identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
FA99492FB85A5140870DE5664C5F03FB.text	FA99492FB85A5140870DE5664C5F03FB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paleothereva Feng, Ren and Wang 2026	<div><p>Genus Paleothereva Feng, Ren and Wang gen. nov.</p><p>Figures 2, 3, 4, 5</p><p>Type species.</p><p>Paleothereva longicoxa Feng, Ren and Wang, sp. nov.</p><p>Diagnosis.</p><p>Scape as wide as pedicel in width; palpus two-segmented; single seta present antero-ventrally on apex of hind femur; R 1 bare; cell m 3 open; A 1 spines elongate and acuminate apically.</p><p>Etymology.</p><p>The generic name is derived from the Greek prefix “ paleo- ” (ancient), combined with Thereva, the type genus of Therevidae .</p></div>	https://treatment.plazi.org/id/FA99492FB85A5140870DE5664C5F03FB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Feng, Qi;Jia, Xisen;Zhou, Chenxin;Ren, Dong;Wang, Yongjie	Feng, Qi, Jia, Xisen, Zhou, Chenxin, Ren, Dong, Wang, Yongjie (2026): The earliest crown therevid lineage: bridging the gap of early evolution of stiletto flies. Arthropod Systematics & Phylogeny 84: 281-292, DOI: 10.3897/asp.84.e185452
D985EB0E96AE541EAE63B70BB4B084AE.text	D985EB0E96AE541EAE63B70BB4B084AE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paleothereva longicoxa Feng, Ren and Wang 2026	<div><p>Paleothereva longicoxa Feng, Ren and Wang, sp. nov. (Figs 2, 3, 4, 5)</p><p>Diagnosis.</p><p>Same as for the genus.</p><p>Locality and horizon.</p><p>Northern Myanmar, Kachin (Hukawng Valley), lowermost Cenomanian, dated 98.79 ± 0.62 Ma (Shi et al. 2012).</p><p>Type material.</p><p>Holotype female, No. CNU–DIP–MA 2016125 .</p><p>Etymology.</p><p>The specific epithet longicoxa is derived from the Latin “ longus ” and “ coxa ”, referring to the distinctly elongate coxa of the species.</p><p>Description.</p><p>Body: Slender, about 8.12 mm in length (Fig. 2 A, B). Wing about 4.61 mm in length. — Head. Semispherical, without macrosetae; eyes dichoptic, bare; occiput with numerous setae; frons wide, sparsely setose (Fig. 3 A, B). Antennae shorter than head height; scape approximately three times length of pedicel, bearing dense stout setae, absent on the medial surface; pedicel as wide as scape; flagellum 3 - segmented, covered with dense pubescence; basal flagellomere bulbous and tapered apically, with numerous sensory pits near base; apical flagellomere longer than flagellomere 2; stylus short (Fig. 3 C, D). Mouthparts short, labrum present, with well-developed fleshy labellum; palpus two-segmented; apical palpomere, gradually tapering to apex (Fig. 3 E). — Thorax. Slightly arched dorsally, with 4 pairs of np, 1 pair of sa, 2 pairs of pa, 1 pair of dc; scutellum well-developed, with a pair of macrosetae (Fig. 4 A, B). Legs slender, and equal length; coxae elongated with sparse macrosetae, 2 × broader than femora; hind coxa knob present; single seta present antero-ventrally on apex of hind femur; tibiae with macrosetae; tarsus 1 equal to tarsi 2-5 combined in length; pulvilli well-developed; empodium setiform (Figs 2 A, 4 C – G). — Wing. Nearly equal to abdomen in length; costal vein circumambient; crossvein h situated near wing base; Sc ending at mid-position of costal margin; R 1 stout and straight, bare dorsally; Rs equal to the distance of R 4 + 5 origin to crossvein r-m; R 2 + 3 slightly sinuous at distal portion, distant to R 1 at wing margin; R 4 + 5 forked at the level of tip of M 3; R 4 and R 5 diverged; R 4 sinuous; cell r 4 encompassing the wing apex; crossvein r-m locating at mid-position of cell d; cell d slender, 2 × longer than M 3; basal portion of M 1 arched; cell m 3 widely opened; crossvein m-cu present; cell cua closed; anal lobe wider than cell cua. Stem of haltere longer the apical knob in length (Fig. 4 H – K). — Abdomen. Slender, densely pubescent; tergites 2–6 well exposed, tergite 2 longest (~ 2 × tergite 1); tergite 7 partly covered by tergite 6; tergite 8 partly overlapping tergite 9 + 10; tergites 6-8 with long pilosity; both acanthophorite A 1 and A 2 spines present, acanthophorite A 1 spines elongate and acuminate apically; cercus broad, one-segmented (Fig. 5).</p><p>Remarks.</p><p>The † Paleothereva gen. nov. is assigned to Asiloidea based on a combination of characters, including reduced flagellomeres, absence of tibial spurs, and wing venations (Yeates 2002). Within Asiloidea, the presence of vein M 3 and a forked R 4 + 5 excludes its affinities with Bombyliidae and Mythicomyiidae, while the configuration of R 2 + 3 and the extension of M 1 beyond the wing apex preclude its assignment to Apioceridae . A two-segmented palpus, together with unfused M 1 and M 2 distinguishes it from Mydidae, and the bare face and non-predatory labellum separate it from Asilidae . The presence of a hind-coxal knob evidently support its inclusion in therevid clade. Among therevid families, † Paleothereva lacks the specialized tergal setae typical of Scenopinidae, and differs from Evocoidae by its short stylus and distinct divergent R 4 and R 5. It is further distinguished from Apsilocephalidae by the termination of R 5 beyond the wing apex. The elongated scape with stout setae, three-segmented flagellomeres, sinuous R 4, and presence of acanthophorite spines collectively provide strong evidence to assign † Paleothereva to Therevidae .</p><p>Within Therevidae, † Paleothereva can be excluded from Phycinae based on several diagnostic characters: in Phycinae, the costal vein terminates at or before CuA, whereas it is circumambient in † Paleothereva (Fig. 4 H – K); all extant and fossil members of Phycinae exhibit a dorsally setulose vein R 1, but R 1 is entirely bare in † Paleothereva (Fig. 4 J); acanthophorite spines are markedly reduced in Phycinae, while they are well developed in † Paleothereva (Fig. 5 C, D). Comparison with Xestomyzinae, the subfamily is most characterized by the presence of modified digging macrosetae on female sternite 8, but it is clearly absent in † Paleothereva . Additional characters further distinguish the genus from Xestomyzinae: cell m 3 is consistently closed in all known species of Xestomyzinae, whereas it is open in † Paleothereva (Fig. 4 H, I); the longitudinal wing fascia characteristic of Xestomyzinae is absent in the new genus (Fig. 4 H, I).</p><p>† Paleothereva is distinguished from the extant Therevinae by combining a single pair of macrosetae on scutellum (Fig. 4 A, B) and lacking lanceolate setae on hind femora (Fig. 4 F) (Hauser et al. 2017). Within Agapophytinae, several key diagnostic characters support a close relationship between † Paleothereva and the Taenogera genus-group: the scape lacks setae along the medial surface (Fig. 3 C), an apical seta is present on the hind femur (Fig. 4 F), and cell m 3 is open (Fig. 4 H, I, K) (Winterton et al. 1999, 2001). Nevertheless, † Paleothereva differs from members of the Taenogera genus group in having a scape as wide as the pedicel and a flagellum lacking setae (Fig. 3 C, D). Therefore, the new genus is erected here as a putative relative of the Taenogera genus-group within Agapophytinae .</p></div>	https://treatment.plazi.org/id/D985EB0E96AE541EAE63B70BB4B084AE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Feng, Qi;Jia, Xisen;Zhou, Chenxin;Ren, Dong;Wang, Yongjie	Feng, Qi, Jia, Xisen, Zhou, Chenxin, Ren, Dong, Wang, Yongjie (2026): The earliest crown therevid lineage: bridging the gap of early evolution of stiletto flies. Arthropod Systematics & Phylogeny 84: 281-292, DOI: 10.3897/asp.84.e185452
