distinct irregular subdiagonal or longitudinal elevations between punctures. Elytra indistinctly broader than long, twice as long as pronotum. Middle and apical portions of each elytron with irregular diagonal elevations between punctures. Metatarsi slightly less than twice as long as metatibia. Male. Posterior margin of abdominal tergite VIII truncate. Posterior margin of abdominal sternite VIII sinuate. Aedeagus with moderately wide basal portion, slightly narrowed toward wide median lobe; median lobe spoon-shaped, from widest middle part gradually narrowed toward rounded apex; accessory plates narrow and relatively short; parameres moderately narrow, distinctly broadened in apical portions, with widely rounded apices, reaching preapical part of median lobe, each with two long and one short apical setae; internal sac wide and moderately short, with two elongate parallel and two shorter transverse sclerotized structures in basal portion (Figs 40, 43). Lateral aspect of the aedeagus as in Fig. 41; apical portion of median lobe (lateral view) wide, with truncate ventrolateral margin (Fig. 42). Female. Posterior margin of abdominal tergite VIII truncate. Posterior margin of abdominal sternite VIII rounded. Accessory sclerite with wide basal portion and median part gradually narrowed toward rounded apex (Fig. 16). Spermatheca as in Fig. 17.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF836C51FF0AFA8AFB79FE79.taxon	discussion	Comparative notes. Based on the length of the body, the presence of moderately strong longitudinal elevations on postocular portions of the head, the shape of the pronotum widest in apical third, and the general shape of the aedeagus, O. cocleare sp. nov. is similar to the Himalayan O. altivagans Bernhauer, 1915, known from Kashmir, India (Shavrin 2023 a) and O. latissimum Shavrin, 2023, described recently from Taiwan (Shavrin 2023 b). From O. altivagans it can be distinguished by the significantly denser punctation of the head and the pronotum, sligthly longer temples, slightly more protruded apical angles of the pronotum with the presence of medioapical depression, longer and narrower parameres, distinctly narrower median lobe without crenulation on lateral margins and the presence of elongate sclerotized structures in basal part of the internal sac. From O. latissimum it differs by the broader pronotum, longer elytra, narrower median lobe and sligthly longer and broader parameres. From both species it differs by the details of the external and internal morphology of the aedeagus, and different shape of the female accessory sclerite.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF836C51FF0AFA8AFB79FE79.taxon	distribution	Distribution. Omalium cocleare sp. nov. is known from several localities in Yunnan and Sichuan, China (Fig. 47).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF836C51FF0AFA8AFB79FE79.taxon	etymology	Etymology. The specific epithet is the Latin noun coclearium, - i (spoon). It alludes to the shape of the median lobe.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF836C51FF0AFA8AFB79FE79.taxon	biology_ecology	Bionomics. Specimens were collected at elevations from 2300 to 3540 m a. s. l. and were sifted from leaf litter, mosses and mushrooms. Some specimens were collected by sifting wet moss and various debris around small seepages in an old, devastated mixed forest (Abies, Larix, Betula, Rhododendron) (C 147) and in remnant of an original Abies, Betula, Rhododendron forest, treading of quite wet, mostly grassy vegetation in a seepage (C 150). The photograph of the locality of one paratype in Diancang Shan Mts. (C 03 - 19 B) as in Fig. 49.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF9F6C53FF0AFE7AFD8DFC71.taxon	description	(Figs 18 – 19, 33, 44 – 46, 48)	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF9F6C53FF0AFE7AFD8DFC71.taxon	materials_examined	Type material examined. Holotype ♂ (Fig. 33): CHINA: SICHUAN: ‘ CHINA, S-Sichuan, road | 56 km E Xichang- Zhaojue, | Mt. WenLin pass, 3200 m, | ~ 27 ° 52´N, 102 ° 30´E, | 5. – 20. VI. 2017, leg. Reuter’ <printed>, ‘ HOLOTYPE | Omalium | contiger sp. nov. | Shavrin A. V. des. 2025 ’ <red, printed> (cF, to be eventually deposited in Naturkundemuseum Erfurt, Germany). Paratypes (15 ♂♂, 11 ♀♀): CHINA: SICHUAN: 2 ♂♂ (dissected; one specimen without right elytron), 1 ♀ (dissected; left elytron missing): same data as the holotype, with additional printed label: ‘ Omalium | sp. | det. Zanetti 2018 ’ (1 ♂: cSh; 1 ♂, 1 ♀: cF); 3 ♂♂ (two specimens dissected): ‘ CHINA, S-Sichuan, | S Yanyuan, Chuandongzi | massif, ~ 27 ° 21`05 `` N, | 101 ° 30´26 ´´ E, 3200 m | 8. – 19. VI. 2017, leg. Reuter’ <printed> (1 ♂: cSh; 2 ♂♂: cF); 4 ♂♂, 3 ♀♀: ‘ CHINA, S-Sichuan, | S Yanyuan, Chuandongzi- | Massif, 27 ° 20`22 `` N, | 101 ° 32´43 ´´ E, 4100 m | 9. – 19. VI. 2017, leg. Reuter’ <printed> ’, ‘ Omalium | sp. | det. Zanetti 2018 ’ <printed> (2 ♂♂, 1 ♀: cSh; 2 ♂♂, 2 ♀♀: cF); YUNNAN: 2 ♂♂ (one specimen dissected), 4 ♀♀: ‘ CHINA: Yunnan | above Dali, 2500 – 2700 m | 8. – 18. IV. 1999 | leg. W. SCHAWALLER’ <printed> (1 ♂, 2 ♀♀: cSh; 1 ♂♂, 2 ♀♀: SMNS); 1 ♂ (dissected): ‘ CHINA: Yunnan, Dali | Cangshan ca 2500 m | 10. IV. 2003 in moss | G. de Rougemont leg. ’ (OUMNH); 3 ♂♂ (two specimens dissected), 3 ♀: ‘ CHINA: Yunnan, Dali | Cangshan ca 2500 m | 10. IV. 2003 stream moss | G. de Rougemont leg. ’ (2 ♂♂, 2 ♀♀: cSh; 1 ♂, 1 ♀: OUMNH). All paratypes with additional red printed label: ‘ PARATYPE | Omalium | contiger sp. nov. | Shavrin A. V. des. 2025 ’.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF9F6C53FF0AFE7AFD8DFC71.taxon	description	Description. Measurements (n = 27): HW: 0.47 – 0.60; HL: 0.35 – 0.41; OL: 0.12 – 0.15; TL: 0.08 – 0.10; AL (holotype): 1.02; PL: 0.40 – 0.51; PWmax: 0.60 – 0.67; PWmin: 0.55 – 0.60; ESL: 0.85 – 0.89; EW: 0.90 – 0.94; MTbL (holotype): 0.38; MTrL (holotype): 0.25 (MTrL 1 – 4: 0.08; MTrL 5: 0.17); AW: 0.85 – 0.89; AedL: 0.67 – 0.77; BL: 2.42 – 3.70 (holotype: 3.00). Habitus as in Fig. 33. Body brown to reddish-brown, usually with slightly darker head and abdomen (lateral parts of pronotum and lateroapical and lateral portions of elytra sometimes slightly paler); antennomeres 6 – 11 brown; mouthparts, antennomeres 1 – 5 and legs yellow-brown; tarsi yellowish. Punctation of head irregular and variable: fine and significantly sparser in middle or distinctly denser, larger and deeper in middle and on infraorbital portions; neck with sparse and fine punctation; punctation of pronotum dense, about as that in middle portion of head or sparser and finer, finer and sparser in middle and / or mediobasal portion, distinctly sparser in lateral parts; punctation of elytra dense, larger and coarser than that on pronotum; abdominal tergites without or with sparse, fine and sometimes indistinct punctation. Apical part of clypeus with distinct fine microsculpture: transverse in apical and in middle, and diagonal on supra-antennal elevations; scutellum with fine isodiametric microreticulation. Anterior and posterior margins of pronotum with row of cuticular setae, sometimes slightly longer on apical margin. Head 1.3 – 1.4 times as broad as long, with broad clypeus usually strongly narrowed posteriad toward level of middle or posterior third of eyes, with deep and wide anteriomedian depressions, reaching anterior third of eyes. Median part and infraorbital portions usually strongly elevated. Latero-apical margin between anterior margin of eyes and clypeus with small semicircular notch. Dorsal surface with irregular elevations between punctures: longitudinal or diagonal in laterobasal portions of clypeus, transverse or diagonal in middle and on infraorbital portions; irregular elevations on postocular portions sometimes forming indisinct three to four longitudinal wrinkles. Anteocellar foveae wide and deep, distinctly convergent latero-apicad toward level of middle length of eyes, sometimes connected with lateral margins of clypeus. Temples 1.5 times as long as longitudinal length of eye, from posterior margins of eyes gradually narrowed toward neck, sometimes with widely rounded hind angles. Apical part of neck narrowly and moderately deeply depressed, with several short and narrow elevations; middle portion sometimes distinctly elevated. Distance between ocelli 1.5 – 1.7 times as long as distance between ocellus and posterior margin of eyes. Apical maxillary palpomere two and a half to three times as long as penultimate segment. Antenna with elongate antennomeres 4 – 6, slightly transverse 7 – 8 and distinctly transverse 9 – 10; 4 about as long as broad, significantly shorter than 3, 5 longer and slightly broader than 4, 6 slightly broader than 5, 7 – 8 slightly longer and broader than 6, 9 – 10 distinctly broader than 8. Pronotum 1.3 – 1.5 times as broad as long, widest in preapical portion, widely rounded apicad and distinctly narrowed posteriad. Apical angles not or slightly protruded anteriad. Anterior margin slightly rounded, not protruded anteriad, sometimes slightly concave in middle. Mediolateral and laterobasal margins widely and deeply impressed. Surface of disc with two long and wide longitudinal depressions distinctly broadened in mediobasal portion; medioapical portion without or with narrow depression; surface between all pronotal depressions narrowly elevated; each lateroapical portion with additional narrow and elongate elevation, from apical part reaching about middle. Middle portion without or with irregular and fine longitudinal and transverse elevations between punctures, more visible in apical and lateral portions. Elytra 1.7 times to twice as long as pronotum, somewhat subparallel or slightly broadened posteriad, some specimens with indistinctly concave lateral margins in about middle. Dorsal surface of each elytron with irregular transverse and subdiagonal elevations between pucntures, usually coarser in mediolateral portions of each elytron. Metatarsi about 1.5 times as long as metatibia. Abdomen slightly narrower than elytra, with two transverse wing-folding patches in middle of abdominal tergite IV and two smaller oval patches in middle of tergite V, invisible in some specimens; abdominal tergites III – VI sometimes with narrow or wide intersegmental membranes. Male. Posterior margin of abdominal tergite VII truncate or rounded. Posterior margin of abdominal sternite VIII widely and slightly sinuate. Aedeagus with wide basal portion, slightly narrowed toward median lobe; median lobe narrow, strongly elongate, from middle slightly broadened apicad and from preapical portion slightly narrowed toward obtuse apex; accessory plates narrow, short, with bilobed apical part; parameres narrow, reaching middle part of median lobe, each with three long apical setae; internal sac wide and moderately long, with two small suboval sclerotized structures in basal portion (Fig. 44). Lateral aspect of the aedeagus as in Fig. 45; apical portion of median lobe (lateral view) elongate, with rounded apex, slightly curved ventrad, with distinctly broadened dorsoventral portion (Fig. 46). Female. Posterior margin of abdominal tergite VIII truncate. Posterior margin of abdominal sternite VIII rounded. Accessory sclerite with moderately wide basal portion and elongate narrow median part, with subacute apex (Fig. 18). Spermatheca as in Fig. 19.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF9F6C53FF0AFE7AFD8DFC71.taxon	discussion	Comparative notes. Based on the body length, proportions of the antennomeres, the shape of the pronotum widest in apical third and the presence of elongate median lobe, O. contiger sp. nov. is similar to the Himalayan O. scapulare Coiffait, 1982, known from India and Nepal (Shavrin 2023 a). It can be distinguished from it by the finer longitudinal wrinkles on infraorbital portions, more visible on postocular parts, slightly more transverse pronotum, different shape of the female accessory sclerite, narrower median lobe, narrower and shorter parameres and other details of the morphology of the aedeagus.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF9F6C53FF0AFE7AFD8DFC71.taxon	distribution	Distribution. Omalium contiger sp. nov. is known from several localities in Sichuan and Yunnan, China (Fig. 48).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF9F6C53FF0AFE7AFD8DFC71.taxon	etymology	Etymology. The specific epithet is the Latin noun contiger (spear-bearer). It alludes to the shape of the median lobe.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF9F6C53FF0AFE7AFD8DFC71.taxon	biology_ecology	Bionomics. Specimens were collected at elevations from 2500 to 4100 m a. s. l. Specimens from Yunnan were sifted from wet mosses near streams.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF9A6C55FF0AFF37FA2EF9DD.taxon	description	(Figs 20, 39, 50, 53 – 55)	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF9A6C55FF0AFF37FA2EF9DD.taxon	materials_examined	Type material examined. Holotype ♂ (dissected): CHINA: ANHUI: ‘ China: S Anhui, | Zinhuashan. 700 – 1200 m. | 16 – 19.05.1998. | S. A. Kurbatov leg. ’ <printed>, ‘ HOLOTYPE | Omalium | kurbatovi sp. nov. | Shavrin A. V. des. 2025 ’ <red, printed> (ZMM). Paratypes: 3 ♀♀ (one specimen dissected): same data as the holotype, with additional red printed label: ‘ PARATYPE | Omalium | kurbatovi sp. nov. | Shavrin A. V. des. 2025 ’ (1 ♀: cSh; 2 ♀♀: ZMM).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF9A6C55FF0AFF37FA2EF9DD.taxon	description	Description. Measurements (n = 4): HW: 0.62 – 0.65; HL: 0.37 – 0.40; OL: 0.17 – 0.20; TL: 0.07; AL (holotype): 1.19; PL: 0.47 – 0.57; PWmax: 0.83 – 0.84; PWmin: 0.72 – 0.76; ESL: 0.97 – 1.10; EW: 1.12 – 1.16; MTbL (holotype): 0.60; MTrL (holotype): 0.26 (MTrL 1 – 4: 0.11; MTrL 5: 0.15); AW: 1.12 – 1.20; AedL: 0.80; BL: 3.47 – 4.35 (holotype). Habitus as in Fig. 50. Body reddish-brown, with darker head and middle of pronotum; antennomeres 7 – 11 brown; mouthparts, antennomeres 1 – 6, lateral and basal portions of pronotum, legs, paratergites and intersegmental membranes of abdomen yellow-brown; tarsi yellow. Punctation of head regular, moderately fine and dense, slightly sparser in middle and denser on infraorbital portion; punctation of neck about as that in middle part of head; punctation of pronotum about as that in middle part of head, slightly denser and larger in apical, distinctly sparser in mediobasal and finer and significantly sparser in lateral portions; elytra with fine and sparse punctation. Anterior part of clypeus with distinct transverse microreticulation; basal portion of neck with fine transverse meshes; clypeus with fine isodiametric microsculpture; abdominal tergites with dense isodiametric sculpture. Anterior and posterior margins of pronotum and apical margins of elytra with row of short cuticular setae. Head 1.6 times as broad as long, with narrow and relatively deep anteriomedian depressions reaching level of anterior margins of eyes. Dorsal surface with elongate wrinkles between anterior margins of eyes and clypeus, and irregular transverse and longitudinal elevations on infraorbital portions. Anteocellar foveae deep and moderately wide, suboval, almost reaching level of middle length of eyes. Temples slightly more than twice shorter than longitudinal length of eyes, from posterior margins of eyes slightly narrowed posteriad, each with widely rounded hind angles. Apical part of neck deeply depressed, with row of short and narrow elevations. Ocelli 1.6 times as long as distance between ocellus and posterior margin of eye. Apical maxillary palpomere about three times as long as penultimate segment. Antenna with slightly transverse antennomere 7 and distinctly transverse 8 – 10; antennomere 5 slightly longer than 4, 6 indistinctly broader than 5, 7 slightly broader than 6, 8 broader than 7, 9 – 10 slightly shorter and distinctly broader than 8, apical antennomere about twice as long as 10. Pronotum 1.4 – 1.7 times as broad as long, 1.2 – 1.3 times as broad as head, from widest middle slightly more narrowed posteriad than anteriad. Anterior angles widely rounded, slightly protruded anteriad. Anterior margin rounded, slightly concave in middle. Laterobasal portions not concave in front of obtuse angles. Surface of disc with two long and wide, shallow or moderately deep longitudinal depressions, slightly broadened basad. Middle portion without elevations between punctures. Elytra about twice as long as pronotum; hind margins rounded. Dorsal surface of each elytron with irregular elevations between punctures: transverse around scutellum, and diagonal and longitudinal in mediolateral and lateral portions. Male. Posterior margin of abdominal tergite VIII truncate. Posterior margins of abdominal sternite VIII widely rounded. Aedeagus with wide basal portion, gradually narrowed toward median lobe; median lobe moderately short and wide, from widest preapical part slightly narrowed toward widely rounded apex; accessory plates narrow and short; parameres relatively narrow, distinctly broadened in preapical portions and narrowed apically, reaching preapical part of median lobe, each with two moderately long and two short apical setae; internal sac long and relatively narrow, with two small oval sclerotized sctructures in prebasal portion (Fig. 53). Lateral aspect of the aedeagus as in Fig. 54; apical portion of median lobe (lateral view) hook-shaped, with rounded apex and strongly curved large ventrolateral tooth in preapical part (Fig. 55). Female. Posterior margin of abdominal tergite VIII truncate. Posterior margin of abdominal sternite VIII rounded. Accessory sclerite leaf-shaped, from basal portion gradually narrowed toward rounded apex (Fig. 20). Spermatheca not recognized.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF9A6C55FF0AFF37FA2EF9DD.taxon	discussion	Comparative notes. Omalium kurbatovi sp. nov. differs from the remaining species of the oxyacanthae group by the larger body and the morphology of the aedeagus. Based on the presence of large hook in median lobe (lateral view, Fig. 55), it is similar to the Himalayan O. marginatum Cameron, 1941, known from Pakistan, India (Kashmir) and Nepal (Shavrin 2023 a), from which it can be easily distinguished by the more transverse antennomeres 8 – 10, longer pronotum and elytra, significantly narrower median lobe, broader parameres and other details of the morphology of the aedeagus.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF9A6C55FF0AFF37FA2EF9DD.taxon	distribution	Distribution. Omalium kurbatovi sp. nov. is known only from the type locality in Huangshan mountain range in Anhui, China (Fig. 39).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF9A6C55FF0AFF37FA2EF9DD.taxon	etymology	Etymology. Patronymic; the species is named to honor Sergey A. Kurbatov (Moscow), expert in Pselaphinae and collector of the type material.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF9A6C55FF0AFF37FA2EF9DD.taxon	biology_ecology	Bionomics. Omalium kurbatovi sp. nov. was collected at elevations from 700 to 1200 m a. s. l. by sifting litter.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF9B6C55FF0AF9DEFF5BF81B.taxon	discussion	Remarks. Omalium oxyacanthae is a species widespread in the Holarctic region (Herman 2001). Schülke & Smetana (2015) recorded it from Helongjiang (China) without providing of the exact locality. Morphological features for this species, including figures of the aedeagus and the female accessory sclerite, can be found in Zanetti (1985, 1987, 2012).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF986C59FF0AF959FAB1FF09.taxon	description	(Figs 21, 48, 51, 56 – 58, 62)	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF986C59FF0AF959FAB1FF09.taxon	materials_examined	Type material examined. Holotype ♂: CHINA: GANSU: ‘ CHINA, Gansu, Xinlong | Shan, cca 70 km S | Lanzhou, 2225 - | 2380 m, 7. VIII. 1994 | A. Smetana [C 32] ’ <printed>, ‘ HOLOTYPE | Omalium | schuelkei sp. nov. | Shavrin A. V. des. 2025 ’ <red, printed> (CNC). Paratypes (13 ♂♂, 15 ♀♀): CHINA: GANSU: 5 ♂♂ (one specimen dissected), 4 ♀♀ (one specimen dissected): same data as that in the holotype (2 ♂♂, 2 ♀♀: cSh; 3 ♂♂, 3 ♀♀: CNC); 7 ♂♂ (two specimens dissected), 7 ♀♀: ‘ CHINA: Gansu Prov. [CH 11 - 27] | Lenglong Ling Mts., 60 km NNW | Honggu, Jin Sha Gorge, mixed forest | (Picea, Populus, Betula) 36 ° 51´56.9 ´´ N, | 102 ° 38´55.3 ´´ E, litter, moss, mushrooms | sifted, 12. VII. 2011, leg. M. Schülke’ <printed>, ‘ Museum für Naturkunde | Berlin | Sammlung M. Schülke’ <printed> (3 ♂♂, 2 ♀♀: cSh; 4 ♂♂, 6 ♀♀: cSch); 1 ♀: ‘ CHINA: Gansu province, | XIAHE (= Labrang) env., | 35 ° 11.3 ´´ N 102 ° 30.6´E, | 3043 m (GPS), 19. – 22. VI. | 2005, J. Hájek, D. Král & | J. Růžička leg. [Ch 2] ’ <printed>, ‘ baited pitfall traps (fish | meat + cheese) with | ethylene glycol. | margin of a wet | coniferous forest ’ <printed>, ‘ Museum für Naturkunde | Berlin | Sammlung M. Schülke’ <printed> (cSch); SICHUAN: 1 ♂ (dissected), 2 ♀♀: ‘ CHINA: N-Sichuan [CH 12 - 21] | 49 km N Songpan, road S 301 km 114, | N Gongangling pass, 33 ° 04´31 ´´ N, | 103 ° 42´38 ´´ E, 3230 m, spruce forest, | litter, moss & mushrooms sifted, | 9. VIII. 2012, M. Schülke’ <printed>, ‘ Museum für Naturkunde | Berlin | Sammlung M. Schülke’ <printed> (cSch); 1 ♀: ‘ CHINA: W-Sichuan 1999 | Ganzi Tibet Aut. Pref., Luding Co. | W Erlangshan-Pass, 2600 m | 7 km SSE Luding, 29 ° 51 N | 102 ° 15 E, Nadelstreu, Pilze | 29. VI., leg. M. Schülke’ <printed>, ‘ Sammlung | M. Schülke | Berlin’ <printed>, ‘ Omalium [handwritten] | sp. [handwritten] | det. M. Schülke 2002 ’ <printed> (cSch). All paratypes with additional red printed label: ‘ PARATYPE | Omalium | schuelkei sp. nov. | Shavrin A. V. des. 2025 ’.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF986C59FF0AF959FAB1FF09.taxon	description	Description. Measurements (n = 29): HW: 0.55 – 0.60; HL: 0.36 – 0.38; OL: 0.15 – 0.16; TL: 0.05 – 0.07; AL (holotype): 0.87; PL: 0.46 – 0.52; PWmax: 0.72 – 0.80; PWmin: 0.65 – 0.70; ESL: 0.85 – 0.97; EW: 0.95 – 1.05; MTbL (holotype): 0.47; MTrL (holotype): 0.27 (MTrL 1 – 4: 0.10; MTrL 5: 0.17); AW: 1.02 – 1.06; AedL: 0.60 – 0.62; BL: 2.65 – 3.50 (holotype: 3.25). Habitus as in Fig. 51. Body reddish-brown, with darker head, pronotum and abdomen (lateral and basal portions of pronotum, elytra, paratergites and intersegmental membranes of abdomen in some paratypes distinctly paler); antennomeres 6 – 11 or 7 – 11 brown; mouthparts, antennomeres 1 – 5 or 1 – 6 and legs yellow-brown; tarsi yellow. Punctation of head dense or sparse, fine, usually denser in middle, denser and coarser on infraorbital portions; punctation of neck dense, about as that in middle of head; punctation of pronotum moderately dense, usually finer and sparser in medioapical, lateral and mediobasal portions and denser in middle; abdominal tergites with indisinct, fine and sparse punctation. Anterior portion of clypeus with distinct transverse microsculpture, basal portions of clypeus with fine transverse and longitudinal meshes, middle portion sometimes with longitudinal meshes between anteocellar foveae, posterior parts of infraorbital portions usually with distinct longitudinal and diagonal microreticulation; middle part of neck without or with fine transverse or isodiametric meshes; scutellum with fine isodiametric sculpture; abdominal tergites with dense isodiametric microsculpture. Anterior and posterior margins of pronotum with row of short cuticular fringe, invisible in some specimens. Head 1.5 times as broad as long, with wide and moderately deep anteriomedian depressions, reaching level of anterior third of eyes; posteriolateral margins of clypeus subparallel, stretching posteriad toward level about middle length of eyes. Latero-apical margin between anterior margin of eyes and clypeus with distinct semicircular notch. Dorsal surface sometimes with fine irregular transverse elevations between punctures in middle, infraorbital portions with irregular elongate wrinkles in postocular portions. Anteocellar foveae suboval and deep, slightly convergent latero-apicad toward level of posterior third or middle length of eyes. Temples very short, from two to three times shorter than longitudinal length of eyes. Apical part of neck sometimes distinctly and narrowly depressed, with irregular short elevations between punctures. Ocelli located slightly below level of posterior margins of eyes; distance between ocelli 1.4 – 1.6 times as long as distance between ocellus and posterior margin of eye. Apical maxillary palpomere about three times as long as penultimate segment. Antenna with slightly transverse antennomeres 6 – 7 and distinctly transverse 8 – 10; 5 indistinctly longer and broader than 4, 6 slightly broader than 5, 7 broader than 6, 8 slightly broader than 7, 9 – 10 slightly longer and distinctly broader than 8, apical antennomere about twice as long as penultimate antennomere. Pronotum 1.5 times as broad as long, 1.3 times as broad as head, widest in or slightly in front of middle, distinctly more narrowed posteriad than anteriad. Apical angles widely rounded, not protruded anteriad. Anterior margin somewaht straight or rounded, not protruded anteriad, sometimes indistinctly concave in middle. Surface of disc with two long, deep and moderately wide depressions. Each mediolateral portion sometimes with distinct narrow elongate elevations, slightly curved posteriad and reaching middle length of pronotum. Middle portion without elevations between punctures. Elytra 1.1 times as broad as long, 1.8 times as long as pronotum. Mediolateral surface of each elytron with irregular longitudinal and diagonal elevations. Metatarsi slightly less than twice shorter than metatibia. Abdomen slightly broader than elytra. Male. Posterior margin of abdominal tergite VIII truncate. Posterior margin of abdominal sternite VIII widely sinuate. Aedeagus with wide basal portion, gradually narrowed toward median lobe; median lobe wide and moderately long, from widest basal part gradually narrowed toward small rounded apex; accessory plates short and relatively narrow, each with bilobed apex; parameres relatively narrow, distinctly broadened in preapical parts, reaching about middle of median lobe, each with two moderately long and two short apical setae; internal sac long and moderately narrow, with two small elongate sclerotized structures in basal part (Fig. 56). Lateral aspect of the aedeagus as in Fig. 57; apical portion of median lobe (lateral view) with distinctly truncate ventrolateral margin (Fig. 58). Female. Posterior margin of abdominal tergite VIII truncate. Posterior margin of abdominal sternite VIII truncate or rounded. Accessory sclerite with wide basal part, gradually anrrowed toward broadened and elongate median part with rounded apex (Fig. 21). Spermatheca not recognized.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF986C59FF0AF959FAB1FF09.taxon	discussion	Comparative notes. Based on the general shape of the body and the aedeagus, the shape of the elliptic anteocellar foveae, O. schuelkei sp. nov. is similar to the Holarctic O. oxyacanthae. The new species can be distinguished from it by the larger body (the body of O. oxyacanthae varies from 2.2 to 2.6 mm), denser and coarser punctation of the pronotum and the elytra, broader and shorter median lobe, slightly broader parameres, and different shape of the female accessory sclerite.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF986C59FF0AF959FAB1FF09.taxon	distribution	Distribution. Omalium schuelkei sp. nov. is known from several localities from Gansu and Sichuan, China (Fig. 48).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF986C59FF0AF959FAB1FF09.taxon	etymology	Etymology. Patronymic; the species is named to honour Michael Schülke (Berlin), one of the collectors of the type specimens.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF986C59FF0AF959FAB1FF09.taxon	biology_ecology	Bionomics. Specimens were collected at elevations from 2225 to 3230 m a. s. l. and were taken by sifting litter, mosses and mushrooms in mixed (Picea, Populus, Betula) and coniferous forests. One specimen from Xiahe was collected using pitfall traps. The photograph of the type localities in Sichuan as in Fig. 62 (CH 12 - 21).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF976C5AFF0AFE32FF5EFBA9.taxon	description	(Figs 22 – 23, 48, 52, 59 – 61)	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF976C5AFF0AFE32FF5EFBA9.taxon	materials_examined	Type material examined. Holotype ♂ (dissected): CHINA: JIANGXI: ‘ CHINA: Jiangxi prov. [MF 10] | Jinggangshan Mts., Songmuping | 26 ° 34.7´N, 114 ° 04.3´E, 1280 m | (stream valley), 27. iv. 2011 | Fikáček, Hájek, Jia & Song’ <printed>, ‘ cut and decaying tops of | bamboo trunks in a sparse | bamboo bush’ <printed>, ‘ HOLOTYPE | Omalium | bambusaphilum sp. nov. | Shavrin A. V. des. 2025 ’ <red, printed> (NMPC). Paratypes: CHINA: JIANGXI: 5 ♀♀ (one specimen dissected): same data as the holotype, with additional red printed label: ‘ PARATYPE | Omalium | bambusaphilum sp. nov. | Shavrin A. V. des. 2025 ’ <red, printed> (2 ♀♀: cSh; 3 ♀♀: NMPC).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF976C5AFF0AFE32FF5EFBA9.taxon	description	Description. Measurements (n = 6): HW: 0.47 – 0.51; HL: 0.28 – 0.30; OL: 0.14 – 0.15; TL: 0.04 – 0.05; AL (holotype): 0.70; PL: 0.34 – 0.36; PWmax: 0.55 – 0.63; PWmin: 0.49 – 0.59; ESL: 0.64 – 0.79; EW: 0.73 – 0.81; MTbL (holotype): 0.27; MTrL (holotype): 0.19 (MTrL 1 – 4: 0.07; MTrL 5: 0.12); AW: 0.72 – 0.85; AedL: 0.55; BL: 2.45 (holotype) – 3.10. Habitus as in Fig. 52. Head, pronotum and abdomen reddish-brown; lateral and basal portions of pronotum, elytra, paratergites (two paratypes) and intersegmental membranes of abdomen yellow-brown (parascutellar portion of elytra slightly darkened); antennomeres 6 – 11 brown; mouthparts, antennomeres 1 – 5 and legs yellowish. Punctation of head irregular, moderately fine and dense, denser in middle, infraorbital portions without punctures except of several fine punctures close to anteocellar foveae; neck with irregular moderately dense and fine punctures in mediobasal part; punctation of pronotum dense, distinctly larger and deeper than that in middle portion of head, with interspaces between pucntures in middle about as long as diameters of one to three nearest punctures, sparser and finer in lateral and mediobasal portion (one paratype with sparse and fine punctation in middle); punctation of elytra dense; abdominal tergites without visible punctation. Head with dense microsculpture: transverse in apical and diagonal in lateral portions of clypeus, transverse between level of anterior third of eyes, longitudinal and diagonal in middle, and longitudinal on infraorbital portions (two paratypes without visible sculpture in middle); neck without meshes; pronotum with indistinct fine isodiametric microreticulation; pronotum with dense isodiametric sculpture. Head 1.6 – 1.7 times as broad as long, with moderately wide and deep anteriomedian depressions, almost reaching level of anterior third of eyes; posteriolateral parts of clypeus slightly narrowed posteriad, reaching level of anterior third of eyes. Latero-apical margin between anterior margin of eyes and clypeus with small semicircular notch. Infraorbital portions with elongate longitudinal wrinkles. Anteocellar foveae somewhat suboval, relatively deep, indistinctly convergent latero-apicad toward level of middle length of eyes. Temples short, about or more than three times shorter than longitudinal length of eyes, from posterior margin strongly narrowed toward neck; postocular ridges present, obtuse, with distance between posterior margin of eye and ridge about as long as one-two nearest ommatidia. Apical part of neck narrowly depressed in apical part, with irregular short and narrow elevations. Ocelli located slightly below level of postocular ridges; distance between ocelli slightly shorter than distance between ocellus and posterior margin of eye. Apical maxillary palpomere about three times as long as penultimate segment. Antenna with slightly transverse antennomeres 7 – 8 and distinctly transverse 9 – 10; 4 – 5 small, about as long as wide, 6 slightly longer and broader than 5, 7 slightly broader than 6, 8 slightly longer and broader than 7, 9 – 10 distinctly broader than 8, apical antennomere slightly less than twice as long as 10. Pronotum 1.6 – 1.7 times as broad as long, 1.1 – 1.2 times as broad as head, from widest middle slightly more narrowed posteriad than anteriad. Apical angles widely rounded, not protruded anteriad. Anterior margin rounded, slightly concave in middle. Laterobasal portions not concave in front of obtuse hind angles. Lateral potions relatively deeply and widely impressed, deeper in middle and broader in laterobasal portions. Surface of disc with two moderately deep longitudinal and narrow, sometimes indistinct, medioapical depressions; surface between all pronotal depressions narrowly elevated. Middle part without elevations between punctures. Elytra 1.1 times broader than long, 1.8 times to twice as long as pronotum. Mediolateral surface of each elytron with irregular longitudinal and diagonal elevations between punctures. Metatarsi 1.4 times as long as metatibia. Abdomen as wide as elytra or slightly broader, with wide intersegmental membranes between abdominal tergites III – V or III – VI. Male. Posterior margin of abdominal tergite VIII slightly and sternite VIII widely and moderately deeply sinuate. Aedeagus with very wide basal portion, gradually narrowed toward median lobe; median lobe narrow, elongate, from basal part slightly broadened toward middle and from middle slightly narrowed toward small rounded apex; accessory plates short, narrow, distinctly curved and rounded apically; internal sac wide and relatively long (Fig. 59). Lateral aspect of the aedeagus as in Fig. 60; apical portion of median lobe (lateral view) hook-shaped, with slightly curved ventrodorsad subacute hind angle and rounded apex (Fig. 61). Female. Posterior margin of abdominal tergite VIII truncate. Posterior margin of abdominal sternite VIII rounded. Accessory sclerite short, triangular, from widest basal portion strongly narrowed toward acute apex (Fig. 22). Spermatheca as in Fig. 23.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF976C5AFF0AFE32FF5EFBA9.taxon	discussion	Comparative notes. Based on the general shape of the body and the aedeagus, the presence of the hook-shaped apical portion of the aedeagus (lateral view), the coloration of the body, with distinctly paler elytra, the short temples and the pronotum widest in middle, and the similar shape of the female accessory sclerite, O. bambusaphilum sp. nov. is similar to the Himalayan O. cariosum Cameron, 1924, known from India (Shavrin 2023 a). The new species can be distinguished from it by the sparser and slightly larger punctation of the slightly more transverse pronotum, slightly narrower elytra, narrower and longer median lobe, the shape of hook-shaped apical part of the median lobe (lateral view), and broader apical parts of the parameres.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF976C5AFF0AFE32FF5EFBA9.taxon	distribution	Distribution. Omalium bambusaphilum sp. nov. is known only from the type locality in Jinggangshan mountain range in Jiangxi, China (Fig. 48).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF976C5AFF0AFE32FF5EFBA9.taxon	etymology	Etymology. The name of the new species is derived from the fact that the type specimens were collected in “ bamboo bush ” (genus Bambusa and related).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF976C5AFF0AFE32FF5EFBA9.taxon	biology_ecology	Bionomics. Specimens were collected at elevation 1280 m a. s. l. by sifting of cut and decaying tops of bamboo trunks.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF946C5CFF0AFB0AFA3AFCC9.taxon	description	(Figs 24 – 25, 39, 63, 66 – 68)	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF946C5CFF0AFB0AFA3AFCC9.taxon	materials_examined	Type material examined. Holotype ♂ (dissected): CHINA: SICHUAN: ‘ CHINA, Sichuan, | NW Heishui, 3428 m | 32 ° 11´29 ´´ N, 102 ° 39´30 ´´ E | 14 – 20. VI. 2018, leg. Reuter’ <printed>, ‘ HOLOTYPE | Omalium | confertum sp. nov. | Shavrin A. V. des. 2025 ’ <red, printed> (cF, to be eventually deposited in Naturkundemuseum Erfurt, Germany). Paratypes: 1 ♀ (Fig. 63; dissected): same data as the holotype, with additional printed label: ‘ HOLOTYPE | Omalium | confertum sp. nov. | Shavrin A. V. des. 2025 ’ <red, printed> (cF).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF946C5CFF0AFB0AFA3AFCC9.taxon	description	Description. Measurements (n = 2): HW: 0.52 – 0.57; HL: 0.42 – 0.43; OL: 0.15; TL: 0.10; AL (holotype): 1.23; PL: 0.45; PWmax: 0.70; PWmin: 0.63 – 0.64; ESL: 0.78 – 0.85; EW: 0.86 – 0.87; MTbL (holotype): 0.51; MTrL (holotype): 0.22 (MTrL 1 – 4: 0.10; MTrL 5: 0.12); AW: 0.87 – 0.90; AedL: 0.55; BL: 2.90 – 2.95 (holotype). Habitus as in Fig. 63. Body dark-brown; antennomeres 7 – 11 brown; mouthparts, antennomeres 1 – 6 and legs yellowish-brown; tarsi yellow. Punctation of head irregular: finer and sparser in basal portions of clypeus, denser and larger in middle, and denser and coarser on infraorbital parts, with portions between ocelli without or with two (paratype) moderately fine punctures; neck with dense and large punctation about as that in middle part of head; pronotum with irregular moderately dense punctation, finer and sparser in medioapical portion, larger, deeper and denser in middle and sparser in mediobasal part, lateral parts without punctures except of latero-apical and laterobasal portions with fine and sparse punctation; elytra with dense and coarse punctation, significantly larger and deeper than that in middle of pronotum, finer around scutellum and along suture; abdomen without visible punctation except of abdominal tergite V with indistinct sparse and fine punctation. Anterior part of clypeus with dense and transverse and laterobasal parts with longitudinal microreticulation, median and posterior portions of head without meshes; neck, pronotum and elytra without microsculpture; abdomen with dense and fine isodiametric sculpture. Head 1.2 – 1.3 times as broad as long, with strongly elevated clypeus and moderately wide and deep anteriomedian depressions; posteriolateral margins of clypeus stretching posteriad toward level of middle length of eyes. Latero-apical portions between eyes and clypeus narrowly and deeply depressed, with distinct semicircular notch between anterior margin of eyes and antennal insertion. Dorsal surface with strong elevations: longitudinal and diagonal in laterobasal parts of clypeus and subdiagonal on infraorbital portions. Anteocellar foveae significantly widened and deep, strongly convergent latero-apicad toward level of middle lengtht of eyes and connected with latero-apical depressions. Temples 1.5 times as long as longitudinal length of eye. Apical part of neck slightly and narrowly depressed. Distance between ocelli about twice as long as distance between ocellus and posterior margin of eye. Apical maxillary palpomere about three times as long as penultimate segment. Antenna with slightly transverse antennomeres 8 – 10; antennomere 4 1.3 times as long as 3, 5 slightly longer than 4, 6 indistinctly broader and longer than 5, 7 slightly shorter than 6, 8 slightly broader than 7, 9 – 10 slightly shorter and distinctly broader than 8. Pronotum 1.5 times as broad as long, 1.2 – 1.3 times as broad as head, widest in middle, with strongly convex middle part. Apical agles widely rounded, not protruded anteriad. Hind angles obtuse. Each lateral portion widely impressed, deeper slightly below middle. Surface of disc with two large and deep longitudinal depression. Elytra 1.1 times as broad as long, 1.7 – 1.8 times as long as pronotum, somewhat subparallel and slightly concave in about middle; lateral portions relatively widely impressed, hind margins rounded. Dorsal surface of each elytron with strong irregular elevations, subdiagonal in mediobasal, transverse in middle and longitudinal in apical portions. Abdomen slightly broader than elytra. Male. Posterior margin of abdominal tergite VIII slightly and sternite VIII widely and deeply sinuate. Aedeagus with wide basal portion, gradually narrowed toward median lobe; median lobe narrow, elongate, from narrower basal part slightly narrowed toward preapical part, with rounded apex; accessory plates narrow, elongate and rounded apically; parameres from narrowest basal parts significantly broadened in middle and narrowed toward apex, reaching level of preapical part of median lobe, each with one long and three short apical setae; internal sac long and wide, with two elongate sclerotized structures in basal part (Fig. 66). Lateral aspect of the aedeagus as in Fig. 67; apical portion of median lobe (lateral view) hook-shaped, with crenulate ventrolateral margin and rounded apex (Fig. 68). Female. Posterior margin of abdominal tergite VIII and sternite VIII truncate. Accessory sclerite with wide basal portion and median part, strongly narrowed toward narrow apical part with acute apex (Fig. 24). Spermatheca as in Fig. 25.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF946C5CFF0AFB0AFA3AFCC9.taxon	discussion	Comparative notes. Based on the proportions of the forebody and antennomeres, coarse punctation of the elytra and the general shape of the aedeagus, O. confertum sp. nov. is similar to O. latocavum Shavrin, 2023, recently described from central Nepal (Shavrin 2023 a). The new species can be distinguished from it by the smaller body, darker coloration, sparser punctation of the head and the pronotum, slightly longer temples more narrowed posteriad, more transverse pronotum with more widely impressed lateral portions and wider preapical part, slightly shorter elytra, different shape of the preapical portion of the median lobe and the female accessory sclerite, and other details of the morphology of the aedeagus.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF946C5CFF0AFB0AFA3AFCC9.taxon	distribution	Distribution. Omalium confertum sp. nov. is known only from the type locality in Sichuan, China (Fig. 39).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF946C5CFF0AFB0AFA3AFCC9.taxon	etymology	Etymology. The specific epithet is the Latin adjective confertus, - a, - um (dense or crowded). It alludes to the dense punctation of the forebody.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF946C5CFF0AFB0AFA3AFCC9.taxon	biology_ecology	Bionomics. Specimens were collected at elevation 3428 m a. s. l. The detailed bionomical data are unknown.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF926C5DFF0AFC2AFC0CF82D.taxon	description	(Figs 26, 64, 69 – 71, 75)	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF926C5DFF0AFC2AFC0CF82D.taxon	materials_examined	Type material examined. Holotype ♂: CHINA: QINGHAI: ‘ CHINA: Qinghai province, | Yunning Si [lamasery], | 2890 m, 36 ° 45.6´N | 102 ° 10.6´E (GPS), 1. – 16. | VII. 2005, J. Hájek, D. Král | & J. Růžička leg. ’ <printed>, ‘ [Ch 10 & 19]; baited pitfall | traps (fish meat) with | ethylene glycol; wet | coniferous forest, close | valley above the village’ <printed>, ‘ Museum für Naturkunde | Berlin | Sammlung M. Schülke’ <printed>, ‘ HOLOTYPE | Omalium | lama sp. nov. | Shavrin A. V. des. 2025 ’ <red, printed> (CNC). Paratypes (12 ♂♂, 26 ♀♀): QINGHAI: 8 ♂♂ (two specimen dissected; four specimens with additional printed label: ‘ Omalium | cf. japonicum Sharp | det. M. Schülke 2007 ’), 19 ♀♀ (one specimen dissected; two specimens without head and pronotum; one specimen without left elytron; six specimens with additional printed label: ‘ Omalium | cf. japonicum Sharp | det. M. Schülke 2007 ’): same data as the holotype (3 ♂♂, 5 ♀♀: cSh; 4 ♂♂, 13 ♀♀: cSch; 1 ♂, 1 ♀: NMPC); 2 ♂♂ (one specimen without left antennomeres 3 – 11; one specimen with additional printed label: ‘ Omalium | cf. japonicum Sharp | det. M. Schülke 2007 ’), 6 ♀♀ (one specimen without left antennomeres 3 – 11; two specimens with additional printed label: ‘ Omalium | cf. japonicum Sharp | det. M. Schülke 2007 ’): ‘ CHINA: Qinghai province, | Yunning Si [lamasery], | 2890 m, 36 ° 45.6´N | 102 ° 10.6´E (GPS), 1. – 2. | VII. 2005, J. Hájek, D. Král | & J. Růžička leg. ’ <printed>, ‘ [Ch 10]; individually under | stones and logs, in | excrements and on | vegetation in | coniferous forest ’ <printed>, ‘ Museum für Naturkunde | Berlin | Sammlung M. Schülke’ <printed> (1 ♂, 1 ♀: cSh; 1 ♂, 5 ♀♀: cSch); SICHUAN: 1 ♂ (dissected): ‘ CHINA Sichuan, Gongga | Shan, Hailuogou, above | Camp 3, 3100 m 8. VII. [19] 96 | 29 ° 35 N 102 ° 00 E C 56 ’ <printed>, ‘ collected by | A. Smetana, J. Farkáč | and P. Kabátek’ <printed> (CNC); 1 ♂ (dissected), 1 ♀: ‘ CHINA, N-Sichuan, | Huanglong Shan, 2611 m | Sanluogou vill. | 32 ° 46´05 ´´ N, 103 ° 55´25 ´´ E | 15 – 17. VI. 2018, leg. Reuter’ <printed> (cF). All paratypes with additional red printed label: ‘ PARATYPE | Omalium | lama sp. nov. | Shavrin A. V. des. 2025 ’.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF926C5DFF0AFC2AFC0CF82D.taxon	description	Description. Measurements (n = 39): HW: 0.48 – 0.59; HL: 0.33 – 0.42; OL: 0.14 – 0.16; TL: 0.07 – 0.10; AL (holotype): 0.94; PL: 0.35 – 0.51; PWmax: 0.60 – 0.74; PWmin: 0.51 – 0.67; ESL: 0.78 – 0.91; EW: 0.83 – 0.99; MTbL (holotype): 0.45; MTrL (holotype): 0.23 (MTrL 1 – 4: 0.08; MTrL 5: 0.15); AW: 0.75 – 0.99; AedL: 0.62 – 0.70; BL: 2.70 – 3.75 (holotype: 2.90). Habitus as in Fig. 64. Head, pronotum and abdomen reddish-brown (lateral and basal portions of pronotum usually paler); elytra brown to reddish-brown; antennomeres 5 – 11 or 6 – 11 brown; mouthparts, antennomeres 1 – 4 or 1 – 5 and legs yellow-brown. Punctation of head variable, moderately fine and sparse or slightly larger and coarser, denser in middle; neck with moderately dense punctation, sometimes with impunctate medioapical part; punctation of pronotum about as that on head, but sometimes larger, denser and deeper, usually finer and sparser in medioapical and mediobasal portions; punctation of elytra denser, larger and coarser than that on pronotum, usually finer and sparser in middle; abdominal tergites without or with indistinct fine and sparse punctation. Anterior part of clypeus with distinct transverse microsculpture, lateral portions of clypeus with fine isodiametric and subdiagonal meshes, portion between apical margin of eyes and scutellum with fine longitudinal microsculpture; neck without or with indistinct transverse or isodiametric sculpture in middle; scutellum with fine transverse or isodiametric meshes; abdominal tergites with dense isodiametric microreticulation. Head 1.4 times as broad as long, with distinctly elevated middle portion and deep and wide anteriomedian depressions, reaching level of anterior third of eyes; posterior part of clypeus gradually narrowed basad, with elongate posteriolateral margins reaching level of middle or posterior third of eyes. Latero-apical parts between anterior margin of eyes and clypeus with moderately wide and deep semicircular notch. Dorsal surface without or with irregular elongate or transverse wrinkles on infraorbital portions; median portion sometimes with irregular elevations between punctures. Anteocellar foveae moderately narrow, deep, distinctly convergent latero-apicad and reaching level of middle length of eyes, usually connected with deep latero-apical depressions between clypeus and eyes. Temples 1.6 times to twice as long as longitudinal length of eyes, from posterior margins of eyes gradually narrowed posteriad toward widely rounded hind angles. Apical part of neck narrowly and moderately deeply depressed, sometimes with short and narrow elevations; mediobasal portion sometimes with a row of short and narrow elevations. Distance between ocelli 1.4 – 1.7 times as long as distance between ocellus and posterior margin of eye. Apical maxillary palpomere about three times as long as penultimate segment. Antenna with slightly transverse antennomeres 6 – 8 and distinctly transverse 9 – 10. Pronotum 1.4 – 1.7 times as broad as long, 1.2 times as broad as head, from widest anterior portion gradually narrowed posteriad. Apical angles widely rounded, not protruded anteriad. Anterior margin rounded, not protruded anteriad, with slightly concave middle. Laterobasal portions not concave in front of obtuse angles. Lateral portions widely depressed, deeper in about middle and significantly broadened in laterobasal portions. Surface of disc with two wide longitudinal depressions, without or with narrow or suboval and shallow depression in medioapical portion; latero-apical parts with narrow and curved elongate elevations, reaching middle. Middle portion sometimes with irregular transverse or diagonal elevations between punctures. Elytra indistinctly broader than long, 1.7 times to slightly more than twice as long as pronotum. Abdomen about as wide as elytra or slightly narrower. Male. Posterior margin of abdominal tergite VII truncate or rounded. Posterior margin of abdominal sternite VIII widely sinuate. Aedeagus with wide basal portion, gradually narrowed toward median lobe; median lobe narrow and elongate, with rounded apex; accessory plates elongate and relatively wide, rounded apically; parameres strongly broadened toward middle and from preapical parts narrowed apically, each with two long and two short preapical setae; internal sac long and wide, without sclerotized structures (Fig. 69). Lateral aspect of the aedeagus as in Fig. 70; apical portion of median lobe (lateral view) narrow, with ventrolateral margin forming disinct five rounded teeth (Fig. 71). Female. Posterior margin of abdominal tergite VII truncate. Posterior margin of abdominal sternite VIII rounded. Accessory sclerite with moderately wide basal portion, gradually narrowed toward wide and elongate median part, rounded apically (Fig. 26). Spermatheca not recognized.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF926C5DFF0AFC2AFC0CF82D.taxon	discussion	Comparative notes. Based on the general shapes of the body and the aedeagus, O. lama sp. nov. is similar to O. smetanai sp. nov. (see below), from which it can be distinguished by the distinctly narrower antennomeres 6 – 10, the shorter temples, denser punctation of the head, slightly broader elytra, the shape of the narrower median lobe, not forming hook in apical portion (lateral view), slightly longer parameres, broader apical part of the female accessory sclerite, and other details of the external and internal morphology of the aedeagus.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF926C5DFF0AFC2AFC0CF82D.taxon	distribution	Distribution. Omalium lama sp. nov. is known from three localities in Yunning Si, Qinghai and Gongga Shan and Huanglong Shan mountain ranges in Sichuan, China (Fig. 75).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF926C5DFF0AFC2AFC0CF82D.taxon	etymology	Etymology. The name of the new species is derived from the fact that most type specimens were collected near monastery of lamas.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF926C5DFF0AFC2AFC0CF82D.taxon	biology_ecology	Bionomics. Specimens were collected at elevations from 2611 to 3100 m a. s. l. Specimens from Qinghai were collected by using pitfall traps in wet coniferous forest, and were collected individually under stones and logs and in excrements. The specimen from Gongga Shan Mts. (C 56) was collected by sifting of fallen leaves, humus and debris under them under rhododendrons at edges of an old Abies forest.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF916C60FF0AFF37FE8BFA15.taxon	description	(Figs 65, 72 – 75)	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF916C60FF0AFF37FE8BFA15.taxon	materials_examined	Type material examined. Holotype ♂ (dissected): CHINA: GANSU: ‘ CHINA, Gansu, Yonghai | cca 20 km SW Yuzhong | 2700 – 2800 m, 9. VIII. 94 | A. Smetana [C 34] ’ <printed>, ‘ HOLOTYPE | Omalium | pseudojaponicum sp. nov. | Shavrin A. V. des. 2025 ’ <red, printed> (CNC). Paratype 1 ♂ (Fig. 65): same data as the holotype, with additional red printed label: ‘ HOLOTYPE | Omalium | pseudojaponicum sp. nov. | Shavrin A. V. des. 2025 ’ (cSh).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF916C60FF0AFF37FE8BFA15.taxon	description	Description. Measurements (n = 2): HW: 0.60 – 0.62; HL: 0.42; OL: 0.17; TL: 0.05; AL (holotype): 0.97; PL: 0.52; PWmax: 0.71 – 0.80; PWmin: 0.65 – 0.72; ESL: 0.95 – 1.01; EW: 1.03 – 1.07; MTbL (holotype): 0.42; MTrL (holotype): 0.26 (MTrL 1 – 4: 0.09; MTrL 5: 0.17); AW: 1.03 – 1.05; AedL: 0.72; BL: 3.30 (holotype) – 3.35. Habitus as in Fig. 65. Head, pronotum and abdomen reddish-brown; elytra yellowish-brown, with slightly darkened mediobasal and lateral portions; antennomeres 6 – 11 brown; mouthparts, antennomeres 1 – 5 and legs yellow. Punctation of head irregular, moderately large and deep, denser in middle part; neck with moderately large and sparse punctation; punctation of pronotum irregular, sparser in middle and lateral portions, mediobasal portion between longitudinal depressions impunctate (paratype); middle part of elytra with fine and sparse punctation, mediolateral and apical parts with distinctly denser, larger and deeper punctation; abdominal tergites with indistinct, fine and sparse punctation. Anterior portion of clypeus with fine transverse microsculpture, basal portions of clypeus with dense transverse meshes; scutellum without sculpture; abdominal tergites with dense transverse microreticulation. Head with wide and deep anteriomedian depressions, reaching level of anterior third of eyes. Latero-apical margin between anterior margin of eyes and clypeus with moderately wide and deep semicircular notch. Dorsal surface with irregular transverse and diagonal elevations between pucntures in middle and in posterior parts of infraorbital portions. Anteocellar foveae deep and elongate, slightly convergent latero-apicad toward level of middle length of eyes and indistinctly connected with lateroapical depressions between apical margin of eyes and clypeus. Temples more than three times shorter than longitudinal length of eyes, from posterior margins of eyes strongly narrowed toward neck. Apical part of neck narrowly depressed, with irregular short and longitudinal elevations. Distance between ocelli about twice as long as distance between ocellus and posterior margin of eye. Apical maxillary palpomere about two and a half times as long as penultimate segment. Antenna with sligthtly transverse antennomeres 6 – 7 and distinctly transverse 8 – 10; 3 slightly longer and distinctly narrower than 2, 4 1.3 times as long as 3, 5 distinctly longer than 4, 6 distinctly broader than 5, 7 slightly longer and broader than 6, 8 slightly longer and broader than 7, 9 – 10 slightly longer and broader than 8. Pronotum 1.3 – 1.5 times as broad as long, from widest middle gradually narrowed both posteriad and anteriad. Apical angles widely rounded, not protruded anteriad. Anterior margin rounded, not protruded anteriad, slightly concave in middle. Laterobasal portions not concave in front of obtuse hind angles. Lateral portions moderately deeply and widely impressed, distinctly deeper in about middle. Surface of disc with two moderately long and deep longitudinal depressions; surface between depressions slightly and widely elevated; latero-apical parts with narrow elongate elevations, reaching middle. Middle portion without elevations between punctures. Elytra 1.8 – 1.9 times as long as pronotum, from basal portion distinctly broadened posteriad; hind margins straight. Dorsal surface of each elytron with irregular elevations between punctures. Metatarsi less than twice as long as than metatibia. Male. Posterior margin of abdominal tergite VIII truncate. Posterior margin of andominal sternite VIII sinuate. Aedeagus with wide and relatively short basal portion, gradually narrowed towad median lobe; median lobe narrow, elongate, from widest middle with widely rounded projections slightly narrowed toward small rounded apex; accessory plates elongate and narrow, rounded apically; each paramere significantly broadened in middle and slightly narrowed toward rounded apex, reaching middle of median lobe, with two long apical and one short preapical setae; internal sac narrow and long, without sclerotized structures (Fig. 72). Lateral aspect of the aedeagus as in Fig. 73; apical portion of median lobe (lateral view) hook-shaped, with distinct moderately wide four ventrolateral teeth, largest ventrolateral tooth relatively long (Fig. 74). Female unknown.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF916C60FF0AFF37FE8BFA15.taxon	discussion	Comparative notes. Based on the coloration of the body, the general shape of the body and the aedeagus, O. pseudojaponicum sp. nov. is similar to O. japonicum Sharp, 1874, known from Japan (Honshu, Kyushu) and South Korea (Watanabe 1990, Kim & Ahn 2014). The new species can be distinguished from it by the longer body, denser and coarser punctation of the head and the pronotum, the longer elytra, longer and narrower median lobe with significantly broader mediolateral projections, narrower parameres and other details of the morphology of the aedeagus.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF916C60FF0AFF37FE8BFA15.taxon	distribution	Distribution. Omalium pseudojaponicum sp. nov. is known only from the type locality in Gansu, China (Fig. 75).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF916C60FF0AFF37FE8BFA15.taxon	etymology	Etymology. The name of the new species is based on the morphological similarity to O. japonicum Sharp, 1874, with the Greek prefix ψευδής (false).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FF916C60FF0AFF37FE8BFA15.taxon	biology_ecology	Bionomics. Specimens were collected at elevations from 2700 to 2800 m a. s. l. The detailed bionomical data are unknown.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FFAE6C63FF0AF996FE99FD91.taxon	description	(Figs 27 – 28, 76, 79 – 81, 88)	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FFAE6C63FF0AF996FE99FD91.taxon	materials_examined	Type material examined. Holotype ♂ (dissected): CHINA: SICHUAN: ‘ P. R. CHINA, Sichuan, | Emei Shan, N 29 ° 33,6´| E 103 ° 20.6´<printed>, 27. vi. – 5. vii. | 2009, 1800 – 2400 m, sifti- | ng 11 – 17, V. Grebennikov’ <printed>, ‘ HOLOTYPE | Omalium | rougemonti sp. nov. | Shavrin A. V. des. 2025 ’ <red, printed> (NMPC). Paratypes (7 ♂♂, 4 ♀♀): CHINA: SICHUAN: 1 ♂ (dissected), 1 ♀: P. R. CHINA, Sichuan, | EmeiShan, N 29 ° 30´46.5 ´´ | E 103 ° 19´57.0 ´´ | 14. vi. 2010, 3035 m. | sifting 28, V. Grebennikov’ <printed> (1 ♂: cSh; 1 ♀: CNC); 1 ♀: ‘ P. R. CHINA, Sichuan, | EmeiShan, N 29 ° 30´54.2 ´´ | E 103 ° 19´50.6 ´´, | 14. iv. 2010, 3055 m, | sifting 29, V. Grebennikov’ <printed> (CNC); 2 ♀♀ (one specimen dissected): ‘ P. R. CHINA, Sichuan, | EmeiShan, N 29 ° 31´36.8 ´´ | E 103 ° 19´52.1 ´´, | 15. iv. 2010, 2926 m, | sifting 31, V. Grebennikov’ <printed> (1 ♀: cSh; 1 ♀: CNC); 1 ♂ (dissected): ‘ CHINA: SICHUAN PROV., | Emeishan Mt., 2820 m, | TAIZIPING TEMPLE, | 29 ° 31´50 ´´ N 103 ° 19`36 `` E’ <printed>, ‘ 8. VI. 2014, sift # 7, | mixed forest with Abies, | bamboo undergrowth, | J. Hájek & J. Růžička leg. ’ <printed> (NMPC); SHAANXI: 1 ♂ (dissected): ‘ China: Shaanxi, Qin Ling Shan | 110.06 E, 34.25 N | Hua Shan Mt., S. - top, 1950 – 2000 m | Forrest [sic], sifted | 19.08.1995, leg. A. Pütz’ <printed>, ‘ Museum für Naturkunde | Berlin | Sammlung M. Schülke’ <printed> (cSch); 1 ♂ (dissected): ‘ China: Shaanxi, Qin Ling Shan | 110.06 E, 34.27 N | Hua Shan Mt. N Valley, 1230 – | 1400 m, 118 km E Xian, sifted | 18. / 20.08.1995, leg. M. Schülke’ <printed>, ‘ Sammlung | M. Schülke | Berlin’ <green, printed> (cSh); HUBEI: 1 ♂: ‘ CHINA: W-Hubei (Daba Shan) | pass E of Mt. Da Shennongjia, | 12 km NW Muyuping, 31 ° 36´N, | 110 ° 21´E, 19. VII. 2001, | leg. M. Schülke [C 01 - 13 C] ’ <printed>, ‘ creek valley 1950 – 2050 m, | mixed deciduous forest, | moss, dead wood, mushrooms | (sifted) [C 01 - 13 C] ’ <printed>, ‘ Omalium [handwritten] | sp. [handwritten] | det. M. Schülke 2001 ’ <printed>, ‘ Museum für Naturkunde | Berlin | Sammlung M. Schülke’ <printed> (cSch); ZHEJIANG: 1 ♂ (dissected): ‘ CHINA Zhejiang | Tienmushan | 29. IV. 1993 | G. de. Rougemont’ <printed>, ‘ Omalium [handwritten] | sp. nov. [handwritten] | det. 2018 | G. de Rougemont’ <printed> (OUMNH); 1 ♂ (dissected): ‘ CHINA: Zhejiang Prov., | Anji County, ca 300 [handwritten] m. | Long Wang Shan N. R. | 12 [handwritten]. v. 1996 J. Cooter’ <printed>, ‘ Rougemont | collection’ (OUMNH). All paratypes with additional red printed label: ‘ PARATYPE | Omalium | rougemonti sp. nov. | Shavrin A. V. des. 2025 ’.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FFAE6C63FF0AF996FE99FD91.taxon	description	Description. Measurements (n = 12): HW: 0.58 – 0.62; HL: 0.35 – 0.42; OL: 0.17 – 0.20; TL: 0.05 – 0.06; AL (holotype): 1.20; PL: 0.45 – 0.52; PWmax: 0.67 – 0.71; PWmin: 0.63 – 0.67; ESL: 0.95 – 1.05; EW: 1.05 – 1.16; MTbL (holotype): 0.52; MTrL (holotype): 0.22 (MTrL 1 – 4: 0.09; MTrL 5: 0.13); AW: 1.00 – 1.06; AedL: 0.57 – 0.68; BL: 2.93 – 3.95 (holotype: 3.65). Habitus as in Fig. 76. Body reddish-brown (pronotum of holotype and three paratypes and abdomen in some specimens distinctly paler); antennomeres 5 – 11 or 6 – 11 brown; mouthparts, antennomeres 1 – 4 or 1 – 5, lateral and basal portions of pronotum an paratergites of abdomen in some specimens yellowish; tarsi yellow. Punctation of head moderately dense and fine, sometimes denser in middle and coarser in basal half of infraorbital portions (two paratypes with denser, larger and coarser punctation in middle and on infraorbital parts); neck with fine and sparse or relatively large and dense punctation; punctation of pronotum slightly larger, deeper and sometimes coarser than that in middle part of head, finer and sparser in medioapical and mediobasal (some specimens) portions, and sparser in lateral parts; punctation of elytra about as that on pronotum, denser and coarser around scutellum and usually finer and sparser in middle portion; abdominal tergites with fine and sparse punctation. Anterior part of clypeus without or with traces of transverse microreticulation, laterobasal portions of clypeus with fine diagonal meshes in some specimens; neck without or with indistinct transverse microsculpture visible in mediobasal portion; scutellum with dense transverse or isodiametric sculpture; abdomen with dense isodiametric microsculpture. Anterior and posterior margins of pronotum with row of short cuticular fringe. Head 1.4 – 1.6 times as broad as long, with strongly elevated lateral parts of clypeus and wide and deep anteriomedian depressions, reaching level of anterior third of eyes; posteriolateral margins of clypeus subparallel or slightly narrowed posteriad, reaching level of anterior third or middle of eyes. Latero-apical margin between anterior margin of eyes and clypeus with indistinct or distinct small semicircular notch. Dorsal surface without or with moderately strong longitudinal and diagonal wrinkles in laterobasal portions of clypeus, without or with transverse irregular elevations in middle, with longitudinal and diagomal elevations on infraorbital portions, usually stronger on postocular parts. Anteocellar foveae narrow and deep, slightly or strongly convergent latero-apicad toward level of middle length of eyes. Temples short, more than three times as long as longitudinal length of eyes, from posterior margins of eyes slightly narrowed posteriad toward widely rounded hind angles. Apical part of neck without or with narrow and deep depressions, with several narrow and short longitudinal elevations. Distance between ocelli 1.5 – 1.8 times as long as distance between ocellus and posterior margin of eye. Apical maxillary palpomere slightly more than three times as long as penultimate segment. Antenna with distinctly elongate antennomeres 5 – 8 and slightly elongate 9 – 10; 4 slightly narrower and 1.3 times as long as 3, 5 distinctly longer than 4, 6 distinctly longer and slightly broader than 5, 7 slightly broader than 6, 8 slightly shorter than 7, 9 – 10 broader than 8, apical antennomere 1.2 – 1.3 times as long as preceding segment, from middle strongly narrowed toward acute apex. Pronotum 1.3 – 1.4 times as broad as long, 1.1 times as broad as head, from widest anterior portion gradually narrowed posteriad. Apical angles widely rounded, not protruded anteriad. Anterior margin rounded, not or distinctly protruded anteriad. Laterobasal portions not concave in front of obtuse or subacute hind angles. Lateral portions deeply and moderately widely impressed, deeper in about middle and broader in laterobasal parts. Surface of disc with two wide and deep longitudinal depressions and deep, small, oval or rounded depression in medioapical portion; surface between all pronotal depressions distinctly elevated, sometimes stronger at sides of medioapical depression. Middle portion without or with irregular fine transverse and diagonal elevations between punctures. Elytra twice as long as pronotum. Dorsal surface of each elytron with strong irregular elevations between punctures in mediobasal, mediolateral and apical portions (some specimens without elevations between punctures in middle parts of each elytron). Abdomen slightly narrower than elytra, sometimes with narrow or wide intersegmental membranes between abdominal tergites III – V or III – VI. Male. Posterior margin of abdominal tergite VIII truncate. Posterior margin of abdominal sternite VIII sinuate. Aedeagus with moderately wide basal portion, gradually narrowed toward median lobe; median lobe narrow and elongate, slightly sinuate in preapical portion, with small rounded apex; accessory plates narrow, short and distinctly convergent lateroapicad, rounded apically; parameres distinctly broadened in middle, and from middle slightly narrowed toward small rounded apices, reaching preapical portion of median lobe, each with one-two moderately long apical and one-two relatively short preapical setae; internal sac long and narrow, with two small oval structures in basal part (Fig. 79). Lateral aspect of the aedeagus as in Fig. 80; apical portion of median lobe (lateral view) with distinctly broadened preapical portion, slightly curved ventrad, with somewhat rounded and disinctly crenulate ventrolateral margin (Fig. 81). Female. Posterior margin of abdominal tergite VIII truncate. Posterior margin of abdominal sternite VIII truncate or rounded. Female accessory sclerite short, from widest basal portion gradually narrowed toward rounded apex (Fig. 27). Spermatheca as in Fig. 28.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FFAE6C63FF0AF996FE99FD91.taxon	discussion	Comparative notes. Based on the general shape of the body and the aedeagus, O. rougemonti sp. nov. is similar to the Himalayan O. secretum Cameron, 1924, known from India and Nepal (Shavrin 2023 a). It can be distinguished from it by the shorter temples, more narrowed posteriad, the presence of small and deep depression in medioapical portion of the more convex pronotum, different shape of the median lobe, with shorter and broader apical portion of the median lobe, longer parameres with broader apical portions, broader apical part of the female accessory sclerite, and details of the morphology of the aedeagus.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FFAE6C63FF0AF996FE99FD91.taxon	distribution	Distribution. Omalium rougemonti sp. nov. is known from several localities in Emei Shan Mts. in Sichuan, Qin Ling Shan Mts. in Shaanxi, Daba Shan Mts. in Hubei and Tienmushan Mts. in Zhejiang, China (Fig. 88).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FFAE6C63FF0AF996FE99FD91.taxon	etymology	Etymology. Patronymic; the species is named in memory of Guillaume de Rougemont † (1945 – 2020), the collector of some type specimens.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FFAE6C63FF0AF996FE99FD91.taxon	biology_ecology	Bionomics. Specimens were collected from 300 to 3055 m a. s. l. by sifting litter in mixed forests. One specimen in Emei Shan Mts. was collected by sifting in mixed forest with Abies and bamboo undergrowth, and one specimen from Daba Shan Mts. (C 01 - 13 C) was collected in mixed deciduous forest by sifting of moss, dead wood and mushrooms.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FFAD6C65FF0AFD12FE41FB51.taxon	description	(Figs 29, 75, 82 – 84)	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FFAD6C65FF0AFD12FE41FB51.taxon	materials_examined	Type material examined. Holotype ♂ (dissected): CHINA: YUNNAN: ‘ CHINA: N-Yunnan Zhong- | dian Co. pass 28 km ESE Zhong- | dian, 27 ° 43.9´N 99 ° 58.2´E | 3700 – 3750 m 22. VIII. 2003 | A. Smetana [C 131] ’ <printed>, ‘ HOLOTYPE | Omalium | smetanai sp. nov. | Shavrin A. V. des. 2025 ’ <red, printed> (CNC). Paratypes: YUNNAN: 4 ♀♀ (one specimen dissected; one specimen without right antennomeres 3 – 11 and abdomen): same data as the holotype, with additional red printed label: ‘ PARATYPE | Omalium | smetanai sp. nov. | Shavrin A. V. des. 2025 ’. (2 ♀♀: cSh; 2 ♀: CNC).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FFAD6C65FF0AFD12FE41FB51.taxon	description	Description. Measurements (n = 5): HW: 0.46 – 0.49; HL: 0.34 – 0.36; OL: 0.13 – 0.15; TL: 0.06 – 0.07; AL (holotype): 0.80; PL: 0.38 – 0.40; PWmax: 0.58 – 0.60; PWmin: 0.52 – 0.54; ESL: 0.75 – 0.79; EW: 0.73 – 0.81; MTbL (holotype): 0.35; MTrL (holotype): 0.20 (MTrL 1 – 4: 0.03; MTrL 5: 0.12); AW: 0.75 – 0.84; AedL: 0.56 – 0.57; BL: 2.40 – 2.86 (holotype: 2.70). Habitus as in Fig. 77. Body dark-brown, with slightly paler elytra; antennomeres 6 – 11 brown; mouthparts, antennomeres 1 – 5 and legs yellow-brown. Punctation of head dense, moderately large and deep, distinctly denser in middle, denser and coarser on infraorbital portions; neck with fine sparse punctation; punctation of pronotum dense, slightly larger, coarser and deeper than that in middle part of head, with interspaces between punctures in mediolateral portion about as long as diameters of one-two nearest puncture, disinctly sparser in middle and mediobasal portions; punctation of elytra about as that on pronotum, denser, larger and coarser around scutellum and in mediolateral portions; abdominal tergites without or with fine and sparse punctation. Anterior part of head with transverse microsculpture, lateral portions of clypeus with irregular transverse and diagonal meshes; scutellum with fine isodiametric sculpture; abdominal tergites with dense isodiametric microreticulation. Head 1.3 times as broad as long, with wide and moderately deep anteriomedian depressions, reaching anterior third of eyes; lateroposterior portions of clypeus distinctly narrowed basad toward level of anterior third or middle of eyes. Latero-apical margin between anterior margin of eyes and clypeus with small semicircular notch. Dorsal surface with fine diagonal elevations in laterobasal portions of clypeus, irregular, fine, transverse and longitudinal elevations in middle and in posterior half of infraorbital portions. Anteocellar fovea narrow, moderately deep, slightly convergent latero-apicad toward level of middle length of eyes. Temples twice as long as longitudinal length of eyes, from posterior margin of eyes gradually narrowed toward widely rounded hind angles. Apical part of neck not or indistinctly and narrowly depressed, with irregular short and narrow elevations. Distance between ocelli 1.3 – 1.5 times as long as distance between ocellus and posterior margin of eye. Apical maxillary palpomere about three times as long as penultimate segment. Antenna with slightly transverse antennomeres 8 – 10; 4 about as long as wide, twice shorter and indistinctly narrower than 3, 5 distinctly longer than 4, 6 slightly longer than 5, 7 slightly longer and indistinctly broader than 6, 8 slightly broader than 7, 9 – 10 distinctly broader than 8. Pronotum 1.5 times as broad as long, 1.2 times as broad as head, from widest middle slightly more narrowed posteriad than anteriad. Apical angles widely rounded, indistinctly protruded anteriad. Anterior margin rounded, slightly concave in middle, not protruded anteriad. Laterobasal portions not concave in front of obtuse hind angles. Lateral portions deeply and moderately widely impressed, deeper slightly below middle. Middle portion without elevations between punctures. Elytra about as long as broad, 1.9 times as long as pronotum; hind margins straight. Dorsal surface of each elytron with irregular diagonal elevations between punctures (holotype without elevations in median part along suture). Abdomen slightly broader than elytra, with moderately wide intersegmental membranes between abdominal tergites III – IV (holotype and one paratype). Male. Posterior margin of abdominal tergite VIII truncate. Posterior margin of abdominal sternite VIII widely sinuate. Aedeagus with wide basal portion, gradually narrowed toward median lobe; median lobe narrow and elongate, broadened in middle, slightly sinuate in lateral portions in front of middle, slightly broadened in posterior portion, with elongate small and rounded apex; accessory plates narrow and elongate, rounded apically; each paramere significantly broadened in middle and strongly narrowed toward small rounded apex, with one long and two short apical setae; internal sac long and narrow, with four small sclerotized structures in basal portion (Fig. 82). Lateral aspect of the aedeagus as in Fig. 83; apical portion of median lobe (lateral view) hook-shaped, with crenulate ventrolateral margin, rounded hind tooth and small rounded apex (Fig. 84). Female. Posterior margin of abdominal tergite VIII truncate. Posterior margin of abdominal sternite VIII widely rounded. Accessory sclerite from widest basal portion strongly narrowed toward significantly elongate narrow median part with subacute apex (Fig. 29). Spermatheca not recognized.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FFAD6C65FF0AFD12FE41FB51.taxon	discussion	Comparative notes. Based on the general shapes of the body and the aedeagus, O. smetanai sp. nov. is similar to O. lama sp. nov. (see above), from which it can be distinguished by distinctly more elongate antennomeres 6 – 10, the longer temples, sparser punctation of the head, slightly narrower elytra, slightly broader apical half of the median lobe distinctly concave in preapical portion, the presence of distinct dorsolateral hook in apical part of the median lobe (lateral view), slightly shorter parameres, narrower apical part of the female accessory sclerite, and other details of the external and internal morphology of the aedeagus.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FFAD6C65FF0AFD12FE41FB51.taxon	distribution	Distribution. Omalium smetanai sp. nov. is known only from the type locality in Zhongdian, northern Yunnan, China (Fig. 75).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FFAD6C65FF0AFD12FE41FB51.taxon	etymology	Etymology. Patronymic; the species is named in the memory of Aleš Smetana † (1931 – 2021), the collector of type specimens.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FFAD6C65FF0AFD12FE41FB51.taxon	biology_ecology	Bionomics. Specimens were collected from altitudes 3700 to 3750 m a. s. l. Specimens were collected by sifting of leaf litter, rotting wood, moss and various floor debris, particularly around and under mushrooms in various stages of development in an degraded original Abies, Sorbus, Larix, Rhododendron forest with various deciduous shrubbery undergrowth [C 131].	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FFAB6C68FF0AFB52FC6FFEB1.taxon	description	(Figs 47, 78, 85 – 87, 89)	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FFAB6C68FF0AFB52FC6FFEB1.taxon	materials_examined	Type material examined. Holotype ♂: CHINA: YUNNAN: ‘ CHINA: N-Yunnan [C 2005 - 03] | Diqing Tibet. Aut. Pref., | Zhongdian Co., 35 km | ESE Zhongdian, 3450 m | 27 ° 41.00´N 100 ° 01.47´E, devas- | tated mixed forest nr. small creek, | litter, moss, dead wood sifted, | 3. VI. 2005, M. Schülke [C 2005 - 3] ’ <printed>, ‘ Museum für Naturkunde | Berlin | Sammlung M. Schülke’ <printed>, ‘ HOLOTYPE | Omalium | tibetanum sp. nov. | Shavrin A. V. des. 2025 ’ <printed> (cSch). Paratypes: CHINA: YUNNAN: 1 ♂ (dissected): same data as the holotype (cSh); 1 ♂ (dissected): ‘ CHINA: N-Yunnan [C 2005 - 09] | Diqing Tibet. Aut. Pref., | Deqin Co., Meili Xue Shan, | E-side, 14 km W Deqin, 2580 m, ’ <printed>, ‘ 28 ° 27.47´N 98 ° 46.35´E, creek | valley below glacier, mixed forest, | leaf litter, moss, dead wood, sifted, | 11. VI. 2005, M. Schülke [C 2005 - 09] ’ <printed>, ‘ Museum für Naturkunde | Berlin | Sammlung M. Schülke’ <printed> (cSch). All paratypes with additional red printed label: ‘ PARATYPE | Omalium | tibetanum sp. nov. | Shavrin A. V. des. 2025 ’.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FFAB6C68FF0AFB52FC6FFEB1.taxon	description	Description. Measurements (n = 3): HW: 0.48 – 0.52; HL: 0.32 – 0.34; OL: 0.14 – 0.15; TL: 0.10; AL (holotype): 1.08; PL: 0.39 – 0.42; PWmax: 0.61 – 0.64; PWmin: 0.57 – 0.59; ESL: 0.79 – 0.88; EW: 0.83 – 0.94; MTbL (holotype): 0.45; MTrL (holotype): 0.23 (MTrL 1 – 4: 0.08; MTrL 5: 0.15); AW: 0.81 – 0.89; AedL: 0.50 – 0.52; BL: 2.87 – 3.20 (holotype). Habitus as in Fig. 78. Body reddish-brown, with darker head and pronotum (lateral and basal portions of pronotum, abdominal paratergites and intersegmental abdomen markedly paler); antennomeres 6 – 11 brown; mouthparts, antennomeres 1 – 5 and legs yellowish. Punctation of head fine and dense, denser in middle and coarser on infraorbital portions; neck with fine sparse or dense punctation; punctation of pronotum dense, finer and sparser in middle and mediobasal portion; punctation of elytra about as that on pronotum, slightly finer and sparser in middle; abdominal tergites with fine and sparse punctation. Anterior portion with distinct transverse microsculpture; neck with fine isodiametric meshes; abdominal tergites with dense isodiametric microreticulation. Anterior and posterior margins of pronotum with row of short cuticular fringe. Head 1.5 times as broad as long, with wide and deep anteriomedian depressions, reaching level of anterior margins of eyes; posteriolateral margins of clypeus reaching level of middle length of eyes. Latero-apical margin between anterior margin of eyes and clypeus with distinct small and moderately deep semicircular notch. Dorsal surface with diagonal longitudinal wrinkles between anterior margin of eyes and scutellum and on laterobasal portions of clypeus, with slightly elevated surface between punctures in middle and irregular longitudinal and diagonal wrinkles on infraorbital portions, denser and coarser on postocular parts. Anteocellar foveae deep and moderately narrow, convergent latero-apicad toward level of middle or anterior third of eyes. Temples 1.4 – 1.5 times as long as longitudinal length of eye, from posterior margin slightly narrowed posteriad toward widely rounded hind angles. Distance between ocelli 1.5 – 1.7 times as long as distance between ocellus and posterior margin of eye. Apical maxillary palpomere about three times as long as penultimate segment. Antenna with slightly transverse antennomeres 8 – 10; antennomere 3 about as long as and distinctly narrower than 2, 4 1.3 times as long as 3, 5 longer and slightly broader than 4, 6 slightly broader than 5, 7 slightly shorter and broader than 6, 8 broader than 7, 9 – 10 broader than 8. Pronotum 1.5 times as broad as long, 1.2 times as broad as head, from widest anterior portion distinctly more narrowed posteriad than anteriad. Anterior margin slightly rounded, not protruded anteriad. Laterobasal portions not concaved in front of obtuse hind angles. Lateral portions moderately widely and deeply impressed, deeper in about middle. Surface of disc with two long and wide longitudinal depressions; mediobasal portion without (holotype) or with shallow oval depression; surface between all pronotal depressions slightly elevated, with indistinct and irregular longitudinal elevations between punctures. Elytra twice as long as pronotum; hind margins straight. Dorsal surface of each elytron with strong irregular transverse and diagonal elevations between punctures (middle part of elytra of holotype without elevations). Metatarsi about twice as long as metatibia. Abdomen slightly narrower than elytra, with wide intersegmental membranes between abdominal tergites III – VI (holotype and one paratype). Male. Posterior margin of abdominal tergite VIII truncate. Posterior margin of abdominal sternite VIII sinuate. Aedeagus with wide basal portion, gradually narrowed toward median lobe; median lobe from basal part gradually broadened toward middle, and from middle strongly narrowed posteriad toward small rounded apex; accessory plates moderately wide, relatively long, curved in apical portions, each with widely rounded apex; parameres reaching preapical part of median lobe, distinctly broadened in about middle, and from middle gradually narrowed apically toward narrow apical parts, each with two long apical and two moderately short preapical setae; internal sac long and moderately narrow, without sclerotized structures (Fig. 85). Lateral aspect of the aedeagus as in Fig. 86; apical portion of median lobe (lateral view) narrow, with slightly crenulate ventrolateral margin and slightly curved small apex, slightly curved laterodorsad (Fig. 87). Female unknown.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FFAB6C68FF0AFB52FC6FFEB1.taxon	discussion	Comparative notes. Based on the general shape of the body and the aedeagus, O. tibetanum sp. nov. is more similar to O. xizangense, described from Xizang (see below), but can be distinguished from it by the more rounded and less protruded anterior angles of the slightly more transverse pronotum widest in anterior portion (pronotum of O. xizangenese broader in the middle) and with slightly denser punctation, and by the shape of longer and narrower median lobe of the aedeagus.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FFAB6C68FF0AFB52FC6FFEB1.taxon	distribution	Distribution. Omalium tibetanum sp. nov. is known from two localities in Zhongdian and Meili Xue Shan Mts., northern Yunnan, China (Fig. 47).	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FFAB6C68FF0AFB52FC6FFEB1.taxon	etymology	Etymology. The specific epithet is the latinized adjective derived from the area where the specimens were collected.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FFAB6C68FF0AFB52FC6FFEB1.taxon	biology_ecology	Bionomics. Specimens were collected from altitudes from 2580 to 3450 m a. s. l. by sifting of litter, moss and dead wood. One specimen was collected in Meili Xue Shan together with O. rutilum sp. nov. and O. sectum sp. nov. The photograph of the type locality in Zhongdian (C 2005 - 3) as in Fig. 89.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FFA66C68FF0AFC9DFE0DFABB.taxon	discussion	Remarks. Omalium chinese was originally described based one the holotype from Heilongjiang, China (“ Holotype female, 1. vi. 1992, Heilongjiang, Shangzhi, Laoyeling, soil layer of a Pinus koraiensis forest, Xue-Ping Zhang leg. ” (Z. - W. Yin, translation from Chinese )). The original description is very short and based on it, it is very difficult to compare O. chinese with other described species. The description of this species: “ Body length 5.2 mm. Body black, antennae, mouthparts, and legs reddish-brown. Anterior margin of clypeus, mouthparts and antennae reddish-brown. Vertex finely punctate and flat. Last segment of maxillary palpus longest. Punctation of pronotum same as that of head; lateral margins with broad reddish-brown band, deflexed, posterior margin slightly emarginate. Elytra with longitudinal row of big punctures; lateral margins, suture and posterior margin reddish-brown, but not distinct. Abdomen finely punctate, glabrous, posterior margin of each segment with microsculpture. Legs reddish-brown; femora thick. ” (translation from Chinese by Z. - W. Yin). To clarify the taxonomic status of this species, the study of the holotype is necessary.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
50613239FFA66C68FF0AFE32FE6CFD54.taxon	discussion	Remarks. Omalium xizangensis was originally described based on the male (holotype) from Xizang (type label (translated from Chinese by Z. - W. Yin): “ Xizang, Mêdog, Gandai, 2150 m, 28.52 ° N, 94.62 ° E, 1998. XI. 25, YAO Jian leg. ”. Based on the morphology of the habitus and the aedeagus (Fig. 1 in Zheng & Pu 2000), O. xizangensis belong to the rivulare group.	en	Shavrin, Alexey V. (2025): Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa 5588 (4): 501-543, DOI: 10.11646/zootaxa.5588.4.1, URL: https://doi.org/10.11646/zootaxa.5588.4.1
